treatments-xml/data/03/A3/0C/03A30C1C162FB964FF1BD2B5FDADCA9D.xml

<document id="A1749BB919D540E78F879EDD135BE06A" ID-CLB-Dataset="27776" ID-DOI="10.11646/zootaxa.4531.3.4" ID-GBIF-Dataset="35b13416-1715-4a8a-a3fb-e03876d9f101" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614748" ID-ZooBank="90C583D1-C656-495E-BDF1-CDB974EFB7C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787106029" checkinUser="plazi" docAuthor="Constantini, Joice P., Carrijo, Tiago Fernandes, Palma-Onetto, Valeria, Scheffrahn, Rudolf, Carnohan, Lucas Paul, Šobotník, Jan &amp; Cancello, Eliana M." docDate="2018" docId="03A30C1C162FB964FF1BD2B5FDADCA9D" docLanguage="en" docName="zootaxa.4531.3.4.pdf" docOrigin="Zootaxa 4531 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonsuritermes Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello 2018, gen. nov." docType="treatment" docVersion="8" lastPageNumber="387" masterDocId="FF9A7464162EB960FF8CD415FFC0CD40" masterDocTitle="Tonsuritermes, a new soldierless termite genus and two new species from South America (Blattaria: Isoptera: Termitidae: Apicotermitinae)" masterLastPageNumber="394" masterPageNumber="383" pageNumber="384" updateTime="1698714892089" updateUser="ExternalLinkService">
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<mods:title id="E9201128F24E364E76AEAA59AE4BD493">Tonsuritermes, a new soldierless termite genus and two new species from South America (Blattaria: Isoptera: Termitidae: Apicotermitinae)</mods:title>
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<mods:namePart id="6B1D993B4074A844C87C0D588DFE6F47">Constantini, Joice P.</mods:namePart>
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<mods:namePart id="E5A5B13957B33F43A295073B8E89D280">Carrijo, Tiago Fernandes</mods:namePart>
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<mods:namePart id="C9F04D73CB0B534D1373979043022B7F">Palma-Onetto, Valeria</mods:namePart>
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<mods:namePart id="953AD4650FAD32BBBFC84BBAFA67C880">Scheffrahn, Rudolf</mods:namePart>
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<mods:namePart id="BE5932435989BE1B835BC7E70C44BA70">Šobotník, Jan</mods:namePart>
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<mods:namePart id="9238F8F45A122FDDA3AF3CFB40FB6F80">Cancello, Eliana M.</mods:namePart>
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<treatment id="03A30C1C162FB964FF1BD2B5FDADCA9D" ID-DOI="http://doi.org/10.5281/zenodo.5963739" ID-GBIF-Taxon="154520923" ID-Zenodo-Dep="5963739" LSID="urn:lsid:plazi:treatment:03A30C1C162FB964FF1BD2B5FDADCA9D" httpUri="http://treatment.plazi.org/id/03A30C1C162FB964FF1BD2B5FDADCA9D" lastPageId="4" lastPageNumber="387" pageId="1" pageNumber="384">
<subSubSection id="C310EE81162FB961FF1BD2B5FD26CBBC" pageId="1" pageNumber="384" type="nomenclature">
<paragraph id="8BB5BD0A162FB961FF1BD2B5FD27CBFA" blockId="1.[151,743,1696,1722]" box="[151,743,1696,1722]" pageId="1" pageNumber="384">
<heading id="D0FD0A66162FB961FF1BD2B5FD27CBFA" bold="true" box="[151,743,1696,1722]" fontSize="11" level="1" pageId="1" pageNumber="384" reason="1">
<emphasis id="B97E6118162FB961FF1BD2B5FD27CBFA" bold="true" box="[151,743,1696,1722]" pageId="1" pageNumber="384">
<taxonomicName id="4C0AC689162FB961FF1BD2B5FDB2CBFA" authority="Cancello &amp; Constantini" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[151,626,1696,1722]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97E6118162FB961FF1BD2B5FE84CBFA" bold="true" box="[151,324,1696,1722]" italics="true" pageId="1" pageNumber="384">Tonsuritermes</emphasis>
Cancello &amp; Constantini
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<taxonomicNameLabel id="A24DDC63162FB961FDF5D2B5FD27CBFA" box="[633,743,1696,1722]" pageId="1" pageNumber="384" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="8BB5BD0A162FB961FF1BD2F3FD26CBBC" blockId="1.[151,742,1766,1789]" box="[151,742,1766,1789]" pageId="1" pageNumber="384">
Type-species. 
<taxonomicName id="4C0AC689162FB961FEABD2FDFE2ECBBC" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[295,494,1766,1789]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="tucki">
<emphasis id="B97E6118162FB961FEABD2FDFE2ECBBC" box="[295,494,1766,1789]" italics="true" pageId="1" pageNumber="384">Tonsuritermes tucki</emphasis>
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, by present designation.
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<subSubSection id="C310EE81162FB961FF1BD33CFAB1CA26" pageId="1" pageNumber="384" type="diagnosis">
<paragraph id="8BB5BD0A162FB961FF1BD33CFAB1CA26" blockId="1.[151,1437,1833,2038]" pageId="1" pageNumber="384">
<emphasis id="B97E6118162FB961FF1BD33CFED1CA02" bold="true" box="[151,273,1833,1858]" pageId="1" pageNumber="384">Diagnosis.</emphasis>
Worker and imago. Fontanelle very large in dorsal view, ranging from 1/4 to 3/4 the diameter of the head capsule (
<figureCitation id="1331A18F162FB961FEBAD358FE58CA26" box="[310,408,1869,1894]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="1" pageNumber="384">Figs. 1B</figureCitation>
; 
<figureCitation id="1331A18F162FB961FE29D35BFDCACA26" box="[421,522,1870,1894]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="1" pageNumber="384">2A, C, E</figureCitation>
); two rows of spine-like bristles on the inner face of the protibia (
<figureCitation id="1331A18F162FB961FB65D358FAEACA26" box="[1257,1322,1869,1894]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="1" pageNumber="384">Fig. 3</figureCitation>
A–C).
</paragraph>
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<subSubSection id="C310EE81162FB964FF4BD364FE80CC25" lastPageId="4" lastPageNumber="387" pageId="1" pageNumber="384" type="description">
<paragraph id="8BB5BD0A162FB962FF4BD364FD66CE54" blockId="1.[151,1437,1833,2038]" lastBlockId="2.[151,1437,151,789]" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384">
<emphasis id="B97E6118162FB961FF4BD364FE99CACA" bold="true" box="[199,345,1905,1930]" pageId="1" pageNumber="384">Description.</emphasis>
Imago (
<figureCitation id="1331A18F162FB961FE3BD364FDC6CACA" box="[439,518,1905,1930]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="1" pageNumber="384">Figs. 1</figureCitation>
; 
<figureCitation id="1331A18F162FB961FD98D367FDF8CACA" box="[532,568,1906,1930]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="1" pageNumber="384">3A</figureCitation>
, E–G, I). Head capsule trapezoidal in dorsal view, flattened dorsoventrally in profile (
<figureCitation id="1331A18F162FB961FF78D380FEB4CAEE" box="[244,372,1941,1966]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="1" pageNumber="384">Fig. 1A, B</figureCitation>
). Two frontal marks located in dorsal view between the postclypeus and ocellus, above the antenna insertion (
<figureCitation id="1331A18F162FB961FEE7D3ACFE05CA92" box="[363,453,1977,2002]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="1" pageNumber="384">Fig. 1B</figureCitation>
, arrows), larger than the ocelli, and two smaller triangular marks between the frontal marks and the fontanelle (
<figureCitation id="1331A18F162FB961FE44D3C8FDE2CAB6" box="[456,546,2013,2038]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="1" pageNumber="384">Fig. 1B</figureCitation>
). Fontanelle massive rounded, slightly depressed, occupying 1/2 of the head capsule in dorsal view (
<figureCitation id="1331A18F162CB962FE2BD482FE3FCDF0" box="[423,511,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1B</figureCitation>
). Eyes rounded, smaller in diameter than the fontanelle. Ocellus small, elliptical, separated from the eye margin by its diameter. Postclypeus moderately inflated, with median line conspicuous (
<figureCitation id="1331A18F162CB962FAE3D4A9FF1ECDB8" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1A, B</figureCitation>
). Labrum with hyaline distal margin. Left mandible (
<figureCitation id="1331A18F162CB962FCCCD4CAFC4DCDB8" box="[832,909,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">
Fig. 
<date id="FFB49BCA162CB962FCFAD4F5FC4DCDB8" box="[886,909,224,248]" pageId="2" pageNumber="385">3I</date>
</figureCitation>
) with apical tooth much more prominent than M1+2, incision conspicuous, M3 triangular with margins forming acute angle with the tip, point of molar tooth not hidden by molar prominence; molar prominence moderately developed; molar region without ridges. Right mandible (
<figureCitation id="1331A18F162CB962FE82D559FE9ACC24" box="[270,346,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">
Fig. 
<date id="FFB49BCA162CB962FECED559FE9ACC24" box="[322,346,332,356]" pageId="2" pageNumber="385">3I</date>
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) with apical tooth much more prominent than M1, M2 triangular with margins forming an obtuse angle with the tip; molar plate moderately developed; molar region without ridges. Pronotum narrower than head without eyes (
<figureCitation id="1331A18F162CB962FEB4D581FE50CCEC" box="[312,400,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1B</figureCitation>
, 
<tableCitation id="C68888B1162CB962FE12D581FE32CCEC" box="[414,498,404,429]" captionStart="TABLE 1" captionStartId="2.[151,239,833,856]" captionText="TABLE 1. Measurements (mm) of imagoes of Tonsuritermes tucki, sp. nov." pageId="2" pageNumber="385">Table 1</tableCitation>
), anterior margin straight (in 
<figureCitation id="1331A18F162CB962FCCBD581FC5ECCEC" box="[839,926,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1B</figureCitation>
, the image became slightly deformed during the stacking process), lateral margins convex, converging posteriorly. Wings ornamented with asteroid micrasters (
<figureCitation id="1331A18F162CB962FF13D5C9FF22CCB4" box="[159,226,476,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3</figureCitation>
F–G). Profemur subcylindrical. Protibia thin, not inflated (
<figureCitation id="1331A18F162CB962FC04D5C9FC25CCB4" box="[904,997,476,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3A</figureCitation>
). Females generally larger than males (
<tableCitation id="C68888B1162CB962FF12D5EAFF32CF58" box="[158,242,511,536]" captionStart="TABLE 1" captionStartId="2.[151,239,833,856]" captionText="TABLE 1. Measurements (mm) of imagoes of Tonsuritermes tucki, sp. nov." pageId="2" pageNumber="385">Table 1</tableCitation>
). Pilosity of head capsule with a dense coverage of uniform bristles and short hairs. Labrum with two long bristles and several hairs. Pronotum with bristles and short hairs, mainly along the margins. Tergites and sternites with short bristles and short hairs covering the center of the plates. Procoxa with 4–5 thick bristles. Profemur densely covered by bristles of different lengths (
<figureCitation id="1331A18F162CB962FD35D679FCD4CFC4" box="[697,788,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3A</figureCitation>
). Protibia with two rows of 10–12 thick bristles, with the femur, resembling a weakly “raptorial” leg (
<figureCitation id="1331A18F162CB962FD06D69AFD25CFE8" box="[650,741,655,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3A</figureCitation>
). Margin and veins of wings densely covered with hairs (
<figureCitation id="1331A18F162CB962FAE3D69AFF18CF8C" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3E, F</figureCitation>
). Coloration of head capsule dark brown, frontal marks and triangular marks slightly lighter than the rest of the head capsule, with poorly defined margins. Fontanelle concolorous with the head capsule (
<figureCitation id="1331A18F162CB962FB4CD6C2FAD7CFB0" box="[1216,1303,727,752]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1B</figureCitation>
). Pronotum slightly lighter than the head capsule (
<figureCitation id="1331A18F162CB962FDCED6E9FD58CE54" box="[578,664,764,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="2" pageNumber="385">Fig. 1B</figureCitation>
).
</paragraph>
<caption id="DF75ED82162CB962FF1BD754FC03CE18" box="[151,963,833,856]" pageId="2" pageNumber="385">
<paragraph id="8BB5BD0A162CB962FF1BD754FC03CE18" blockId="2.[151,963,833,856]" box="[151,963,833,856]" pageId="2" pageNumber="385">
<emphasis id="B97E6118162CB962FF1BD754FECACE18" bold="true" box="[151,266,833,856]" pageId="2" pageNumber="385">TABLE 1.</emphasis>
Measurements (mm) of imagoes of 
<taxonomicName id="4C0AC689162CB962FD02D754FCA0CE18" authority="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello, 2018" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[654,864,833,856]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="tucki" status="sp. nov.">
<emphasis id="B97E6118162CB962FD02D754FCA0CE18" box="[654,864,833,856]" italics="true" pageId="2" pageNumber="385">Tonsuritermes tucki</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24DDC63162CB962FCE1D754FC03CE18" box="[877,963,833,856]" pageId="2" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB5BD0A162CB962FD78D761FBAECB5F" pageId="2" pageNumber="385">
<table id="F90A4FAA162C469FFF13D761FAA4CB5F" box="[159,1380,884,1567]" gridcols="5" gridrows="17" pageId="2" pageNumber="385">
<tr id="353ABF48162C469FFF13D761FAA4CECA" box="[159,1380,884,906]" gridrow="0" pageId="2" pageNumber="385" rowspan-0="1" rowspan-2="1" rowspan-4="1">
<th id="76EBD634162C469FFD78D761FCB4CECA" box="[756,884,884,906]" gridcol="1" gridrow="0" pageId="2" pageNumber="385">female</th>
<th id="76EBD634162C469FFBC1D761FB0BCECA" box="[1101,1227,884,906]" gridcol="3" gridrow="0" pageId="2" pageNumber="385">male</th>
</tr>
<tr id="353ABF48162C469FFF13D78BFAA4CEF3" box="[159,1380,926,947]" gridrow="1" pageId="2" pageNumber="385" rowspan-0="1">
<td id="76EBD634162C469FFD78D78BFCB4CEF3" box="[756,884,926,947]" gridcol="1" gridrow="1" pageId="2" pageNumber="385">range</td>
<td id="76EBD634162C469FFC58D78BFBCDCEF3" box="[980,1037,926,947]" gridcol="2" gridrow="1" pageId="2" pageNumber="385">mean</td>
<td id="76EBD634162C469FFBC1D78BFB0BCEF3" box="[1101,1227,926,947]" gridcol="3" gridrow="1" pageId="2" pageNumber="385">range</td>
<td id="76EBD634162C469FFAA7D78BFAA4CEF3" box="[1323,1380,926,947]" gridcol="4" gridrow="1" pageId="2" pageNumber="385">mean</td>
</tr>
<tr id="353ABF48162C469FFF13D7D3FAA4CE9C" box="[159,1380,966,988]" gridrow="2" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D7D3FDDACE9C" box="[159,538,966,988]" gridcol="0" gridrow="2" pageId="2" pageNumber="385">length of the head</th>
<td id="76EBD634162C469FFD78D7D3FCB4CE9C" box="[756,884,966,988]" gridcol="1" gridrow="2" pageId="2" pageNumber="385">0.81–0.90</td>
<td id="76EBD634162C469FFC58D7D3FBCDCE9C" box="[980,1037,966,988]" gridcol="2" gridrow="2" pageId="2" pageNumber="385">0.88</td>
<td id="76EBD634162C469FFBC1D7D3FB0BCE9C" box="[1101,1227,966,988]" gridcol="3" gridrow="2" pageId="2" pageNumber="385">0.90–0.95</td>
<td id="76EBD634162C469FFAA7D7D3FAA4CE9C" box="[1323,1380,966,988]" gridcol="4" gridrow="2" pageId="2" pageNumber="385">0.92</td>
</tr>
<tr id="353ABF48162C469FFF13D7E5FAA4C946" box="[159,1380,1008,1030]" gridrow="3" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D7E5FDDAC946" box="[159,538,1008,1030]" gridcol="0" gridrow="3" pageId="2" pageNumber="385">width of head with eyes</th>
<td id="76EBD634162C469FFD78D7E5FCB4C946" box="[756,884,1008,1030]" gridcol="1" gridrow="3" pageId="2" pageNumber="385">1.25–1.36</td>
<td id="76EBD634162C469FFC58D7E5FBCDC946" box="[980,1037,1008,1030]" gridcol="2" gridrow="3" pageId="2" pageNumber="385">1.32</td>
<td id="76EBD634162C469FFBC1D7E5FB0BC946" box="[1101,1227,1008,1030]" gridcol="3" gridrow="3" pageId="2" pageNumber="385">1.20–1.25</td>
<td id="76EBD634162C469FFAA7D7E5FAA4C946" box="[1323,1380,1008,1030]" gridcol="4" gridrow="3" pageId="2" pageNumber="385">1.21</td>
</tr>
<tr id="353ABF48162C469FFF13D00CFAA4C96F" box="[159,1380,1049,1071]" gridrow="4" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D00CFDDAC96F" box="[159,538,1049,1071]" gridcol="0" gridrow="4" pageId="2" pageNumber="385">maximum diameter of compound eye</th>
<td id="76EBD634162C469FFD78D00CFCB4C96F" box="[756,884,1049,1071]" gridcol="1" gridrow="4" pageId="2" pageNumber="385">0.31</td>
<td id="76EBD634162C469FFC58D00CFBCDC96F" box="[980,1037,1049,1071]" gridcol="2" gridrow="4" pageId="2" pageNumber="385">0.31</td>
<td id="76EBD634162C469FFBC1D00CFB0BC96F" box="[1101,1227,1049,1071]" gridcol="3" gridrow="4" pageId="2" pageNumber="385">0.27–0.29</td>
<td id="76EBD634162C469FFAA7D00CFAA4C96F" box="[1323,1380,1049,1071]" gridcol="4" gridrow="4" pageId="2" pageNumber="385">0.29</td>
</tr>
<tr id="353ABF48162C469FFF13D057FAA4C918" box="[159,1380,1090,1112]" gridrow="5" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D057FDDAC918" box="[159,538,1090,1112]" gridcol="0" gridrow="5" pageId="2" pageNumber="385">inter-eye distance</th>
<td id="76EBD634162C469FFD78D057FCB4C918" box="[756,884,1090,1112]" gridcol="1" gridrow="5" pageId="2" pageNumber="385">1.06–1.11</td>
<td id="76EBD634162C469FFC58D057FBCDC918" box="[980,1037,1090,1112]" gridcol="2" gridrow="5" pageId="2" pageNumber="385">1.08</td>
<td id="76EBD634162C469FFBC1D057FB0BC918" box="[1101,1227,1090,1112]" gridcol="3" gridrow="5" pageId="2" pageNumber="385">0.95–0.97</td>
<td id="76EBD634162C469FFAA7D057FAA4C918" box="[1323,1380,1090,1112]" gridcol="4" gridrow="5" pageId="2" pageNumber="385">0.96</td>
</tr>
<tr id="353ABF48162C469FFF13D079FAA4C9C2" box="[159,1380,1132,1154]" gridrow="6" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D079FDDAC9C2" box="[159,538,1132,1154]" gridcol="0" gridrow="6" pageId="2" pageNumber="385">maximum diameter of ocellus</th>
<td id="76EBD634162C469FFD78D079FCB4C9C2" box="[756,884,1132,1154]" gridcol="1" gridrow="6" pageId="2" pageNumber="385">0.11–0.13</td>
<td id="76EBD634162C469FFC58D079FBCDC9C2" box="[980,1037,1132,1154]" gridcol="2" gridrow="6" pageId="2" pageNumber="385">0.12</td>
<td id="76EBD634162C469FFBC1D079FB0BC9C2" box="[1101,1227,1132,1154]" gridcol="3" gridrow="6" pageId="2" pageNumber="385">0.11–0.13</td>
<td id="76EBD634162C469FFAA7D079FAA4C9C2" box="[1323,1380,1132,1154]" gridcol="4" gridrow="6" pageId="2" pageNumber="385">0.11</td>
</tr>
<tr id="353ABF48162C469FFF13D080FAA4C9EB" box="[159,1380,1173,1195]" gridrow="7" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D080FDDAC9EB" box="[159,538,1173,1195]" gridcol="0" gridrow="7" pageId="2" pageNumber="385">minimum diameter of ocellus</th>
<td id="76EBD634162C469FFD78D080FCB4C9EB" box="[756,884,1173,1195]" gridcol="1" gridrow="7" pageId="2" pageNumber="385">0.09</td>
<td id="76EBD634162C469FFC58D080FBCDC9EB" box="[980,1037,1173,1195]" gridcol="2" gridrow="7" pageId="2" pageNumber="385">0.09</td>
<td id="76EBD634162C469FFBC1D080FB0BC9EB" box="[1101,1227,1173,1195]" gridcol="3" gridrow="7" pageId="2" pageNumber="385">0.06–0.09</td>
<td id="76EBD634162C469FFAA7D080FAA4C9EB" box="[1323,1380,1173,1195]" gridcol="4" gridrow="7" pageId="2" pageNumber="385">0.07</td>
</tr>
<tr id="353ABF48162C469FFF13D0ABFAA4C994" box="[159,1380,1214,1236]" gridrow="8" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D0ABFDDAC994" box="[159,538,1214,1236]" gridcol="0" gridrow="8" pageId="2" pageNumber="385">eye-ocellus distance</th>
<td id="76EBD634162C469FFD78D0ABFCB4C994" box="[756,884,1214,1236]" gridcol="1" gridrow="8" pageId="2" pageNumber="385">0.09–0.11</td>
<td id="76EBD634162C469FFC58D0ABFBCDC994" box="[980,1037,1214,1236]" gridcol="2" gridrow="8" pageId="2" pageNumber="385">0.09</td>
<td id="76EBD634162C469FFBC1D0ABFB0BC994" box="[1101,1227,1214,1236]" gridcol="3" gridrow="8" pageId="2" pageNumber="385">0.90</td>
<td id="76EBD634162C469FFAA7D0ABFAA4C994" box="[1323,1380,1214,1236]" gridcol="4" gridrow="8" pageId="2" pageNumber="385">0.90</td>
</tr>
<tr id="353ABF48162C469FFF13D0FDFAA4C9BE" box="[159,1380,1256,1278]" gridrow="9" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D0FDFDDAC9BE" box="[159,538,1256,1278]" gridcol="0" gridrow="9" pageId="2" pageNumber="385">length of pronotum</th>
<td id="76EBD634162C469FFD78D0FDFCB4C9BE" box="[756,884,1256,1278]" gridcol="1" gridrow="9" pageId="2" pageNumber="385">0.54–0.65</td>
<td id="76EBD634162C469FFC58D0FDFBCDC9BE" box="[980,1037,1256,1278]" gridcol="2" gridrow="9" pageId="2" pageNumber="385">0.61</td>
<td id="76EBD634162C469FFBC1D0FDFB0BC9BE" box="[1101,1227,1256,1278]" gridcol="3" gridrow="9" pageId="2" pageNumber="385">0.52–0.54</td>
<td id="76EBD634162C469FFAA7D0FDFAA4C9BE" box="[1323,1380,1256,1278]" gridcol="4" gridrow="9" pageId="2" pageNumber="385">0.53</td>
</tr>
<tr id="353ABF48162C469FFF13D104FAA4C867" box="[159,1380,1297,1319]" gridrow="10" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D104FDDAC867" box="[159,538,1297,1319]" gridcol="0" gridrow="10" pageId="2" pageNumber="385">width of pronotum</th>
<td id="76EBD634162C469FFD78D104FCB4C867" box="[756,884,1297,1319]" gridcol="1" gridrow="10" pageId="2" pageNumber="385">1.08–1.16</td>
<td id="76EBD634162C469FFC58D104FBCDC867" box="[980,1037,1297,1319]" gridcol="2" gridrow="10" pageId="2" pageNumber="385">1.15</td>
<td id="76EBD634162C469FFBC1D104FB0BC867" box="[1101,1227,1297,1319]" gridcol="3" gridrow="10" pageId="2" pageNumber="385">0.96–1.00</td>
<td id="76EBD634162C469FFAA7D104FAA4C867" box="[1323,1380,1297,1319]" gridcol="4" gridrow="10" pageId="2" pageNumber="385">0.97</td>
</tr>
<tr id="353ABF48162C469FFF13D12FFAA4C810" box="[159,1380,1338,1360]" gridrow="11" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D12FFDDAC810" box="[159,538,1338,1360]" gridcol="0" gridrow="11" pageId="2" pageNumber="385">length of forewing with scale</th>
<td id="76EBD634162C469FFD78D12FFCB4C810" box="[756,884,1338,1360]" gridcol="1" gridrow="11" pageId="2" pageNumber="385">13.10–15.10</td>
<td id="76EBD634162C469FFC58D12FFBCDC810" box="[980,1037,1338,1360]" gridcol="2" gridrow="11" pageId="2" pageNumber="385">13.89</td>
<td id="76EBD634162C469FFBC1D12FFB0BC810" box="[1101,1227,1338,1360]" gridcol="3" gridrow="11" pageId="2" pageNumber="385">11.06–12.10</td>
<td id="76EBD634162C469FFAA7D12FFAA4C810" box="[1323,1380,1338,1360]" gridcol="4" gridrow="11" pageId="2" pageNumber="385">11.54</td>
</tr>
<tr id="353ABF48162C469FFF13D171FAA4C83A" box="[159,1380,1380,1402]" gridrow="12" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D171FDDAC83A" box="[159,538,1380,1402]" gridcol="0" gridrow="12" pageId="2" pageNumber="385">width of the protibia</th>
<td id="76EBD634162C469FFD78D171FCB4C83A" box="[756,884,1380,1402]" gridcol="1" gridrow="12" pageId="2" pageNumber="385">0.12–0.14</td>
<td id="76EBD634162C469FFC58D171FBCDC83A" box="[980,1037,1380,1402]" gridcol="2" gridrow="12" pageId="2" pageNumber="385">0.13</td>
<td id="76EBD634162C469FFBC1D171FB0BC83A" box="[1101,1227,1380,1402]" gridcol="3" gridrow="12" pageId="2" pageNumber="385">0.10–0.12</td>
<td id="76EBD634162C469FFAA7D171FAA4C83A" box="[1323,1380,1380,1402]" gridcol="4" gridrow="12" pageId="2" pageNumber="385">0.11</td>
</tr>
<tr id="353ABF48162C469FFF13D198FAA4C8E3" box="[159,1380,1421,1443]" gridrow="13" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D198FDDAC8E3" box="[159,538,1421,1443]" gridcol="0" gridrow="13" pageId="2" pageNumber="385">length of the protibia</th>
<td id="76EBD634162C469FFD78D198FCB4C8E3" box="[756,884,1421,1443]" gridcol="1" gridrow="13" pageId="2" pageNumber="385">0.92–0.94</td>
<td id="76EBD634162C469FFC58D198FBCDC8E3" box="[980,1037,1421,1443]" gridcol="2" gridrow="13" pageId="2" pageNumber="385">0.92</td>
<td id="76EBD634162C469FFBC1D198FB0BC8E3" box="[1101,1227,1421,1443]" gridcol="3" gridrow="13" pageId="2" pageNumber="385">0.80–0.86</td>
<td id="76EBD634162C469FFAA7D198FAA4C8E3" box="[1323,1380,1421,1443]" gridcol="4" gridrow="13" pageId="2" pageNumber="385">0.83</td>
</tr>
<tr id="353ABF48162C469FFF13D1A3FAA4C88C" box="[159,1380,1462,1484]" gridrow="14" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D1A3FDDAC88C" box="[159,538,1462,1484]" gridcol="0" gridrow="14" pageId="2" pageNumber="385">length of the metatibia</th>
<td id="76EBD634162C469FFD78D1A3FCB4C88C" box="[756,884,1462,1484]" gridcol="1" gridrow="14" pageId="2" pageNumber="385">1.25–1.34</td>
<td id="76EBD634162C469FFC58D1A3FBCDC88C" box="[980,1037,1462,1484]" gridcol="2" gridrow="14" pageId="2" pageNumber="385">1.26</td>
<td id="76EBD634162C469FFBC1D1A3FB0BC88C" box="[1101,1227,1462,1484]" gridcol="3" gridrow="14" pageId="2" pageNumber="385">1.18–1.36</td>
<td id="76EBD634162C469FFAA7D1A3FAA4C88C" box="[1323,1380,1462,1484]" gridcol="4" gridrow="14" pageId="2" pageNumber="385">1.22</td>
</tr>
<tr id="353ABF48162C469FFF13D1F5FAA4C8B6" box="[159,1380,1504,1526]" gridrow="15" pageId="2" pageNumber="385">
<th id="76EBD634162C469FFF13D1F5FDDAC8B6" box="[159,538,1504,1526]" gridcol="0" gridrow="15" pageId="2" pageNumber="385">maximum diameter of fontanelle</th>
<td id="76EBD634162C469FFD78D1F5FCB4C8B6" box="[756,884,1504,1526]" gridcol="1" gridrow="15" pageId="2" pageNumber="385">0.34–0.36</td>
<td id="76EBD634162C469FFC58D1F5FBCDC8B6" box="[980,1037,1504,1526]" gridcol="2" gridrow="15" pageId="2" pageNumber="385">0.36</td>
<td id="76EBD634162C469FFBC1D1F5FB0BC8B6" box="[1101,1227,1504,1526]" gridcol="3" gridrow="15" pageId="2" pageNumber="385">0.22</td>
<td id="76EBD634162C469FFAA7D1F5FAA4C8B6" box="[1323,1380,1504,1526]" gridcol="4" gridrow="15" pageId="2" pageNumber="385">0.22</td>
</tr>
<tr id="353ABF48162C469FFF13D21CFAA4CB5F" box="[159,1380,1545,1567]" gridrow="16" pageId="2" pageNumber="385" rowspan-2="1" rowspan-4="1">
<th id="76EBD634162C469FFF13D21CFDDACB5F" box="[159,538,1545,1567]" gridcol="0" gridrow="16" pageId="2" pageNumber="385">specimens/colonies</th>
<td id="76EBD634162C469FFD78D21CFCB4CB5F" box="[756,884,1545,1567]" gridcol="1" gridrow="16" pageId="2" pageNumber="385">5/1</td>
<td id="76EBD634162C469FFBC1D21CFB0BCB5F" box="[1101,1227,1545,1567]" gridcol="3" gridrow="16" pageId="2" pageNumber="385">5/1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB5BD0A162CB963FF4BD24AFBF1CF7C" blockId="2.[151,1436,1631,2017]" lastBlockId="3.[151,1436,151,573]" lastPageId="3" lastPageNumber="386" pageId="2" pageNumber="385">
Worker. Mono- or dimorphic (
<figureCitation id="1331A18F162CB962FDBDD24AFD47CB38" box="[561,647,1631,1656]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="2" pageNumber="385">Figs. 2</figureCitation>
; 
<figureCitation id="1331A18F162CB962FD14D275FD64CB38" box="[664,676,1632,1656]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">3</figureCitation>
B–D, H, 4, 5). Head capsule flattened dorsoventrally in profile, rounded in dorsal view. Fontanelle margins well delineated, depressed within vertex, occupying 1/4 to 3/4 of the head capsule (
<figureCitation id="1331A18F162CB962FEB4D2B2FE0CCBFF" box="[312,460,1703,1728]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="2" pageNumber="385">Fig. 2A,C, E</figureCitation>
, see histological discussion below). Antenna with 14 articles. Postclypeus moderately to highly inflated. Left mandible (
<figureCitation id="1331A18F162CB962FD90D2D9FDB4CBA4" box="[540,628,1740,1765]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3H</figureCitation>
) with apical tooth longer than M1+2, cutting edge between M1+2 and incision, incision conspicuous, M3 triangular with lateral sides forming an acute angle with the tip; molar tooth not hidden by molar prominence; molar prominence well developed; molar region concave, without ridges. Right mandible (
<figureCitation id="1331A18F162CB962FE83D322FEA7CA10" box="[271,359,1847,1872]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="2" pageNumber="385">Fig. 3H</figureCitation>
) with apical tooth longer than M1; M2 triangular with margins forming an acute angle with the tip; molar plate well developed; molar region concave, without ridges. Pronotum in lateral view (
<figureCitation id="1331A18F162CB962FB7BD349FA54CA34" box="[1271,1428,1884,1909]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="2" pageNumber="385">Fig. 2B, D, F</figureCitation>
) with anterior lobe much longer and forming right angle with posterior lobe. Mesonotum and metanotum subretangular. Thoraco-abdominal glands or dehiscent organs absent. Profemur with ventral surface forming a groove between the two rows of bristles. Protibia with ventral face strongly flattened, resembling an interlocking raptorial leg (
<figureCitation id="1331A18F162DB963FEBED482FE6ECDF0" box="[306,430,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="3" pageNumber="386">Fig. 3B, C</figureCitation>
). Digestive tube (
<figureCitation id="1331A18F162DB963FDF5D482FD7ECDF0" box="[633,702,151,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1360,1382]" captionTargetBox="[151,1436,215,1329]" captionTargetId="figure@7.[151,1436,215,1329]" captionTargetPageId="7" captionText="FIGURE 4. Tonsuritermes tucki sp. nov., worker. Digestive tube in A—dorsal; B—right; C—ventral; D—left views; Eenteric valve armature; F—enlarged view of the a cushion. Specimens from lots MZUSP6840 (A–E) and MZUSP 10367 (F). Gut regions indicated in figs. A–D: C=crop, G=gizzard, M=mesenteron, MS= mixed segment, P1= ileum, P2= enteric valve, P3= paunch, P4= colon, P5=rectum." httpUri="https://zenodo.org/record/2614756/files/figure.png" pageId="3" pageNumber="386">Fig. 4</figureCitation>
) with inner mixed segment vestigial (
<figureCitation id="1331A18F162DB963FBFDD482FB0BCDF0" box="[1137,1227,151,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1360,1382]" captionTargetBox="[151,1436,215,1329]" captionTargetId="figure@7.[151,1436,215,1329]" captionTargetPageId="7" captionText="FIGURE 4. Tonsuritermes tucki sp. nov., worker. Digestive tube in A—dorsal; B—right; C—ventral; D—left views; Eenteric valve armature; F—enlarged view of the a cushion. Specimens from lots MZUSP6840 (A–E) and MZUSP 10367 (F). Gut regions indicated in figs. A–D: C=crop, G=gizzard, M=mesenteron, MS= mixed segment, P1= ileum, P2= enteric valve, P3= paunch, P4= colon, P5=rectum." httpUri="https://zenodo.org/record/2614756/files/figure.png" pageId="3" pageNumber="386">Fig. 4C</figureCitation>
); P1 with uniform diameter throughout, forming an inverted 'C' in ventral view; P2 not armed, composed of six symmetrical cushions covered with faint polygonal scales (
<figureCitation id="1331A18F162DB963FDB8D4CAFDB6CDB8" box="[564,630,223,248]" captionStart="FIGURE 4" captionStartId="7.[151,250,1360,1382]" captionTargetBox="[151,1436,215,1329]" captionTargetId="figure@7.[151,1436,215,1329]" captionTargetPageId="7" captionText="FIGURE 4. Tonsuritermes tucki sp. nov., worker. Digestive tube in A—dorsal; B—right; C—ventral; D—left views; Eenteric valve armature; F—enlarged view of the a cushion. Specimens from lots MZUSP6840 (A–E) and MZUSP 10367 (F). Gut regions indicated in figs. A–D: C=crop, G=gizzard, M=mesenteron, MS= mixed segment, P1= ileum, P2= enteric valve, P3= paunch, P4= colon, P5=rectum." httpUri="https://zenodo.org/record/2614756/files/figure.png" pageId="3" pageNumber="386">Fig. 4</figureCitation>
E–F) and 3–5 small short triangular spines in the proximal region of the cushions (
<figureCitation id="1331A18F162DB963FE85D511FE9FCC5C" box="[265,351,260,285]" captionStart="FIGURE 4" captionStartId="7.[151,250,1360,1382]" captionTargetBox="[151,1436,215,1329]" captionTargetId="figure@7.[151,1436,215,1329]" captionTargetPageId="7" captionText="FIGURE 4. Tonsuritermes tucki sp. nov., worker. Digestive tube in A—dorsal; B—right; C—ventral; D—left views; Eenteric valve armature; F—enlarged view of the a cushion. Specimens from lots MZUSP6840 (A–E) and MZUSP 10367 (F). Gut regions indicated in figs. A–D: C=crop, G=gizzard, M=mesenteron, MS= mixed segment, P1= ileum, P2= enteric valve, P3= paunch, P4= colon, P5=rectum." httpUri="https://zenodo.org/record/2614756/files/figure.png" pageId="3" pageNumber="386">Fig. 4F</figureCitation>
). Enteric valve seating tubular, without lobes. Paunch with P3a pyriform and P3b forming an Sshaped, isthmus conspicuous in almost all the specimens; P4 of uniform width, passing under mesenteron in the sagital line and making a 180° loop in the left side of body right before the P5. Pilosity of head capsule covered by bristles of variable orientation and length. Pronotum with long bristles with variable orientation, in greater number on margin of the anterior lobe and in the rounded regions in the posterior lobe. Tergites and sternites with short bristles in the center of the plates, in variable orientations. Procoxa with 7–11 thick bristles. Profemur with two rows of bristles less striking and organized than the protibia (
<figureCitation id="1331A18F162DB963FCDED5C9FC0ECCB4" box="[850,974,476,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="3" pageNumber="386">Fig. 3B, C</figureCitation>
). Protibia with two well marked rows of spine-like bristles (thick and short) on inner margin; number of spine-like bristles varying from six to 16 (
<figureCitation id="1331A18F162DB963FAB3D5EAFF6DCF7C" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="3" pageNumber="386">Fig. 3B, C</figureCitation>
). Head capsule coloration whitish yellow; fontanelle lighter (
<figureCitation id="1331A18F162DB963FCD6D631FC33CF7C" box="[858,1011,548,573]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="3" pageNumber="386">Fig. 2A, C, E</figureCitation>
, 
<figureCitation id="1331A18F162DB963FC73D631FBE4CF7C" box="[1023,1060,548,572]" captionStart="FIGURE 3" captionStartId="6.[151,250,1268,1290]" captionTargetBox="[151,1436,193,1246]" captionTargetId="figure@6.[151,1436,193,1246]" captionTargetPageId="6" captionText="FIGURE 3. Tonsuritermes, workers and imago. A—foreleg of the imago of T. tucki, sp. nov.; B—foreleg of the worker of T. tucki; C—foreleg of the worker of T. mathewsi, sp. nov.; D—detail of the texture and surrounding pilosity of the glandular opening, worker type 1 of T. tucki; E—detail of external margin of the wing of T. tucki; F—detail of inner margin of the wing; G—micrasters on membranous area; H—worker mandibles; I—imago mandibles. Specimens from lots MZUSP6480 (A–B, E– I), MZUSP12268 (C) and UF PU1104 (D)." httpUri="https://zenodo.org/record/2614754/files/figure.png" pageId="3" pageNumber="386">3D</figureCitation>
).
</paragraph>
<caption id="DF75ED82162DB963FF1BD0AEFDF3C9AF" httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="3" pageNumber="386" startId="3.[151,250,1211,1233]" targetBox="[151,1436,665,1146]" targetPageId="3">
<paragraph id="8BB5BD0A162DB963FF1BD0AEFDF3C9AF" blockId="3.[151,1436,1211,1263]" pageId="3" pageNumber="386">
<emphasis id="B97E6118162DB963FF1BD0AEFED3C990" bold="true" box="[151,275,1211,1233]" pageId="3" pageNumber="386">FIGURE 1.</emphasis>
<taxonomicName id="4C0AC689162DB963FE94D0A9FE1FC991" authority="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello, 2018" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[280,479,1211,1233]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="tucki" status="sp. nov.">
<emphasis id="B97E6118162DB963FE94D0A9FE1FC991" box="[280,479,1211,1233]" italics="true" pageId="3" pageNumber="386">Tonsuritermes tucki</emphasis>
</taxonomicName>
<emphasis id="B97E6118162DB963FE6AD0AEFDF6C990" bold="true" box="[486,566,1211,1232]" pageId="3" pageNumber="386">
<taxonomicNameLabel id="A24DDC63162DB963FE6AD0AEFDF6C990" box="[486,566,1211,1232]" pageId="3" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480.
</paragraph>
</caption>
<paragraph id="8BB5BD0A162DB963FF4BD103FD51CB9F" blockId="3.[151,1437,1302,2011]" pageId="3" pageNumber="386">
<emphasis id="B97E6118162DB963FF4BD103FEACC86F" bold="true" box="[199,364,1302,1327]" pageId="3" pageNumber="386">Comparisons.</emphasis>
<taxonomicName id="4C0AC689162DB963FEF4D10DFDD7C86F" authority="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello, 2018" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[376,535,1304,1327]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97E6118162DB963FEF4D10DFDD7C86F" box="[376,535,1304,1327]" italics="true" pageId="3" pageNumber="386">Tonsuritermes</emphasis>
</taxonomicName>
, with its massive worker and imago fontanelle, differs from all known New World Apicotermitinae. 
<taxonomicName id="4C0AC689162DB963FE3BD129FD96C813" authority="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello, 2018" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[439,598,1340,1363]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97E6118162DB963FE3BD129FD96C813" box="[439,598,1340,1363]" italics="true" pageId="3" pageNumber="386">Tonsuritermes</emphasis>
</taxonomicName>
is close to 
<taxonomicName id="4C0AC689162DB963FD79D12EFBEDC813" authority="Fontes, 1986" authorityName="Fontes" authorityYear="1986" box="[757,1069,1338,1363]" class="Insecta" family="Termitidae" genus="Aparatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="B97E6118162DB963FD79D12EFC46C812" box="[757,902,1339,1362]" italics="true" pageId="3" pageNumber="386">Aparatermes</emphasis>
<bibRefCitation id="EF9BC0FB162DB963FC18D12EFBEDC813" author="Fontes, L. R." box="[916,1069,1338,1363]" pageId="3" pageNumber="386" pagination="285 - 297" refId="ref5360" refString="Fontes, L. R. (1986) Two new genera of soldierless Apicotermitinae from the Neotropical region (Isoptera, Termitidae). Sociobiology, 12, 285 - 297." type="journal article" year="1986">Fontes, 1986</bibRefCitation>
</taxonomicName>
(worker width of head of 
<taxonomicName id="4C0AC689162DB963FA08D12EFEDAC837" baseAuthorityName="Silvestri" baseAuthorityYear="1901" class="Insecta" family="Termitidae" genus="Aparatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="abbreviatus">
<emphasis id="B97E6118162DB963FA08D12EFEDAC837" italics="true" pageId="3" pageNumber="386">A. abbreviatus</emphasis>
</taxonomicName>
= 
<quantity id="4CF210EF162DB963FEB5D14BFE5EC837" box="[313,414,1374,1399]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="3" pageNumber="386" unit="mm" value="1.16">1.16 mm</quantity>
(mean), 
<bibRefCitation id="EF9BC0FB162DB963FD8FD175FD4EC837" author="Fontes, L. R." box="[515,654,1374,1399]" pageId="3" pageNumber="386" pagination="285 - 297" refId="ref5360" refString="Fontes, L. R. (1986) Two new genera of soldierless Apicotermitinae from the Neotropical region (Isoptera, Termitidae). Sociobiology, 12, 285 - 297." type="journal article" year="1986">Fontes 1986</bibRefCitation>
) and 
<taxonomicName id="4C0AC689162DB963FD42D175FBCAC837" authority="Mathews, 1977" authorityName="Mathews" authorityYear="1977" box="[718,1034,1374,1399]" class="Insecta" family="Termitidae" genus="Ruptitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="B97E6118162DB963FD42D175FC95C837" box="[718,853,1376,1399]" italics="true" pageId="3" pageNumber="386">Ruptitermes</emphasis>
<bibRefCitation id="EF9BC0FB162DB963FCD0D14BFBCAC837" author="Mathews, A. G. A." box="[860,1034,1374,1399]" pageId="3" pageNumber="386" refId="ref5509" refString="Mathews, A. G. A. (1977) Studies on Termites from the Mato Grosso State, Brazil. Academia Brasileira de Ciencias, Rio de Janeiro, 267 pp." type="book" year="1977">
Mathews, 
<quantity id="4CF210EF162DB963FC5ED14AFBCAC837" box="[978,1034,1375,1399]" metricMagnitude="1" metricUnit="m" metricValue="5.02158" pageId="3" pageNumber="386" unit="in" value="1977.0">1977</quantity>
</bibRefCitation>
</taxonomicName>
in size (worker width of head of 
<taxonomicName id="4C0AC689162DB963FA08D175FE0FC8DB" authority="(Silvestri, 1901)" authorityName="Silvestri" authorityYear="1901" baseAuthorityName="Silvestri" baseAuthorityYear="1901" class="Insecta" family="Termitidae" genus="Ruptitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="reconditus">
<emphasis id="B97E6118162DB963FA08D175FECCC8DB" italics="true" pageId="3" pageNumber="386">R. reconditus</emphasis>
(
<bibRefCitation id="EF9BC0FB162DB963FE92D197FE06C8DB" author="Silvestri, F." box="[286,454,1410,1435]" pageId="3" pageNumber="386" pagination="1 - 8" refId="ref6029" refString="Silvestri, F. (1901) Nota preliminare sui Termitidi sud-americani. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino, 16, 1 - 8." type="journal article" year="1901">Silvestri, 1901</bibRefCitation>
)
</taxonomicName>
= 
<quantity id="4CF210EF162DB963FE7ED196FD99C8DA" box="[498,601,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="3" pageNumber="386" unit="mm" value="1.13">1.13 mm</quantity>
(mean), 
<bibRefCitation id="EF9BC0FB162DB963FD4FD197FBC1C8DB" author="Acioli, A. N. &amp; Constantino, R." box="[707,1025,1410,1435]" pageId="3" pageNumber="386" pagination="451 - 492" refId="ref4588" refString="Acioli, A. N. &amp; Constantino, R. (2015) A taxonomic revision of the neotropical termite genus Ruptitermes (Isoptera, Termitidae, Apicotermitinae). Zootaxa, 4032 (5), 451 - 492. https: // doi. org / 10.11646 / zootaxa. 4032.5.1" type="journal article" year="2015">Acioli &amp; Constantino 2015</bibRefCitation>
). In 
<bibRefCitation id="EF9BC0FB162DB963FBB4D197FB2BC8DB" author="Snyder, T. E." box="[1080,1259,1410,1435]" pageId="3" pageNumber="386" pagination="1 - 76" refId="ref6062" refString="Snyder, T. E. (1926) Termites collected on the Mulford Biological Exploration to the Amazon Basin, 1921 - 1922. Proceedings of the United States National Museum, 68, 1 - 76. https: // doi. org / 10.5479 / si. 00963801.68 - 2615.1" type="journal article" year="1926">Snyder's (1926)</bibRefCitation>
description for 
<taxonomicName id="4C0AC689162DB963FF1BD1B3FE76C8FF" box="[151,438,1446,1471]" class="Insecta" family="Termitidae" genus="Anoplotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="grandifons">
<emphasis id="B97E6118162DB963FF1BD1B3FE76C8FF" box="[151,438,1446,1471]" italics="true" pageId="3" pageNumber="386">Anoplotermes grandifons</emphasis>
</taxonomicName>
, he highlights the fontanelle as a diagnostic feature of the species with a description based only on a wingless female: 
<emphasis id="B97E6118162DB963FD99D1DFFDF6CB47" italics="true" pageId="3" pageNumber="386">
'Fontanelle a very prominent hyaline oblong depression, slightly on a bias, 
<quantity id="4CF210EF162DB963FAE7D1DEFF00CB47" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="386" unit="mm" value="0.25">0.25 mm</quantity>
in length and 
<quantity id="4CF210EF162DB963FEEED1FAFE02CB47" box="[354,450,1519,1543]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="386" unit="mm" value="0.2">0.20 mm</quantity>
in width’.
</emphasis>
The fontanelle of the imago of 
<taxonomicName id="4C0AC689162DB963FC17D1E5FBB7CB47" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[923,1143,1518,1543]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="tucki">
<emphasis id="B97E6118162DB963FC17D1E5FBB7CB47" box="[923,1143,1518,1543]" italics="true" pageId="3" pageNumber="386">Tonsuritermes tucki</emphasis>
</taxonomicName>
is not hyaline and has the maximum diameter between 
<quantity id="4CF210EF162DB963FE52D207FD44CB6A" box="[478,644,1554,1579]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="3.5999999999999996" metricValueMin="3.4" pageId="3" pageNumber="386" unit="mm" value="0.35" valueMax="0.36" valueMin="0.34">0.34–0.36 mm</quantity>
. Snyder does not make any mention of bristles or spine-like bristles in the legs. Efforts were made to examine the 
<typeStatus id="54B103A8162DB963FD10D22DFD0CCB10" box="[668,716,1592,1616]" pageId="3" pageNumber="386">type</typeStatus>
material of 
<taxonomicName id="4C0AC689162DB963FCECD222FC3DCB0F" box="[864,1021,1590,1615]" class="Insecta" family="Termitidae" genus="Anoplotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="grandifons">
<emphasis id="B97E6118162DB963FCECD222FC3DCB0F" box="[864,1021,1590,1615]" italics="true" pageId="3" pageNumber="386">A. grandifons</emphasis>
</taxonomicName>
, but without success. Thus, for the difficulty of comparisons using only the description of the imagoes we considered our samples different from 
<taxonomicName id="4C0AC689162DB963FA08D24EFED0CBD7" class="Insecta" family="Termitidae" genus="Anoplotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="grandifons">
<emphasis id="B97E6118162DB963FA08D24EFED0CBD7" italics="true" pageId="3" pageNumber="386">A. grandifons</emphasis>
</taxonomicName>
. The protibia and profemur with thick spines, although also a very distinct characteristic in relation to the other described termites in the subfamily, has been observed in other groups of termites not related to the genus and not yet described (JC, unpublished data).
</paragraph>
<paragraph id="8BB5BD0A162DB964FF4BD2FFFE80CC25" blockId="3.[151,1437,1302,2011]" lastBlockId="4.[151,1437,151,536]" lastPageId="4" lastPageNumber="387" pageId="3" pageNumber="386">
<emphasis id="B97E6118162DB963FF4BD2FFFE1CCA43" bold="true" box="[199,476,1770,1795]" pageId="3" pageNumber="386">Histological discussion.</emphasis>
The large region of frons in 
<taxonomicName id="4C0AC689162DB963FC92D2F9FC3ACA43" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[798,1018,1770,1795]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="tucki">
<emphasis id="B97E6118162DB963FC92D2F9FC3ACA43" box="[798,1018,1770,1795]" italics="true" pageId="3" pageNumber="386">Tonsuritermes tucki</emphasis>
</taxonomicName>
workers is formed by a frontal gland of unique structure, even compared to other members of the 
<taxonomicName id="4C0AC689162DB963FCC6D31BFC26CA67" box="[842,998,1806,1831]" class="Insecta" family="Termitidae" genus="Anoplotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="B97E6118162DB963FCC6D31BFC26CA67" box="[842,998,1806,1831]" italics="true" pageId="3" pageNumber="386">Anoplotermes</emphasis>
</taxonomicName>
-group studied previously (
<bibRefCitation id="EF9BC0FB162DB963FA94D31BFEC2CA0B" author="Sobotnik, J. &amp; Sillam-Dusses, D. &amp; Weyda, F. &amp; Dejean, A. &amp; Roisin, Y. &amp; Hanus, R. &amp; Bourguignon, T." pageId="3" pageNumber="386" pagination="495 - 503" refId="ref6206" refString="Sobotnik, J., Sillam-Dusses, D., Weyda, F., Dejean, A., Roisin, Y., Hanus, R. &amp; Bourguignon, T. (2010 a) The frontal gland in workers of Neotropical soldierless termites. Naturwissenschaften, 97, 495 - 503. https: // doi. org / 10.1007 / s 00114 - 010 - 0664 - 0" type="journal article" year="2010">
Šobotník 
<emphasis id="B97E6118162DB963FA04D305FF6DCA0B" italics="true" pageId="3" pageNumber="386">et al</emphasis>
. 2010a
</bibRefCitation>
) (see 
<figureCitation id="1331A18F162DB963FECAD327FE48CA0B" box="[326,392,1842,1867]" captionStart="FIGURE 5" captionStartId="8.[151,250,1454,1476]" captionTargetBox="[152,1436,745,1441]" captionTargetId="figure@8.[151,1436,745,1441]" captionTargetPageId="8" captionText="FIGURE 5. Tonsuritermes tucki sp. nov., structure of the frontal gland in worker type 1, A—Sagittal section of the head of Tonsuritermes tucki sp. nov. worker. The frontal gland is stretched in-between the two arrows. B—Transition between the head cuticle and highly modified glandular cuticle overlying the secretory cells. C—Remains of secretory cell ultrastructures. Abbreviations: br, brain (supraoesophageal ganglium); cl, clypeus; gc1, inner glandular cuticle; gc2, outer glandular cuticle; hc, head cuticle; hp, hypopharynx; lb, labrum; m, mitochondria; mm, mandibular muscles; n, nucleus; ph, pharynx; sc, remains of a secretory cell; sv, secretory vesicle." httpUri="https://zenodo.org/record/2614758/files/figure.png" pageId="3" pageNumber="386">Fig. 5</figureCitation>
A–C). This observation was corroborated by transmission electron microscopy that revealed the glandular nature of the tissue. The most unusual feature is the cuticle overlying the frontal gland, which is made of two discrete layers of modified cuticle, both being very different from normal head cuticle (see 
<figureCitation id="1331A18F162DB963FA8DD36FFA9ACAD3" box="[1281,1370,1914,1939]" captionStart="FIGURE 5" captionStartId="8.[151,250,1454,1476]" captionTargetBox="[152,1436,745,1441]" captionTargetId="figure@8.[151,1436,745,1441]" captionTargetPageId="8" captionText="FIGURE 5. Tonsuritermes tucki sp. nov., structure of the frontal gland in worker type 1, A—Sagittal section of the head of Tonsuritermes tucki sp. nov. worker. The frontal gland is stretched in-between the two arrows. B—Transition between the head cuticle and highly modified glandular cuticle overlying the secretory cells. C—Remains of secretory cell ultrastructures. Abbreviations: br, brain (supraoesophageal ganglium); cl, clypeus; gc1, inner glandular cuticle; gc2, outer glandular cuticle; hc, head cuticle; hp, hypopharynx; lb, labrum; m, mitochondria; mm, mandibular muscles; n, nucleus; ph, pharynx; sc, remains of a secretory cell; sv, secretory vesicle." httpUri="https://zenodo.org/record/2614758/files/figure.png" pageId="3" pageNumber="386">Fig. 5B</figureCitation>
). The two-layered glandular cuticle is a unique character not shared by any other termite gland studied so far, just slightly reminding the two cuticles occurring in presoldiers of 
<taxonomicName id="4C0AC689162DB964FC95D3D7FF1CCDF0" authority="(Hagen, 1858) (Sobotnik et al. 2004)" authorityName="Sobotnik et al." authorityYear="2004" baseAuthorityName="Hagen" baseAuthorityYear="1858" class="Insecta" family="Rhinotermitidae" genus="Prorhinotermes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="4" lastPageNumber="387" order="Blattodea" pageId="3" pageNumber="386" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B97E6118162DB963FC95D3D7FBEACA9B" box="[793,1066,1986,2011]" italics="true" pageId="3" pageNumber="386">Prorhinotermes simplex</emphasis>
(
<bibRefCitation id="EF9BC0FB162DB963FBB0D3D6FB12CA9B" author="Hagen, H. A." box="[1084,1234,1986,2011]" pageId="3" pageNumber="386" pagination="4 - 342" refId="ref5424" refString="Hagen, H. A. (1858) Monographie der Termiten. Linnaea Entomologica, 12, i-iii + 4 - 342 + 459." type="journal article" year="1858">Hagen, 1858</bibRefCitation>
) (
<bibRefCitation id="EF9BC0FB162DB964FB61D3D7FF14CDF0" author="Sobotnik, J. &amp; Weyda, F. &amp; Hanus, R. &amp; Kyjakova, P. &amp; Doubsky, J." lastPageId="4" lastPageNumber="387" pageId="3" pageNumber="386" pagination="153 - 163" refId="ref6279" refString="Sobotnik, J., Weyda, F., Hanus, R., Kyjakova, P. &amp; Doubsky, J. (2004) Ultrastructure of the frontal gland in Prorhinotermes simplex (Isoptera: Rhinotermitidae) and quantity of the defensive substance. European Journal of Entomology, 101, 153 - 163. https: // doi. org / 10.14411 / eje. 2004.020" type="journal article" year="2004">
Šobotník 
<emphasis id="B97E6118162DB963FAECD3D1FA55CA9B" box="[1376,1429,1986,2011]" italics="true" pageId="3" pageNumber="386">et al</emphasis>
. 2004
</bibRefCitation>
)
</taxonomicName>
. The glandular cells were severely damaged by the 80% alcohol, and the specialised secretory organelles (e.g. microvilli, endoplasmic reticulum or Golgi apparatus) could not be observed. We also cannot confirm the existence of the second cell layer and the envelope cells observed in 
<taxonomicName id="4C0AC689162AB964FC6ED4F5FA58CDB8" authority="(Sobotnik et al. 2010 a)" authorityName="Sobotnik" authorityYear="2010" baseAuthorityName="Sobotnik" baseAuthorityYear="2010" box="[994,1432,223,248]" class="Insecta" family="Termitidae" genus="Aparatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="B97E6118162AB964FC6ED4F5FBB3CDB7" box="[994,1139,224,247]" italics="true" pageId="4" pageNumber="387">Aparatermes</emphasis>
(
<bibRefCitation id="EF9BC0FB162AB964FB0BD4CAFA51CDB8" author="Sobotnik, J. &amp; Sillam-Dusses, D. &amp; Weyda, F. &amp; Dejean, A. &amp; Roisin, Y. &amp; Hanus, R. &amp; Bourguignon, T." box="[1159,1425,223,248]" pageId="4" pageNumber="387" pagination="495 - 503" refId="ref6206" refString="Sobotnik, J., Sillam-Dusses, D., Weyda, F., Dejean, A., Roisin, Y., Hanus, R. &amp; Bourguignon, T. (2010 a) The frontal gland in workers of Neotropical soldierless termites. Naturwissenschaften, 97, 495 - 503. https: // doi. org / 10.1007 / s 00114 - 010 - 0664 - 0" type="journal article" year="2010">
Šobotník 
<emphasis id="B97E6118162AB964FB70D4F4FAF3CDB8" box="[1276,1331,223,248]" italics="true" pageId="4" pageNumber="387">et al</emphasis>
. 2010a
</bibRefCitation>
)
</taxonomicName>
. However, locally the cell remanent contained electron-lucent secretory vesicles and abundant mitochondria. We hope to acquire the living material in the future, allowing us to describe the ultrastructure of this peculiar secretory organ in detail.
</paragraph>
</subSubSection>
<subSubSection id="C310EE81162AB964FF4BD57AFD88CCB4" pageId="4" pageNumber="387" type="etymology">
<paragraph id="8BB5BD0A162AB964FF4BD57AFD88CCB4" blockId="4.[151,1437,151,536]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD57AFE8ECCC8" bold="true" box="[199,334,367,392]" pageId="4" pageNumber="387">Etymology.</emphasis>
From Latin 
<emphasis id="B97E6118162AB964FE54D564FDEECCC8" box="[472,558,369,392]" italics="true" pageId="4" pageNumber="387">tonsura</emphasis>
(“a clipping, trimming”). “Tonsure” is named after Franciscan monks’ haircut, which the fontanelle resembles, particularly in the workers. The idea for the name came from a note written by Filippo Silvestri (MZUSP 1199, 
<date id="FFB49BCA162AB964FD87D5A2FDB5CC90" box="[523,629,439,464]" pageId="4" pageNumber="387" value="1909-01-03">03.i.1909</date>
) for a sample of this genus he examined where he indicated a possible name for the species: &quot;magnotonsura&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C310EE81162AB964FF4BD5EAFDE1CFE8" pageId="4" pageNumber="387" type="distribution">
<paragraph id="8BB5BD0A162AB964FF4BD5EAFB14CF58" blockId="4.[151,1437,151,536]" box="[199,1236,511,536]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD5EAFEA0CF58" bold="true" box="[199,352,511,536]" pageId="4" pageNumber="387">Distribution.</emphasis>
Neotropical region: 
<collectingCountry id="F31DFD9A162AB964FDC5D5EAFD50CF58" box="[585,656,511,536]" name="Brazil" pageId="4" pageNumber="387">Brazil</collectingCountry>
, 
<collectingCountry id="F31DFD9A162AB964FD17D5EAFCCCCF58" box="[667,780,511,536]" name="Colombia" pageId="4" pageNumber="387">Colombia</collectingCountry>
, 
<collectingCountry id="F31DFD9A162AB964FC9BD5EAFC00CF58" box="[791,960,511,536]" name="French Guiana" pageId="4" pageNumber="387">French Guiana</collectingCountry>
, 
<collectingCountry id="F31DFD9A162AB964FC47D615FBF4CF58" box="[971,1076,512,536]" name="Paraguay" pageId="4" pageNumber="387">Paraguay</collectingCountry>
, 
<collectingCountry id="F31DFD9A162AB964FBCCD615FBB3CF57" box="[1088,1139,512,535]" name="Peru" pageId="4" pageNumber="387">Peru</collectingCountry>
(
<figureCitation id="1331A18F162AB964FB0ED5EAFB07CF58" box="[1154,1223,511,536]" captionStart="FIGURE 6" captionStartId="9.[151,250,1536,1558]" captionTargetBox="[151,1434,194,1512]" captionTargetId="figure@9.[150,1436,193,1515]" captionTargetPageId="9" captionText="FIGURE 6. Distribution map of the species of Tonsuritermes gen. nov." httpUri="https://zenodo.org/record/2614760/files/figure.png" pageId="4" pageNumber="387">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BB5BD0A162AB964FF1BD679FD3DCFC4" blockId="4.[151,765,620,680]" box="[151,765,620,645]" pageId="4" pageNumber="387">
<taxonomicName id="4C0AC689162AB964FF1BD679FD5DCFC5" authority="Cancello &amp; Constantini" authorityName="Constantini &amp; Carrijo &amp; Palma-Onetto &amp; Scheffrahn &amp; Carnohan &amp; Šobotník &amp; Cancello" authorityYear="2018" box="[151,669,620,645]" class="Insecta" family="Termitidae" genus="Tonsuritermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="tucki" status="sp. nov.">
<emphasis id="B97E6118162AB964FF1BD679FD5DCFC5" bold="true" box="[151,669,620,645]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF1BD679FEBDCFC5" bold="true" box="[151,381,620,645]" italics="true" pageId="4" pageNumber="387">Tonsuritermes tucki</emphasis>
Cancello &amp; Constantini
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DDC63162AB964FD28D679FD3DCFC4" box="[676,765,620,645]" pageId="4" pageNumber="387" rank="species">
<emphasis id="B97E6118162AB964FD28D679FD37CFC5" bold="true" box="[676,759,620,645]" pageId="4" pageNumber="387">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB5BD0A162AB964FF1BD69AFDE1CFE8" blockId="4.[151,765,620,680]" box="[151,545,655,680]" pageId="4" pageNumber="387">
(
<figureCitation id="1331A18F162AB964FF2CD69AFF28CFE8" box="[160,232,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1211,1233]" captionTargetBox="[151,1436,665,1146]" captionTargetId="figure@3.[151,1436,665,1146]" captionTargetPageId="3" captionText="FIGURE 1. Tonsuritermes tucki sp. nov., imago head and thorax. A—lateral view, B—dorsal view. Arrows point to the frontal marks. Specimen from lot MZUSP6480." httpUri="https://zenodo.org/record/2614750/files/figure.png" pageId="4" pageNumber="387">Figs.1</figureCitation>
; 
<figureCitation id="1331A18F162AB964FF78D685FEC2CFE8" box="[244,258,656,680]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="4" pageNumber="387">2</figureCitation>
A–D, 3A–B, D–I, 4, 5, 6)
</paragraph>
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<paragraph id="8BB5BD0A162AB964FF1BD6C2FB06CFB0" blockId="4.[151,1437,727,2013]" box="[151,1222,727,752]" pageId="4" pageNumber="387">
<materialsCitation id="3B62B757162AB964FF1BD6C2FB06CFB0" ID-GBIF-Occurrence="2116190228" box="[151,1222,727,752]" collectionCode="MZUSP" pageId="4" pageNumber="387" specimenCode="MZUSP 6480" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B103A8162AB964FF1BD6C2FEC6CFB0" box="[151,262,727,752]" pageId="4" pageNumber="387" type="holotype">Holotype</typeStatus>
.
</emphasis>
Worker type 1 from lot 
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(in a separate vial with the remaining sample).
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<emphasis id="B97E6118162AB964FF4BD6E9FEA8CE55" bold="true" box="[199,360,764,789]" pageId="4" pageNumber="387">Type-locality.</emphasis>
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. 
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<emphasis id="B97E6118162AB964FF4BD70AFE4BCE78" bold="true" box="[199,395,799,824]" pageId="4" pageNumber="387">Type-repository.</emphasis>
MZUSP
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<paragraph id="8BB5BD0A162AB964FF4BD751FE77C880" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
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<typeStatus id="54B103A8162AB964FF4BD751FE81CE1D" box="[199,321,836,861]" pageId="4" pageNumber="387" type="paratype">Paratypes</typeStatus>
.
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<collectingCountry id="F31DFD9A162AB964FEDDD751FE78CE1C" box="[337,440,836,860]" name="Brazil" pageId="4" pageNumber="387">BRASIL</collectingCountry>
. 
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: 
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, lat 
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coll., 
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: Caldas Novas, 
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de
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coll., UFG 1740, 1741
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<collectorName id="26FFD8DC162AB964FB9CD798FB93CEE4" box="[1040,1107,909,932]" pageId="4" pageNumber="387">Poços</collectorName>
de
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<collectingMunicipality id="6BD12770162AB964FB68D798FA5CCEE4" box="[1252,1436,908,933]" pageId="4" pageNumber="387">Morro do Ferro</collectingMunicipality>
(
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), lat 
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, 
<date id="FFB49BCA162AB964FDFED7BAFD3BCE88" box="[626,763,943,968]" pageId="4" pageNumber="387" value="1967-09-18">
<collectingDate id="EFF06222162AB964FDFED7BAFD3BCE88" box="[626,763,943,968]" pageId="4" pageNumber="387" value="1967-09-18">18.ix. 1967</collectingDate>
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, 
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coll., 
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; 
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: 
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coll., 
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<collectingDate id="EFF06222162AB964FC5DD7E2FB8BC950" box="[977,1099,1015,1040]" pageId="4" pageNumber="387" value="2016-11-24">24.xi.2016</collectingDate>
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<collectorName id="26FFD8DC162AB964FBF4D7E2FB39C950" box="[1144,1273,1015,1040]" pageId="4" pageNumber="387">Constantini</collectorName>
coll., 
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; 
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<collectingRegion id="49CE73E8162AB964FF62D009FE9FC975" box="[238,351,1052,1077]" country="Brazil" name="Rondonia" pageId="4" pageNumber="387">Rondônia</collectingRegion>
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<date id="FFB49BCA162AB964FC3ED009FBEEC975" box="[946,1070,1052,1077]" pageId="4" pageNumber="387" value="2010-03-09">
<collectingDate id="EFF06222162AB964FC3ED009FBEEC975" box="[946,1070,1052,1077]" pageId="4" pageNumber="387" value="2010-03-09">09.iii.2010</collectingDate>
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Carrijo &amp; 
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Carrijo &amp; 
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; Nova Mutum 
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sample)
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coll., 
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, (imago, workers)
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; 
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: 
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coll., 
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coll.
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<collectingRegion id="49CE73E8162AB964FEF7D14AFE31C838" box="[379,497,1375,1400]" country="Paraguay" name="Cordillera" pageId="4" pageNumber="387">Cordillera</collectingRegion>
: 
<location id="8ED5EBD1162AB964FE71D14AFD43C838" LSID="urn:lsid:plazi:treatment:03A30C1C162FB964FF1BD2B5FDADCA9D:8ED5EBD1162AB964FE71D14AFD43C838" box="[509,643,1375,1400]" country="Paraguay" longitude="-56.8667" name="Vallenzuela" pageId="4" pageNumber="387" stateProvince="Cordillera">Vallenzuela</location>
, lat 25.5833, long 
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, 
<date id="FFB49BCA162AB964FC50D14AFB88C838" box="[988,1096,1375,1400]" pageId="4" pageNumber="387" value="1992-01-04">
<collectingDate id="EFF06222162AB964FC50D14AFB88C838" box="[988,1096,1375,1400]" pageId="4" pageNumber="387" value="1992-01-04">04.i.1992</collectingDate>
</date>
, L Cabello &amp; 
<collectorName id="26FFD8DC162AB964FB62D175FA9CC838" box="[1262,1372,1375,1400]" pageId="4" pageNumber="387">B Barrios</collectorName>
coll., 
<specimenCode id="DBAC1571162AB964FF1BD191FE86C8DD" box="[151,326,1412,1437]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="387">MZUSP 10855</specimenCode>
</materialsCitation>
. 
<materialsCitation id="3B62B757162AB964FEDCD190FE73C880" ID-GBIF-Occurrence="2116188962" collectingDate="2015-09-09" collectionCode="UF" collectorName="L Carnohan" country="Peru" latitude="-13.137" location="Research Lodge" longLatPrecision="1" longitude="-69.612" municipality="Tambopata" pageId="4" pageNumber="387" specimenCode="PU1104" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
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. 
<collectingRegion id="49CE73E8162AB964FE2AD191FD91C8DD" box="[422,593,1412,1437]" country="Peru" name="Madre de Dios" pageId="4" pageNumber="387">Madre de Dios</collectingRegion>
: 
<collectingMunicipality id="6BD12770162AB964FDD0D191FD1BC8DD" box="[604,731,1412,1437]" pageId="4" pageNumber="387">Tambopata</collectingMunicipality>
, 
<location id="8ED5EBD1162AB964FD6BD191FC5FC8DD" LSID="urn:lsid:plazi:treatment:03A30C1C162FB964FF1BD2B5FDADCA9D:8ED5EBD1162AB964FD6BD191FC5FC8DD" box="[743,927,1412,1437]" country="Peru" latitude="-13.137" longLatPrecision="1" longitude="-69.612" municipality="Tambopata" name="Research Lodge" pageId="4" pageNumber="387" stateProvince="Madre de Dios">Research Lodge</location>
, lat 
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, 
<date id="FFB49BCA162AB964FA91D191FA57C8DD" box="[1309,1431,1412,1437]" pageId="4" pageNumber="387" value="2015-09-09">
<collectingDate id="EFF06222162AB964FA91D191FA57C8DD" box="[1309,1431,1412,1437]" pageId="4" pageNumber="387" value="2015-09-09">09.ix.2015</collectingDate>
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, 
<collectorName id="26FFD8DC162AB964FF1BD1BCFEDFC880" box="[151,287,1447,1472]" pageId="4" pageNumber="387">L Carnohan</collectorName>
, 
<collectionCode id="ED1B25CF162AB964FEA6D1BCFE8EC880" box="[298,334,1449,1472]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="4" pageNumber="387" type="Museum">UF</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id="C310EE81162AB964FF4BD1D9FD75CBFD" pageId="4" pageNumber="387" type="diagnosis">
<paragraph id="8BB5BD0A162AB964FF4BD1D9FC8BCB48" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD1D9FE81C8A5" bold="true" box="[199,321,1484,1509]" pageId="4" pageNumber="387">Diagnosis.</emphasis>
Dimorphic worker with fontanelle of two sizes; head capsule with two lengths of well marked bristles and bristles on tergites uniformly oriented backwards.
</paragraph>
<paragraph id="8BB5BD0A162AB964FF4BD200FD8ECB6D" blockId="4.[151,1437,727,2013]" box="[199,590,1556,1581]" pageId="4" pageNumber="387">Imago. As described for the genus.</paragraph>
<paragraph id="8BB5BD0A162AB964FF4BD222FD75CBFD" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
Worker. Dimorphic (
<figureCitation id="1331A18F162AB964FE3FD222FE37CB10" box="[435,503,1591,1616]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[151,1436,205,1884]" captionTargetId="figure@5.[151,1436,205,1884]" captionTargetPageId="5" captionText="FIGURE 2. Tonsuritermes, workers head and thorax. A—worker type 1 of T. tucki, sp. nov., dorsal view; B—same as A in lateral view; C—worker type 2 of T. tucki, sp. nov., dorsal view; D—same as C in lateral view. E—T. mathewsi sp. nov., dorsal view; F—same as E in lateral view. Specimens from lots MZUSP6480 (A–D) and MZUSP12268 (E–F)." httpUri="https://zenodo.org/record/2614752/files/figure.png" pageId="4" pageNumber="387">Fig. 2</figureCitation>
A–D). Head capsule with two lengths of well marked bristles. Bristles on tergites uniformly oriented backwards. Postclypeus moderately inflated. Protibia inflated. Worker (
<typeStatus id="54B103A8162AB964FB46D249FB3CCB34" box="[1226,1276,1628,1652]" pageId="4" pageNumber="387">type</typeStatus>
1, W1) with fontanelle occupying 3/4 of the cephalic capsule in dorsal view. Worker 2 (
<typeStatus id="54B103A8162AB964FC6AD295FBD8CBD8" box="[998,1048,1664,1688]" pageId="4" pageNumber="387">type</typeStatus>
2, W2) with fontanelle occupying 1/4 to 1/2 of the cephalic capsule in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C310EE81162AB964FF4BD2D2FE9ACA0D" pageId="4" pageNumber="387" type="etymology">
<paragraph id="8BB5BD0A162AB964FF4BD2D2FF1DCA45" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD2D2FE8ECBA0" bold="true" box="[199,334,1735,1760]" pageId="4" pageNumber="387">Etymology.</emphasis>
In reference to the character Friar Tuck, the supposed Franciscan monk of the legend of Robin Hood.
</paragraph>
<paragraph id="8BB5BD0A162AB964FF4BD31AFE9ACA0D" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD31AFE4CCA68" bold="true" box="[199,396,1807,1832]" pageId="4" pageNumber="387">Biological notes.</emphasis>
Collected foraging in galleries in the ground, at base of trees, among litter and sticks and under rotting log.
</paragraph>
</subSubSection>
<subSubSection id="C310EE81162AB964FF4BD342FDADCA9D" pageId="4" pageNumber="387" type="discussion">
<paragraph id="8BB5BD0A162AB964FF4BD342FDADCA9D" blockId="4.[151,1437,727,2013]" pageId="4" pageNumber="387">
<emphasis id="B97E6118162AB964FF4BD342FE8FCA30" bold="true" box="[199,335,1879,1904]" pageId="4" pageNumber="387">Comments.</emphasis>
We described as two 
<typeStatus id="54B103A8162AB964FDC6D34CFD45CA31" box="[586,645,1881,1905]" pageId="4" pageNumber="387">types</typeStatus>
of workers because there are two sizes of fontanelle. To know if it is the same worker with the fontanelle changing over time, it would be necessary to do a developmental analysis, and this is out of the scope of the present paper. For taxonomic purposes, it is important to know that two 
<typeStatus id="54B103A8162AB964FACCD3B4FABBCAF9" box="[1344,1403,1953,1977]" pageId="4" pageNumber="387">types</typeStatus>
of workers can be found in the same sample.
</paragraph>
</subSubSection>
</treatment>
</document>