166 lines
19 KiB
XML
166 lines
19 KiB
XML
<document id="79D5C8EED506C20B2D76D8A2E788F7BF" ID-DOI="10.11646/zootaxa.3641.3.5" ID-GBIF-Dataset="26137e83-060f-42f8-82c9-011d9a86c15b" ID-ISSN="1175-5326" ID-Zenodo-Dep="221272" ID-ZooBank="E5849A06-1F26-47ED-91E4-433A1F57B0BE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460595528880" checkinUser="plazi" docAuthor="Marshall, Stephen A." docDate="2013" docId="039B878E44249C0DDFE6FB86FCBB3AE9" docLanguage="en" docName="zt03641p270.pdf" docOrigin="Zootaxa 3641 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bregmosina obunca Marshall, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="268" masterDocId="FFA2FFF644229C05DF71FF98FFF33B6E" masterDocTitle="Bregmosina, a new Neotropical genus of Limosininae (Diptera: Sphaeroceridae)" masterLastPageNumber="270" masterPageNumber="260" pageNumber="266" updateTime="1698326706864" updateUser="plazi">
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<mods:titleInfo id="D361BDAEF07312C522FA3FCB44884151">
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<mods:title id="FAE7A5298E5871E9BBF33C59C5135B85">Bregmosina, a new Neotropical genus of Limosininae (Diptera: Sphaeroceridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0E966DB8C7714C615CBD354F159AC63A" type="personal">
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<mods:roleTerm id="DA254601BC3C691840877C0917D40CB6">Author</mods:roleTerm>
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<mods:namePart id="9E6B6A7EB39646205B13A5FDFA026771">Marshall, Stephen A.</mods:namePart>
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<mods:typeOfResource id="ADE65747CF02BD27BA69B005E6FC2BC8">text</mods:typeOfResource>
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<mods:title id="981D66628261E473779B9285C1BE2B80">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="95AF398E395261351D41A4AE6E308381">
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<mods:date id="55CC00D4FE2C917BED4182B04D74697A">2013</mods:date>
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<mods:detail id="A4570D9AA84FB9464F4216E4A7E13AF7" type="volume">
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<mods:number id="445508B2A260D4DA1C6DF4002E2758ED">3641</mods:number>
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<mods:detail id="7877D8E0A6BF534A882E5AC88AA33C9A" type="issue">
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<mods:number id="9A750CF9FB6D487AABF91517A7431605">3</mods:number>
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<mods:classification id="F1CDF6B1D9F5EBEBDF96A52BFF325C21">journal article</mods:classification>
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<mods:identifier id="499D12FFDC5D5B3D0271A3B52A2C92AF" type="DOI">10.11646/zootaxa.3641.3.5</mods:identifier>
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<mods:identifier id="F24BB69E9EFD3087A25B6C35E21AB3C1" type="GBIF-Dataset">26137e83-060f-42f8-82c9-011d9a86c15b</mods:identifier>
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<mods:identifier id="5732B09E3CC058C74B5744F8BFDDDBCF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="02A7B1B175572DBDAA9DDC603281CEC3" type="Zenodo-Dep">221272</mods:identifier>
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<treatment id="039B878E44249C0DDFE6FB86FCBB3AE9" ID-DOI="http://doi.org/10.5281/zenodo.6165132" ID-GBIF-Taxon="119566828" ID-Zenodo-Dep="6165132" LSID="urn:lsid:plazi:treatment:039B878E44249C0DDFE6FB86FCBB3AE9" httpUri="http://treatment.plazi.org/id/039B878E44249C0DDFE6FB86FCBB3AE9" lastPageId="8" lastPageNumber="268" pageId="6" pageNumber="266">
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<subSubSection id="C328651344249C03DFE6FB86FEE83F37" pageId="6" pageNumber="266" type="nomenclature">
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<paragraph id="8B8D369844249C03DFE6FB86FDD23F56" blockId="6.[151,545,1054,1113]" box="[151,545,1054,1080]" pageId="6" pageNumber="266">
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<heading id="D0C581F444249C03DFE6FB86FDD23F56" bold="true" box="[151,545,1054,1080]" fontSize="11" level="1" pageId="6" pageNumber="266" reason="1">
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<emphasis id="B946EA8A44249C03DFE6FB86FDD23F56" bold="true" box="[151,545,1054,1080]" pageId="6" pageNumber="266">
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<taxonomicName id="4C324D1B44249C03DFE6FB86FE783F56" ID-CoL="8B9DW" box="[151,395,1054,1080]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="obunca" status="sp. nov.">
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<emphasis id="B946EA8A44249C03DFE6FB86FE783F56" bold="true" box="[151,395,1054,1080]" italics="true" pageId="6" pageNumber="266">Bregmosina obunca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27557F144249C03DEE0FB86FDD23F56" box="[401,545,1054,1080]" pageId="6" pageNumber="266" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8D369844249C03DFE6FBD9FEE83F37" blockId="6.[151,545,1054,1113]" box="[151,283,1089,1113]" pageId="6" pageNumber="266">
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<figureCitation id="13092A1D44249C03DFE6FBD9FEE83F37" box="[151,283,1089,1113]" captionStart="FIGURES 18 – 22" captionStartId="7.[151,264,1888,1911]" captionTargetBox="[157,1428,199,1870]" captionTargetId="figure@7.[151,1436,193,1881]" captionTargetPageId="7" captionText="FIGURES 18 – 22. Bregmosina obunca and B. howdeni. 18, B. obunca female terminalia; 19, B. obunca aedeagus and associated structures left lateral; 20, B. obunca male terminalia ventral; 21, B. obunca epandrium and associated structures left lateral; 22, B. howdeni epandrium and associated structures left lateral. Abbreviations: SP, spermatheca; T 7, T 8, S 5, S 7, S 8, sternites and tergites; P, phallapodeme; PG, postgonite; H, hypandrium; C, cercus; B, basiphallus; D, distiphallus; SU, surstylus." httpUri="https://zenodo.org/record/221276/files/figure.png" pageId="6" pageNumber="266">Figs. 18–22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328651344249C03DFE6FB11FAD93D8A" pageId="6" pageNumber="266" type="description">
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<paragraph id="8B8D369844249C03DFE6FB11FCEB3E3B" blockId="6.[151,1437,1161,2016]" pageId="6" pageNumber="266">
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<emphasis id="B946EA8A44249C03DFE6FB11FEDA3FCC" bold="true" box="[151,297,1161,1186]" pageId="6" pageNumber="266">Description.</emphasis>
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Size (antennal base to wing tip) 1.9–2.0 mm. Eye height 2.5–2.6X genal height. Lower orbital bristle thin, 0.3–0.4X as long as upper orbital bristle. Tarsomeres 2–5 of fore leg short, slightly flattened and widened; tarsomeres 2, 3 and 5 equal in length, tarsomere 4 shorter. Tarsomeres 1 and 2 of hind leg weakly swollen, second tarsomere as long as tarsomeres 3, 4, and 5 combined. Mid tibia with 3 distal dorsal bristle, including a long anterodorsal, a long dorsal, and a short posterodorsal (length equals tibial width). Wing: Second costal sector 0.6– 0.7 times as long as third; R2+3 evenly bowed, not sinuate.
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</paragraph>
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<paragraph id="8B8D369844249C03DFB8FAFCFD013E86" blockId="6.[151,1437,1161,2016]" pageId="6" pageNumber="266">Female abdomen: Tergite 8 shortened dorsomedially, broad laterally. Cercus as long as middle part of tergite 8, with a long apical bristle. Spermathecae large (diameter greater than length of sternite 7), spherical with a small conical base; ducts thin and sclerotized, longer than spermatheca. Sternite 8 pale anteromedially. Sternite 10 divided posteromedially, forming two posterior lobes.</paragraph>
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<paragraph id="8B8D369844249C03DFB8FA6CFAD93D8A" blockId="6.[151,1437,1161,2016]" pageId="6" pageNumber="266">Male abdomen: Sternite 4 extensively overlapping sternite 5, posteromedial part with two short, rounded lobes. Sternite 5 desclerotized medially, made up mostly of two long posterolateral lobes and their broad bases, lobes as long as rest of sternite and pointed and slightly incurved distally. Posteromedial point of sternite 5 weakly ridged. Ventral part of sternite 6 relatively broad, concave posteromedially. Ring sclerite complete. Cerci separate, elongate-rectangular. Surstylus elongate and sinuate, ending in a conspicuously long, anteroventrally pointed apex. Postgonite "can opener"-shaped, posterior lobe conspicuously longer than broader anterior lobe. Basiphallus with a short, broad epiphallus. Distiphallus narrow, dorsal surface with inconspicuous spiked surface near base.</paragraph>
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</subSubSection>
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<subSubSection id="C328651344249C03DFB8F968FD633C8E" pageId="6" pageNumber="266" type="materials_examined">
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<paragraph id="8B8D369844249C03DFB8F968FD633C8E" blockId="6.[151,1437,1161,2016]" pageId="6" pageNumber="266">
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<emphasis id="B946EA8A44249C03DFB8F968FE8B3C66" bold="true" box="[201,376,1775,1800]" pageId="6" pageNumber="266">
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<typeStatus id="5489883A44249C03DFB8F968FEF73C66" box="[201,260,1776,1800]" pageId="6" pageNumber="266">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="5489883A44249C03DEF0F968FE1A3C66" box="[385,489,1776,1800]" pageId="6" pageNumber="266" type="holotype">Holotype</typeStatus>
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male (INBIO) and 3
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<typeStatus id="5489883A44249C03DD90F969FCBE3C67" box="[737,845,1777,1801]" pageId="6" pageNumber="266" type="paratype">paratypes</typeStatus>
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(13, 2Ƥ, DEBU):
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<collectingCountry id="F325760844249C03DB58F968FB263C66" box="[1065,1237,1776,1800]" name="Costa Rica" pageId="6" pageNumber="266">COSTA RICA</collectingCountry>
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. Alajuela. Volcan Tenorio, N slope nr. Bijagua Biological Station,
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<quantity id="4CCA9B7D44249C03DDBAF88CFCE43C42" box="[715,791,1812,1836]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="266" unit="m" value="700.0">700 m</quantity>
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, pans in treefall,
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<date id="FF8C105844249C03DC94F88CFB933C42" box="[997,1120,1812,1836]" pageId="6" pageNumber="266" value="2000-06-18">18.vi.2000</date>
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, Buck and
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<collectingCountry id="F325760844249C03DB96F88CFABD3C42" box="[1255,1358,1812,1836]" name="Marshall Islands" pageId="6" pageNumber="266">Marshall</collectingCountry>
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. Other
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<typeStatus id="5489883A44249C03DFE6F8A1FEF43C3F" box="[151,263,1849,1873]" pageId="6" pageNumber="266" type="paratype">paratypes</typeStatus>
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:
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<collectingCountry id="F325760844249C03DE60F8A0FE493C3E" box="[273,442,1848,1872]" name="Costa Rica" pageId="6" pageNumber="266">COSTA RICA</collectingCountry>
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. Heredia. Pico Viejo, La Selva Biological Station,
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<date id="FF8C105844249C03DC9FF8A0FB893C3E" box="[1006,1146,1848,1872]" pageId="6" pageNumber="266" value="1989-05-08" valueMax="1989-05-15" valueMin="1989-05-08">8–15.v.1989</date>
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, B. Brown and D. Feener, Malaise trap, CC100 (13, DEBU); Puntarenas, R.F. Golfo Dulce,
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<quantity id="4CCA9B7D44249C03DCF7F8C4FC333C1A" box="[902,960,1884,1908]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="266" unit="km" value="3.0">3 km</quantity>
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SW Rincon,
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<quantity id="4CCA9B7D44249C03DB2BF8C4FB653C1A" box="[1114,1174,1884,1908]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="266" unit="m" value="10.0">10 m</quantity>
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,
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<date id="FF8C105844249C03DBD2F8C4FAED3C1A" box="[1187,1310,1884,1908]" pageId="6" pageNumber="266" value="1990-10" valueMax="1990-12-31" valueMin="1990-10-00">x–xii.1990</date>
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, P. Hanson (13, 1Ƥ DEBU); Braulio Carrillo National Park,
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<date id="FF8C105844249C03DD89F818FC9C3CF6" box="[760,879,1920,1944]" pageId="6" pageNumber="266" value="1985-04-10">
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10.iv.
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<quantity id="4CCA9B7D44249C03DC42F818FC9C3CF6" box="[819,879,1920,1944]" metricMagnitude="6" metricUnit="m" metricValue="1.9855" pageId="6" pageNumber="266" unit="m" value="1985500.0">1985</quantity>
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</date>
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, 500 m, H. Goulet, L. Masner (1Ƥ, DEBU).
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<collectingCountry id="F325760844249C03DFE6F83CFEE23CD2" box="[151,273,1956,1980]" name="Guyana" pageId="6" pageNumber="266">GUYANA</collectingCountry>
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. Mazaruni-Potaro Distr., Tukelt Creek on E. side Potaro, 300’, rainforest, Malaise,
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<date id="FF8C105844249C03DBA1F83CFA873CD2" box="[1232,1396,1956,1980]" pageId="6" pageNumber="266" value="1990-09-26" valueMax="1990-09-30" valueMin="1990-09-26">26–30.ix.1990</date>
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, B. Hubley, L.D. Coote (1Ƥ, 13, ROM 905019).
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</paragraph>
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</subSubSection>
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<caption id="DF4D661044259C02DFE6F8F8FA6E3CBC" httpUri="https://zenodo.org/record/221276/files/figure.png" pageId="7" pageNumber="267" targetBox="[157,1428,199,1870]" targetPageId="7">
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<paragraph id="8B8D369844259C02DFE6F8F8FA6E3CBC" blockId="7.[151,1437,1888,2002]" pageId="7" pageNumber="267">
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<emphasis id="B946EA8A44259C02DFE6F8F8FEAA3C1B" bold="true" box="[151,345,1888,1911]" pageId="7" pageNumber="267">FIGURES 18–22.</emphasis>
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<taxonomicName id="4C324D1B44259C02DE14F8F9FDC73C18" box="[357,564,1888,1910]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="obunca">
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<emphasis id="B946EA8A44259C02DE14F8F9FDC73C18" box="[357,564,1888,1910]" italics="true" pageId="7" pageNumber="267">Bregmosina obunca</emphasis>
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</taxonomicName>
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and
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<emphasis id="B946EA8A44259C02DD03F8F9FD023C18" box="[626,753,1888,1910]" italics="true" pageId="7" pageNumber="267">
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<taxonomicName id="4C324D1B44259C02DD03F8F9FD1E3C18" box="[626,749,1888,1910]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="howdeni">B. howdeni</taxonomicName>
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.
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</emphasis>
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18,
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<taxonomicName id="4C324D1B44259C02DC56F8F9FC603C18" box="[807,915,1888,1910]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="obunca">B. obunca</taxonomicName>
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female terminalia; 19,
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<taxonomicName id="4C324D1B44259C02DBE7F8F9FAF13C18" box="[1174,1282,1888,1910]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="obunca">
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<emphasis id="B946EA8A44259C02DBE7F8F9FAF13C18" box="[1174,1282,1888,1910]" italics="true" pageId="7" pageNumber="267">B. obunca</emphasis>
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</taxonomicName>
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aedeagus and associated structures left lateral; 20,
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<taxonomicName id="4C324D1B44259C02DD7BF818FD823CFB" box="[522,625,1919,1941]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="obunca">
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<emphasis id="B946EA8A44259C02DD7BF818FD823CFB" box="[522,625,1919,1941]" italics="true" pageId="7" pageNumber="267">B. obunca</emphasis>
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</taxonomicName>
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male terminalia ventral; 21,
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<taxonomicName id="4C324D1B44259C02DCE6F818FC0D3CFB" box="[919,1022,1919,1941]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="obunca">
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<emphasis id="B946EA8A44259C02DCE6F818FC0D3CFB" box="[919,1022,1919,1941]" italics="true" pageId="7" pageNumber="267">B. obunca</emphasis>
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</taxonomicName>
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epandrium and associated structures left lateral; 22,
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<taxonomicName id="4C324D1B44259C02DE7CF807FE713CDA" box="[269,386,1950,1972]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="howdeni">
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<emphasis id="B946EA8A44259C02DE7CF807FE713CDA" box="[269,386,1950,1972]" italics="true" pageId="7" pageNumber="267">B. howdeni</emphasis>
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</taxonomicName>
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epandrium and associated structures left lateral. Abbreviations: SP, spermatheca; T7, T8, S5, S7, S8, sternites and tergites; P, phallapodeme; PG, postgonite; H, hypandrium; C, cercus; B, basiphallus; D, distiphallus; SU, surstylus.
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</paragraph>
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</caption>
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<subSubSection id="C3286513442A9C0DDFB8FF00FF093BBA" pageId="8" pageNumber="268" type="etymology">
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<paragraph id="8B8D3698442A9C0DDFB8FF00FF093BBA" blockId="8.[151,1436,151,391]" pageId="8" pageNumber="268">
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<emphasis id="B946EA8A442A9C0DDFB8FF00FEBC3BDE" bold="true" box="[201,335,152,176]" pageId="8" pageNumber="268">Etymology.</emphasis>
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<taxonomicName id="4C324D1B442A9C0DDE2DFF01FDB23BDE" box="[348,577,152,176]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
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||
<emphasis id="B946EA8A442A9C0DDE2DFF01FDB23BDE" box="[348,577,152,176]" italics="true" pageId="8" pageNumber="268">Bregmosina obunca</emphasis>
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||
</taxonomicName>
|
||
is named for the hooked surstylus (from the Latin
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<emphasis id="B946EA8A442A9C0DDBC5FF00FAE23BDE" box="[1204,1297,152,176]" italics="true" pageId="8" pageNumber="268">obuncus</emphasis>
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||
, bent in or hooked).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3286513442A9C0DDFB8FF47FCBB3AE9" pageId="8" pageNumber="268" type="discussion">
|
||
<paragraph id="8B8D3698442A9C0DDFB8FF47FCBB3AE9" blockId="8.[151,1436,151,391]" pageId="8" pageNumber="268">
|
||
<emphasis id="B946EA8A442A9C0DDFB8FF47FEA23B96" bold="true" box="[201,337,223,248]" pageId="8" pageNumber="268">Comments.</emphasis>
|
||
<taxonomicName id="4C324D1B442A9C0DDE29FF79FDC43B96" box="[344,567,224,248]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
|
||
<emphasis id="B946EA8A442A9C0DDE29FF79FDC43B96" box="[344,567,224,248]" italics="true" pageId="8" pageNumber="268">Bregmosina obunca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D1B442A9C0DDD1FFF79FD1F3B96" box="[622,748,224,248]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="howdeni">
|
||
<emphasis id="B946EA8A442A9C0DDD1FFF79FD1F3B96" box="[622,748,224,248]" italics="true" pageId="8" pageNumber="268">B. howdeni</emphasis>
|
||
</taxonomicName>
|
||
are obviously closely related but
|
||
<taxonomicName id="4C324D1B442A9C0DDB17FF79FB2B3B96" box="[1126,1240,224,248]" class="Insecta" family="Sphaeroceridae" genus="Bregmosina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="obunca">
|
||
<emphasis id="B946EA8A442A9C0DDB17FF79FB2B3B96" box="[1126,1240,224,248]" italics="true" pageId="8" pageNumber="268">B. obunca</emphasis>
|
||
</taxonomicName>
|
||
is distinct for the long, pointed anteroventral surstylus apex and the bilobed posterior margin of male sternite 4. Both species differ from congeners by their greatly reduced lower orbital bristle and by the strikingly large posterolateral processes of the male fifth sternite. The
|
||
<typeStatus id="5489883A442A9C0DDEA4FED4FDC43A0A" box="[469,567,332,356]" pageId="8" pageNumber="268" type="holotype">holotype</typeStatus>
|
||
was collected at the same time and place as B. buck, but these species are extremely different in male and female abdominal characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |