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<document id="9251E6291EA8DC93E10DDD0592ADEF8C" ID-CLB-Dataset="39051" ID-DOI="10.5281/zenodo.46289" ID-GBIF-Dataset="05d72676-d09d-442f-a042-b9cd3fe7ade2" ID-ISSN="0035-418" ID-Zenodo-Dep="46289" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463490966785" checkinUser="plazi" docAuthor="Katsuyuki Eguchi, Bui Tuan Viet, Emiko Oguri &amp; Seiki Yamane" docDate="2016" docId="039B87BCFFAE8672FEF6FA5D580EF9CC" docLanguage="en" docName="rsz_123_1_5.pdf" docOrigin="Revue suisse de Zoologie 123 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Pheidole leloi Eguchi &amp; Bui, sp. n." docType="treatment" docVersion="12" lastPageNumber="49" masterDocId="FFA2FFC4FFAC8676FFF3FFE95C7EFFAF" masterDocTitle="The first discovery of the “ Pheidole quadricuspis group ” in the Indo-Chinese Peninsula (Insecta: Hymenoptera: Formicidae: Myrmicinae)" masterLastPageNumber="55" masterPageNumber="45" pageNumber="47" updateTime="1698662530584" updateUser="ExternalLinkService">
<mods:mods id="4EE42568D7CD3F6192091601360BFBD7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="67ADFBFB08097EBAAFBB1C5AB3CF50F6">The first discovery of the “ Pheidole quadricuspis group ” in the Indo-Chinese Peninsula (Insecta: Hymenoptera: Formicidae: Myrmicinae)</mods:title>
</mods:titleInfo>
<mods:name id="905F472F07856CE92A3CC4087A80F595" type="personal">
<mods:role id="A1792062C13E5F15CEFB50934E8F022F">
<mods:roleTerm id="7C905CBAD736CC44E76C530436A4B0A3">Author</mods:roleTerm>
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<mods:namePart id="D6A0EBF851ABA2E11932E0D5354AF5F3">Katsuyuki Eguchi</mods:namePart>
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<mods:namePart id="4E331F29B9118D72994ED5499F3745D4">Bui Tuan Viet</mods:namePart>
</mods:name>
<mods:name id="E09732B03FE5C234EF08C721C4F41AFD" type="personal">
<mods:role id="4E30B760CAE76DC09EE73E66111F5345">
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<mods:namePart id="CA15F2208CA8DB75DC4FEFC61F85F3DD">Emiko Oguri</mods:namePart>
</mods:name>
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<mods:role id="FF5DF71C6DE5C4532A2E0964DD602E5E">
<mods:roleTerm id="5E169BD6C126110DF7E34EFF9FE51414">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3573BADDEF15A114610A10AE4A95F29">Seiki Yamane</mods:namePart>
</mods:name>
<mods:typeOfResource id="A48466FD0C268D598C55F182B7E60990">text</mods:typeOfResource>
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<mods:title id="1411D845EB3A05C186567405D3F812FC">Revue suisse de Zoologie</mods:title>
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<mods:part id="6ED73019DB56CC9A60AB9492DCDC580A">
<mods:date id="510D7335D67306EEF77EA33C4E9F25EC">2016</mods:date>
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<treatment id="039B87BCFFAE8672FEF6FA5D580EF9CC" ID-DOI="http://doi.org/10.5281/zenodo.5684612" ID-Zenodo-Dep="5684612" LSID="urn:lsid:plazi:treatment:039B87BCFFAE8672FEF6FA5D580EF9CC" httpUri="http://treatment.plazi.org/id/039B87BCFFAE8672FEF6FA5D580EF9CC" lastPageId="4" lastPageNumber="49" pageId="2" pageNumber="47">
<paragraph id="8B8D36AAFFAE8674FEF6FA5D5EF8FA62" blockId="2.[261,646,1460,1518]" box="[261,646,1460,1485]" pageId="2" pageNumber="47">
<heading id="D0C581C6FFAE8674FEF6FA5D5EF8FA62" bold="true" box="[261,646,1460,1485]" centered="true" fontSize="10" level="3" pageId="2" pageNumber="47" reason="4">
<emphasis id="B946EAB8FFAE8674FEF6FA5D5EF8FA62" bold="true" box="[261,646,1460,1485]" pageId="2" pageNumber="47">
<taxonomicName id="4C324D29FFAE8674FEF6FA5D5E40FA63" authority="Eguchi &amp; Bui" authorityName="Eguchi &amp; Bui" box="[261,574,1460,1484]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="leloi" status="sp. nov.">
<emphasis id="B946EAB8FFAE8674FEF6FA5D5DE5FA63" bold="true" box="[261,411,1460,1484]" italics="true" pageId="2" pageNumber="47">Pheidole leloi</emphasis>
Eguchi &amp; Bui
</taxonomicName>
,
<taxonomicNameLabel id="A27557C3FFAE8674FDB9FA5C5EF8FA62" box="[586,646,1461,1485]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFAE8674FE60FA3C5D87FA41" blockId="2.[261,646,1460,1518]" box="[403,505,1493,1518]" pageId="2" pageNumber="47">
<figureCitation id="13092A2FFFAE8674FE60FA3C5D87FA41" box="[403,505,1493,1518]" captionStart-0="Figs 1 - 5" captionStart-1="Figs 6 - 8" captionStart-2="Figs 9 - 11" captionStart-3="Figs 12 - 13" captionStartId-0="3.[151,192,1872,1893]" captionStartId-1="5.[151,192,1913,1934]" captionStartId-2="6.[157,198,1454,1475]" captionStartId-3="7.[151,192,1433,1454]" captionTargetBox-0="[153,1434,259,1797]" captionTargetBox-1="[151,1436,257,1843]" captionTargetBox-2="[151,1435,257,1414]" captionTargetBox-3="[151,1435,259,1385]" captionTargetId-0="figure@3.[151,1436,257,1800]" captionTargetId-1="figure@5.[151,1436,257,1843]" captionTargetId-2="figure@6.[151,1436,257,1414]" captionTargetId-3="figure@7.[151,1436,257,1385]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Figs 1 - 5. Pheidole leloi sp. nov., major. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view." captionText-1="Figs 6 - 8. Pheidole leloi sp. nov., minor. (6) Head in full-face view. (7) Right antenna. (8) Mesosoma and waist in lateral view." captionText-2="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." captionText-3="Figs 12 - 13. Pheidole leloi sp. nov., queen. (12) Mesosoma and waist in lateral view. (13) Mesosoma and waist in dorsal view." httpUri-0="https://zenodo.org/record/306895/files/figure.png" httpUri-1="https://zenodo.org/record/306901/files/figure.png" httpUri-2="https://zenodo.org/record/306904/files/figure.png" httpUri-3="https://zenodo.org/record/306906/files/figure.png" pageId="2" pageNumber="47">Figs 1-13</figureCitation>
</paragraph>
<subSubSection id="C3286521FFAE8674FF64F9FF5D23F880" pageId="2" pageNumber="47" type="materials_examined">
<paragraph id="8B8D36AAFFAE8674FF64F9FF5D23F880" blockId="2.[151,757,1557,1839]" pageId="2" pageNumber="47">
<emphasis id="B946EAB8FFAE8674FF64F9FF5DBEF983" bold="true" box="[151,448,1557,1582]" pageId="2" pageNumber="47">
<typeStatus id="54898808FFAE8674FF64F9FF5CB1F981" box="[151,207,1558,1582]" pageId="2" pageNumber="47">Type</typeStatus>
material examined:
</emphasis>
<collectionCode id="ED23AE6FFFAE8674FE3FF9FF5E5FF982" box="[460,545,1558,1581]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="2" pageNumber="47">VNMN</collectionCode>
;
<typeStatus id="54898808FFAE8674FDC2F9FC5EF1F983" box="[561,655,1557,1580]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
(major);
<collectingCountry id="F325763AFFAE8674FF64F9DC5C88F9E3" box="[151,246,1589,1612]" name="Vietnam" pageId="2" pageNumber="47">Vietnam</collectingCountry>
, Khanh Hoa, Hon Ba Nature Reserve, 12˚0724”-38”N, 108˚5824”-28”E, ca.
<quantity id="4CCA9B4FFFAE8674FDAFF9BC5E8AF9C2" box="[604,756,1621,1645]" metricMagnitude="3" metricUnit="m" metricValue="1.04" metricValueMax="1.05" metricValueMin="1.03" pageId="2" pageNumber="47" unit="m" value="1040.0" valueMax="1050.0" valueMin="1030.0">1030-1050 m</quantity>
alt.; K. Eguchi leg.;
<date id="FF8C106AFFAE8674FE8BF99C5D93F923" box="[376,493,1653,1676]" pageId="2" pageNumber="47" value="2013-03-16">16/iii/2013</date>
[colony: Eg
<date id="FF8C106AFFAE8674FD8BF99C5E94F923" box="[632,746,1653,1676]" pageId="2" pageNumber="47" value="1916-03-13" valueMax="1916-03-24" valueMin="1916-03-13">16iii13-24</date>
].
<collectionCode id="ED23AE6FFFAE8674FF5FF97E5D7EF901" box="[172,256,1687,1710]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="2" pageNumber="47">VNMN</collectionCode>
,
<collectionCode id="ED23AE6FFFAE8674FEF8F97F5D27F901" box="[267,345,1686,1710]" pageId="2" pageNumber="47">MCZC</collectionCode>
,
<collectionCode id="ED23AE6FFFAE8674FE96F97F5DC7F901" box="[357,441,1686,1710]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="47">MHNG</collectionCode>
,
<collectionCode id="ED23AE6FFFAE8674FE30F97F5E71F901" box="[451,527,1686,1710]" pageId="2" pageNumber="47">ACEG</collectionCode>
;
<typeStatus id="54898808FFAE8674FDE8F97E5EFBF900" box="[539,645,1687,1711]" pageId="2" pageNumber="47" type="paratype">paratypes</typeStatus>
; 8 majors, 9 minors,
<specimenCount id="9D34FD23FFAE8674FEF8F95F5D1DF961" box="[267,355,1718,1742]" pageId="2" pageNumber="47" type="queen">1 queen</specimenCount>
from the same colony as
<typeStatus id="54898808FFAE8674FD7CF95C5E8EF963" box="[655,752,1717,1740]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
; Accession No.
<accessionNumber id="9461AB49FFAE8674FEA4F93F5DB0F942" box="[343,462,1750,1773]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC020537" pageId="2" pageNumber="47">
LC
<date id="FF8C106AFFAE8674FE89F93F5DB0F942" box="[378,462,1750,1773]" pageId="2" pageNumber="47" value="37-02-05">020537</date>
</accessionNumber>
; voucher specimen No. Ext20130410-1 (
<typeStatus id="54898808FFAE8674FEA6F91F5DC8F8A1" box="[341,438,1782,1806]" pageId="2" pageNumber="47" type="paratype">paratype</typeStatus>
) from the same colony as
<typeStatus id="54898808FFAE8674FF64F8FF5C8AF882" box="[151,244,1814,1837]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED23AE6FFFAE8674FEF0F8FE5D2CF880" box="[259,338,1815,1839]" pageId="2" pageNumber="47">ACEG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3286521FFAE8674FF64F8AC586EFDD1" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="8B8D36AAFFAE8674FF64F8AC586EFDD1" blockId="2.[151,756,1861,2077]" lastBlockId="2.[831,1437,258,638]" pageId="2" pageNumber="47">
<emphasis id="B946EAB8FFAE8674FF64F8AC5D6EF8F3" bold="true" box="[151,272,1861,1884]" pageId="2" pageNumber="47">Diagnosis:</emphasis>
In the major, vertex as well as dorsal, lateral and ventrolateral faces of vertexal lobe strongly and coarsely reticulate; promesonotal dome with an inconspicuous transverse ridge on its posterior slope; pronotal spine almost straight; posterior slope and lateral face of the dome, mesopleuron and metapleuron irregularly rugoso-reticulate; propodeal spine somewhat digitiform, with blunt apex; ventral face of petiole with a conspicuous longitudinal carina; anterior half to two thirds of first gastral tergite finely rugoso-punctate. In the minor, posterior part of frons and vertex reticulate; mesosoma relatively densely covered with standing hairs; dorsal face of promesonotal dome weakly punctured, overlain with several rugae, and lateral face of the dome, mesopleuron, metapleuron and dorsal face of propodeum punctate; propodeal spine in lateral view elongate-triangular; petiolar node in lateral view acute at apex, and in posterior view widely and shallowly emarginate at apex.
</paragraph>
</subSubSection>
<paragraph id="8B8D36AAFFAE8674FCCCFD7C5FBDFD03" blockId="2.[831,1437,661,2078]" box="[831,963,661,684]" pageId="2" pageNumber="47">
<heading id="D0C581C6FFAE8674FCCCFD7C5FBDFD03" bold="true" box="[831,963,661,684]" fontSize="10" level="2" pageId="2" pageNumber="47" reason="6">
<emphasis id="B946EAB8FFAE8674FCCCFD7C5FBDFD03" bold="true" box="[831,963,661,684]" pageId="2" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFAE8674FCCCFD5E58FFF811" blockId="2.[831,1437,661,2078]" pageId="2" pageNumber="47">
<emphasis id="B946EAB8FFAE8674FCCCFD5E5FFCFD61" box="[831,898,695,718]" italics="true" pageId="2" pageNumber="47">Major</emphasis>
(
<figureCitation id="13092A2FFFAE8674FC61FD5F5F90FD62" box="[914,1006,694,717]" captionStart="Figs 1 - 5" captionStartId="3.[151,192,1872,1893]" captionTargetBox="[153,1434,259,1797]" captionTargetId="figure@3.[151,1436,257,1800]" captionTargetPageId="3" captionText="Figs 1 - 5. Pheidole leloi sp. nov., major. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view." httpUri="https://zenodo.org/record/306895/files/figure.png" pageId="2" pageNumber="47">Figs 1-5</figureCitation>
): Body dark reddish brown with darker gaster, relatively densely covered with standing hairs (
<figureCitation id="13092A2FFFAE8674FCB4FD1E5F9BFCA1" box="[839,997,759,783]" captionStart="Figs 1 - 5" captionStartId="3.[151,192,1872,1893]" captionTargetBox="[153,1434,259,1797]" captionTargetId="figure@3.[151,1436,257,1800]" captionTargetPageId="3" captionText="Figs 1 - 5. Pheidole leloi sp. nov., major. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view." httpUri="https://zenodo.org/record/306895/files/figure.png" pageId="2" pageNumber="47">Figs 1, 2, 4, 5</figureCitation>
). Head in full-face view subrectangular, with its posterior margin narrowly and shallowly emarginated medially, in lateral view relatively thick, faintly impressed on vertex; frons longitudinally rugoso-reticulate; vertex, and dorsal, lateral and ventrolateral faces of vertexal lobe strongly and coarsely reticulate; gena longitudinally rugose; frontal carina almost absent or present just as rugula(e); antennal scrobe absent; median portion of clypeus almost smooth or faintly rugose longitudinally, with a few longitudinal rugae laterally; median longitudinal carina of clypeus relatively distinct; hypostoma with a pair of stout submedian processes (“SM” in
<figureCitation id="13092A2FFFAE8674FAA5FBB259EDFBDC" box="[1366,1427,1115,1139]" captionStart="Figs 1 - 5" captionStartId="3.[151,192,1872,1893]" captionTargetBox="[153,1434,259,1797]" captionTargetId="figure@3.[151,1436,257,1800]" captionTargetPageId="3" captionText="Figs 1 - 5. Pheidole leloi sp. nov., major. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view." httpUri="https://zenodo.org/record/306895/files/figure.png" pageId="2" pageNumber="47">Fig. 3</figureCitation>
) but without median process; lateral processes of hypostoma present but tiny (“L” in
<figureCitation id="13092A2FFFAE8674FB80FB7258CBFB1B" box="[1139,1205,1179,1204]" captionStart="Figs 1 - 5" captionStartId="3.[151,192,1872,1893]" captionTargetBox="[153,1434,259,1797]" captionTargetId="figure@3.[151,1436,257,1800]" captionTargetPageId="3" captionText="Figs 1 - 5. Pheidole leloi sp. nov., major. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view." httpUri="https://zenodo.org/record/306895/files/figure.png" pageId="2" pageNumber="47">Fig. 3</figureCitation>
); masticatory margin of mandible with apical and preapical teeth, and a tooth in front of basal angle; outer surface of mandible smooth except its base, scattering relatively long decumbent/appressed hairs; antenna with 3-segmented club; maximal diameter of eye longer than or almost as long as antennal segment X. Promesonotal dome with an inconspicuous transverse ridge on its posterior slope; pronotal spine long, slender, almost straight, with pointed apex; anterior slope of promesonotal dome largely smooth with several transverse rugae; posterior slope and lateral face of the dome, mesopleuron and metapleuron irregularly rugoso-reticulate; propodeum irregularly rugoso-reticulate dorsally, longitudinally rugose laterally, and transversely rugose posteriorly; propodeal spine somewhat digitiform, with blunt apex. Ventral surface of midcoxa and hindcoxa smooth. Petiole longer than postpetiole (excluding helcium); petiolar node in lateral view blunt at apex, in posterior view widely and shallowly emarginate at apex, in dorsal view laterally with a narrow flange; ventral face of petiole with a conspicuous longitudinal carina; postpetiole in dorsal view much broader than long, somewhat spindle-shaped; its anteroventral part forming a sharp transverse ridge. Anterior half to two thirds of first gastral tergite finely rugoso-punctate.
</paragraph>
<paragraph id="8B8D36AAFFAE8672FCCCF82C5E0CFE76" blockId="2.[831,1437,661,2078]" lastBlockId="4.[151,756,258,2078]" lastPageId="4" lastPageNumber="49" pageId="2" pageNumber="47">
<typeStatus id="54898808FFAE8674FCCCF82C5FDCF873" box="[831,930,1989,2012]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
(major): HL,
<quantity id="4CCA9B4FFFAE8674FBBAF82C58CFF872" box="[1097,1201,1989,2013]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="2" pageNumber="47" unit="mm" value="2.36">2.36 mm</quantity>
; HW,
<quantity id="4CCA9B4FFFAE8674FB0DF82C5918F872" box="[1278,1382,1989,2013]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" pageId="2" pageNumber="47" unit="mm" value="2.62">2.62 mm</quantity>
; SL,
<quantity id="4CCA9B4FFFAE8674FCCCF80F5FDDF852" box="[831,931,2022,2046]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="47" unit="mm" value="1.22">1.22 mm</quantity>
; ML,
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; PW,
<quantity id="4CCA9B4FFFAE8674FB78F80F5891F852" box="[1163,1263,2022,2046]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="47" unit="mm" value="1.18">1.18 mm</quantity>
; HFL
<quantity id="4CCA9B4FFFAE8674FAC7F80C59E6F852" box="[1332,1432,2021,2045]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="2" pageNumber="47" unit="mm" value="1.76">1.76 mm</quantity>
; PtL
<quantity id="4CCA9B4FFFAE8674FC81F7EF5FA5F7B1" box="[882,987,2054,2078]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="47" unit="mm" value="0.58">0.58 mm</quantity>
; PtW
<quantity id="4CCA9B4FFFAE8674FBDAF7EF58ECF7B1" box="[1065,1170,2054,2078]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="2" pageNumber="47" unit="mm" value="0.47">0.47 mm</quantity>
; PPtL
<quantity id="4CCA9B4FFFAE8674FB17F7EF5933F7B1" box="[1252,1357,2054,2078]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="47" unit="mm" value="0.45">0.45 mm</quantity>
; PPtW
<quantity id="4CCA9B4FFFA88672FF64FEEA5C82FEB5" box="[151,252,259,283]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="49" unit="mm" value="0.77">0.77 mm</quantity>
; CI, 111; SI, 46; MI, 193; HFI, 67; PtI1, 130; PtI2, 62.
<typeStatus id="54898808FFA88672FF04FECA5D2AFE94" box="[247,340,291,315]" pageId="4" pageNumber="49" type="paratype">Paratype</typeStatus>
majors (n=8): HL,
<quantity id="4CCA9B4FFFA88672FDD2FECA5EC7FE95" box="[545,697,291,315]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" metricValueMax="2.49" metricValueMin="2.39" pageId="4" pageNumber="49" unit="mm" value="2.44" valueMax="2.49" valueMin="2.39">2.39-2.49 mm</quantity>
, HW,
<quantity id="4CCA9B4FFFA88672FF64FEAB5D4AFEF5" box="[151,308,322,346]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" metricValueMax="2.71" metricValueMin="2.55" pageId="4" pageNumber="49" unit="mm" value="2.63" valueMax="2.71" valueMin="2.55">2.55-2.71 mm</quantity>
; SL,
<quantity id="4CCA9B4FFFA88672FE82FEAA5E70FEF5" box="[369,526,323,347]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.24" metricValueMin="1.18" pageId="4" pageNumber="49" unit="mm" value="1.21" valueMax="1.24" valueMin="1.18">1.18-1.24 mm</quantity>
; ML,
<quantity id="4CCA9B4FFFA88672FDA0FEAB5E8EFEF5" box="[595,752,322,346]" metricMagnitude="-3" metricUnit="m" metricValue="2.29" metricValueMax="2.35" metricValueMin="2.23" pageId="4" pageNumber="49" unit="mm" value="2.29" valueMax="2.35" valueMin="2.23">2.23-2.35 mm</quantity>
; PW,
<quantity id="4CCA9B4FFFA88672FF21FE8B5D0DFED5" box="[210,371,354,378]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.2" metricValueMin="1.16" pageId="4" pageNumber="49" unit="mm" value="1.18" valueMax="1.2" valueMin="1.16">1.16-1.20 mm</quantity>
; HFL,
<quantity id="4CCA9B4FFFA88672FE3EFE8A5E11FED5" box="[461,623,355,379]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" metricValueMax="1.8" metricValueMin="1.74" pageId="4" pageNumber="49" unit="mm" value="1.77" valueMax="1.8" valueMin="1.74">1.74-1.80 mm</quantity>
; PtL,
<quantity id="4CCA9B4FFFA88672FD4FFE8B5D7EFE35" metricMagnitude="-4" metricUnit="m" metricValue="6.4" metricValueMax="6.7" metricValueMin="6.1" pageId="4" pageNumber="49" unit="mm" value="0.64" valueMax="0.67" valueMin="0.61">0.61- 0.67 mm</quantity>
; PtW,
<quantity id="4CCA9B4FFFA88672FEA0FE6B5D8AFE35" box="[339,500,386,410]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.2" metricValueMin="4.6" pageId="4" pageNumber="49" unit="mm" value="0.49" valueMax="0.52" valueMin="0.46">0.46-0.52 mm</quantity>
; PPtL,
<quantity id="4CCA9B4FFFA88672FDBCFE6B5E8EFE35" box="[591,752,386,410]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.9" metricValueMin="4.5" pageId="4" pageNumber="49" unit="mm" value="0.47" valueMax="0.49" valueMin="0.45">0.45-0.49 mm</quantity>
; PPtW,
<quantity id="4CCA9B4FFFA88672FF11FE4B5D00FE15" box="[226,382,418,442]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.6" metricValueMin="7.7" pageId="4" pageNumber="49" unit="mm" value="0.815" valueMax="0.86" valueMin="0.77">0.77-0.86 mm</quantity>
; CI, 107-110; SI, 45-47; MI, 188- 198; HFI, 65-68; PtI1, 128-145; PtI2, 60-63.
</paragraph>
<caption id="DF4D6622FFAF8675FF64F8B95896F82E" httpUri="https://zenodo.org/record/306895/files/figure.png" pageId="3" pageNumber="48" targetBox="[153,1434,259,1797]" targetPageId="3">
<paragraph id="8B8D36AAFFAF8675FF64F8B95896F82E" blockId="3.[151,1436,1872,1923]" pageId="3" pageNumber="48">
Figs 1-5.
<taxonomicName id="4C324D29FFAF8675FF05F8B95D02F8CA" box="[246,380,1872,1893]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="leloi" status="sp. nov.">
<emphasis id="B946EAB8FFAF8675FF05F8B95D02F8CA" box="[246,380,1872,1893]" italics="true" pageId="3" pageNumber="48">Pheidole leloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557C3FFAF8675FE72F8B85DB0F8C9" box="[385,462,1873,1894]" pageId="3" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C324D29FFAF8675FE25F8B95E6EF8CA" box="[470,528,1872,1893]" pageId="3" pageNumber="48" rank="species" species="major">major</taxonomicName>
. (1) Head in full-face view. (2) Head in lateral view. (3) Hypostoma in ventral view: (SM) submedian processes; (L) lateral processes. (4) Mesosoma and waist in lateral view. (5) Body in dorsal view.
</paragraph>
</caption>
<paragraph id="8B8D36AAFFA88672FF64FE0D5E0FFB54" blockId="4.[151,756,258,2078]" pageId="4" pageNumber="49">
<emphasis id="B946EAB8FFA88672FF64FE0D5CA4FE54" box="[151,218,484,507]" italics="true" pageId="4" pageNumber="49">Minor</emphasis>
(
<figureCitation id="13092A2FFFA88672FF1FFE0B5D32FE56" box="[236,332,482,505]" captionStart-0="Figs 6 - 8" captionStart-1="Figs 9 - 11" captionStartId-0="5.[151,192,1913,1934]" captionStartId-1="6.[157,198,1454,1475]" captionTargetBox-0="[151,1436,257,1843]" captionTargetBox-1="[151,1435,257,1414]" captionTargetId-0="figure@5.[151,1436,257,1843]" captionTargetId-1="figure@6.[151,1436,257,1414]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs 6 - 8. Pheidole leloi sp. nov., minor. (6) Head in full-face view. (7) Right antenna. (8) Mesosoma and waist in lateral view." captionText-1="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." httpUri-0="https://zenodo.org/record/306901/files/figure.png" httpUri-1="https://zenodo.org/record/306904/files/figure.png" pageId="4" pageNumber="49">Figs 6-9</figureCitation>
): Body dark reddish brown, relatively densely covered with standing hairs (
<figureCitation id="13092A2FFFA88672FDC1FDEB5EE9FDB4" box="[562,663,514,539]" captionStart="Figs 6 - 8" captionStartId="5.[151,192,1913,1934]" captionTargetBox="[151,1436,257,1843]" captionTargetId="figure@5.[151,1436,257,1843]" captionTargetPageId="5" captionText="Figs 6 - 8. Pheidole leloi sp. nov., minor. (6) Head in full-face view. (7) Right antenna. (8) Mesosoma and waist in lateral view." httpUri="https://zenodo.org/record/306901/files/figure.png" pageId="4" pageNumber="49">Figs 6, 8</figureCitation>
,
<figureCitation id="13092A2FFFA88672FD55FDEA5EC8FDB4" box="[678,694,515,539]" captionStart="Figs 9 - 11" captionStartId="6.[157,198,1454,1475]" captionTargetBox="[151,1435,257,1414]" captionTargetId="figure@6.[151,1436,257,1414]" captionTargetPageId="6" captionText="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." httpUri="https://zenodo.org/record/306904/files/figure.png" pageId="4" pageNumber="49">9</figureCitation>
). Anteromedian part of frons largely smooth; posterior part of frons and vertex reticulate; area between antennal insertion and eye, and gena rugose; preoccipital carina conspicuous dorsally and laterally; median portion of clypeus slightly punctate, with a median longitudinal carina; masticatory margin of mandible with apical and preapical teeth followed by several small teeth; 1 or 2 small denticles between the preapical tooth and 3rd tooth; outer surface of mandible longitudinally rugose, scattered with long decumbent hairs; antenna with 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome with a pair of spines that are long, slender and pointed apically; posterior slope of the dome with an inconspicuous transverse ridge; dorsal face of the dome weakly punctate, overlain with several rugae; lateral face of the dome, mesopleuron, metapleuron and dorsal face of propodeum punctate; propodeal spine in lateral view elongate-triangular. Petiole a little longer than postpetiole (excluding helcium); petiolar node in lateral view acute at apex, and in posterior view widely and shallowly emarginate at apex; ventral face of petiole without any process and longitudinal carina.
</paragraph>
<paragraph id="8B8D36AAFFA88672FF64FAEC5D68FA14" blockId="4.[151,756,258,2078]" pageId="4" pageNumber="49">
HL,
<quantity id="4CCA9B4FFFA88672FF21FAEC5D08FAB3" box="[210,374,1285,1309]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="10.1" metricValueMin="9.7" pageId="4" pageNumber="49" unit="mm" value="0.99" valueMax="1.01" valueMin="0.97">0.97-1.01 mm</quantity>
, HW,
<quantity id="4CCA9B4FFFA88672FE3FFAEC5E0FFAB3" box="[460,625,1285,1309]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" metricValueMax="9.3" metricValueMin="8.9" pageId="4" pageNumber="49" unit="mm" value="0.91" valueMax="0.93" valueMin="0.89">0.89-0.93 mm</quantity>
; SL,
<quantity id="4CCA9B4FFFA88672FD4FFAEC5C84FA93" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.27" metricValueMin="1.21" pageId="4" pageNumber="49" unit="mm" value="1.24" valueMax="1.27" valueMin="1.21">1.21- 1.27 mm</quantity>
; ML,
<quantity id="4CCA9B4FFFA88672FECFFACD5DA8FA93" box="[316,470,1316,1340]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" metricValueMax="1.5" metricValueMin="1.42" pageId="4" pageNumber="49" unit="mm" value="1.46" valueMax="1.5" valueMin="1.42">1.42-1.50 mm</quantity>
; PW,
<quantity id="4CCA9B4FFFA88672FDE6FACD5ECEFA93" box="[533,688,1316,1340]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="6.5" metricValueMin="6.1" pageId="4" pageNumber="49" unit="mm" value="0.63" valueMax="0.65" valueMin="0.61">0.61-0.65 mm</quantity>
; HFL,
<quantity id="4CCA9B4FFFA88672FF64FAAD5D4CFAF3" box="[151,306,1348,1372]" metricMagnitude="-3" metricUnit="m" metricValue="1.395" metricValueMax="1.43" metricValueMin="1.36" pageId="4" pageNumber="49" unit="mm" value="1.395" valueMax="1.43" valueMin="1.36">1.36-1.43 mm</quantity>
; PtL,
<quantity id="4CCA9B4FFFA88672FE80FAAC5E70FAF3" box="[371,526,1349,1373]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="4" pageNumber="49" unit="mm" value="0.325" valueMax="0.34" valueMin="0.31">0.31-0.34 mm</quantity>
; PtW,
<quantity id="4CCA9B4FFFA88672FDA6FAAD5E8EFAF3" box="[597,752,1348,1372]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="4" pageNumber="49" unit="mm" value="0.15" valueMax="0.16" valueMin="0.14">0.14-0.16 mm</quantity>
; PPtL,
<quantity id="4CCA9B4FFFA88672FF2DFA8C5D05FAD3" box="[222,379,1381,1405]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="4" pageNumber="49" unit="mm" value="0.285" valueMax="0.29" valueMin="0.28">0.28-0.29 mm</quantity>
; PPtW,
<quantity id="4CCA9B4FFFA88672FE24FA8C5E0AFAD3" box="[471,628,1381,1405]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="4" pageNumber="49" unit="mm" value="0.325" valueMax="0.34" valueMin="0.31">0.31-0.34 mm</quantity>
; CI, 91-93; SI, 134-138; MI, 227-235; HFI, 152-155; PtI1, 111-119; PtI2, 45-49.
</paragraph>
<paragraph id="8B8D36AAFFA88672FF64FA2C5FBCFB2E" blockId="4.[151,756,258,2078]" lastBlockId="4.[831,1437,258,1282]" pageId="4" pageNumber="49">
<emphasis id="B946EAB8FFA88672FF64FA2C5CA3FA73" box="[151,221,1477,1500]" italics="true" pageId="4" pageNumber="49">Queen</emphasis>
(
<figureCitation id="13092A2FFFA88672FF19FA2D5D1FFA72" box="[234,353,1476,1501]" captionStart-0="Figs 9 - 11" captionStart-1="Figs 12 - 13" captionStartId-0="6.[157,198,1454,1475]" captionStartId-1="7.[151,192,1433,1454]" captionTargetBox-0="[151,1435,257,1414]" captionTargetBox-1="[151,1435,259,1385]" captionTargetId-0="figure@6.[151,1436,257,1414]" captionTargetId-1="figure@7.[151,1436,257,1385]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." captionText-1="Figs 12 - 13. Pheidole leloi sp. nov., queen. (12) Mesosoma and waist in lateral view. (13) Mesosoma and waist in dorsal view." httpUri-0="https://zenodo.org/record/306904/files/figure.png" httpUri-1="https://zenodo.org/record/306906/files/figure.png" pageId="4" pageNumber="49">Figs 10-13</figureCitation>
): Body dark reddish brown with dark- er gaster, relatively densely covered with standing hairs (
<figureCitation id="13092A2FFFA88672FF6CF9EC5C8CF9B2" box="[159,242,1541,1565]" captionStart="Figs 9 - 11" captionStartId="6.[157,198,1454,1475]" captionTargetBox="[151,1435,257,1414]" captionTargetId="figure@6.[151,1436,257,1414]" captionTargetPageId="6" captionText="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." httpUri="https://zenodo.org/record/306904/files/figure.png" pageId="4" pageNumber="49">Figs 10</figureCitation>
,
<figureCitation id="13092A2FFFA88672FF0DF9EC5D3AF9B2" box="[254,324,1541,1565]" captionStart="Figs 12 - 13" captionStartId="7.[151,192,1433,1454]" captionTargetBox="[151,1435,259,1385]" captionTargetId="figure@7.[151,1436,257,1385]" captionTargetPageId="7" captionText="Figs 12 - 13. Pheidole leloi sp. nov., queen. (12) Mesosoma and waist in lateral view. (13) Mesosoma and waist in dorsal view." httpUri="https://zenodo.org/record/306906/files/figure.png" pageId="4" pageNumber="49">12, 13</figureCitation>
). Head in full-face view subtrapezoidal, with its posterior margin broadly and shallowly concave, in lateral view relatively thick, not impressed on vertex, with its posterior margin narrowly and shallowly emarginated medially; frons longitudinally rugoso-reticulate; vertex, and dorsal, lateral and ventrolateral faces of vertexal lobe strongly and coarsely reticulate; gena longitudinally rugose; frontal carina present just as rugula(e); antennal scrobe absent; median portion of clypeus faintly rugose longitudinally, with a few longitudinal rugae laterally; median longitudinal carina of clypeus relatively conspicuous; hypostoma with a pair of stout submedian processes but without median process; lateral processes of hypostoma present but tiny; masticatory margin of mandible with apical and preapical teeth, and a tooth in front of basal angle; outer surface of mandible smooth except its base, scattered with relatively long decumbent to appressed hairs; antenna with 3-segmented club; maximal diameter of eye longer than or almost as long as antennal segment X; median ocellus in full-face view located a little behind the level of the posterior margin of compound eye; maximum diameter of median ocellus a little longer than the distance between the median and lateral ocelli (
<figureCitation id="13092A2FFFA88672FC26FE2A585AFE73" box="[981,1060,451,476]" captionStart="Figs 9 - 11" captionStartId="6.[157,198,1454,1475]" captionTargetBox="[151,1435,257,1414]" captionTargetId="figure@6.[151,1436,257,1414]" captionTargetPageId="6" captionText="Figs 9 - 11. Pheidole leloi sp. nov. (9) Minor, mesosoma and waist in dorsal view. (10) Queen, head in full-face view. (11) Queen, ocelli." httpUri="https://zenodo.org/record/306904/files/figure.png" pageId="4" pageNumber="49">Fig. 11</figureCitation>
). Mesosoma fully segmented; pronotum almost smooth anteriorly and reticulate laterally; pronotal spine stout and straight, directing anterolaterad; mesoscutum longitudinally rugose medially and irregularly rugoso-reticulate laterally; parapsidal line weakly recognized; scuto-scutellar suture present as a deep and broad impression; mesoscutellum largely smooth dorsally; mesopleuron irregularly rugoso-reticulate, subdivid- ed into anepisternum and katepisternum; metapleuron irregularly rugoso-reticulate; propodeum longitudinally rugose laterally, and transversely rugose posteriorly; propodeal spine somewhat digitiform, with blunt apex. Ventral surface of midcoxa and hindcoxa smooth. Petiole longer than postpetiole (excluding helcium); petiolar node in lateral view blunt at apex, in posterior view widely and shallowly emarginate at apex, in dorsal view laterally with a narrow flange; ventral face of petiole with a conspicuous longitudinal carina; postpetiole in dorsal view much broader than long, somewhat spindle-shaped; its anteroventral part forming a sharp transverse ridge. Anterior three fifths of first gastral tergite finely rugoso-punctate.
</paragraph>
<paragraph id="8B8D36AAFFA88672FCCCFB635887FAAD" blockId="4.[831,1437,258,1282]" pageId="4" pageNumber="49">
HL,
<quantity id="4CCA9B4FFFA88672FC9DFB635FB3FB0E" box="[878,973,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="49" unit="mm" value="2.1">2.10 mm</quantity>
, HW,
<quantity id="4CCA9B4FFFA88672FBF8FB605815FB0E" box="[1035,1131,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="49" unit="mm" value="2.5">2.50 mm</quantity>
; SL,
<quantity id="4CCA9B4FFFA88672FB6EFB635883FB0E" box="[1181,1277,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="49" unit="mm" value="1.21">1.21 mm</quantity>
; ML,
<quantity id="4CCA9B4FFFA88672FACBFB6359E6FB0E" box="[1336,1432,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="3.19" pageId="4" pageNumber="49" unit="mm" value="3.19">3.19 mm</quantity>
; MSW,
<quantity id="4CCA9B4FFFA88672FC7DFB435F8AFB6E" box="[910,1012,1194,1218]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="4" pageNumber="49" unit="mm" value="2.03">2.03 mm</quantity>
; HFL,
<quantity id="4CCA9B4FFFA88672FBB4FB4358D3FB6E" box="[1095,1197,1194,1218]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="49" unit="mm" value="1.9">1.90 mm</quantity>
; PtL,
<quantity id="4CCA9B4FFFA88672FB07FB435924FB6E" box="[1268,1370,1194,1218]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="49" unit="mm" value="0.82">0.82 mm</quantity>
; PtW,
<quantity id="4CCA9B4FFFA88672FCCCFB235FDDFB4D" box="[831,931,1226,1250]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="4" pageNumber="49" unit="mm" value="0.72">0.72 mm</quantity>
; PPtL,
<quantity id="4CCA9B4FFFA88672FC07FB205829FB4D" box="[1012,1111,1225,1250]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="49" unit="mm" value="0.6">0.60 mm</quantity>
; PPtW,
<quantity id="4CCA9B4FFFA88672FB5DFB23596CFB4D" box="[1198,1298,1226,1250]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="49" unit="mm" value="0.91">0.91 mm</quantity>
; CI, 119; SI, 48; MI, 157; HFI, 76; PtI1, 136; PtI2, 79.
</paragraph>
<paragraph id="8B8D36AAFFA88672FCCCFAF05849FA3D" blockId="4.[831,1436,1305,1427]" pageId="4" pageNumber="49">
<emphasis id="B946EAB8FFA88672FCCCFAF05FBFFA9F" bold="true" box="[831,961,1305,1328]" pageId="4" pageNumber="49">Bionomics:</emphasis>
The
<typeStatus id="54898808FFA88672FBF8FAF35844FA9D" box="[1035,1082,1306,1330]" pageId="4" pageNumber="49">type</typeStatus>
series (a single colony) was collected from rotten wood on the forest floor of a relatively disturbed patch of an evergreen forest at an altitude of ca.
<quantity id="4CCA9B4FFFA88672FC2BFA92584DFA3D" box="[984,1075,1403,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="49" unit="m" value="1000.0">1,000 m</quantity>
.
</paragraph>
<paragraph id="8B8D36AAFFA88672FCCCFA335854FA5D" blockId="4.[831,1066,1498,1522]" box="[831,1066,1498,1522]" pageId="4" pageNumber="49">
<heading id="D0C581C6FFA88672FCCCFA335854FA5D" allCaps="true" bold="true" box="[831,1066,1498,1522]" fontSize="10" level="1" pageId="4" pageNumber="49" reason="2">
<emphasis id="B946EAB8FFA88672FCCCFA335854FA5D" bold="true" box="[831,1066,1498,1522]" pageId="4" pageNumber="49">SIMILAR SPECIES</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFA88672FCCCF9E3580EF9CC" blockId="4.[831,1436,1546,1635]" pageId="4" pageNumber="49">
<taxonomicName id="4C324D29FFA88672FCCCF9E35FADF98E" box="[831,979,1546,1569]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="leloi">
<emphasis id="B946EAB8FFA88672FCCCF9E35FADF98E" box="[831,979,1546,1569]" italics="true" pageId="4" pageNumber="49">Pheidole leloi</emphasis>
</taxonomicName>
can be morphologically well distinguished from
<taxonomicName id="4C324D29FFA88672FC89F9C55860F9EE" box="[890,1054,1578,1603]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="quadricuspis">
<emphasis id="B946EAB8FFA88672FC89F9C55860F9EE" box="[890,1054,1578,1603]" italics="true" pageId="4" pageNumber="49">P. quadricuspis</emphasis>
</taxonomicName>
and its named allies as follows (see also “Diagnosis” of
<taxonomicName id="4C324D29FFA88672FBE4F9A5581EF9CE" box="[1047,1120,1610,1635]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="leloi">
<emphasis id="B946EAB8FFA88672FBE4F9A5581EF9CE" box="[1047,1120,1610,1635]" italics="true" pageId="4" pageNumber="49">P. leloi</emphasis>
</taxonomicName>
).
</paragraph>
</treatment>
</document>