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<mods:title id="88943D275B1C1EA3AC07D75C3F63029D">The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae)</mods:title>
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<treatment id="039B87D1FFEFB115169BFF5BC49AFEC1" ID-DOI="http://doi.org/10.5281/zenodo.5987855" ID-GBIF-Taxon="142616420" ID-Zenodo-Dep="5987855" LSID="urn:lsid:plazi:treatment:039B87D1FFEFB115169BFF5BC49AFEC1" httpUri="http://treatment.plazi.org/id/039B87D1FFEFB115169BFF5BC49AFEC1" lastPageId="15" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="C328654CFFEFB111169BFF5BC556FF17" box="[151,527,151,178]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B8D36C7FFEFB111169BFF5BC556FF17" blockId="11.[151,527,151,178]" box="[151,527,151,178]" pageId="11" pageNumber="12">
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Genus
<taxonomicName id="4C324D44FFEFB11116E1FF54C556FF17" ID-CoL="32TD" ID-ENA="593438" authority="Audouin, 1826" authorityName="Audouin" authorityYear="1826" box="[237,527,151,178]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB11116E1FF54C608FF14" bold="true" box="[237,337,151,177]" italics="true" pageId="11" pageNumber="12">Ariadna</emphasis>
<bibRefCitation id="EFA34B36FFEFB1111755FF5BC556FF17" author="Audouin, V." box="[345,527,152,178]" journalOrPublisher="Description de l'Egypte ... &quot;. Histoire Naturelle, 1" pageId="11" pageNumber="12" pagination="1 - 339" part="4" refId="ref87124" refString="Audouin, V. (1826) Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in &quot; Description de l'Egypte ... &quot;. Histoire Naturelle, 1 (4), 1 - 339. [arachnids, pp. 99 - 186]" title="Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in" type="journal article" year="1826">Audouin, 1826</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8B8D36C7FFEFB111169BFF1DC2D4FF51" blockId="11.[151,1436,222,398]" box="[151,1421,222,244]" pageId="11" pageNumber="12">
<taxonomicName id="4C324D44FFEFB111169BFF1DC6E7FF51" ID-CoL="32TD" authority="Audouin, 1826: 109" authorityName="Audouin" authorityPageNumber="109" authorityYear="1826" box="[151,446,222,244]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB111169BFF1DC7B3FF51" box="[151,234,222,244]" italics="true" pageId="11" pageNumber="12">Ariadna</emphasis>
<bibRefCitation id="EFA34B36FFEFB11116FCFF1DC6D2FF51" author="Audouin, V." box="[240,395,222,244]" journalOrPublisher="Description de l'Egypte ... &quot;. Histoire Naturelle, 1" pageId="11" pageNumber="12" pagination="1 - 339" part="4" refId="ref87124" refString="Audouin, V. (1826) Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in &quot; Description de l'Egypte ... &quot;. Histoire Naturelle, 1 (4), 1 - 339. [arachnids, pp. 99 - 186]" title="Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in" type="journal article" year="1826">Audouin, 1826</bibRefCitation>
: 109
</taxonomicName>
. Type species by monotypy,
<taxonomicName id="4C324D44FFEFB11114EDFF1DC366FF51" ID-CoL="GLLN" authority="Audouin, 1826" authorityName="Audouin" authorityYear="1826" box="[737,1087,222,244]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="B946EAD5FFEFB11114EDFF1DC4FAFF51" box="[737,931,222,244]" italics="true" pageId="11" pageNumber="12">Ariadna insidiatrix</emphasis>
<bibRefCitation id="EFA34B36FFEFB11115A5FF1DC366FF51" author="Audouin, V." box="[937,1087,222,244]" journalOrPublisher="Description de l'Egypte ... &quot;. Histoire Naturelle, 1" pageId="11" pageNumber="12" pagination="1 - 339" part="4" refId="ref87124" refString="Audouin, V. (1826) Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in &quot; Description de l'Egypte ... &quot;. Histoire Naturelle, 1 (4), 1 - 339. [arachnids, pp. 99 - 186]" title="Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie publiees ... in" type="journal article" year="1826">Audouin, 1826</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="EFA34B36FFEFB1111251FF1DC2D0FF51" author="World Spider Catalog" box="[1117,1417,222,244]" journalOrPublisher="Avaliable from: http: // wsc. nmbe. ch / (accessed on" pageId="11" pageNumber="12" pagination="2018" part="16" refId="ref91516" refString="World Spider Catalog (2018) World Spider Catalog. Natural History Museum Bern, version 19.0. Avaliable from: http: // wsc. nmbe. ch / (accessed on 16 January 2018)" title="World Spider Catalog. Natural History Museum Bern, version 19.0" type="journal article" year="2018">World Spider Catalog (2018)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8D36C7FFEFB111169BFF3EC65DFE97" blockId="11.[151,1436,222,398]" pageId="11" pageNumber="12">
<taxonomicName id="4C324D44FFEFB111169BFF3EC6F8FEB6" authority="Hentz, 1842: 225" authorityName="Hentz" authorityPageNumber="225" authorityYear="1842" box="[151,417,253,275]" class="Arachnida" family="Curculionidae" genus="Pylarus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB111169BFF3EC7BFFEB6" box="[151,230,253,275]" italics="true" pageId="11" pageNumber="12">Pylarus</emphasis>
<bibRefCitation id="EFA34B36FFEFB11116E1FF3DC637FEB6" author="Hentz, N. M." box="[237,366,253,275]" journalOrPublisher="Boston Journal of Natural History" pageId="11" pageNumber="12" pagination="54 - 57" part="4" publicationUrl="https://doi.org/10.5962/bhl.part.16996" refId="ref88990" refString="Hentz, N. M. (1842) Descriptions and figures of the araneides of the United States. Boston Journal of Natural History, 4, 54 - 57 + 223 - 231. https: // doi. org / 10.5962 / bhl. part. 16996" title="Descriptions and figures of the araneides of the United States" type="journal article" year="1842">Hentz, 1842</bibRefCitation>
: 225
</taxonomicName>
. Type species,
<taxonomicName id="4C324D44FFEFB1111430FF3EC407FEB6" authority="Hentz, 1842" authorityName="Hentz" authorityYear="1842" box="[572,862,253,275]" class="Arachnida" family="Curculionidae" genus="Pylarus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B946EAD5FFEFB1111430FF3EC582FEB6" box="[572,731,253,275]" italics="true" pageId="11" pageNumber="12">Pylarus bicolor</emphasis>
<bibRefCitation id="EFA34B36FFEFB11114EEFF3DC407FEB6" author="Hentz, N. M." box="[738,862,253,275]" journalOrPublisher="Boston Journal of Natural History" pageId="11" pageNumber="12" pagination="54 - 57" part="4" publicationUrl="https://doi.org/10.5962/bhl.part.16996" refId="ref88990" refString="Hentz, N. M. (1842) Descriptions and figures of the araneides of the United States. Boston Journal of Natural History, 4, 54 - 57 + 223 - 231. https: // doi. org / 10.5962 / bhl. part. 16996" title="Descriptions and figures of the araneides of the United States" type="journal article" year="1842">Hentz, 1842</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="EFA34B36FFEFB1111570FF3EC367FEB6" author="Beatty, J. A." box="[892,1086,253,275]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="11" pageNumber="12" pagination="433 - 518" part="139" refId="ref87290" refString="Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518." title="The spider genus Ariadna in the Americas (Araneae, Dysderidae)" type="journal article" year="1970">Beatty (1970: 458)</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFEFB1111250FF3EC3A5FEB6" author="Emerton, J. H." box="[1116,1276,253,275]" editor="Hentz, N. M." journalOrPublisher="Occasional Papers of the Boston Society of Natural History" pageId="11" pageNumber="12" pagination="15 - 164" part="2" refId="ref88270" refString="Emerton, J. H. (1875) Notes and descriptions. In: Hentz, N. M. (1842) Descriptions and figures of the Araneides of the United States. Occasional Papers of the Boston Society of Natural History, 2, 15 - 164." title="Notes and descriptions" type="journal article" volumeTitle="(1842) Descriptions and figures of the Araneides of the United States" year="1875">Emerton (1875</bibRefCitation>
: 20; 1876: 131, Syn.).
</paragraph>
<paragraph id="8B8D36C7FFEFB111169BFEF9C6A2FECA" blockId="11.[151,1436,222,398]" pageId="11" pageNumber="12">
<emphasis id="B946EAD5FFEFB111169BFEF9C650FEF5" box="[151,265,314,336]" italics="true" pageId="11" pageNumber="12">Macedonia</emphasis>
<bibRefCitation id="EFA34B36FFEFB1111717FEF8C6FBFEF5" author="Hogg, H. R." box="[283,418,314,336]" journalOrPublisher="Proceedings of the Royal Society of Victoria, New Series" pageId="11" pageNumber="12" pagination="68 - 123" part="13" refId="ref89067" refString="Hogg, H. R. (1900) A contribution to our knowledge of the spiders of Victoria: including some new species and genera. Proceedings of the Royal Society of Victoria, New Series, 13, 68 - 123." title="A contribution to our knowledge of the spiders of Victoria: including some new species and genera" type="journal article" year="1900">Hogg, 1900</bibRefCitation>
: 85. Type species by monotypy,
<emphasis id="B946EAD5FFEFB111153FFEF9C348FEF5" box="[819,1041,314,336]" italics="true" pageId="11" pageNumber="12">Macedonia burchelli</emphasis>
<bibRefCitation id="EFA34B36FFEFB1111228FEF8C3FFFEF5" author="Hogg, H. R." box="[1060,1190,314,336]" journalOrPublisher="Proceedings of the Royal Society of Victoria, New Series" pageId="11" pageNumber="12" pagination="68 - 123" part="13" refId="ref89067" refString="Hogg, H. R. (1900) A contribution to our knowledge of the spiders of Victoria: including some new species and genera. Proceedings of the Royal Society of Victoria, New Series, 13, 68 - 123." title="A contribution to our knowledge of the spiders of Victoria: including some new species and genera" type="journal article" year="1900">Hogg, 1900</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFEFB11112C9FEF9C7A0FECA" author="Simon, E." journalOrPublisher="Jena, 1" pageId="11" pageNumber="12" pagination="359 - 446" part="12" refId="ref91108" refString="Simon, E. (1908) Araneae. 1 re partie. In: Michaelsen &amp; Hartmeyer (Eds.), Die Fauna Sudwest-Australiens. Jena, 1 (12), 359 - 446." title="Araneae. 1 re partie" type="journal article" volumeTitle="Michaelsen &amp; Hartmeyer (Eds.), Die Fauna Sudwest-Australiens" year="1908">Simon (1908: 380, Syn.)</bibRefCitation>
.—
<bibRefCitation id="EFA34B36FFEFB111171BFE9AC6AEFECA" author="Rainbow, W. J." box="[279,503,345,367]" journalOrPublisher="Records of the Australian Museum" pageId="11" pageNumber="12" pagination="107 - 319" part="9" publicationUrl="https://doi.org/10.3853/j.0067-1975.9.1911.928" refId="ref90434" refString="Rainbow, W. J. (1911) A census of Australian Araneidae. Records of the Australian Museum, 9, 107 - 319. https: // doi. org / 10.3853 / j. 0067 - 1975.9.1911.928" title="A census of Australian Araneidae" type="journal article" year="1911">Rainbow (1911: 136)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8D36C7FFEFB111169BFEBBC2C8FE2B" blockId="11.[151,1436,222,398]" box="[151,1425,376,398]" pageId="11" pageNumber="12">
<taxonomicName id="4C324D44FFEFB111169BFEBBC68BFE2B" authority="Purcell, 1904: 165" authorityName="Purcell" authorityPageNumber="165" authorityYear="1904" box="[151,466,376,398]" class="Arachnida" family="Segestriidae" genus="Segestriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB111169BFEBBC656FE2B" box="[151,271,376,398]" italics="true" pageId="11" pageNumber="12">Segestriella</emphasis>
<bibRefCitation id="EFA34B36FFEFB1111718FEBBC6C6FE2B" author="Purcell, W. F." box="[276,415,376,398]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="11" pageNumber="12" pagination="115 - 173" part="15" publicationUrl="https://doi.org/10.1080/21560382.1904.9626437" refId="ref90390" refString="Purcell, W. F. (1904) Descriptions of new genera and species of South African spiders. Transactions of the South African Philosophical Society, 15, 115 - 173. https: // doi. org / 10.1080 / 21560382.1904.9626437" title="Descriptions of new genera and species of South African spiders" type="journal article" year="1904">Purcell, 1904</bibRefCitation>
: 165
</taxonomicName>
. Type species by monotypy,
<taxonomicName id="4C324D44FFEFB11114F9FEBBC329FE2B" authority="Purcell, 1904" authorityName="Purcell" authorityYear="1904" box="[757,1136,376,398]" class="Arachnida" family="Segestriidae" genus="Segestriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gryllotalpa">
<emphasis id="B946EAD5FFEFB11114F9FEBBC4BDFE2B" box="[757,996,376,398]" italics="true" pageId="11" pageNumber="12">Segestriella gryllotalpa</emphasis>
<bibRefCitation id="EFA34B36FFEFB11115E6FEBBC329FE2B" author="Purcell, W. F." box="[1002,1136,376,398]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="11" pageNumber="12" pagination="115 - 173" part="15" publicationUrl="https://doi.org/10.1080/21560382.1904.9626437" refId="ref90390" refString="Purcell, W. F. (1904) Descriptions of new genera and species of South African spiders. Transactions of the South African Philosophical Society, 15, 115 - 173. https: // doi. org / 10.1080 / 21560382.1904.9626437" title="Descriptions of new genera and species of South African spiders" type="journal article" year="1904">Purcell, 1904</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="EFA34B36FFEFB1111282FEBBC2D4FE2B" author="Beatty, J. A." box="[1166,1421,376,398]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="11" pageNumber="12" pagination="433 - 518" part="139" refId="ref87290" refString="Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518." title="The spider genus Ariadna in the Americas (Araneae, Dysderidae)" type="journal article" year="1970">Beatty (1970: 454, Syn.)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328654CFFEFB111169BFE79C4C0FD56" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8B8D36C7FFEFB111169BFE79C4C0FD56" blockId="11.[151,1436,442,755]" pageId="11" pageNumber="12">
<emphasis id="B946EAD5FFEFB111169BFE79C648FE76" bold="true" box="[151,273,442,467]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
The American
<taxonomicName id="4C324D44FFEFB11117C7FE79C57EFE76" authorityName="Audouin" authorityYear="1826" box="[459,551,442,467]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB11117C7FE79C57EFE76" box="[459,551,442,467]" italics="true" pageId="11" pageNumber="12">Ariadna</emphasis>
</taxonomicName>
differ from the other
<taxonomicName id="4C324D44FFEFB1111521FE79C4EFFE76" authorityName="Simon" authorityYear="1893" box="[813,950,442,467]" class="Arachnida" family="Segestriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Segestriidae</taxonomicName>
genera by the following characters: from
<taxonomicName id="4C324D44FFEFB111169BFE1CC7A6FE53" authorityName="Latreille" authorityYear="1804" box="[151,255,479,502]" class="Arachnida" family="Segestriidae" genus="Segestria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB111169BFE1CC7A6FE53" box="[151,255,479,502]" italics="true" pageId="11" pageNumber="12">Segestria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D44FFEFB1111738FE1DC6F6FE52" authorityName="Hogg" authorityYear="1900" box="[308,431,478,503]" class="Arachnida" family="Segestriidae" genus="Gippsicola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB1111738FE1DC6F6FE52" box="[308,431,478,503]" italics="true" pageId="11" pageNumber="12">Gippsicola</emphasis>
</taxonomicName>
by the labium narrowed in its distal portion (
<figureCitation id="13092A42FFEFB11115A9FE1DC4A3FE52" box="[933,1018,478,503]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="11" pageNumber="12">Fig. 3E</figureCitation>
), labrum well-developed, longer than the labium, and without dorsal median setae (
<figureCitation id="13092A42FFEFB1111499FDC1C5B2FDBE" box="[661,747,514,539]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="11" pageNumber="12">Figs 3F</figureCitation>
,
<figureCitation id="13092A42FFEFB11114FBFDC0C471FDBE" box="[759,808,515,539]" captionStart="FIGURE 18" captionStartId="30.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@30.[151,1436,193,1803]" captionTargetPageId="30" captionText="FIGURE 18. Ariadna crassipalpa (Blackwall, 1863). Male from Porto Alegre (MCN 3577) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E), arrowhead = curved macroseta. Female from Canela (MCN 0551) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207679/files/figure.png" pageId="11" pageNumber="12">18B</figureCitation>
,
<figureCitation id="13092A42FFEFB1111538FDC0C43FFDBE" box="[820,870,515,539]" captionStart="FIGURE 20" captionStartId="34.[151,250,1894,1916]" captionTargetBox="[151,1436,299,1866]" captionTargetId="figure@34.[151,1436,299,1872]" captionTargetPageId="34" captionText="FIGURE 20. Ariadna mollis (Holmberg, 1876). Male from Parque Estadual do Jaraguá (IBSP 166961) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E). Female from Instituto Butantan (IBSP 72885) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207685/files/figure.png" pageId="11" pageNumber="12">20B</figureCitation>
), presence of a lateral basal transverse ridge in the chelicerae (
<figureCitation id="13092A42FFEFB1111714FDE5C62BFD9A" box="[280,370,550,575]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
;
<bibRefCitation id="EFA34B36FFEFB1111772FDE4C578FD9A" author="Lipke, E. &amp; Ramirez, M. J. &amp; Michalik, P." box="[382,545,551,575]" journalOrPublisher="Journal of Morphology" pageId="11" pageNumber="12" pagination="1238 - 1257" part="275" publicationUrl="https://doi.org/10.1002/jmor.20298" refId="ref89380" refString="Lipke, E., Ramirez, M. J., Michalik, P. (2014) Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea. Journal of Morphology, 275, 1238 - 1257. https: // doi. org / 10.1002 / jmor. 20298" title="Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
: 40, fig. 18H) posterior eyes in a straight or slightly recurved line (
<figureCitation id="13092A42FFEFB1111315FDE5C2C1FD9A" box="[1305,1432,550,575]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="11" pageNumber="12">Figs 3A‒B</figureCitation>
,
<figureCitation id="13092A42FFEFB111169BFD89C790FDC6" box="[151,201,586,611]" captionStart="FIGURE64" captionStartId="101.[151,250,1781,1803]" captionTargetBox="[151,1435,301,1760]" captionTargetId="figure@101.[151,1436,301,1760]" captionTargetPageId="101" captionText="FIGURE64. Aridna caparaon. sp.. Male from Trilha do Terreirão (IBSP 166970) (A‒F):habitus, dorsal (A) and ventral view (B); cheliceral, dorsal view (C), arrowheads = lameliform distal dorso-prolateral projections; left leg I, prolateral (D) and retrolateral view (E); left metatarsus I. ventral view (F). Female (Paratype) from Trilha do Terreirão (IBSP 38345) (G‒J): habitus, dorsal (G) and ventral view (H); right leg I, prolateral (I) and retrolateral view (J). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207775/files/figure.png" pageId="11" pageNumber="12">64C</figureCitation>
), absence of tenent setae on the legs (
<figureCitation id="13092A42FFEFB111147BFD89C599FDC7" box="[631,704,586,611]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="11" pageNumber="12">Figs 7</figureCitation>
,
<figureCitation id="13092A42FFEFB11114C2FD88C400FDC6" box="[718,857,587,611]" captionStart="FIGURE 12" captionStartId="17.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1796]" captionTargetId="figure@17.[151,1436,193,1796]" captionTargetPageId="17" captionText="FIGURE 12. Ariadna bicolor (Hentz, 1842). Male from Rockville (USNM) (A‒E): Habitus, dorsal (A) and ventral view (B); Left leg I, prolateral view (C) and retrolateral (D), arrowhead = lanceolated macroseta; Left metatarsus I, ventral view (E). Female from 2 mi W. Cedar Crk. (USNM) (F‒I): Habitus, dorsal (F) and ventral view (G); Right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207667/files/figure.png" pageId="11" pageNumber="12">12C‒E, H‒I</figureCitation>
), tarsus of the female palps with a group of five or more macrosetae (
<figureCitation id="13092A42FFEFB111177DFDADC694FD22" box="[369,461,622,647]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="11" pageNumber="12">Fig. 4F</figureCitation>
), and posterior receptaculum with tubular invaginations (
<figureCitation id="13092A42FFEFB1111275FDADC25FFD22" box="[1145,1286,622,647]" captionStart="FIGURE 10" captionStartId="13.[151,250,1897,1919]" captionTargetBox="[151,1436,841,1875]" captionTargetId="figure@13.[151,1436,841,1875]" captionTargetPageId="13" captionText="FIGURE 10. SEM images of Ariadna boliviana Simon, 1907, female from Parauapebas (IBSP 193499): anterior receptaculum, lateral (A) and frontal view (B); internal genitalia, frontal view (C); posterior wall of bursa copulatrix, lateral view, origin of the posterior receptaculum (D), arrowhead = median membranous crack; pore of a tubular invagination in the membranous cuticle of the posterior receptaculum (E); posterior receptaculum, internal view, tubular invaginations (F‒G)." httpUri="https://zenodo.org/record/1207663/files/figure.png" pageId="11" pageNumber="12">Figs 10E‒G</figureCitation>
,
<figureCitation id="13092A42FFEFB1111314FDACC213FD22" box="[1304,1354,623,647]" captionStart="FIGURE 11" captionStartId="14.[151,250,1105,1127]" captionTargetBox="[151,1436,193,1083]" captionTargetId="figure@14.[151,1436,193,1083]" captionTargetPageId="14" captionText="FIGURE 11. Ariadna maxima (Nicolet, 1849), female from Santiago (IBSP 161292), after enzime prepatarion: habitus, dorsal view, without carapaceand dorsal part of the abdomen (A); cephalothorax and anterior region of the abdomen, dorsal view (B); anterior region of the abdomen, dorsal view (C); posterior receptaculum, dorsal view, tubular invaginations (D)." httpUri="https://zenodo.org/record/1207665/files/figure.png" pageId="11" pageNumber="12">11D</figureCitation>
); from
<taxonomicName id="4C324D44FFEFB111169BFD51C646FD0E" authorityName="Chamberlin" authorityYear="1924" box="[151,287,658,683]" class="Arachnida" family="Segestriidae" genus="Citharoceps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAD5FFEFB111169BFD51C646FD0E" box="[151,287,658,683]" italics="true" pageId="11" pageNumber="12">Citharoceps</emphasis>
</taxonomicName>
by the absence of the stridulatory apparatus (
<bibRefCitation id="EFA34B36FFEFB1111516FD51C268FD0E" author="Giroti, A. M. &amp; Brescovit, A. D." box="[794,1329,658,683]" journalOrPublisher="ZooKeys" pageId="11" pageNumber="12" pagination="1 - 19" part="495" publicationUrl="https://doi.org/10.3897/zookeys.495.8950" refId="ref88701" refString="Giroti, A. M. &amp; Brescovit, A. D. (2015) Revalidation of the spider genus Citharoceps Chamberlin, 1924 (Araneae, Segestriidae). ZooKeys, 495, 1 - 19. https: // doi. org / 10.3897 / zookeys. 495.8950" title="Revalidation of the spider genus Citharoceps Chamberlin, 1924 (Araneae, Segestriidae)" type="journal article" year="2015">Giroti and Brescovit 2015: 4, figs 1A‒C, 2A‒B</bibRefCitation>
), labiumsternum junction larger than the length of the endite-sternum junction (
<figureCitation id="13092A42FFEFB11115E0FD75C315FD6A" box="[1004,1100,694,719]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="11" pageNumber="12">Figs 3E</figureCitation>
,
<figureCitation id="13092A42FFEFB1111252FD74C3D6FD6A" box="[1118,1167,695,719]" captionStart="FIGURE 18" captionStartId="30.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@30.[151,1436,193,1803]" captionTargetPageId="30" captionText="FIGURE 18. Ariadna crassipalpa (Blackwall, 1863). Male from Porto Alegre (MCN 3577) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E), arrowhead = curved macroseta. Female from Canela (MCN 0551) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207679/files/figure.png" pageId="11" pageNumber="12">18B</figureCitation>
), and females with an interpulmonary fold lacking a conspicuous median flap (
<figureCitation id="13092A42FFEFB111151DFD19C4D4FD56" box="[785,909,730,755]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="11" pageNumber="12">Fig. 9A‒B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4D664FFFEFB111169BF8B4C3E1F84C" httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1911,1933]" targetBox="[161,1425,797,1898]" targetPageId="11">
<paragraph id="8B8D36C7FFEFB111169BF8B4C3E1F84C" blockId="11.[151,1436,1911,2025]" pageId="11" pageNumber="12">
<emphasis id="B946EAD5FFEFB111169BF8B4C64AF829" bold="true" box="[151,275,1911,1933]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
SEM images of
<taxonomicName id="4C324D44FFEFB11117CCF8B4C45CF828" authority="Simon, 1907" authorityName="Simon" authorityYear="1907" box="[448,773,1911,1933]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="B946EAD5FFEFB11117CCF8B4C523F828" box="[448,634,1911,1933]" italics="true" pageId="11" pageNumber="12">Ariadna boliviana</emphasis>
Simon, 1907
</taxonomicName>
, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F).
</paragraph>
</caption>
<caption id="DF4D664FFFE8B116169BFB2AC2C4FAFE" httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1257,1279]" targetBox="[151,1435,193,1236]" targetPageId="12">
<paragraph id="8B8D36C7FFE8B116169BFB2AC2C4FAFE" blockId="12.[151,1437,1257,1371]" pageId="12" pageNumber="13">
<emphasis id="B946EAD5FFE8B116169BFB2AC64FFB5B" bold="true" box="[151,278,1257,1279]" pageId="12" pageNumber="13">FIGURE 9.</emphasis>
SEM images of
<taxonomicName id="4C324D44FFE8B11617C1FB2AC47AFB5A" authority="(Nicolet, 1849)" authorityName="Nicolet" authorityYear="1849" baseAuthorityName="Nicolet" baseAuthorityYear="1849" box="[461,803,1257,1279]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maxima">
<emphasis id="B946EAD5FFE8B11617C1FB2AC523FB5A" box="[461,634,1257,1279]" italics="true" pageId="12" pageNumber="13">Ariadna maxima</emphasis>
(Nicolet, 1849)
</taxonomicName>
, females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct.
</paragraph>
</caption>
<subSubSection id="C328654CFFE8B11516CBFA44C49AFEC1" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="8B8D36C7FFE8B11516CBFA44C49AFEC1" blockId="12.[151,1436,1415,2016]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<emphasis id="B946EAD5FFE8B11616CBFA44C600FA05" bold="true" box="[199,345,1415,1440]" pageId="12" pageNumber="13">Description.</emphasis>
<emphasis id="B946EAD5FFE8B1161753FA44C6BBFA05" box="[351,482,1415,1440]" italics="true" pageId="12" pageNumber="13">Coloration:</emphasis>
Carapace color ranging from light orange to dark reddish tones (
<figureCitation id="13092A42FFE8B11612BFFA44C241FA05" box="[1203,1304,1415,1440]" captionStart="FIGURE14" captionStartId="23.[151,250,1945,1967]" captionTargetBox="[151,1436,267,1908]" captionTargetId="figure@23.[151,1436,267,1923]" captionTargetPageId="23" captionText="FIGURE14. Ariadna maxima (Nicolet,1849). Male from Puerto Montt (IBSP 166780) (A‒E): Habitus, dorsal (A) andventral view (B); Left leg I, prolateral (C) and retrolateral view (D); left metatarsus I, ventral view (E). Female from Santiago (IBSP 161292) (F‒I): Habitus, dorsal (F) and ventral view (G); left leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207671/files/figure.png" pageId="12" pageNumber="13">Figs 14F</figureCitation>
,
<figureCitation id="13092A42FFE8B116132EFA44C20EFA05" box="[1314,1367,1415,1440]" captionStart="FIGURE 56" captionStartId="90.[151,250,1900,1922]" captionTargetBox="[151,1436,337,1878]" captionTargetId="figure@90.[151,1436,337,1878]" captionTargetPageId="90" captionText="FIGURE 56. Ariadna aurea n. sp.. Male (Holotype) from Gruta dos Brejões (IBSP 151451) (A‒E): habitus, dorsal (A) and ventral view (B); rightleg I, prolateral (C) and retrolateral view (D); rightmetatarsus I, ventral view (E), arrowhead= truncated macroseta.Female (Paratype) from Grutasdos Brejões (IBSP 151451) (F‒I):habitus, dorsal (F) and ventral view (G); rightleg I, prolateral (H) and retrolateral (I). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207759/files/figure.png" pageId="12" pageNumber="13">56A</figureCitation>
); eyes bounded by dark outlines (
<figureCitation id="13092A42FFE8B11617C4FA6FC56BFA60" box="[456,562,1452,1477]" captionStart="FIGURE54" captionStartId="87.[151,250,1862,1884]" captionTargetBox="[151,1436,193,1841]" captionTargetId="figure@87.[151,1436,193,1841]" captionTargetPageId="87" captionText="FIGURE54. Ariadna reginae n. sp.. Male (Paratype) from 11km ESE La Virgen (INBIO 4407641) (A‒E): habitus, dorsal (A) and ventral view (B); rightleg I, prolateral(C) andretrolateral view (D); right metatarsus I,ventral view (E).Female(Paratype) from 4km SE Puerto Viejo de Sarapiqui (CAS 9046549) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207755/files/figure.png" pageId="12" pageNumber="13">Fig. 54A</figureCitation>
); legs and palps colors ranging from light yellow to dark reddish tones (
<figureCitation id="13092A42FFE8B1161366FA6FC7B3FA4D" captionStart="FIGURE17" captionStartId="27.[151,250,1446,1468]" captionTargetBox="[151,1436,193,1408]" captionTargetId="figure@27.[151,1436,193,1425]" captionTargetPageId="27" captionText="FIGURE17. Ariadna obscura(Blackwall, 1858). Male from Mata São João (IBSP 85421) (A‒B):left palp, prolateral (A) and retrolateral view (B). Female from Una (IBSP 18283) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D).Scale bar:1mm." httpUri="https://zenodo.org/record/1207677/files/figure.png" pageId="12" pageNumber="13">Figs 17A‒B</figureCitation>
,
<figureCitation id="13092A42FFE8B11616FBFA0CC615FA4D" box="[247,332,1487,1512]" captionStart="FIGURE 65" captionStartId="102.[151,250,1521,1543]" captionTargetBox="[151,1436,193,1500]" captionTargetId="figure@102.[151,1436,193,1515]" captionTargetPageId="102" captionText="FIGURE 65. Aridna caparao n. sp.. Male from Trilha do Terreirão (IBSP 166970) (A‒B): left palp, prolateral (A) and retrolateral view (B). Female (Paratype) from Trilha do Terreirão (IBSP 38345) (C‒D): internal genitalia, lateral view (E); anterior receptaculum, ventral view (F).Scale bar:1mm." httpUri="https://zenodo.org/record/1207777/files/figure.png" pageId="12" pageNumber="13">65A‒B</figureCitation>
); abdomen color ranging from light gray with a conspicuous dark dorsal pattern (
<figureCitation id="13092A42FFE8B11612E4FA0CC22BFA4D" box="[1256,1394,1487,1512]" captionStart="FIGURE 18" captionStartId="30.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@30.[151,1436,193,1803]" captionTargetPageId="30" captionText="FIGURE 18. Ariadna crassipalpa (Blackwall, 1863). Male from Porto Alegre (MCN 3577) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E), arrowhead = curved macroseta. Female from Canela (MCN 0551) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207679/files/figure.png" pageId="12" pageNumber="13">Fig. 18A‒B</figureCitation>
), to dark grey (
<figureCitation id="13092A42FFE8B1161710FA37C6F5F9A8" box="[284,428,1524,1549]" captionStart="FIGURE64" captionStartId="101.[151,250,1781,1803]" captionTargetBox="[151,1435,301,1760]" captionTargetId="figure@101.[151,1436,301,1760]" captionTargetPageId="101" captionText="FIGURE64. Aridna caparaon. sp.. Male from Trilha do Terreirão (IBSP 166970) (A‒F):habitus, dorsal (A) and ventral view (B); cheliceral, dorsal view (C), arrowheads = lameliform distal dorso-prolateral projections; left leg I, prolateral (D) and retrolateral view (E); left metatarsus I. ventral view (F). Female (Paratype) from Trilha do Terreirão (IBSP 38345) (G‒J): habitus, dorsal (G) and ventral view (H); right leg I, prolateral (I) and retrolateral view (J). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207775/files/figure.png" pageId="12" pageNumber="13">Fig. 64A‒B</figureCitation>
). Total length: 2.8‒20.25; carapace 1.38‒5.6 long, 1.02‒3.8 wide.
<emphasis id="B946EAD5FFE8B11612C8FA36C26BF9A9" box="[1220,1330,1525,1548]" italics="true" pageId="12" pageNumber="13">Carapace</emphasis>
: oblong, cephalic region narrower than thoracic region, posterior border straight or slightly concave (
<figureCitation id="13092A42FFE8B11612EEF9D4C20BF995" box="[1250,1362,1559,1584]" captionStart="FIGURE14" captionStartId="23.[151,250,1945,1967]" captionTargetBox="[151,1436,267,1908]" captionTargetId="figure@23.[151,1436,267,1923]" captionTargetPageId="23" captionText="FIGURE14. Ariadna maxima (Nicolet,1849). Male from Puerto Montt (IBSP 166780) (A‒E): Habitus, dorsal (A) andventral view (B); Left leg I, prolateral (C) and retrolateral view (D); left metatarsus I, ventral view (E). Female from Santiago (IBSP 161292) (F‒I): Habitus, dorsal (F) and ventral view (G); left leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207671/files/figure.png" pageId="12" pageNumber="13">Figs 14A</figureCitation>
<figureCitation id="13092A42FFE8B1161352F9DBC2D6F995" box="[1374,1423,1560,1584]" captionStart="FIGURE31" captionStartId="51.[151,250,1917,1939]" captionTargetBox="[213,1370,193,1878]" captionTargetId="figure@51.[213,1370,193,1894]" captionTargetPageId="51" captionText="FIGURE31. Ariadna perkinsiSimon, 1900.Male (Holotype) from Kauai (NHM 1904.X.24.3) (A‒D): habitus, dorsal (A) and ventral view (B); left palp, retrolateral view (C); right metatarsi I, ventral view (D). Female from Oahu (IBSP 209956) (E‒J): habitus, dorsal (E) and ventral view (F); right leg I, prolateral (G) and retrolateral view (H); internal genitalia, lateral view (I); anterior receptaculum, ventral view (J). Scale bar:1mm." httpUri="https://zenodo.org/record/1207709/files/figure.png" pageId="12" pageNumber="13">31E</figureCitation>
); sparsely distributed setae usually concentrated in its edge and in the cephalic region; thoracic fovea smooth/ inconspicuous with only a shallow depression for muscular insertion (
<figureCitation id="13092A42FFE8B11615A7F99CC371F9DD" box="[939,1064,1631,1656]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3A‒B</figureCitation>
).
<emphasis id="B946EAD5FFE8B1161237F9A2C329F9DD" box="[1083,1136,1633,1656]" italics="true" pageId="12" pageNumber="13">Eyes</emphasis>
: six indirect eyes, anterior median eyes absent, lateral eyes arising from a single tubercle (
<figureCitation id="13092A42FFE8B116157AF947C4ADF939" box="[886,1012,1668,1693]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3A‒B</figureCitation>
).
<emphasis id="B946EAD5FFE8B1161205F947C3DDF938" box="[1033,1156,1668,1693]" italics="true" pageId="12" pageNumber="13">Chelicerae</emphasis>
: conical, frontal surface flattened, wider basally in lateral view, usually projected forwardly in males; lateral basal transverse ridge prominent (
<figureCitation id="13092A42FFE8B1161717F90FC62FF941" box="[283,374,1740,1765]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3C</figureCitation>
); frontal basal longitudinal band of ridges and/or tubercles present, without insterstitial setae (
<figureCitation id="13092A42FFE8B1161693F92CC7A7F8AD" box="[159,254,1775,1800]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3C</figureCitation>
); setae scattered distally on the paturon frontal and lateral region (
<figureCitation id="13092A42FFE8B1161225F92CC3D3F8AD" box="[1065,1162,1775,1800]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Figs 3C</figureCitation>
,
<figureCitation id="13092A42FFE8B1161291F92CC388F8AD" box="[1181,1233,1775,1800]" captionStart="FIGURE 36" captionStartId="62.[151,250,1692,1714]" captionTargetBox="[151,1436,195,1669]" captionTargetId="figure@62.[151,1436,195,1670]" captionTargetPageId="62" captionText="FIGURE 36. Ariadna boliviana Simon, 1907. Male from Morrinhos (IBSP 152775) (A‒F): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsi I, ventral view (E); left metatarsi IV, retrolateral view (F). Female from Parauapebas (IBSP 193500) (G‒J): habitus, dorsal (G) and ventral view (H); right leg I, prolateral (I) and retrolateral view (J). Scale bar:1mm." httpUri="https://zenodo.org/record/1207719/files/figure.png" pageId="12" pageNumber="13">36G</figureCitation>
); cheliceral teeth composed by three promarginal and one retromarginal, all conical, and with ridged cuticle (
<figureCitation id="13092A42FFE8B11612B1F8D7C241F889" box="[1213,1304,1812,1837]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3D</figureCitation>
, left inset); cheliceral gland with scattered pores (
<figureCitation id="13092A42FFE8B1161443F8F4C5F2F8F5" box="[591,683,1847,1872]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3D</figureCitation>
, right inset); posterior distal region with a depression, where the endites distal end fits (
<figureCitation id="13092A42FFE8B1161794F89FC6AAF8D1" box="[408,499,1884,1909]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3D</figureCitation>
); fangs short, almost two times the diameter of the distal part of the paturon, shaft with an internal serrula (
<figureCitation id="13092A42FFE8B11617B5F8BCC54EF83D" box="[441,535,1919,1944]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3D</figureCitation>
).
<emphasis id="B946EAD5FFE8B1161422F8BCC5E8F83D" box="[558,689,1919,1944]" italics="true" pageId="12" pageNumber="13">Mouthparts</emphasis>
: labrum well-developed, longer than the labium, without dorsal median setae (
<figureCitation id="13092A42FFE8B1161734F867C6D6F819" box="[312,399,1956,1981]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Figs 3F</figureCitation>
,
<figureCitation id="13092A42FFE8B1161790F867C695F819" box="[412,460,1956,1980]" captionStart="FIGURE 18" captionStartId="30.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@30.[151,1436,193,1803]" captionTargetPageId="30" captionText="FIGURE 18. Ariadna crassipalpa (Blackwall, 1863). Male from Porto Alegre (MCN 3577) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E), arrowhead = curved macroseta. Female from Canela (MCN 0551) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207679/files/figure.png" pageId="12" pageNumber="13">18B</figureCitation>
,
<figureCitation id="13092A42FFE8B11617D5F867C553F819" box="[473,522,1956,1980]" captionStart="FIGURE19" captionStartId="31.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1583]" captionTargetId="figure@31.[151,1436,193,1598]" captionTargetPageId="31" captionText="FIGURE19.Ariadna crassipalpa (Blackwall, 1863).Male fromPorto Alegre (MCN 3577) (A‒B):left palp,prolateral (A) and retrolateral view (B). Female from Canela (MCN 0552) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D).Scale bar:1mm." httpUri="https://zenodo.org/record/1207681/files/figure.png" pageId="12" pageNumber="13">19B</figureCitation>
); endites spatulate, obliquely depressed, narrowed medianly (
<bibRefCitation id="EFA34B36FFE8B11612CCF867C2D6F818" author="Lipke, E. &amp; Ramirez, M. J. &amp; Michalik, P." box="[1216,1423,1956,1981]" journalOrPublisher="Journal of Morphology" pageId="12" pageNumber="13" pagination="1238 - 1257" part="275" publicationUrl="https://doi.org/10.1002/jmor.20298" refId="ref89380" refString="Lipke, E., Ramirez, M. J., Michalik, P. (2014) Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea. Journal of Morphology, 275, 1238 - 1257. https: // doi. org / 10.1002 / jmor. 20298" title="Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea" type="journal article" year="2014">Ramírez 2014: 60</bibRefCitation>
), and rounded proximally (
<figureCitation id="13092A42FFE8B11617BAF804C572F845" box="[438,555,1991,2016]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Fig. 3E‒F</figureCitation>
), hyaline distal prolateral end bearing a setae tuft (
<figureCitation id="13092A42FFE8B1161265F804C3B9F845" box="[1129,1248,1991,2016]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="12" pageNumber="13">Figs 3E‒F</figureCitation>
,
<figureCitation id="13092A42FFE8B11612E0F80BC279F845" box="[1260,1312,1992,2016]" captionStart="FIGURE 12" captionStartId="17.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1796]" captionTargetId="figure@17.[151,1436,193,1796]" captionTargetPageId="17" captionText="FIGURE 12. Ariadna bicolor (Hentz, 1842). Male from Rockville (USNM) (A‒E): Habitus, dorsal (A) and ventral view (B); Left leg I, prolateral view (C) and retrolateral (D), arrowhead = lanceolated macroseta; Left metatarsus I, ventral view (E). Female from 2 mi W. Cedar Crk. (USNM) (F‒I): Habitus, dorsal (F) and ventral view (G); Right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207667/files/figure.png" pageId="12" pageNumber="13">12G</figureCitation>
); setae tuft composed by cylindrical setae with one row of conical barbs (
<figureCitation id="13092A42FFE9B1171545FF54C4C3FF15" box="[841,922,151,176]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="13" pageNumber="14">Fig. 3F</figureCitation>
, inset); serrula with a single row of teeth (
<figureCitation id="13092A42FFE9B1171363FF54C7EEFF71" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="13" pageNumber="14">Fig. 3F</figureCitation>
); labium longer than wider, narrowed in its distal portion, 2/3 of the length of the endite, separated from the sternum by a partially membranous suture (
<figureCitation id="13092A42FFE9B117149CFF1CC5B4FF5D" box="[656,749,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,1882,1904]" captionTargetBox="[197,1383,257,1869]" captionTargetId="figure@7.[197,1383,257,1869]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Ariadna boliviana Simon, 1907, male from Estação Ecológica de Jataí (IBSP 166947) (A, C‒F) and female from Paranaíba (IBSP 14084) (B): cephalothorax, dorsal view (A, B); chelicerae, frontal view (C), arrowheads = basal transverse ridge, asterisks = basal longitudinal band of ridges and/or tubercles; left chelicera, ventro-prolateral view (D), arrowhead = internal serrula, asterisk = left inset = cheliceral tooth, right inset = cheliceral gland; endites and labium, ventral view (E); endites and labrum, frontal view (F), inset = setae that compose the distal setae tuft." httpUri="https://zenodo.org/record/1207647/files/figure.png" pageId="13" pageNumber="14">Figs 3E</figureCitation>
,
<figureCitation id="13092A42FFE9B11714F0FF23C477FF5D" box="[764,814,224,248]" captionStart="FIGURE38" captionStartId="65.[151,250,1919,1941]" captionTargetBox="[182,1406,195,1894]" captionTargetId="figure@65.[182,1406,195,1894]" captionTargetPageId="65" captionText="FIGURE38. Ariadna arthuri Petrunkevitch, 1926.Male from Sierra de Cubitas (IBSP 166771) (A‒E):habitus, dorsal (A) and ventral view (B); left leg I, prolateral (C) and retrolateral view (D); left metatarsus I, ventral view (E). Female from Sierra de Cubitas (IBSP 166771) (F‒I): habitus, dorsal (F) andventral view (G); rightleg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207723/files/figure.png" pageId="13" pageNumber="14">38B</figureCitation>
).
<emphasis id="B946EAD5FFE9B1171548FF22C352FF5D" box="[836,1035,225,248]" italics="true" pageId="13" pageNumber="14">Digestive system:</emphasis>
labial plate with lateral transverse walls, slightly narrowed distally, labrum duct straight, connected to the esophagus through a short pharynx which leads to the sucking stomach (
<figureCitation id="13092A42FFE9B11717F3FEE4C5D5FEE5" box="[511,652,295,320]" captionStart="FIGURE 11" captionStartId="14.[151,250,1105,1127]" captionTargetBox="[151,1436,193,1083]" captionTargetId="figure@14.[151,1436,193,1083]" captionTargetPageId="14" captionText="FIGURE 11. Ariadna maxima (Nicolet, 1849), female from Santiago (IBSP 161292), after enzime prepatarion: habitus, dorsal view, without carapaceand dorsal part of the abdomen (A); cephalothorax and anterior region of the abdomen, dorsal view (B); anterior region of the abdomen, dorsal view (C); posterior receptaculum, dorsal view, tubular invaginations (D)." httpUri="https://zenodo.org/record/1207665/files/figure.png" pageId="13" pageNumber="14">Fig. 11A‒B</figureCitation>
).
<emphasis id="B946EAD5FFE9B11714A8FEEBC5A6FE9A" box="[676,767,296,319]" italics="true" pageId="13" pageNumber="14">Sternum</emphasis>
: longer than wider, anterior region truncated, straight or slightly concave, not fused with the carapace or the pedicel ventral sclerite (
<figureCitation id="13092A42FFE9B117120DFE8FC332FEC0" box="[1025,1131,332,357]" captionStart="FIGURE 36" captionStartId="62.[151,250,1692,1714]" captionTargetBox="[151,1436,195,1669]" captionTargetId="figure@62.[151,1436,195,1670]" captionTargetPageId="62" captionText="FIGURE 36. Ariadna boliviana Simon, 1907. Male from Morrinhos (IBSP 152775) (A‒F): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsi I, ventral view (E); left metatarsi IV, retrolateral view (F). Female from Parauapebas (IBSP 193500) (G‒J): habitus, dorsal (G) and ventral view (H); right leg I, prolateral (I) and retrolateral view (J). Scale bar:1mm." httpUri="https://zenodo.org/record/1207719/files/figure.png" pageId="13" pageNumber="14">Fig. 36H</figureCitation>
).
<emphasis id="B946EAD5FFE9B117128CFE8FC3ACFEC0" box="[1152,1269,332,357]" italics="true" pageId="13" pageNumber="14">Male palp</emphasis>
: tibiae basally swollen, longer than the cymbium (
<figureCitation id="13092A42FFE9B1171427FEACC5D3FE2D" box="[555,650,367,392]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Figs 4A</figureCitation>
,
<figureCitation id="13092A42FFE9B117149BFEB3C591FE2D" box="[663,712,368,392]" captionStart="FIGURE17" captionStartId="27.[151,250,1446,1468]" captionTargetBox="[151,1436,193,1408]" captionTargetId="figure@27.[151,1436,193,1425]" captionTargetPageId="27" captionText="FIGURE17. Ariadna obscura(Blackwall, 1858). Male from Mata São João (IBSP 85421) (A‒B):left palp, prolateral (A) and retrolateral view (B). Female from Una (IBSP 18283) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D).Scale bar:1mm." httpUri="https://zenodo.org/record/1207677/files/figure.png" pageId="13" pageNumber="14">17B</figureCitation>
), and with one or two dorsal trichobothria; cymbium forming a distal notch with the prolateral portion slightly longer than the retrolateral one, forming a distal notch (
<figureCitation id="13092A42FFE9B1171335FE57C2CEFE09" box="[1337,1431,404,429]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Figs 4B</figureCitation>
,
<figureCitation id="13092A42FFE9B117169BFE74C791FE75" box="[151,200,439,464]" captionStart="FIGURE 51" captionStartId="82.[151,250,1516,1538]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@82.[151,1436,193,1494]" captionTargetPageId="82" captionText="FIGURE 51. Ariadna levii Grismado, 2008. Male from R. N. Los Ruiles (MACN-Ar 29452) (A‒B): left palp, prolateral view (A) (arrowhead = chemosensory patch), and retrolateral view (B). Female from Valdivia (MHNS 836) (C‒D): internal genitalia, lateral view (E); anterior receptaculum, ventral view (F). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207749/files/figure.png" pageId="13" pageNumber="14">51B</figureCitation>
), sometimes bearing a dorsal chemosensory setae patch (
<figureCitation id="13092A42FFE9B117156FFE74C48BFE75" box="[867,978,439,464]" captionStart="FIGURE 15" captionStartId="24.[151,250,1507,1529]" captionTargetBox="[151,1436,193,1467]" captionTargetId="figure@24.[151,1436,193,1486]" captionTargetPageId="24" captionText="FIGURE 15. Ariadna maxima (Nicolet, 1849). Male from Illapel (IBSP 166832) (A‒B): left palp, prolateral view (A) (arrowhead = chemosensory patch), and retrolateral view (B). Female from Santiago (IBSP 161292) (C‒D): internal genitalia, lateral view (C); anterior receptaculum, ventral view (D). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207673/files/figure.png" pageId="13" pageNumber="14">Figs 15A</figureCitation>
,
<figureCitation id="13092A42FFE9B11715EDFE7BC34CFE75" box="[993,1045,440,464]" captionStart="FIGURE 48" captionStartId="78.[151,250,1561,1583]" captionTargetBox="[151,1436,193,1521]" captionTargetId="figure@78.[151,1436,193,1540]" captionTargetPageId="78" captionText="FIGURE 48. Ariadna changellkuk Grismado, 2008. Male from Renca (AMNH) (A‒B): left palp, prolateral view (A) (arrowhead = chemosensory patch), and retrolateral view (B). Female(Paratype) from East of Curicó (AMNH) (C‒D):internal genitalia, lateral view (C); anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207743/files/figure.png" pageId="13" pageNumber="14">48A</figureCitation>
); bulb rounded or subrectangular, abruptly narrowed forming a mid-portion between the bulb and the embolus, the latter curved forwards (
<figureCitation id="13092A42FFE9B1171335FE1FC2CEFE51" box="[1337,1431,476,501]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Figs 4A</figureCitation>
,
<figureCitation id="13092A42FFE9B117169BFDC3C7B5FDBD" box="[151,236,512,536]" captionStart="FIGURE 21" captionStartId="35.[151,250,1612,1634]" captionTargetBox="[151,1436,193,1575]" captionTargetId="figure@35.[151,1436,193,1590]" captionTargetPageId="35" captionText="FIGURE 21. Ariadna mollis (Holmberg, 1876). Male from Parque Estadual do Jaraguá (IBSP 166961) (A‒B): left palp, prolateral (A) and retrolateral view (B). Female from Perdizes (MZSP 1709) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D). Scale bar:1mm." httpUri="https://zenodo.org/record/1207687/files/figure.png" pageId="13" pageNumber="14">21A‒B</figureCitation>
); spermatic duct with an internal porous stripe with one convolution inside the bulb (
<figureCitation id="13092A42FFE9B11712B5FE3CC26EFDBD" box="[1209,1335,511,536]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Figs 4D‒E</figureCitation>
,
<figureCitation id="13092A42FFE9B1171348FDC3C2C1FDBD" box="[1348,1432,512,536]" captionStart="FIGURE17" captionStartId="27.[151,250,1446,1468]" captionTargetBox="[151,1436,193,1408]" captionTargetId="figure@27.[151,1436,193,1425]" captionTargetPageId="27" captionText="FIGURE17. Ariadna obscura(Blackwall, 1858). Male from Mata São João (IBSP 85421) (A‒B):left palp, prolateral (A) and retrolateral view (B). Female from Una (IBSP 18283) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D).Scale bar:1mm." httpUri="https://zenodo.org/record/1207677/files/figure.png" pageId="13" pageNumber="14">17A‒B</figureCitation>
; following
<bibRefCitation id="EFA34B36FFE9B1171701FDE7C6C5FD98" author="Cooke, J. A. L." box="[269,412,548,573]" journalOrPublisher="Senckenbergiana Biologica" pageId="13" pageNumber="14" pagination="35 - 43" part="47" refId="ref88019" refString="Cooke, J. A. L. (1966) Synopsis of the structure and function of the genitalia in Dysdera crocata (Araneae, Dysderidae). Senckenbergiana Biologica, 47, 35 - 43." title="Synopsis of the structure and function of the genitalia in Dysdera crocata (Araneae, Dysderidae)" type="journal article" year="1966">Cooke 1966</bibRefCitation>
: 37;
<bibRefCitation id="EFA34B36FFE9B11717DBFDE7C53FFD99" author="Cooke, J. A. L." box="[471,614,548,573]" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle de Paris" pageId="13" pageNumber="14" pagination="142 - 146" part="41" refId="ref88057" refString="Cooke, J. A. L. (1970) Spider genitalia and phylogeny. Bulletin du Museum National d'Histoire Naturelle de Paris, 41 (Supplement 1), 142 - 146." title="Spider genitalia and phylogeny" type="journal article" year="1970">Cooke 1970</bibRefCitation>
: 143, 145), becoming narrow and hyaline between the mid-portion and embolus (
<figureCitation id="13092A42FFE9B1171709FD84C606FDC5" box="[261,351,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,250,747,769]" captionTargetBox="[151,1436,235,680]" captionTargetId="figure@9.[151,1436,235,680]" captionTargetPageId="9" captionText="FIGURE 5. Ariadna boliviana Simon, 1907, male from Morrinhos (IBSP 152775) (A‒B): left palp, retrolateral view, midportion showing the less sclerotized region in detail (A); tarsus-metatarsus IV junction, retrolateral view (B), arrowheads = ventral brush of putative chemosensory setae,asterisk = preening comb of macrosetae." httpUri="https://zenodo.org/record/1207651/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
, following
<bibRefCitation id="EFA34B36FFE9B11717D2FD84C5FAFDC5" author="Lipke, E. &amp; Ramirez, M. J. &amp; Michalik, P." box="[478,675,583,608]" journalOrPublisher="Journal of Morphology" pageId="13" pageNumber="14" pagination="1238 - 1257" part="275" publicationUrl="https://doi.org/10.1002/jmor.20298" refId="ref89380" refString="Lipke, E., Ramirez, M. J., Michalik, P. (2014) Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea. Journal of Morphology, 275, 1238 - 1257. https: // doi. org / 10.1002 / jmor. 20298" title="Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea" type="journal article" year="2014">
Lipke
<emphasis id="B946EAD5FFE9B117142BFD8AC539FDC5" box="[551,608,583,608]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2014
</bibRefCitation>
); embolus tubular (
<figureCitation id="13092A42FFE9B1171570FD84C4AEFDC5" box="[892,1015,583,608]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Fig. 4A‒C</figureCitation>
).
<emphasis id="B946EAD5FFE9B1171207FD84C3C1FDC5" box="[1035,1176,583,608]" italics="true" pageId="13" pageNumber="14">Female palp</emphasis>
: short and robust, with one articulated smooth claw at the tip, diaxially orientated (
<figureCitation id="13092A42FFE9B117154BFDAFC4FAFD21" box="[839,931,620,645]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Fig. 4G</figureCitation>
), and scattered macrosetae on the prolateral region of the tarsi and tibiae (sometimes patellae) (
<figureCitation id="13092A42FFE9B11714F5FD4CC40DFD0D" box="[761,852,655,680]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Fig. 4F</figureCitation>
); tarsi surrounded almost entirely by tactile setae (interstitial chemosensory setae is also found), the ventral ones thickened, strongly serrated, and gradually decreasing in size in distal direction (
<figureCitation id="13092A42FFE9B1171433FD14C5E1FD55" box="[575,696,727,752]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="13" pageNumber="14">Fig. 4F‒G</figureCitation>
); tibia with one dorsal trichobothria.
<emphasis id="B946EAD5FFE9B1171251FD1AC3CBFD55" box="[1117,1170,729,752]" italics="true" pageId="13" pageNumber="14">Legs</emphasis>
: robust, prograde, with the third pair directed forward (
<figureCitation id="13092A42FFE9B1171409FD3FC5DAFCB1" box="[517,643,764,789]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="13" pageNumber="14">Figs 7A‒C</figureCitation>
,
<figureCitation id="13092A42FFE9B1171483FD3FC58CFCB1" box="[655,725,764,788]" captionStart="FIGURE 18" captionStartId="30.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1802]" captionTargetId="figure@30.[151,1436,193,1803]" captionTargetPageId="30" captionText="FIGURE 18. Ariadna crassipalpa (Blackwall, 1863). Male from Porto Alegre (MCN 3577) (A‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D); right metatarsus I, ventral view (E), arrowhead = curved macroseta. Female from Canela (MCN 0551) (F‒I): habitus, dorsal (F) and ventral view (G); right leg I, prolateral (H) and retrolateral view (I). Scale bar:1mm." httpUri="https://zenodo.org/record/1207679/files/figure.png" pageId="13" pageNumber="14">18F‒I</figureCitation>
), usually with pairs I‒II with more setae and macrosetae than pairs III‒IV; metatarsi and tarsi IV with a ventral brush of thickened and serrated setae and metatarsi IV with a distal ventro-retrolateral preening comb of macrosetae (
<figureCitation id="13092A42FFEAB1141505FB2CC43CFAAD" box="[777,869,1263,1288]" captionStart="FIGURE 5" captionStartId="9.[151,250,747,769]" captionTargetBox="[151,1436,235,680]" captionTargetId="figure@9.[151,1436,235,680]" captionTargetPageId="9" captionText="FIGURE 5. Ariadna boliviana Simon, 1907, male from Morrinhos (IBSP 152775) (A‒B): left palp, retrolateral view, midportion showing the less sclerotized region in detail (A); tarsus-metatarsus IV junction, retrolateral view (B), arrowheads = ventral brush of putative chemosensory setae,asterisk = preening comb of macrosetae." httpUri="https://zenodo.org/record/1207651/files/figure.png" pageId="14" pageNumber="15">Figs 5B</figureCitation>
,
<figureCitation id="13092A42FFEAB114157EFB33C4EAFAAD" box="[882,947,1264,1288]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="14" pageNumber="15">7E‒F</figureCitation>
); leg formula I-II-IV-III or II-I-IV-III; three tarsal claws, superior ones pectinated, and inferior one with only a small tooth (
<figureCitation id="13092A42FFEAB1141244FAD0C3FFFA89" box="[1096,1190,1299,1324]" captionStart="FIGURE6" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[151,1436,924,1868]" captionTargetId="figure@9.[151,1436,924,1868]" captionTargetPageId="9" captionText="FIGURE6. SEM imagesof Ariadna tarsalisBanks, 1902, male from Barbones (MUSM 0504979)(A‒F):right metatarsus and tarsus I, ventral view, showing examples of modifications in the male metatarsus I; modified macrosetae (B‒D); tarsal claws, prolateral view (E), arrowhead = denticle in the inferior claw, asterisk = tarsal organ; tarsal organ, dorsal view (F), arrowheads = rimmedreceptors." httpUri="https://zenodo.org/record/1207655/files/figure.png" pageId="14" pageNumber="15">Figs 6E</figureCitation>
,
<figureCitation id="13092A42FFEAB11412BAFAD7C3A4FA89" box="[1206,1277,1300,1324]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="14" pageNumber="15">7C‒D</figureCitation>
); tarsal organ exposed with edges, and usually two rimmed receptors (
<figureCitation id="13092A42FFEAB114151CFAF4C4DCFAF5" box="[784,901,1335,1360]" captionStart="FIGURE6" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[151,1436,924,1868]" captionTargetId="figure@9.[151,1436,924,1868]" captionTargetPageId="9" captionText="FIGURE6. SEM imagesof Ariadna tarsalisBanks, 1902, male from Barbones (MUSM 0504979)(A‒F):right metatarsus and tarsus I, ventral view, showing examples of modifications in the male metatarsus I; modified macrosetae (B‒D); tarsal claws, prolateral view (E), arrowhead = denticle in the inferior claw, asterisk = tarsal organ; tarsal organ, dorsal view (F), arrowheads = rimmedreceptors." httpUri="https://zenodo.org/record/1207655/files/figure.png" pageId="14" pageNumber="15">Fig. 6E‒F</figureCitation>
; Platnick
<emphasis id="B946EAD5FFEAB11415F8FAFBC374FAF5" box="[1012,1069,1335,1360]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2012: figs 36‒45); trichobothria on the dorsal subdistal region of metatarsi I‒IV, dorsal subproximal region of tibia I‒IV; trichobothrial bases with a transverse ridge, and without a well-delimitation between proximal and distal plate (
<figureCitation id="13092A42FFEAB114126CFABCC3E0FA3D" box="[1120,1209,1407,1432]" captionStart="FIGURE4" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,211,1776]" captionTargetId="figure@8.[151,1436,211,1776]" captionTargetPageId="8" captionText="FIGURE4. SEM imagensof Ariadna bolivianaSimon, 1907, males from Parque Estadual do Rio Doce (UFMG 10501) (A‒C) and Estação Ecológica de Jataí (IBSP 166947) (D‒E), and female from Paranaíba (IBSP 14084) (F‒G): left palp, retrolateral (A) and frontal view (B), arrowhead = cymbium distal notch, inset = trichobothrium from palp tibia; embolus tip, retrolateral view (C); bulb, internal view of spermatic duct (D), arrowhead = internal porous stripe; internal porous stripe in detail, internal view (E); right palp, ventro-prolateral view (F), asterisk = thickened and serrated tactile setae; apical region of palp tarsus, arrowheads = chemosensory setae, asterisks = thickened and serrated tactile setae." httpUri="https://zenodo.org/record/1207649/files/figure.png" pageId="14" pageNumber="15">Fig. 4B</figureCitation>
, inset); tenent setae absent in all legs; males with a tarsal brush of putative chemosensory setae located in the ventral region (
<figureCitation id="13092A42FFEAB1141333FA60C2CEFA19" box="[1343,1431,1443,1468]" captionStart="FIGURE 5" captionStartId="9.[151,250,747,769]" captionTargetBox="[151,1436,235,680]" captionTargetId="figure@9.[151,1436,235,680]" captionTargetPageId="9" captionText="FIGURE 5. Ariadna boliviana Simon, 1907, male from Morrinhos (IBSP 152775) (A‒B): left palp, retrolateral view, midportion showing the less sclerotized region in detail (A); tarsus-metatarsus IV junction, retrolateral view (B), arrowheads = ventral brush of putative chemosensory setae,asterisk = preening comb of macrosetae." httpUri="https://zenodo.org/record/1207651/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
, following
<bibRefCitation id="EFA34B36FFEAB1141706FA04C688FA45" author="Foelix, R. F. &amp; Erb, B. &amp; Michalik, P." box="[266,465,1479,1504]" journalOrPublisher="The Journal of Arachnology" pageId="14" pageNumber="15" pagination="599 - 603" part="38" publicationUrl="https://doi.org/10.1636/B10-42.1" refId="ref88484" refString="Foelix, R. F., Erb, B. &amp; Michalik, P. (2010) Scopulate hairs in male Liphistius spiders: probable contact chemoreceptors. The Journal of Arachnology, 38, 599 - 603. https: // doi. org / 10.1636 / B 10 - 42.1" title="Scopulate hairs in male Liphistius spiders: probable contact chemoreceptors" type="journal article" year="2010">
Foelix
<emphasis id="B946EAD5FFEAB1141754FA0BC6C9FA45" box="[344,400,1479,1504]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2010
</bibRefCitation>
, figs 1‒2;
<bibRefCitation id="EFA34B36FFEAB1141448FA0BC5BFFA45" author="Lipke, E. &amp; Ramirez, M. J. &amp; Michalik, P." box="[580,742,1480,1504]" journalOrPublisher="Journal of Morphology" pageId="14" pageNumber="15" pagination="1238 - 1257" part="275" publicationUrl="https://doi.org/10.1002/jmor.20298" refId="ref89380" refString="Lipke, E., Ramirez, M. J., Michalik, P. (2014) Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea. Journal of Morphology, 275, 1238 - 1257. https: // doi. org / 10.1002 / jmor. 20298" title="Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
: 125); male metatarsi I usually bearing projections, apophyses or modified macrosetae (
<figureCitation id="13092A42FFEAB11417B0FA28C564F9A1" box="[444,573,1515,1540]" captionStart="FIGURE6" captionStartId="9.[151,250,1922,1944]" captionTargetBox="[151,1436,924,1868]" captionTargetId="figure@9.[151,1436,924,1868]" captionTargetPageId="9" captionText="FIGURE6. SEM imagesof Ariadna tarsalisBanks, 1902, male from Barbones (MUSM 0504979)(A‒F):right metatarsus and tarsus I, ventral view, showing examples of modifications in the male metatarsus I; modified macrosetae (B‒D); tarsal claws, prolateral view (E), arrowhead = denticle in the inferior claw, asterisk = tarsal organ; tarsal organ, dorsal view (F), arrowheads = rimmedreceptors." httpUri="https://zenodo.org/record/1207655/files/figure.png" pageId="14" pageNumber="15">Fig. 6A‒D</figureCitation>
); female tibiae and metatarsi without ventral and dorsal macrosetae, only ventro-prolateral, ventro-retrolateral, and sometimes prolateral and retrolateral (
<figureCitation id="13092A42FFEAB1141217F9CCC3CEF98D" box="[1051,1175,1551,1576]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="14" pageNumber="15">Fig. 7A‒C</figureCitation>
); female tarsi I‒II with ventro-prolateral lines of thickened tactile setae, interspersed with lines of chemosensory setae (
<figureCitation id="13092A42FFEAB1141305F9F0C2C9F9E9" box="[1289,1424,1587,1612]" captionStart="FIGURE 7" captionStartId="10.[151,250,1681,1703]" captionTargetBox="[151,1436,211,1647]" captionTargetId="figure@10.[151,1436,211,1648]" captionTargetPageId="10" captionText="FIGURE 7. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒G):left leg I, prolateral view (A); leftmetatarsus and tarsus I, prolateralview (B);left metatarsus andtarsus,ventral view (C); left tarsus I,ventral view (D), thickened tactile setae in blue, chemosensory setae in yellow; right metatarsus and tarsus IV, prolateral view (E), arrowheads = brush of thickened and serrated setae; right metatarsus IV, ventro-retrolateral view, distal preening comb of macrosetae (F), asterisks = thickened and serrated setae; left tarsus I, ventral view, detail of thickened setae." httpUri="https://zenodo.org/record/1207657/files/figure.png" pageId="14" pageNumber="15">Fig. 7D, G</figureCitation>
).
<emphasis id="B946EAD5FFEAB114169BF994C658F9D5" box="[151,257,1623,1648]" italics="true" pageId="14" pageNumber="15">Abdomen</emphasis>
: longer than wide, uniformly hairy, coated with short setae, and scattered long setae among them (
<figureCitation id="13092A42FFEAB1141362F994C7B0F931" captionStart="FIGURE 16" captionStartId="26.[151,250,1825,1847]" captionTargetBox="[151,1436,193,1803]" captionTargetId="figure@26.[151,1436,193,1803]" captionTargetPageId="26" captionText="FIGURE 16. Ariadna obscura (Blackwall, 1858). Males from Fortaleza (MPEG 11628) (A‒B) and Mata São João (IBSP 85421) (C‒E): habitus, dorsal (A) and ventral view (B); right leg I, prolateral (C) and retrolateral view (D), arrowhead = lanceolated macroseta; right metatarsus I, ventral view (E), arrowhead = group of short distinct setae. Female from Fortaleza (IBSP 165434) (F‒I): habitus, dorsal (F) and ventral (G) view; left leg I, prolateral (H) and retrolateral view (I). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207675/files/figure.png" pageId="14" pageNumber="15">Fig. 16F‒G</figureCitation>
); book lung cover weakly sclerotized, and ventral anterior sigillae distinct, with slit sensillae (
<figureCitation id="13092A42FFEAB1141328F9B8C2D6F931" box="[1316,1423,1659,1684]" captionStart="FIGURE 22" captionStartId="38.[151,250,1743,1765]" captionTargetBox="[151,1436,301,1722]" captionTargetId="figure@38.[151,1436,301,1722]" captionTargetPageId="38" captionText="FIGURE 22. Ariadna boesenbergi Keyserling, 1877. Male from Reserva Biológica São Donato (IBSP 166778) (A‒E): Habitus, dorsal (A) and ventral view (B); left leg I, prolateral view (C) and retrolateral view (D); left metatarsi I, ventral view (E). Female from Parque Natural INIA Las Brujas (IBSP 166779) (F‒J): habitus, dorsal (F) and ventral view (G); epigastric furrow (H), arrowheads = ventral anterior sigillae; right leg I, prolateral (I) and retrolateral view (J). Scale bar:1mm." httpUri="https://zenodo.org/record/1207689/files/figure.png" pageId="14" pageNumber="15">Fig. 22H</figureCitation>
); tracheal system dysderoidlike, with tracheoles spreading through the cephalothorax, including all the appendages, and the abdomen (
<figureCitation id="13092A42FFEAB1141762F900C694F979" box="[366,461,1731,1756]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="14" pageNumber="15">Figs 9A</figureCitation>
,
<figureCitation id="13092A42FFEAB11417D0F907C569F979" box="[476,560,1732,1756]" captionStart="FIGURE 11" captionStartId="14.[151,250,1105,1127]" captionTargetBox="[151,1436,193,1083]" captionTargetId="figure@14.[151,1436,193,1083]" captionTargetPageId="14" captionText="FIGURE 11. Ariadna maxima (Nicolet, 1849), female from Santiago (IBSP 161292), after enzime prepatarion: habitus, dorsal view, without carapaceand dorsal part of the abdomen (A); cephalothorax and anterior region of the abdomen, dorsal view (B); anterior region of the abdomen, dorsal view (C); posterior receptaculum, dorsal view, tubular invaginations (D)." httpUri="https://zenodo.org/record/1207665/files/figure.png" pageId="14" pageNumber="15">11A‒C</figureCitation>
;
<bibRefCitation id="EFA34B36FFEAB1141432F907C5BDF979" author="Ramirez, M. J." box="[574,740,1732,1756]" journalOrPublisher="The Journal of Arachnology" pageId="14" pageNumber="15" pagination="149 - 157" part="28" publicationUrl="https://doi.org/10.1636/0161-8202(2000)028[0149:RSMATP]2.0.CO;2" refId="ref90515" refString="Ramirez, M. J. (2000) Respiratory system morphology and the phylogeny of haplogyne spiders (Araneae, Araneomorphae). The Journal of Arachnology, 28, 149 - 157. https: // doi. org / 10.1636 / 0161 - 8202 (2000) 028 [0149: RSMATP] 2.0. CO; 2" title="Respiratory system morphology and the phylogeny of haplogyne spiders (Araneae, Araneomorphae)" type="journal article" year="2000">Ramírez 2000</bibRefCitation>
: 153;
<bibRefCitation id="EFA34B36FFEAB1141520F900C346F979" author="Griswold, C. E. &amp; Ramirez, M. J. &amp; Coddington, J. A. &amp; Platnick, N. I." box="[812,1055,1731,1756]" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="14" pageNumber="15" pagination="1 - 324" part="56" refId="ref88922" refString="Griswold, C. E., Ramirez, M. J., Coddington, J. A. &amp; Platnick, N. I. (2005) Atlas of Phylogenetic Data for Entelegynae Spiders (Araneae: Araneomorphae: Entelegynae) with Comments on their Phylogeny. Proceedings of the California Academy of Sciences, 56 (Supplement II), 1 - 324." title="Atlas of Phylogenetic Data for Entelegynae Spiders (Araneae: Araneomorphae: Entelegynae) with Comments on their Phylogeny" type="journal article" year="2005">
Griswold
<emphasis id="B946EAD5FFEAB1141592F906C480F979" box="[926,985,1731,1756]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2005
</bibRefCitation>
: 39).
<emphasis id="B946EAD5FFEAB114126FF907C381F97E" box="[1123,1240,1732,1755]" italics="true" pageId="14" pageNumber="15">Spinnerets</emphasis>
: ALS with three segments, the basal segment crossed by a diagonal membranous stripe (
<figureCitation id="13092A42FFEAB11415B5F924C34BF8A5" box="[953,1042,1767,1792]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1933]" captionTargetBox="[161,1425,797,1898]" captionTargetId="figure@11.[161,1425,797,1899]" captionTargetPageId="11" captionText="FIGURE 8. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F)." httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="14" pageNumber="15">Fig. 8A</figureCitation>
), one MAP spigot with a prolateral field of about 15 strain sensillae, divided in two rounded and one canoe-shaped area (
<figureCitation id="13092A42FFEAB1141264F8C8C3BAF881" box="[1128,1251,1803,1828]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1933]" captionTargetBox="[161,1425,797,1898]" captionTargetId="figure@11.[161,1425,797,1899]" captionTargetPageId="11" captionText="FIGURE 8. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F)." httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="14" pageNumber="15">Fig. 8D‒E</figureCitation>
); PMS with one mAP and one AC spigot (
<figureCitation id="13092A42FFEAB11417C6F8ECC57FF8ED" box="[458,550,1839,1864]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1933]" captionTargetBox="[161,1425,797,1898]" captionTargetId="figure@11.[161,1425,797,1899]" captionTargetPageId="11" captionText="FIGURE 8. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F)." httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="14" pageNumber="15">Fig. 8C</figureCitation>
); PLS usually with 3‒4 AC (
<figureCitation id="13092A42FFEAB114158CF8ECC482F8ED" box="[896,987,1839,1864]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1933]" captionTargetBox="[161,1425,797,1898]" captionTargetId="figure@11.[161,1425,797,1899]" captionTargetPageId="11" captionText="FIGURE 8. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F)." httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="14" pageNumber="15">Fig. 8B</figureCitation>
); colulus distinct and pilose (
<figureCitation id="13092A42FFEAB114133AF8ECC2D6F8ED" box="[1334,1423,1839,1864]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1933]" captionTargetBox="[161,1425,797,1898]" captionTargetId="figure@11.[161,1425,797,1899]" captionTargetPageId="11" captionText="FIGURE 8. SEM images of Ariadna boliviana Simon, 1907, female from Paranaíba (IBSP 14084) (A‒F): spinnerets, ventroposterior view (A), arrowheads = diagnal membranous stripe from ALS; right PLS, posterior view (B); left PMS, posterior view (C); right ALS, posterior view (D), asterisk = prolateral field of strain sensillae; left ALS, detail of the prolateral field of strain sensillae (E), arrowhead = canoe-shaped area, asterisks = rounded areas; colulus, ventral view (F)." httpUri="https://zenodo.org/record/1207659/files/figure.png" pageId="14" pageNumber="15">Fig. 8F</figureCitation>
).
<emphasis id="B946EAD5FFEAB114169BF890C605F8C9" box="[151,348,1875,1900]" italics="true" pageId="14" pageNumber="15">Female genitalia</emphasis>
: external plate distinct and sclerotized (
<figureCitation id="13092A42FFEAB114154EF890C4E8F8C9" box="[834,945,1875,1900]" captionStart="FIGURE 22" captionStartId="38.[151,250,1743,1765]" captionTargetBox="[151,1436,301,1722]" captionTargetId="figure@38.[151,1436,301,1722]" captionTargetPageId="38" captionText="FIGURE 22. Ariadna boesenbergi Keyserling, 1877. Male from Reserva Biológica São Donato (IBSP 166778) (A‒E): Habitus, dorsal (A) and ventral view (B); left leg I, prolateral view (C) and retrolateral view (D); left metatarsi I, ventral view (E). Female from Parque Natural INIA Las Brujas (IBSP 166779) (F‒J): habitus, dorsal (F) and ventral view (G); epigastric furrow (H), arrowheads = ventral anterior sigillae; right leg I, prolateral (I) and retrolateral view (J). Scale bar:1mm." httpUri="https://zenodo.org/record/1207689/files/figure.png" pageId="14" pageNumber="15">Fig. 22H</figureCitation>
); internal genitalia composed by anterior receptaculum, uterus externus, interpulmonary fold, and posterior receptaculum (
<figureCitation id="13092A42FFEAB114122EF8B4C332F835" box="[1058,1131,1911,1936]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="14" pageNumber="15">Figs 9</figureCitation>
,
<figureCitation id="13092A42FFEAB114127BF8BBC3CCF835" box="[1143,1173,1912,1936]" captionStart="FIGURE 10" captionStartId="13.[151,250,1897,1919]" captionTargetBox="[151,1436,841,1875]" captionTargetId="figure@13.[151,1436,841,1875]" captionTargetPageId="13" captionText="FIGURE 10. SEM images of Ariadna boliviana Simon, 1907, female from Parauapebas (IBSP 193499): anterior receptaculum, lateral (A) and frontal view (B); internal genitalia, frontal view (C); posterior wall of bursa copulatrix, lateral view, origin of the posterior receptaculum (D), arrowhead = median membranous crack; pore of a tubular invagination in the membranous cuticle of the posterior receptaculum (E); posterior receptaculum, internal view, tubular invaginations (F‒G)." httpUri="https://zenodo.org/record/1207663/files/figure.png" pageId="14" pageNumber="15">10</figureCitation>
,
<figureCitation id="13092A42FFEAB11412ACF8BBC3AAF835" box="[1184,1267,1912,1936]" captionStart="FIGURE 11" captionStartId="14.[151,250,1105,1127]" captionTargetBox="[151,1436,193,1083]" captionTargetId="figure@14.[151,1436,193,1083]" captionTargetPageId="14" captionText="FIGURE 11. Ariadna maxima (Nicolet, 1849), female from Santiago (IBSP 161292), after enzime prepatarion: habitus, dorsal view, without carapaceand dorsal part of the abdomen (A); cephalothorax and anterior region of the abdomen, dorsal view (B); anterior region of the abdomen, dorsal view (C); posterior receptaculum, dorsal view, tubular invaginations (D)." httpUri="https://zenodo.org/record/1207665/files/figure.png" pageId="14" pageNumber="15">11C‒D</figureCitation>
,
<figureCitation id="13092A42FFEAB11412F2F8BBC269F835" box="[1278,1328,1912,1936]" captionStart="FIGURE33" captionStartId="55.[151,250,1553,1575]" captionTargetBox="[151,1436,193,1514]" captionTargetId="figure@55.[151,1436,193,1531]" captionTargetPageId="55" captionText="FIGURE33.Ariadna tarsalis Banks,1902.Male from Playa Ibbetson (RBINS) (A‒B):left palp, prolateral(A) and retrolateral view (B).Female from DarwinIsland (CAS9039512)(C‒D):internal genitalia,lateral view (C);anterior receptaculum,ventral view (D). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207713/files/figure.png" pageId="14" pageNumber="15">33C</figureCitation>
); anterior receptaculum usually billobulated, with a dorsal and a ventral lobe, both being dead-end structures, and a glandular ducts plate that fills the median space between the two lobes (
<figureCitation id="13092A42FFEAB1141563F87CC497F87D" box="[879,974,1983,2008]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="14" pageNumber="15">Figs 9B</figureCitation>
,
<figureCitation id="13092A42FFEAB11415D1F803C368F87D" box="[989,1073,1984,2008]" captionStart="FIGURE 10" captionStartId="13.[151,250,1897,1919]" captionTargetBox="[151,1436,841,1875]" captionTargetId="figure@13.[151,1436,841,1875]" captionTargetPageId="13" captionText="FIGURE 10. SEM images of Ariadna boliviana Simon, 1907, female from Parauapebas (IBSP 193499): anterior receptaculum, lateral (A) and frontal view (B); internal genitalia, frontal view (C); posterior wall of bursa copulatrix, lateral view, origin of the posterior receptaculum (D), arrowhead = median membranous crack; pore of a tubular invagination in the membranous cuticle of the posterior receptaculum (E); posterior receptaculum, internal view, tubular invaginations (F‒G)." httpUri="https://zenodo.org/record/1207663/files/figure.png" pageId="14" pageNumber="15">10A‒B</figureCitation>
,
<figureCitation id="13092A42FFEAB114124CF803C3CCF87D" box="[1088,1173,1984,2008]" captionStart="FIGURE17" captionStartId="27.[151,250,1446,1468]" captionTargetBox="[151,1436,193,1408]" captionTargetId="figure@27.[151,1436,193,1425]" captionTargetPageId="27" captionText="FIGURE17. Ariadna obscura(Blackwall, 1858). Male from Mata São João (IBSP 85421) (A‒B):left palp, prolateral (A) and retrolateral view (B). Female from Una (IBSP 18283) (C‒D): Internal genitalia, lateral view (C); Anterior receptaculum, ventral view (D).Scale bar:1mm." httpUri="https://zenodo.org/record/1207677/files/figure.png" pageId="14" pageNumber="15">17C‒D</figureCitation>
); glandular ducts plate heavily sclerotized (i.e. anterior receptaculum most sclerotized region), composed by cuticular ducts with expanded bases that allows glandular cells to discharge secretion in the lumen of the anterior receptaculum through pores in its cuticle (
<figureCitation id="13092A42FFEBB115177EFF1CC6B6FF5D" box="[370,495,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="15" pageNumber="16">Figs 9B‒D</figureCitation>
,
<figureCitation id="13092A42FFEBB11517F7FF23C517FF5D" box="[507,590,224,248]" captionStart="FIGURE 10" captionStartId="13.[151,250,1897,1919]" captionTargetBox="[151,1436,841,1875]" captionTargetId="figure@13.[151,1436,841,1875]" captionTargetPageId="13" captionText="FIGURE 10. SEM images of Ariadna boliviana Simon, 1907, female from Parauapebas (IBSP 193499): anterior receptaculum, lateral (A) and frontal view (B); internal genitalia, frontal view (C); posterior wall of bursa copulatrix, lateral view, origin of the posterior receptaculum (D), arrowhead = median membranous crack; pore of a tubular invagination in the membranous cuticle of the posterior receptaculum (E); posterior receptaculum, internal view, tubular invaginations (F‒G)." httpUri="https://zenodo.org/record/1207663/files/figure.png" pageId="15" pageNumber="16">10A‒B</figureCitation>
,
<figureCitation id="13092A42FFEBB1151455FF1CC5F7FF5D" box="[601,686,223,248]" captionStart="FIGURE 35" captionStartId="58.[151,250,1462,1484]" captionTargetBox="[151,1436,193,1423]" captionTargetId="figure@58.[151,1436,193,1441]" captionTargetPageId="58" captionText="FIGURE 35. Ariadna cephalotes Simon, 1907. Male from Vale do Zongo (MCN 24097) (A‒B): left palp, prolateral view (A). and retrolateral view (B) (arrowhead = basal anterior keel). Female from La Cumbre (MCN 24072) (C‒D): internal genitalia, lateral view (C); anterior receptaculum, ventral view (D). Scale bar: 1mm." httpUri="https://zenodo.org/record/1207717/files/figure.png" pageId="15" pageNumber="16">35C‒D</figureCitation>
); interpulmonary fold with dorsal smooth curvature, with a median membranous crack (absent in
<taxonomicName id="4C324D44FFEBB11517EAFEC7C528FEB9" authorityName="Grismado" authorityYear="2008" box="[486,625,260,284]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="araucana">
<emphasis id="B946EAD5FFEBB11517EAFEC7C528FEB9" box="[486,625,260,284]" italics="true" pageId="15" pageNumber="16">A. araucana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D44FFEBB11514ABFEC7C5A7FEB8" authority="Giroti &amp; Brescovit, 2018" authorityName="Giroti &amp; Brescovit" authorityYear="2018" box="[679,766,260,285]" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="lalen" status="sp. nov.">
<emphasis id="B946EAD5FFEBB11514ABFEC7C5A7FEB8" box="[679,766,260,285]" italics="true" pageId="15" pageNumber="16">A. lalen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557AEFFEBB1151509FEC7C41DFEB8" box="[773,836,260,285]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B946EAD5FFEBB1151509FEC7C41DFEB8" bold="true" box="[773,836,260,285]" pageId="15" pageNumber="16">n. sp.</emphasis>
</taxonomicNameLabel>
) (
<figureCitation id="13092A42FFEBB1151550FEC7C481FEB9" box="[860,984,260,285]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="15" pageNumber="16">Fig. 9A‒B</figureCitation>
); posterior receptaculum composed by a membranous sac that originates from the posterior wall of the bursa copulatrix, with pores connected with tubular invaginations that goes towards the lumen (
<figureCitation id="13092A42FFEBB1151472FE8FC5A5FEC1" box="[638,764,332,357]" captionStart="FIGURE 9" captionStartId="12.[151,250,1257,1279]" captionTargetBox="[151,1435,193,1236]" captionTargetId="figure@12.[151,1436,193,1236]" captionTargetPageId="12" captionText="FIGURE 9. SEM images of Ariadna maxima (Nicolet, 1849), females from Santiago (IBSP 166664) (A‒B) and Doñihue (IBSP 166795) (C‒D): internal genitalia, frontal view (A); internal genitalia, lateral view (B), arrowhead = median membranous crack; internal genitalia, frontal view, anterior receptaculum broken (C), asterisk = opening of the anterior receptaculum connected to the bursa copulatrix; internal wall of the anterior receptaculum (D), inset = pore of a glandular duct." httpUri="https://zenodo.org/record/1207661/files/figure.png" pageId="15" pageNumber="16">Figs 9A‒C</figureCitation>
,
<figureCitation id="13092A42FFEBB1151504FE8FC401FEC1" box="[776,856,332,356]" captionStart="FIGURE 10" captionStartId="13.[151,250,1897,1919]" captionTargetBox="[151,1436,841,1875]" captionTargetId="figure@13.[151,1436,841,1875]" captionTargetPageId="13" captionText="FIGURE 10. SEM images of Ariadna boliviana Simon, 1907, female from Parauapebas (IBSP 193499): anterior receptaculum, lateral (A) and frontal view (B); internal genitalia, frontal view (C); posterior wall of bursa copulatrix, lateral view, origin of the posterior receptaculum (D), arrowhead = median membranous crack; pore of a tubular invagination in the membranous cuticle of the posterior receptaculum (E); posterior receptaculum, internal view, tubular invaginations (F‒G)." httpUri="https://zenodo.org/record/1207663/files/figure.png" pageId="15" pageNumber="16">10C‒G</figureCitation>
,
<figureCitation id="13092A42FFEBB115156FFE8FC4EEFEC1" box="[867,951,332,356]" captionStart="FIGURE 11" captionStartId="14.[151,250,1105,1127]" captionTargetBox="[151,1436,193,1083]" captionTargetId="figure@14.[151,1436,193,1083]" captionTargetPageId="14" captionText="FIGURE 11. Ariadna maxima (Nicolet, 1849), female from Santiago (IBSP 161292), after enzime prepatarion: habitus, dorsal view, without carapaceand dorsal part of the abdomen (A); cephalothorax and anterior region of the abdomen, dorsal view (B); anterior region of the abdomen, dorsal view (C); posterior receptaculum, dorsal view, tubular invaginations (D)." httpUri="https://zenodo.org/record/1207665/files/figure.png" pageId="15" pageNumber="16">11C‒D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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