treatments-xml/data/03/9B/87/039B878E442A9C0DDFE6FE44FE0D3F88.xml
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<document id="36B23AB3992229F48B033C1D0FD20F9B" ID-DOI="10.11646/zootaxa.3641.3.5" ID-GBIF-Dataset="26137e83-060f-42f8-82c9-011d9a86c15b" ID-ISSN="1175-5326" ID-Zenodo-Dep="221272" ID-ZooBank="E5849A06-1F26-47ED-91E4-433A1F57B0BE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460595528880" checkinUser="plazi" docAuthor="Marshall, Stephen A." docDate="2013" docId="039B878E442A9C0DDFE6FE44FE0D3F88" docLanguage="en" docName="zt03641p270.pdf" docOrigin="Zootaxa 3641 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bregmosina howdeni Marshall, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="268" masterDocId="FFA2FFF644229C05DF71FF98FFF33B6E" masterDocTitle="Bregmosina, a new Neotropical genus of Limosininae (Diptera: Sphaeroceridae)" masterLastPageNumber="270" masterPageNumber="260" pageNumber="268" updateTime="1698326706864" updateUser="plazi">
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<mods:title id="053B885E2EBB41E85D9A293F996B3F24">Bregmosina, a new Neotropical genus of Limosininae (Diptera: Sphaeroceridae)</mods:title>
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<mods:namePart id="BB363657C51764C419C6F4D5327D7C65">Marshall, Stephen A.</mods:namePart>
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<mods:title id="BA73DEE849612B110E6749A3AAE21CAF">Zootaxa</mods:title>
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<mods:part id="3A9D0C0E06662348C525DA9E175B9361">
<mods:date id="5D8E25F8F8F8EAD5724CF62AAC56E834">2013</mods:date>
<mods:detail id="E5455EE47C566556A43800BE2A719DA9" type="volume">
<mods:number id="C71A47B962E074738A6A0D0CBA2F1464">3641</mods:number>
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<mods:classification id="4B5FB17B528FD061737CE2F61E39E2F6">journal article</mods:classification>
<mods:identifier id="B4B4C024919C4F8FBC6894A1D6F65788" type="DOI">10.11646/zootaxa.3641.3.5</mods:identifier>
<mods:identifier id="50B19B6F01D97427B4F9B6B20DD8A599" type="GBIF-Dataset">26137e83-060f-42f8-82c9-011d9a86c15b</mods:identifier>
<mods:identifier id="918C2180B7147A2697A412DF2B9A11CA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D4E997DD0A3A2590EA56507F62D6DB71" type="Zenodo-Dep">221272</mods:identifier>
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<treatment id="039B878E442A9C0DDFE6FE44FE0D3F88" ID-DOI="http://doi.org/10.5281/zenodo.6165134" ID-GBIF-Taxon="119566825" ID-Zenodo-Dep="6165134" LSID="urn:lsid:plazi:treatment:039B878E442A9C0DDFE6FE44FE0D3F88" httpUri="http://treatment.plazi.org/id/039B878E442A9C0DDFE6FE44FE0D3F88" lastPageNumber="268" pageId="8" pageNumber="268">
<subSubSection id="C3286513442A9C0DDFE6FE44FF153978" pageId="8" pageNumber="268" type="nomenclature">
<paragraph id="8B8D3698442A9C0DDFE6FE44FDDD3A9B" blockId="8.[151,558,475,534]" box="[151,558,475,502]" pageId="8" pageNumber="268">
<heading id="D0C581F4442A9C0DDFE6FE44FDDD3A9B" bold="true" box="[151,558,475,502]" fontSize="11" level="1" pageId="8" pageNumber="268" reason="1">
<emphasis id="B946EA8A442A9C0DDFE6FE44FDDD3A9B" bold="true" box="[151,558,475,502]" pageId="8" pageNumber="268">
<taxonomicName id="4C324D1B442A9C0DDFE6FE44FE6B3A98" ID-CoL="8B9DV" box="[151,408,476,502]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="B946EA8A442A9C0DDFE6FE44FE6B3A98" bold="true" box="[151,408,476,502]" italics="true" pageId="8" pageNumber="268">Bregmosina howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557F1442A9C0DDEEFFE43FDDD3A9B" box="[414,558,475,501]" pageId="8" pageNumber="268" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D3698442A9C0DDFE6FE66FF153978" blockId="8.[151,558,475,534]" box="[151,230,510,534]" pageId="8" pageNumber="268">
<figureCitation id="13092A1D442A9C0DDFE6FE66FF153978" box="[151,230,510,534]" captionStart="FIGURES 18 22" captionStartId="7.[151,264,1888,1911]" captionTargetBox="[157,1428,199,1870]" captionTargetId="figure@7.[151,1436,193,1881]" captionTargetPageId="7" captionText="FIGURES 18 22. Bregmosina obunca and B. howdeni. 18, B. obunca female terminalia; 19, B. obunca aedeagus and associated structures left lateral; 20, B. obunca male terminalia ventral; 21, B. obunca epandrium and associated structures left lateral; 22, B. howdeni epandrium and associated structures left lateral. Abbreviations: SP, spermatheca; T 7, T 8, S 5, S 7, S 8, sternites and tergites; P, phallapodeme; PG, postgonite; H, hypandrium; C, cercus; B, basiphallus; D, distiphallus; SU, surstylus." httpUri="https://zenodo.org/record/221276/files/figure.png" pageId="8" pageNumber="268">Fig. 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3286513442A9C0DDFE6FDDEFD4E3834" pageId="8" pageNumber="268" type="description">
<paragraph id="8B8D3698442A9C0DDFE6FDDEFC853931" blockId="8.[151,1437,582,1254]" box="[151,886,582,607]" pageId="8" pageNumber="268">
<emphasis id="B946EA8A442A9C0DDFE6FDDEFEDB3931" bold="true" box="[151,296,582,607]" pageId="8" pageNumber="268">Description.</emphasis>
Externally as described for
<taxonomicName id="4C324D1B442A9C0DDD12FDD0FD273930" box="[611,724,582,607]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
<emphasis id="B946EA8A442A9C0DDD12FDD0FD273930" box="[611,724,582,607]" italics="true" pageId="8" pageNumber="268">B. obunca</emphasis>
</taxonomicName>
. Size 2.0 mm.
</paragraph>
<paragraph id="8B8D3698442A9C0DDFB8FDF2FD4E3834" blockId="8.[151,1437,582,1254]" pageId="8" pageNumber="268">
Male abdomen: Sternite 4 extensively overlapping sternite 5 but otherwise unmodified. Sternite 5 as illustrated for
<taxonomicName id="4C324D1B442A9C0DDFB1FD08FEC039C8" box="[192,307,654,679]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
<emphasis id="B946EA8A442A9C0DDFB1FD08FEC039C8" box="[192,307,654,679]" italics="true" pageId="8" pageNumber="268">B. obunca</emphasis>
</taxonomicName>
(
<figureCitation id="13092A1D442A9C0DDE35FD16FE6B39C8" box="[324,408,654,678]" captionStart="FIGURES 18 22" captionStartId="7.[151,264,1888,1911]" captionTargetBox="[157,1428,199,1870]" captionTargetId="figure@7.[151,1436,193,1881]" captionTargetPageId="7" captionText="FIGURES 18 22. Bregmosina obunca and B. howdeni. 18, B. obunca female terminalia; 19, B. obunca aedeagus and associated structures left lateral; 20, B. obunca male terminalia ventral; 21, B. obunca epandrium and associated structures left lateral; 22, B. howdeni epandrium and associated structures left lateral. Abbreviations: SP, spermatheca; T 7, T 8, S 5, S 7, S 8, sternites and tergites; P, phallapodeme; PG, postgonite; H, hypandrium; C, cercus; B, basiphallus; D, distiphallus; SU, surstylus." httpUri="https://zenodo.org/record/221276/files/figure.png" pageId="8" pageNumber="268">Fig. 18</figureCitation>
); pale medially, made up almost entirely of two long lobes and their broad bases, lobes as long as rest of sternite and pointed and incurved distally. Posteromedial point of sternite 5 weakly ridged. Ventral part of sternite 6 narrow but unmodified. Cerci separate, elongate-rectangular. Surstylus elongate, mostly parallel sided but ending in a weakly bilobed club. Postgonite canopener-shaped, circular distal notch about as broad as anterodistal lobe. Basiphallus with a prominent hook-like epiphallus. Distiphallus narrow, distal dorsal surface bilobed and with a conspicuously spiked surface.
</paragraph>
</subSubSection>
<subSubSection id="C3286513442A9C0DDFB8FCFFFA9138A8" pageId="8" pageNumber="268" type="materials_examined">
<paragraph id="8B8D3698442A9C0DDFB8FCFFFA9138A8" blockId="8.[151,1437,582,1254]" pageId="8" pageNumber="268">
<emphasis id="B946EA8A442A9C0DDFB8FCFFFE8E3811" bold="true" box="[201,381,870,895]" pageId="8" pageNumber="268">
<typeStatus id="5489883A442A9C0DDFB8FCFFFEF73811" box="[201,260,871,895]" pageId="8" pageNumber="268">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5489883A442A9C0DDEFBFCFEFE013810" box="[394,498,870,894]" pageId="8" pageNumber="268" type="holotype">Holotype</typeStatus>
(male, DEBU).
<collectingCountry id="F3257608442A9C0DDDCFFCFEFCBB3810" box="[702,840,870,894]" name="Ecuador" pageId="8" pageNumber="268">ECUADOR</collectingCountry>
. Pichincha Province,
<quantity id="4CCA9B7D442A9C0DDB3DFCFEFB6A3810" box="[1100,1177,870,894]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="268" unit="km" value="47.0">47 km</quantity>
S Sto
<collectingCountry id="F3257608442A9C0DDB87FCFEFA913810" box="[1270,1378,870,894]" name="Dominican Republic" pageId="8" pageNumber="268">Domingo</collectingCountry>
, Rio Palenque Biological Station,
<date id="FF8C1058442A9C0DDE87FC12FD6738CC" box="[502,660,906,930]" pageId="8" pageNumber="268" value="1979-02-17" valueMax="1979-02-25" valueMin="1979-02-17">1725.ii.1979</date>
, S.A.
<collectingCountry id="F3257608442A9C0DDD9BFC12FCA238CC" box="[746,849,906,930]" name="Marshall Islands" pageId="8" pageNumber="268">Marshall</collectingCountry>
.
<typeStatus id="5489883A442A9C0DDC15FC13FC3938CD" box="[868,970,907,931]" pageId="8" pageNumber="268" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F3257608442A9C0DDCACFC12FB9438CC" box="[989,1127,906,930]" name="Ecuador" pageId="8" pageNumber="268">ECUADOR</collectingCountry>
. Rio Palenque Biological Station,
<date id="FF8C1058442A9C0DDF82FC36FE6E38A8" box="[243,413,942,966]" pageId="8" pageNumber="268" value="1987-04-30" valueMax="1987-05-05" valueMin="1987-04-30">
30.iv5.v.
<quantity id="4CCA9B7D442A9C0DDE10FC36FE6E38A8" box="[353,413,942,966]" metricMagnitude="6" metricUnit="m" metricValue="1.9871599999999998" pageId="8" pageNumber="268" unit="m" value="1987160.0">1987</quantity>
</date>
, 160 m, primary rain forest Malaise trap, L. Coote and B. Brown (1Ƥ, ROM 870002).
</paragraph>
</subSubSection>
<subSubSection id="C3286513442A9C0DDFB8FC4AFE773F14" pageId="8" pageNumber="268" type="discussion">
<paragraph id="8B8D3698442A9C0DDFB8FC4AFE773F14" blockId="8.[151,1437,582,1254]" pageId="8" pageNumber="268">
<emphasis id="B946EA8A442A9C0DDFB8FC4AFEA23885" bold="true" box="[201,337,978,1003]" pageId="8" pageNumber="268">Comments.</emphasis>
This species is closely related to
<taxonomicName id="4C324D1B442A9C0DDD95FC4CFCA93885" box="[740,858,979,1003]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
<emphasis id="B946EA8A442A9C0DDD95FC4CFCA93885" box="[740,858,979,1003]" italics="true" pageId="8" pageNumber="268">B. obunca</emphasis>
</taxonomicName>
, known from
<collectingCountry id="F3257608442A9C0DDB75FC4AFBA83884" box="[1028,1115,978,1002]" name="Guyana" pageId="8" pageNumber="268">Guyana</collectingCountry>
and
<collectingCountry id="F3257608442A9C0DDBEBFC4AFAEF3884" box="[1178,1308,978,1002]" name="Costa Rica" pageId="8" pageNumber="268">Costa Rica</collectingCountry>
. Both
<typeStatus id="5489883A442A9C0DDA1DFC4BFA6F3885" box="[1388,1436,979,1003]" pageId="8" pageNumber="268">type</typeStatus>
specimens were collected in one of the few remaining patches of primary rainforest in the western lowlands on the Pacific side of the Ecuadorian Andes. This species joins a growing list of fly taxa known only from the remarkable Rio Palenque Biological Station, a reserve now threatened as an increasingly isolated island of forest in a sea of intensive agriculture.
</paragraph>
</subSubSection>
<subSubSection id="C3286513442A9C0DDFB8FB1FFE0D3F88" pageId="8" pageNumber="268" type="etymology">
<paragraph id="8B8D3698442A9C0DDFB8FB1FFE0D3F88" blockId="8.[151,1437,582,1254]" pageId="8" pageNumber="268">
<emphasis id="B946EA8A442A9C0DDFB8FB1FFEA33FF1" bold="true" box="[201,336,1159,1183]" pageId="8" pageNumber="268">Etymology.</emphasis>
<taxonomicName id="4C324D1B442A9C0DDE27FB1FFDB13FF0" box="[342,578,1158,1182]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B946EA8A442A9C0DDE27FB1FFDB13FF0" box="[342,578,1158,1182]" italics="true" pageId="8" pageNumber="268">Bregmosina howdeni</emphasis>
</taxonomicName>
is a patronym in honour of my MSc supervisor, coleopterist Henry Howden. Dr. Howden organized the 1979 trip to Rio Palenque during which I collected this and many other species still known only from Rio Palenque.
</paragraph>
</subSubSection>
</treatment>
</document>