treatments-xml/data/03/9B/87/039B87AB004EFFC037E776EBFC081896.xml
2024-06-21 12:22:17 +02:00

184 lines
21 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="07C98C503295A87B2A78912E980BFA43" ID-DOI="10.11646/zootaxa.3686.2.1" ID-GBIF-Dataset="4d62d5da-3890-4af3-9758-674a13b43a79" ID-ISSN="1175-5326" ID-Zenodo-Dep="221213" ID-ZooBank="1AF48240-6A27-452E-9D32-9C6D45C715E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460646465142" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Kurina, Olavi" docDate="2013" docId="039B87AB004EFFC037E776EBFC081896" docLanguage="en" docName="zt03686p139.pdf" docOrigin="Zootaxa 3686 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manota papillosa Hippa &amp; Kurina, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="124" masterDocId="FFA2FFD3005BFFD73770753FFF811A31" masterDocTitle="New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region" masterLastPageNumber="139" masterPageNumber="101" pageNumber="122" updateTime="1698337046318" updateUser="plazi">
<mods:mods id="3946642DA7A1A1B06CE866704ABB9826" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9DA1E2ADD91E52A7C0C8A7EE1920740">
<mods:title id="583C0431E636AE94371A9B561FD30423">New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region</mods:title>
</mods:titleInfo>
<mods:name id="BAB05DD05B0BA92698E0896EE64C6276" type="personal">
<mods:role id="271E4F0A74F50CB3F9292E3328446E90">
<mods:roleTerm id="8FEBCB51C472D8A13B9B709D8BA0343B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE6AAD855F9646F298A203B7364704FB">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:name id="75008E06DE8BF3C30211976FC216DE47" type="personal">
<mods:role id="CC6040282A729FD5E26E67FA6FDD1E86">
<mods:roleTerm id="4068F8A3F5DFE7161B3F749A794A4DC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A86FEC6B147BE3CB3AB4AACD0B29D65">Kurina, Olavi</mods:namePart>
</mods:name>
<mods:typeOfResource id="08B6728CF11D0CD79E513E2FB5122871">text</mods:typeOfResource>
<mods:relatedItem id="B97C21D5867643865A261D413FC8EC00" type="host">
<mods:titleInfo id="46709F6BFE6F6024F7C1E82F78D8E60B">
<mods:title id="102BCE89D7B06C163960DC278DC1C6B8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6FC9DE1180C2231919892A9E7E2926B2">
<mods:date id="4B3F3EB11D1A1B3A01280BFCF0F04352">2013</mods:date>
<mods:detail id="2B2A7792220578C75B68781FEB75D790" type="volume">
<mods:number id="E9FBDD5D17E5F37802CEB3CAD40FC708">3686</mods:number>
</mods:detail>
<mods:detail id="758917AD2F5BB413383D0E10800BF693" type="issue">
<mods:number id="65651140E5595C4271E99D470CB4DC34">2</mods:number>
</mods:detail>
<mods:extent id="59A06772A43C5C7B5573872BEA3B0656" unit="page">
<mods:start id="2EBAADDF2B55960A9DA56419CE7218AC">101</mods:start>
<mods:end id="14F1E331899529BE818599AE3CBD8325">139</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="823E447CA3403DB96FBC1D1CEFB285BC">journal article</mods:classification>
<mods:identifier id="C79A44A98DCF77639692EA02B83BEA2C" type="DOI">10.11646/zootaxa.3686.2.1</mods:identifier>
<mods:identifier id="CB64CDCC8872E08DBDEB9EAD553F3ABF" type="GBIF-Dataset">4d62d5da-3890-4af3-9758-674a13b43a79</mods:identifier>
<mods:identifier id="0C081A7BFAD3F12D6E081AEB5E1F23C5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="973E656AAF10815B3B0AB2FD998B1B68" type="Zenodo-Dep">221213</mods:identifier>
<mods:identifier id="137465E86F02796952618437018761EE" type="ZooBank">1AF48240-6A27-452E-9D32-9C6D45C715E3</mods:identifier>
</mods:mods>
<treatment id="039B87AB004EFFC037E776EBFC081896" ID-DOI="http://doi.org/10.5281/zenodo.6155078" ID-GBIF-Taxon="119572321" ID-Zenodo-Dep="6155078" LSID="urn:lsid:plazi:treatment:039B87AB004EFFC037E776EBFC081896" httpUri="http://treatment.plazi.org/id/039B87AB004EFFC037E776EBFC081896" lastPageId="23" lastPageNumber="124" pageId="21" pageNumber="122">
<subSubSection id="C3286536004EFFC237E776EBFE1C1DEB" pageId="21" pageNumber="122" type="nomenclature">
<paragraph id="8B8D36BD004EFFC237E776EBFE3319DC" blockId="21.[151,434,979,1038]" box="[151,434,979,1006]" pageId="21" pageNumber="122">
<heading id="D0C581D1004EFFC237E776EBFE3319DC" bold="true" box="[151,434,979,1006]" fontSize="11" level="1" pageId="21" pageNumber="122" reason="1">
<emphasis id="B946EAAF004EFFC237E776EBFE3319DC" bold="true" box="[151,434,979,1006]" pageId="21" pageNumber="122">
<taxonomicName id="4C324D3E004EFFC237E776EBFEE819DF" ID-CoL="8GTZ6" box="[151,361,980,1006]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="papillosa" status="sp. nov.">
<emphasis id="B946EAAF004EFFC237E776EBFEE819DF" bold="true" box="[151,361,980,1006]" italics="true" pageId="21" pageNumber="122">Manota papillosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557D4004EFFC2360076ECFE3319DC" box="[368,434,979,1005]" pageId="21" pageNumber="122" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36BD004EFFC237E776C9FEAC1E3F" blockId="21.[151,434,979,1038]" box="[151,301,1014,1038]" pageId="21" pageNumber="122">
<figureCitation id="13092A38004EFFC237E776C9FF701E3F" box="[151,241,1014,1038]" captionStart="FIGURE 12" captionStartId="22.[151,250,1875,1898]" captionTargetBox="[170,1402,206,1839]" captionTargetId="figure@22.[151,1436,195,1851]" captionTargetPageId="22" captionText="FIGURE 12. Manota papillosa sp. n. (holotype). A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221224/files/figure.png" pageId="21" pageNumber="122">Figs. 12</figureCitation>
AC
</paragraph>
<paragraph id="8B8D36BD004EFFC237E77101FE251D87" blockId="21.[151,1437,1086,2010]" pageId="21" pageNumber="122">
Male.
<emphasis id="B946EAAF004EFFC237927101FEB51E66" bold="true" box="[226,308,1086,1111]" pageId="21" pageNumber="122">Colour</emphasis>
. The specimen is extremely faded and the colours are difficult to estimate. Head brownish, face yellow. Antenna pale brownish. Clypeus and mouthparts pale yellowish. Thorax yellow, medial part of scutum and scutellum brownish. Legs unicolorous yellowish. Wing pale brownish; haltere yellow with dark brown knob. Abdominal tergites yellowish brown, sternites paler than tergites. All setosity pale, yellowish or brownish.
<emphasis id="B946EAAF004EFFC232257194FA141EF2" bold="true" box="[1365,1429,1195,1219]" pageId="21" pageNumber="122">Head</emphasis>
. Antennal flagellomere 4 similar to
<figureCitation id="13092A38004EFFC2354571F1FD081ED7" box="[565,649,1230,1254]" captionStart="FIGURE 10" captionStartId="18.[151,250,1982,2005]" captionTargetBox="[162,1416,218,1953]" captionTargetId="figure@18.[151,1436,195,1964]" captionTargetPageId="18" captionText="FIGURE 10. Manota micula sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221222/files/figure.png" pageId="21" pageNumber="122">Fig. 10</figureCitation>
A (
<taxonomicName id="4C324D3E004EFFC235CD71EFFCB21ED7" box="[701,819,1230,1255]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="micula">
<emphasis id="B946EAAF004EFFC235CD71EFFCB21ED7" box="[701,819,1230,1255]" italics="true" pageId="21" pageNumber="122">M. micula</emphasis>
</taxonomicName>
). Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.2 times longer than palpomere 4. Number of strong postocular setae 9.
<emphasis id="B946EAAF004EFFC234FA7028FC621F1E" bold="true" box="[906,995,1303,1327]" pageId="21" pageNumber="122">Thorax</emphasis>
. Anepisternum setose, with 28 setae; anterior basalare non-setose; preepisternum 2 non-setose; laterotergite non-setose, episternum 3 setose, with 8 setae.
<emphasis id="B946EAAF004EFFC237AC7060FE951F46" bold="true" box="[220,276,1375,1399]" pageId="21" pageNumber="122">Legs</emphasis>
. Mid and hind tibial organs absent.
<emphasis id="B946EAAF004EFFC235D67061FD661F46" bold="true" box="[678,743,1374,1399]" pageId="21" pageNumber="122">Wing</emphasis>
. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 extending to the level of the tip of R1; wing length
<quantity id="4CCA9B58004EFFC2348A70BDFBD71FAB" box="[1018,1110,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="122" unit="mm" value="1.5">1.5 mm</quantity>
.
<emphasis id="B946EAAF004EFFC2331370BDFB7F1FAA" bold="true" box="[1123,1278,1410,1435]" pageId="21" pageNumber="122">Hypopygium</emphasis>
(stained with Chlorasol black).
<figureCitation id="13092A38004EFFC236167099FE451F8F" box="[358,452,1446,1470]" captionStart="FIGURE 12" captionStartId="22.[151,250,1875,1898]" captionTargetBox="[170,1402,206,1839]" captionTargetId="figure@22.[151,1436,195,1851]" captionTargetPageId="22" captionText="FIGURE 12. Manota papillosa sp. n. (holotype). A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221224/files/figure.png" pageId="21" pageNumber="122">Figs. 12</figureCitation>
AC: Sternite 9 laterally fused with gonocoxa, posterior margin transverse, not just reaching the middle length of gonocoxa, anterior margin deeply incised, the setae similar to the adjacent ventral setae of gonocoxa. Ventral mesial margin of gonocoxa angled; there is no posterolateral lobe on gonocoxa, the ventral setae unmodified. Parastylar lobe membranous, covered by the gonocoxa, with 1 seta. No paraapodemal lobe observable. Dorsal mesial margin of gonocoxa lobe-like broadly expanded towards the medial line, otherwise simple. Number of juxtagonostylar setae two, the ventral one a simple seta, the dorsal one flattened megaseta with a whip-like branch, both arising from a common basal body which is shorter than the setae. Anteriorly from the juxtagonostylar megasetae there is a plate-like lobe bearing anteriorly a strong megaseta and posteriorly three shorter pointed megasetae. Otherwise the dorsal setae of gonocoxa similar to those on the ventral side except for a row of very long setae in the middle of the posterior half. Inside the gonocoxa there is a conspicuous membranous part bearing numerous finger-like papillae and which is concave towards the medial line. Gonostylus flat, subtriangular, the apicomesial and apicolateral corners lobe-like, the former with two setae, the latter with one dorsally curved seta. Aedeagus narrowly subtriangular, the sides concave, the apex curved ventrad. Hypoproct extending posteriorly just over the base of gonostyli, the number of ventral setae (sternite 10) ca. 15 on each half. Cerci mesially separate.
</paragraph>
<paragraph id="8B8D36BD004EFFC237B772FDFE1C1DEB" blockId="21.[151,1437,1086,2010]" box="[199,413,1986,2010]" pageId="21" pageNumber="122">Female. Unknown.</paragraph>
</subSubSection>
<caption id="DF4D6635004DFFC137E7726CFDD11DB9" httpUri="https://zenodo.org/record/221224/files/figure.png" pageId="22" pageNumber="123" targetBox="[170,1402,206,1839]" targetPageId="22">
<paragraph id="8B8D36BD004DFFC137E7726CFDD11DB9" blockId="22.[151,1436,1875,1928]" pageId="22" pageNumber="123">
<emphasis id="B946EAAF004DFFC137E7726CFEA01D59" bold="true" box="[151,289,1875,1898]" pageId="22" pageNumber="123">FIGURE 12.</emphasis>
<taxonomicName id="4C324D3E004DFFC13659726BFE5C1D58" box="[297,477,1875,1897]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="123" phylum="Arthropoda" rank="species" species="papillosa" status="sp. nov.">
<emphasis id="B946EAAF004DFFC13659726BFE5C1D58" box="[297,477,1875,1897]" italics="true" pageId="22" pageNumber="123">Manota papillosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557D4004DFFC13696726BFDA11D58" box="[486,544,1876,1897]" pageId="22" pageNumber="123" rank="species">
<emphasis id="B946EAAF004DFFC13696726BFDA11D58" bold="true" box="[486,544,1876,1897]" pageId="22" pageNumber="123">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype).
<emphasis id="B946EAAF004DFFC135D0726CFD381D59" bold="true" box="[672,697,1875,1896]" pageId="22" pageNumber="123">A.</emphasis>
Hypopygium, dorsal view.
<emphasis id="B946EAAF004DFFC134A8726BFC711D58" bold="true" box="[984,1008,1876,1897]" pageId="22" pageNumber="123">B.</emphasis>
Hypopygium, ventral view.
<emphasis id="B946EAAF004DFFC13267726CFAB11D5B" bold="true" box="[1303,1328,1875,1898]" pageId="22" pageNumber="123">C.</emphasis>
Aedeagus and hypoproct, ventral view. Scale 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C3286536004CFFC037B775A8FB381829" pageId="23" pageNumber="124" type="discussion">
<paragraph id="8B8D36BD004CFFC037B775A8FCC01B0E" blockId="23.[151,1436,151,679]" pageId="23" pageNumber="124">
<emphasis id="B946EAAF004CFFC037B775A8FECD1A81" bold="true" box="[199,332,151,176]" pageId="23" pageNumber="124">Discussion.</emphasis>
The exact character of the papillose membrane inside the hypopygium of
<taxonomicName id="4C324D3E004CFFC033E075A6FAD71A81" box="[1168,1366,152,176]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="papillosa">
<emphasis id="B946EAAF004CFFC033E075A6FAD71A81" box="[1168,1366,152,176]" italics="true" pageId="23" pageNumber="124">Manota papillosa</emphasis>
</taxonomicName>
is not known to us. Similar membrane in the same position is also seen in
<taxonomicName id="4C324D3E004CFFC034C87582FB001AE5" box="[952,1153,188,212]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="appendiculata">
<emphasis id="B946EAAF004CFFC034C87582FB001AE5" box="[952,1153,188,212]" italics="true" pageId="23" pageNumber="124">M. appendiculata</emphasis>
</taxonomicName>
(
<figureCitation id="13092A38004CFFC033E37583FB591AE5" box="[1171,1240,188,212]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1932]" captionTargetBox="[264,1347,213,1846]" captionTargetId="figure@7.[250,1348,195,1883]" captionTargetPageId="7" captionText="FIGURE 4. Manota appendiculata sp. n. (A paratype from French Guyana, B, C, D holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. E. Gonostylus, dorsal view. F. Juxtagonostylar megasetae, dorsal view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221217/files/figure.png" pageId="23" pageNumber="124">Fig. 4</figureCitation>
C). Reminiscent membranous structures are also seen in many other
<taxonomicName id="4C324D3E004CFFC035A975DEFCB11AC9" box="[729,816,225,248]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAAF004CFFC035A975DEFCB11AC9" box="[729,816,225,248]" italics="true" pageId="23" pageNumber="124">Manota</emphasis>
</taxonomicName>
even if the size of the papillae are smaller and can look more like scales (see e.g. under
<taxonomicName id="4C324D3E004CFFC03572743AFD2D1B2D" box="[514,684,260,284]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="squamulata">
<emphasis id="B946EAAF004CFFC03572743AFD2D1B2D" box="[514,684,260,284]" italics="true" pageId="23" pageNumber="124">M. squamulata</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFA34B4C004CFFC035A1743BFC631B2D" author="Jaschhof" box="[721,994,260,284]" pageId="23" pageNumber="124" refString="Jaschhof, M. &amp; Hippa, H. (2005) The genus Manota in Costa Rica (Diptera: Mycetophilidae). Zootaxa, 1011, 1 - 54." type="journal article" year="2005">Jaschhof &amp; Hippa 2005</bibRefCitation>
). Often they are only vaguely seen and not mentioned in the descriptions or marked in the drawings.
</paragraph>
<paragraph id="8B8D36BD004CFFC037B77472FB381829" blockId="23.[151,1436,151,679]" pageId="23" pageNumber="124">
In the non-setose anterior basalare and laterotergite, sternite 9 fused laterally with the gonocoxa, the membranous parastylar lobe with one seta, the 1+3 megasetae situated anteriorly from the juxtagonostylar megasetae on the dorsal side of the gonocoxa, the subtriangular gonostylus with a long seta at the apicomesial angle and with a curved seta at the apicolateral angle,
<taxonomicName id="4C324D3E004CFFC034737486FC4B1BE1" box="[771,970,440,464]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="papillosa">
<emphasis id="B946EAAF004CFFC034737486FC4B1BE1" box="[771,970,440,464]" italics="true" pageId="23" pageNumber="124">Manota papillosa</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C324D3E004CFFC0333B7486FB421BE1" box="[1099,1219,440,464]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="bisulca">
<emphasis id="B946EAAF004CFFC0333B7486FB421BE1" box="[1099,1219,440,464]" italics="true" pageId="23" pageNumber="124">M. bisulca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E004CFFC033A27486FAEF1BE1" box="[1234,1390,440,464]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="corcovado">
<emphasis id="B946EAAF004CFFC033A27486FAEF1BE1" box="[1234,1390,440,464]" italics="true" pageId="23" pageNumber="124">M. corcovado</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E004CFFC0320D7486FE901BC5" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="diversiseta">
<emphasis id="B946EAAF004CFFC0320D7486FE901BC5" italics="true" pageId="23" pageNumber="124">M. diversiseta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D3E004CFFC0363D74E2FE661BC5" box="[333,487,476,500]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="vexillifera">
<emphasis id="B946EAAF004CFFC0363D74E2FE661BC5" box="[333,487,476,500]" italics="true" pageId="23" pageNumber="124">M. vexillifera</emphasis>
</taxonomicName>
. It differs from all by having the dorsal mesial margin of the gonocoxa lobe-like widened towards the medial line and by lacking a distinct posterolateral lobe on the gonocoxa.
</paragraph>
</subSubSection>
<subSubSection id="C3286536004CFFC037B7771BFC081896" pageId="23" pageNumber="124" type="etymology">
<paragraph id="8B8D36BD004CFFC037B7771BFEAD186E" blockId="23.[151,1436,151,679]" pageId="23" pageNumber="124">
<emphasis id="B946EAAF004CFFC037B7771BFEC7180D" bold="true" box="[199,326,548,572]" pageId="23" pageNumber="124">Etymology</emphasis>
. The name is Latin,
<taxonomicName id="4C324D3E004CFFC03533771BFD2A180D" box="[579,683,548,572]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="papillosa">
<emphasis id="B946EAAF004CFFC03533771BFD2A180D" box="[579,683,548,572]" italics="true" pageId="23" pageNumber="124">papillosa</emphasis>
</taxonomicName>
, full of papillae, referring to the papillose membrane inside the hypopygium.
</paragraph>
<paragraph id="8B8D36BD004CFFC037B77753FC081896" blockId="23.[151,1436,151,679]" pageId="23" pageNumber="124">
<emphasis id="B946EAAF004CFFC037B77753FE9018B5" bold="true" box="[199,273,620,644]" pageId="23" pageNumber="124">
<typeStatus id="5489881F004CFFC037B77753FE8B18B5" box="[199,266,620,644]" pageId="23" pageNumber="124">Types</typeStatus>
.
</emphasis>
<emphasis id="B946EAAF004CFFC0366A7753FE0918B5" box="[282,392,620,644]" italics="true" pageId="23" pageNumber="124">
<typeStatus id="5489881F004CFFC0366A7753FE0518B5" box="[282,388,620,644]" pageId="23" pageNumber="124" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingCountry id="F325762D004CFFC036A57753FDDE18B5" box="[469,607,620,644]" name="Ecuador" pageId="23" pageNumber="124">ECUADOR</collectingCountry>
, Nape Prov., Yasuni NP, Yasuni Res. Station,
<geoCoordinate id="EE06507A004CFFC0332B7753FB3E18B5" box="[1115,1215,620,644]" direction="south" orientation="latitude" pageId="23" pageNumber="124" precision="925" value="-0.6333333">00°38 S</geoCoordinate>
,
<geoCoordinate id="EE06507A004CFFC033BB7753FAB818B5" box="[1227,1337,620,644]" direction="west" orientation="longitude" pageId="23" pageNumber="124" precision="925" value="-76.6">76°36 W</geoCoordinate>
, Malaise trap,
<date id="FF8C107D004CFFC037A077B0FEE61896" box="[208,359,655,679]" pageId="23" pageNumber="124" value="1998-11-03" valueMax="1998-11-20" valueMin="1998-11-03">320.xi.1998</date>
, leg. T. Pape &amp; B. Viklund (on slide, in SMNH).
</paragraph>
</subSubSection>
</treatment>
</document>