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<document id="75F466435D7F422E758A2B8669207877" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. &amp; Congdon, Bradley C." docDate="2021" docId="039B87838757FFFDFF1EF2181749FBDC" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Seraphs mayeri Jung 1974" docType="treatment" docVersion="7" lastPageNumber="414" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="414" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8F423BEA7A4FCAD4B542CAB3718B9E02">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="2F3E52B8C038DF0EC2189499F85C5E53">Maxwell, Stephen J.</mods:namePart>
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<mods:affiliation id="5BA00DE5464AA34D7545982DFDF1BE31">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="BB819D2B35AFD38B49F92BE9983CF375">Rymer, Tasmin L.</mods:namePart>
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<mods:affiliation id="1C4DE95C59EBC0E3D7DE980B4860CF21">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
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<mods:namePart id="3CEF8B2DE54C2586FBE96EBE71AE4258">Congdon, Bradley C.</mods:namePart>
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<mods:affiliation id="05739DA11F66D45E5AD6B16B737C24F7">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
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<treatment id="039B87838757FFFDFF1EF2181749FBDC" ID-DOI="http://doi.org/10.5281/zenodo.5088682" ID-GBIF-Taxon="183411020" ID-Zenodo-Dep="5088682" LSID="urn:lsid:plazi:treatment:039B87838757FFFDFF1EF2181749FBDC" httpUri="http://treatment.plazi.org/id/039B87838757FFFDFF1EF2181749FBDC" lastPageNumber="414" pageId="13" pageNumber="414">
<subSubSection id="C328651E8757FFFDFF1EF21812CCFDAE" box="[151,482,546,573]" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="8B8D36958757FFFDFF1EF21812CCFDAE" blockId="13.[151,482,546,608]" box="[151,482,546,573]" pageId="13" pageNumber="414">
<heading id="D0C581F98757FFFDFF1EF21812CCFDAE" bold="true" box="[151,482,546,573]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
<taxonomicName id="4C324D168757FFFDFF1EF21812CCFDAE" authority="Jung, 1974" authorityName="Jung" authorityYear="1974" box="[151,482,546,573]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="mayeri">
<emphasis id="B946EA878757FFFDFF1EF21812CCFDAE" bold="true" box="[151,482,546,573]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF1EF218127DFDAE" bold="true" box="[151,339,546,573]" italics="true" pageId="13" pageNumber="414">Seraphs mayeri</emphasis>
<bibRefCitation id="EFA34B648757FFFDFED2F21812CCFDAE" author="Jung, P." box="[347,482,546,572]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung, 1974</bibRefCitation>
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<subSubSection id="C328651E8757FFFDFF1EF27C1033FD53" pageId="13" pageNumber="414" type="description">
<paragraph id="8B8D36958757FFFDFF1EF27C1227FDF2" blockId="13.[151,482,546,608]" box="[151,265,582,608]" pageId="13" pageNumber="414">
<figureCitation id="13092A108757FFFDFF1EF27C1227FDF2" box="[151,265,582,608]" captionStart="FIGURE 4" captionStartId="8.[151,250,1766,1790]" captionTargetBox="[240,1350,188,1732]" captionTargetId="figure-19@8.[226,1361,181,1742]" captionTargetPageId="8" captionText="FIGURE 4. Seraphs Montfort, 1810. A. Seraphs andersoni (Dickerson, 1917), holotype, CAS Department of Geology 434, length 38 mm (Jung 1974, pl. 9, fig. 5). B. Seraphs belemnitus (Palmer, 1953), PRI 29236, length 48.2 mm (Jung 1974, pl. 6, figs. 1214). C. Seraphs caracassense (Leymerie, 1846), BMNH G50180, length 51.3mm (Jung 1974, pl. 4, figs. 10, 11). D. Seraphs chilophorus (Cossmann, 1889), MNHN A28779, Collection de Paleontologie, Paris, length 29 mm (Caze et al. 2010, figs. 15A, B, D). E. Seraphs hernandoensis (Mansfield, 1937), holotype, USNM 495948, length 31.4 mm (Jung 1974, pl. 9, figs. 79). F. Seraphs kaindraperi new sp., holotype, USNM 135098, length 50.6 mm (Jung 1974, pl. 5, figs. 3, 4). G. Seraphs leukoleptus Jung, 1974, neotype, MNHN A28781, Collection de Paleontologie, Paris, length 27.6 mm (Caze et al. 2010, figs. 11F, H, I). H. Seraphs mayeri Jung, 1974, holotype, NMB H16579, length 21.4 mm (Jung 1974, pl. 8, figs. 1618)." figureDoi="http://doi.org/10.5281/zenodo.5026787" httpUri="https://zenodo.org/record/5026787/files/figure.png" pageId="13" pageNumber="414">Figure 4H</figureCitation>
</paragraph>
<paragraph id="8B8D36958757FFFDFF1EF2B1104CFD31" blockId="13.[151,866,651,705]" box="[151,866,651,675]" pageId="13" pageNumber="414">
1974
<taxonomicName id="4C324D168757FFFDFF59F2B1113EFD31" authority="Jung" authorityName="Jung" authorityYear="1974" box="[208,528,651,675]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="mayeri" subGenus="Seraphs">
<emphasis id="B946EA878757FFFDFF59F2B1120CFD31" box="[208,290,651,675]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
(
<emphasis id="B946EA878757FFFDFEB8F2B112ADFD31" box="[305,387,651,675]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
)
<emphasis id="B946EA878757FFFDFE18F2B112F9FD30" box="[401,471,651,674]" italics="true" pageId="13" pageNumber="414">mayeri</emphasis>
Jung
</taxonomicName>
, p. 27, pl. 8, figs. 13 to 21, tf. 18.
</paragraph>
<paragraph id="8B8D36958757FFFDFF1EF2931033FD53" blockId="13.[151,866,651,705]" box="[151,797,681,705]" pageId="13" pageNumber="414">
2008
<taxonomicName id="4C324D168757FFFDFF59F2931161FD53" authority="Quaggioto &amp; Mellini" authorityName="Quaggioto &amp; Mellini" box="[208,591,681,705]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="sopitus">
<emphasis id="B946EA878757FFFDFF59F293125EFD53" box="[208,368,681,705]" italics="true" pageId="13" pageNumber="414">Seraphs sopitus</emphasis>
Quaggioto &amp; Mellini
</taxonomicName>
, p. 48, pl. V, fig. 28.
</paragraph>
</subSubSection>
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<paragraph id="8B8D36958757FFFDFF1EF2CA13CEFCBC" blockId="13.[151,1437,751,1102]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF1EF2CA1269FC9B" bold="true" box="[151,327,751,778]" pageId="13" pageNumber="414">Type material.</emphasis>
<materialsCitation id="3B5A3CC88757FFFDFEC6F2D513CEFCBC" ID-GBIF-Occurrence="3320081328" accessionNumber="H16578" location="NMB" pageId="13" pageNumber="414" specimenCount="1" stateProvince="Namibe" typeStatus="holotype">
<typeStatus id="548988378757FFFDFEC6F2D51299FC9B" box="[335,439,751,777]" pageId="13" pageNumber="414" type="holotype">Holotype</typeStatus>
<collectingRegion id="49F6F8778757FFFDFE5DF2D5113DFC9B" box="[468,531,751,777]" country="Angola" name="Namibe" pageId="13" pageNumber="414">NMB</collectingRegion>
H16578. The type measures
<quantity id="4CCA9B708757FFFDFCEDF2CA10E7FC98" box="[868,969,752,778]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" pageId="13" pageNumber="414" unit="mm" value="21.4">21.4 mm</quantity>
in length, with a width of
<quantity id="4CCA9B708757FFFDFB77F2CA1678FC98" box="[1278,1366,752,778]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="414" unit="mm" value="7.3">7.3 mm</quantity>
(
<bibRefCitation id="EFA34B648757FFFDFAEFF2CA13FDFCBC" author="Jung, P." pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
).
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</paragraph>
<paragraph id="8B8D36958757FFFDFF4EF30210B4FCC0" blockId="13.[151,1437,751,1102]" box="[199,922,823,850]" pageId="13" pageNumber="414">
<materialsCitation id="3B5A3CC88757FFFDFF4EF30210B9FCC0" ID-GBIF-Occurrence="3320081365" box="[199,919,823,850]" country="Italy" location="Monte Grumi" municipality="Castelgomberto" pageId="13" pageNumber="414" specimenCount="1" typeStatus="holotype">
<emphasis id="B946EA878757FFFDFF4EF3021248FCC3" bold="true" box="[199,358,823,850]" pageId="13" pageNumber="414">
<typeStatus id="548988378757FFFDFF4EF302122CFCC0" box="[199,258,824,850]" pageId="13" pageNumber="414">Type</typeStatus>
locality.
</emphasis>
<location id="8EED604E8757FFFDFEE5F3021126FCC3" LSID="urn:lsid:plazi:treatment:039B87838757FFFDFF1EF2181749FBDC:8EED604E8757FFFDFEE5F3021126FCC3" box="[364,520,823,850]" country="Italy" municipality="Castelgomberto" name="Monte Grumi" pageId="13" pageNumber="414">Monte Grumi</location>
,
<collectingMunicipality id="6BE9ACEF8757FFFDFD9DF30D11E6FCC3" box="[532,712,823,849]" pageId="13" pageNumber="414">Castelgomberto</collectingMunicipality>
,
<collectingCountry id="F32576058757FFFDFD5AF30D1028FCC3" box="[723,774,823,849]" name="Italy" pageId="13" pageNumber="414">Italy</collectingCountry>
[Oligocene]
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.
</paragraph>
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<subSubSection id="C328651E8757FFFDFF4EF36111ADFC2F" pageId="13" pageNumber="414" type="diagnosis">
<paragraph id="8B8D36958757FFFDFF4EF36111ADFC2F" blockId="13.[151,1437,751,1102]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF361126CFCE7" bold="true" box="[199,322,859,885]" pageId="13" pageNumber="414">Diagnosis.</emphasis>
The shell is moderately slender with a moderately acute apex. The labium is thin and continues above the adapical end of the aperture to the apex. The labrum is thin but thickens forming a posterior canal. The columella is basally slightly bent backward.
</paragraph>
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<subSubSection id="C328651E8757FFFDFF4EF3FD12DFFB94" pageId="13" pageNumber="414" type="distribution">
<paragraph id="8B8D36958757FFFDFF4EF3FD12DFFB94" blockId="13.[151,1437,751,1102]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF3FD124FFC73" bold="true" box="[199,353,967,993]" pageId="13" pageNumber="414">Distribution.</emphasis>
OLIGOCENE—
<emphasis id="B946EA878757FFFDFDADF3FD1179FC73" box="[548,599,967,993]" italics="true" pageId="13" pageNumber="414">
<collectingCountry id="F32576058757FFFDFDADF3FD1179FC73" box="[548,599,967,993]" name="Italy" pageId="13" pageNumber="414">Italy</collectingCountry>
</emphasis>
Monte Grumi, near Castelgomberto (
<bibRefCitation id="EFA34B648757FFFDFC75F3F2175BFC70" author="Jung, P." box="[1020,1141,968,994]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
); Santa Trinità, near Mon- tecchio Maggiore (
<bibRefCitation id="EFA34B648757FFFDFEE2F3D612CAFB94" author="Jung, P." box="[363,484,1004,1030]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF4EF42A1749FBDC" pageId="13" pageNumber="414" type="discussion">
<paragraph id="8B8D36958757FFFDFF4EF42A1749FBDC" blockId="13.[151,1437,751,1102]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF42A1215FBB8" bold="true" box="[199,315,1040,1066]" pageId="13" pageNumber="414">Remarks.</emphasis>
A small
<taxonomicName id="4C324D168757FFFDFE15F43512D8FBBB" authorityName="Montfort" authorityYear="1810" box="[412,502,1039,1065]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878757FFFDFE15F43512D8FBBB" box="[412,502,1039,1065]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
</taxonomicName>
similar to
<taxonomicName id="4C324D168757FFFDFDE4F42A1021FBBB" box="[621,783,1039,1066]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="chilophorus">
<emphasis id="B946EA878757FFFDFDE4F42A1021FBBB" box="[621,783,1039,1066]" italics="true" pageId="13" pageNumber="414">S. chilophorus</emphasis>
</taxonomicName>
, but the shoulder of that species is more defined, and there is no reabsorption of the shell behind the posterior canal in
<taxonomicName id="4C324D168757FFFDFCB9F40E174CFBDC" authority="(Jung 1974)" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[816,1122,1075,1102]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="chilophorus">
<emphasis id="B946EA878757FFFDFCB9F40E10FCFBDF" box="[816,978,1075,1102]" italics="true" pageId="13" pageNumber="414">S. chilophorus</emphasis>
(
<bibRefCitation id="EFA34B648757FFFDFC69F40E1774FBDC" author="Jung, P." box="[992,1114,1076,1102]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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