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<document id="6AE8DB1AFF12195CB8907517235C0A77" ID-DOI="10.11646/zootaxa.3686.2.1" ID-GBIF-Dataset="4d62d5da-3890-4af3-9758-674a13b43a79" ID-ISSN="1175-5326" ID-Zenodo-Dep="221213" ID-ZooBank="1AF48240-6A27-452E-9D32-9C6D45C715E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460646465142" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Kurina, Olavi" docDate="2013" docId="039B87AB0047FFC937E77397FC081E91" docLanguage="en" docName="zt03686p139.pdf" docOrigin="Zootaxa 3686 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manota quantilla Hippa &amp; Kurina, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="131" masterDocId="FFA2FFD3005BFFD73770753FFF811A31" masterDocTitle="New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region" masterLastPageNumber="139" masterPageNumber="101" pageNumber="129" updateTime="1698337046318" updateUser="plazi">
<mods:mods id="6873412D8289DD860523C17E41B6FDCB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CE897D868E16D33380D257E4A14F3B24">
<mods:title id="EF96A46F0CDF57445FF62D3C606E2756">New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region</mods:title>
</mods:titleInfo>
<mods:name id="E38B823C3BADD4AAE94EED820B4D8404" type="personal">
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<mods:namePart id="7BB738D8B7CFE9A684B2AF65941AA609">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:name id="DE518875902948ECA24CA5B7ADEC3E23" type="personal">
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<mods:namePart id="9BCB796F52688E370492A2365D4F7700">Kurina, Olavi</mods:namePart>
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<mods:title id="395956AA3EB2916817F033DE2AA50C43">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4EA1549FE5C9E2541385FF046565B89A">
<mods:date id="4D616C663B3F67108F7C5139DCE8364D">2013</mods:date>
<mods:detail id="BE33C8B394D8EDFA42C6DD2E02804EF5" type="volume">
<mods:number id="969DA611D3C562FD7D99F8AE86E2B6FC">3686</mods:number>
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<mods:detail id="84BFFAEE8B0FFF031C51D1B8F77F91E2" type="issue">
<mods:number id="D2FCF93B76E0882C10EE276B905D65FE">2</mods:number>
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<mods:start id="52A23D50106B6476668765530C44B9FE">101</mods:start>
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<mods:classification id="BF3063A789A54CE6CCB9ACDE0E359EFE">journal article</mods:classification>
<mods:identifier id="B757E7AFF32FBD31815905E22CA78D6B" type="DOI">10.11646/zootaxa.3686.2.1</mods:identifier>
<mods:identifier id="825CFE47EAD4906DC6994A00C4C6C546" type="GBIF-Dataset">4d62d5da-3890-4af3-9758-674a13b43a79</mods:identifier>
<mods:identifier id="842C31A19E5BABEFAC61D9ECCB185766" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="48E14E7B635A01F91D07790EED7ADA01" type="Zenodo-Dep">221213</mods:identifier>
<mods:identifier id="FB76DEA7F46DF4874CC23C4170289490" type="ZooBank">1AF48240-6A27-452E-9D32-9C6D45C715E3</mods:identifier>
</mods:mods>
<treatment id="039B87AB0047FFC937E77397FC081E91" ID-DOI="http://doi.org/10.5281/zenodo.6155088" ID-GBIF-Taxon="119572330" ID-Zenodo-Dep="6155088" LSID="urn:lsid:plazi:treatment:039B87AB0047FFC937E77397FC081E91" httpUri="http://treatment.plazi.org/id/039B87AB0047FFC937E77397FC081E91" lastPageId="30" lastPageNumber="131" pageId="28" pageNumber="129">
<subSubSection id="C32865360047FFC937E77397FE1C1906" lastPageId="30" lastPageNumber="131" pageId="28" pageNumber="129" type="nomenclature">
<paragraph id="8B8D36BD0047FFCB37E77397FE321CF3" blockId="28.[151,435,1704,1763]" box="[151,435,1704,1730]" pageId="28" pageNumber="129">
<heading id="D0C581D10047FFCB37E77397FE321CF3" bold="true" box="[151,435,1704,1730]" fontSize="11" level="1" pageId="28" pageNumber="129" reason="1">
<emphasis id="B946EAAF0047FFCB37E77397FE321CF3" bold="true" box="[151,435,1704,1730]" pageId="28" pageNumber="129">
<taxonomicName id="4C324D3E0047FFCB37E77397FEE81CF3" ID-CoL="8CPZP" box="[151,361,1704,1730]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="quantilla" status="sp. nov.">
<emphasis id="B946EAAF0047FFCB37E77397FEE81CF3" bold="true" box="[151,361,1704,1730]" italics="true" pageId="28" pageNumber="129">Manota quantilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557D40047FFCB36017397FE321CF3" box="[369,435,1704,1730]" pageId="28" pageNumber="129" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36BD0047FFCB37E773F4FEAE1CD2" blockId="28.[151,435,1704,1763]" box="[151,303,1739,1763]" pageId="28" pageNumber="129">
<figureCitation id="13092A380047FFCB37E773F4FF701CD2" box="[151,241,1739,1763]" captionStart="FIGURE 16" captionStartId="29.[151,250,1945,1968]" captionTargetBox="[165,1398,216,1915]" captionTargetId="figure@29.[151,1436,195,1921]" captionTargetPageId="29" captionText="FIGURE 16. Manota quantilla sp. n. (holotype). A. Antennal flagellomere 4, latewral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221228/files/figure.png" pageId="28" pageNumber="129">Figs. 16</figureCitation>
A–D
</paragraph>
<paragraph id="8B8D36BD0047FFCB37E7722CFAFC1DEE" blockId="28.[151,1437,1810,2016]" pageId="28" pageNumber="129">
Male.
<emphasis id="B946EAAF0047FFCB37AF722DFEB01D1A" bold="true" box="[223,305,1810,1835]" pageId="28" pageNumber="129">Colour</emphasis>
. Head dark brown, face brown. Antenna pale brown. Clypeus brownish, mouthparts pale yellowish. Thorax pale brown, medial part of scutum and scutellum at least seemingly darker brownish. Legs unicolorous yellowish. Wing pale brown; haltere yellow with dark brown knob. Abdominal tergites yellowish brown, sternites paler than tergites. All setosity pale, yellowish or brownish.
<emphasis id="B946EAAF0047FFCB344D7240FCFC1DA6" bold="true" box="[829,893,1919,1943]" pageId="28" pageNumber="129">Head</emphasis>
. Antennal flagellomere 4,
<figureCitation id="13092A380047FFCB33DA7240FB7B1DA6" box="[1194,1274,1919,1943]" captionStart="FIGURE 16" captionStartId="29.[151,250,1945,1968]" captionTargetBox="[165,1398,216,1915]" captionTargetId="figure@29.[151,1436,195,1921]" captionTargetPageId="29" captionText="FIGURE 16. Manota quantilla sp. n. (holotype). A. Antennal flagellomere 4, latewral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221228/files/figure.png" pageId="28" pageNumber="129">Fig. 16</figureCitation>
A. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.3 times longer than palpomere 4. Number of strong postocular setae 9.
<emphasis id="B946EAAF0047FFCB326D72F8FAF41DEE" bold="true" box="[1309,1397,1991,2015]" pageId="28" pageNumber="129">Thorax</emphasis>
.
</paragraph>
<caption id="DF4D66350046FFCA37E772A6FC751DE2" httpUri="https://zenodo.org/record/221228/files/figure.png" pageId="29" pageNumber="130" targetBox="[165,1398,216,1915]" targetPageId="29">
<paragraph id="8B8D36BD0046FFCA37E772A6FC751DE2" blockId="29.[151,1436,1945,2003]" pageId="29" pageNumber="130">
<emphasis id="B946EAAF0046FFCA37E772A6FEA11D9E" bold="true" box="[151,288,1945,1968]" pageId="29" pageNumber="130">FIGURE 16.</emphasis>
<taxonomicName id="4C324D3E0046FFCA365772A4FE561D81" box="[295,471,1946,1968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="quantilla" status="sp. nov.">
<emphasis id="B946EAAF0046FFCA365772A4FE561D81" box="[295,471,1946,1968]" italics="true" pageId="29" pageNumber="130">Manota quantilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557D40046FFCA36AF72A5FD981D9E" box="[479,537,1946,1967]" pageId="29" pageNumber="130" rank="species">
<emphasis id="B946EAAF0046FFCA36AF72A5FD981D9E" bold="true" box="[479,537,1946,1967]" pageId="29" pageNumber="130">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype).
<emphasis id="B946EAAF0046FFCA35E672A5FD311D9E" bold="true" box="[662,688,1946,1967]" pageId="29" pageNumber="130">A.</emphasis>
Antennal flagellomere 4, latewral view.
<emphasis id="B946EAAF0046FFCA332072A5FBE91D9E" bold="true" box="[1104,1128,1946,1967]" pageId="29" pageNumber="130">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="B946EAAF0046FFCA32F372A6FA1D1D81" bold="true" box="[1411,1436,1945,1968]" pageId="29" pageNumber="130">C.</emphasis>
Hypopygium, ventral view.
<emphasis id="B946EAAF0046FFCA36C27282FE4A1DE3" bold="true" box="[434,459,1981,2002]" pageId="29" pageNumber="130">D.</emphasis>
Aedeagus and hypoproct, ventral view. Scale 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8D36BD0045FFC937E775A8FD561925" blockId="30.[151,1436,151,1184]" pageId="30" pageNumber="131">
Anepisternum setose, with 32 setae; anterior basalare non-setose; preepisternum 2 non-setose; laterotergite nonsetose, episternum 3 setose, with 7 setae.
<emphasis id="B946EAAF0045FFC935167583FD1F1AE5" bold="true" box="[614,670,188,212]" pageId="30" pageNumber="131">Legs</emphasis>
. Mid and hind tibial organs absent.
<emphasis id="B946EAAF0045FFC9335F7584FBF11AE5" bold="true" box="[1071,1136,187,212]" pageId="30" pageNumber="131">Wing</emphasis>
. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 not extending to the level of the tip of R1; wing length
<quantity id="4CCA9B580045FFC9320975E0FF471B2D" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="30" pageNumber="131" unit="mm" value="1.4">1.4 mm</quantity>
.
<emphasis id="B946EAAF0045FFC937A6743CFEF91B2D" bold="true" box="[214,376,259,284]" pageId="30" pageNumber="131">Hypopygium.</emphasis>
<figureCitation id="13092A380045FFC936F5743BFE641B2D" box="[389,485,260,284]" captionStart="FIGURE 16" captionStartId="29.[151,250,1945,1968]" captionTargetBox="[165,1398,216,1915]" captionTargetId="figure@29.[151,1436,195,1921]" captionTargetPageId="29" captionText="FIGURE 16. Manota quantilla sp. n. (holotype). A. Antennal flagellomere 4, latewral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm." httpUri="https://zenodo.org/record/221228/files/figure.png" pageId="30" pageNumber="131">Figs. 16</figureCitation>
B–D: Sternite 9 laterally fused with gonocoxa, the posterior margin transverse, slightly convex, extending to the middle of gonocoxa, anterior margin deeply incised, the setae few, similar to the adjacent ventral setae of gonocoxa. Ventral mesial margin of gonocoxa curved, the ventral setae of gonocoxa unmodified; posterolateral part of gonocoxa is drawn into a lobe. Parastylar lobe membranous or lacking, nonsetose. No paraapodemal lobe identifiable. Dorsal mesial margin of gonocoxa slightly lobe-like produced mesiad on the posterior half, with an aggregation of setae at the margin of the lobe, otherwise the setae on the dorsal side of gonostylus similar to those on the ventral side. Number of juxtagonostylar setae two, the ventral one flattened but otherwise rather unmodified megaseta, the dorsal one flattened with a whip-like branch, both arising from a common basal body which is about half of the length of the megasetae. Anteriorly from the juxtagonostylar megasetae there is a plate-like lobe bearing 1 large megaseta. Posteriorly from the juxtagonostylar megasetae there is a narrow transverse lobe bearing two or three strong curved setae. Gonostylus flat, bilobed, the smaller apicolateral lobe with two short setae, the larger apicomesial lobe with a few dorsal setae and with two long apical setae. Aedeagus broadly subtriangular, the lateral sides slightly concave, the apex curved ventrad. Hypoproct extending posteriorly to middle of gonostyli, the number of ventral setae (sternite 10) ca. 10 on each half. Cerci, except shortly separate apical parts, mesially fused.
</paragraph>
<paragraph id="8B8D36BD0045FFC937B77620FE1C1906" blockId="30.[151,1436,151,1184]" box="[199,413,799,823]" pageId="30" pageNumber="131">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C32865360045FFC937B7767CFD7A1E05" pageId="30" pageNumber="131" type="discussion">
<paragraph id="8B8D36BD0045FFC937B7767CFD7A1E05" blockId="30.[151,1436,151,1184]" pageId="30" pageNumber="131">
<emphasis id="B946EAAF0045FFC937B7767CFECD196D" bold="true" box="[199,332,835,860]" pageId="30" pageNumber="131">Discussion.</emphasis>
By the non-setose anterior basalare, non-setose laterotergite, laterally with the gonocoxa fused sternite 9, membranous or lacking parastylar lobe, posterolateral lobe on the gonocoxa, subtriangular gonostylus with a long seta at the apicomesial angle and a curved seta at the apicolateral angle
<taxonomicName id="4C324D3E0045FFC9332E76B2FAA21995" box="[1118,1315,908,932]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="quantilla">
<emphasis id="B946EAAF0045FFC9332E76B2FAA21995" box="[1118,1315,908,932]" italics="true" pageId="30" pageNumber="131">Manota quantilla</emphasis>
</taxonomicName>
resembles several species, viz.
<taxonomicName id="4C324D3E0045FFC936F4768EFE7C19F9" box="[388,509,944,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="bisulca">
<emphasis id="B946EAAF0045FFC936F4768EFE7C19F9" box="[388,509,944,968]" italics="true" pageId="30" pageNumber="131">M. bisulca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC9357E768EFD4919F9" box="[526,712,945,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="B946EAAF0045FFC9357E768EFD4919F9" box="[526,712,945,968]" italics="true" pageId="30" pageNumber="131">M. costaricensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC935A9768EFCCA19F9" box="[729,843,944,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="depilis">
<emphasis id="B946EAAF0045FFC935A9768EFCCA19F9" box="[729,843,944,968]" italics="true" pageId="30" pageNumber="131">M. depilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC9342C768EFC6119F9" box="[860,992,944,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B946EAAF0045FFC9342C768EFC6119F9" box="[860,992,944,968]" italics="true" pageId="30" pageNumber="131">M. fraterna</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC93481768EFBDE19F9" box="[1009,1119,944,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="panda">
<emphasis id="B946EAAF0045FFC93481768EFBDE19F9" box="[1009,1119,944,968]" italics="true" pageId="30" pageNumber="131">M. panda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC93300768EFA8119F9" box="[1136,1280,944,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="papillosa">
<emphasis id="B946EAAF0045FFC93300768EFA8119F9" box="[1136,1280,944,968]" italics="true" pageId="30" pageNumber="131">M. papillosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D3E0045FFC93261768EFAFB19F9" box="[1297,1402,945,968]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B946EAAF0045FFC93261768EFAFB19F9" box="[1297,1402,945,968]" italics="true" pageId="30" pageNumber="131">M. parva</emphasis>
</taxonomicName>
. It differs from all of them by having anteriorly from the juxtagonostylar megasetae, at the dorsal mesial margin of the gonocoxa, a lobe with only one large megaseta and without no other prominent setae or lobes. The row of 3 curved setae on the dorsal surface of the gonostylus is unique.
</paragraph>
</subSubSection>
<subSubSection id="C32865360045FFC937B7717FFC081E91" pageId="30" pageNumber="131" type="etymology">
<paragraph id="8B8D36BD0045FFC937B7717FFB621E66" blockId="30.[151,1436,151,1184]" box="[199,1251,1087,1113]" pageId="30" pageNumber="131">
<emphasis id="B946EAAF0045FFC937B7717FFEC71E69" bold="true" box="[199,326,1088,1112]" pageId="30" pageNumber="131">Etymology</emphasis>
. The name is Latin,
<taxonomicName id="4C324D3E0045FFC9355D717FFD131E69" box="[557,658,1088,1112]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="quantilla">
<emphasis id="B946EAAF0045FFC9355D717FFD131E69" box="[557,658,1088,1112]" italics="true" pageId="30" pageNumber="131">quantilla</emphasis>
</taxonomicName>
, how very little, referring to the small size of the fly.
</paragraph>
<paragraph id="8B8D36BD0045FFC937B7715BFC081E91" blockId="30.[151,1436,151,1184]" pageId="30" pageNumber="131">
<emphasis id="B946EAAF0045FFC937B7715BFE901E4D" bold="true" box="[199,273,1124,1148]" pageId="30" pageNumber="131">
<typeStatus id="5489881F0045FFC937B7715BFE8B1E4D" box="[199,266,1124,1148]" pageId="30" pageNumber="131">Types</typeStatus>
.
</emphasis>
<emphasis id="B946EAAF0045FFC9366A715BFE091E4D" box="[282,392,1124,1148]" italics="true" pageId="30" pageNumber="131">
<typeStatus id="5489881F0045FFC9366A715BFE051E4D" box="[282,388,1124,1148]" pageId="30" pageNumber="131" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingCountry id="F325762D0045FFC936A5715BFDDE1E4D" box="[469,607,1124,1148]" name="Ecuador" pageId="30" pageNumber="131">ECUADOR</collectingCountry>
, Nape Prov., Yasuni NP, Yasuni Res. Station,
<geoCoordinate id="EE06507A0045FFC9332B715BFB3E1E4D" box="[1115,1215,1124,1148]" direction="south" orientation="latitude" pageId="30" pageNumber="131" precision="925" value="-0.6333333">00°38’ S</geoCoordinate>
,
<geoCoordinate id="EE06507A0045FFC933BB715BFAB81E4D" box="[1227,1337,1124,1148]" direction="west" orientation="longitude" pageId="30" pageNumber="131" precision="925" value="-76.6">76°36’ W</geoCoordinate>
, Malaise trap,
<date id="FF8C107D0045FFC937A071B7FEE61E91" box="[208,359,1160,1184]" pageId="30" pageNumber="131" value="1998-11-03" valueMax="1998-11-20" valueMin="1998-11-03">3–20.xi.1998</date>
, leg. T. Pape &amp; B. Viklund (on slide, in SMNH).
</paragraph>
</subSubSection>
</treatment>
</document>
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