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<document id="0D9E03B04F941CF8B8C8245BF1E261C8" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. &amp; Congdon, Bradley C." docDate="2021" docId="039B87838754FFFFFF1EF62112F0FED3" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Seraphs peterjungi Caze, Merle, Pacaud" docType="treatment" docVersion="7" lastPageNumber="416" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="415" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1ADDC31045E72DBF5BDE7BA9854B16AD">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="D8C5E33B02D7C5B442D29DE56E14DE1C">Maxwell, Stephen J.</mods:namePart>
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<mods:affiliation id="A4D3BBD8C8B53236854022E7A9CF1642">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="625D7D4F88DEEF949D2F16829418C000">Rymer, Tasmin L.</mods:namePart>
<mods:nameIdentifier id="E4B85ADA1DF13DFB5CF3E9C9B798D584" type="ORCID">0000-0002-9963-6345</mods:nameIdentifier>
<mods:affiliation id="0B48B839BEE1DBED768F8B293A7573CB">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
<mods:nameIdentifier id="0D540F03C4F924F4230897F48256D1A4" type="email">amsin.rymer@jcu.edu.au</mods:nameIdentifier>
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<mods:namePart id="BFDF30F608255D8D7F6276AEACC2EA0E">Congdon, Bradley C.</mods:namePart>
<mods:nameIdentifier id="EDB65652FF95A981F664A89ABDA142E8" type="ORCID">0000-0002-8751-0892</mods:nameIdentifier>
<mods:affiliation id="433E0E295CDE43DD52A4F550AD79B3BF">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
<mods:nameIdentifier id="DBDCB27B5A0A410CF88928114F0DFB06" type="email">brad.congdon@jcu.edu.au</mods:nameIdentifier>
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<mods:title id="65C485F44EF38B9B2E025CD2800FC447">Zootaxa</mods:title>
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<mods:date id="47EC92DA444A196203DC2032C6AEB5B3">2021</mods:date>
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<mods:number id="0CECB4B4FBB6C550F3267E63B1D55546">2021-06-22</mods:number>
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<treatment id="039B87838754FFFFFF1EF62112F0FED3" ID-DOI="http://doi.org/10.5281/zenodo.5088694" ID-GBIF-Taxon="183411029" ID-Zenodo-Dep="5088694" LSID="urn:lsid:plazi:treatment:039B87838754FFFFFF1EF62112F0FED3" httpUri="http://treatment.plazi.org/id/039B87838754FFFFFF1EF62112F0FED3" lastPageId="15" lastPageNumber="416" pageId="14" pageNumber="415">
<subSubSection id="C328651E8754FFFEFF1EF62110B7F9A4" box="[151,921,1563,1590]" pageId="14" pageNumber="415" type="nomenclature">
<paragraph id="8B8D36958754FFFEFF1EF62110B7F9A4" blockId="14.[151,921,1563,1627]" box="[151,921,1563,1590]" pageId="14" pageNumber="415">
<heading id="D0C581F98754FFFEFF1EF62110B7F9A4" bold="true" box="[151,921,1563,1590]" fontSize="11" level="1" pageId="14" pageNumber="415" reason="1">
<emphasis id="B946EA878754FFFEFF1EF62110B7F9A4" bold="true" box="[151,921,1563,1590]" pageId="14" pageNumber="415">
<taxonomicName id="4C324D168754FFFEFF1EF62111A9F9A4" authority="Caze, Merle, Pacaud" authorityName="Caze, Merle, Pacaud" box="[151,647,1563,1590]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="415" phylum="Mollusca" rank="species" species="peterjungi">
<emphasis id="B946EA878754FFFEFF1EF6211252F9A4" bold="true" box="[151,380,1563,1590]" italics="true" pageId="14" pageNumber="415">Seraphs peterjungi</emphasis>
Caze, Merle, Pacaud
</taxonomicName>
&amp;
<collectingCountry id="F32576058754FFFEFD27F626107FF9A4" box="[686,849,1564,1590]" name="Sint Martin" pageId="14" pageNumber="415">Saint Martin</collectingCountry>
, 2010
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C328651E8754FFFEFF1EF67A11D2F90F" pageId="14" pageNumber="415" type="description">
<paragraph id="8B8D36958754FFFEFF1EF67A1227F9C9" blockId="14.[151,921,1563,1627]" box="[151,265,1600,1627]" pageId="14" pageNumber="415">
<figureCitation id="13092A108754FFFEFF1EF67A1227F9C9" box="[151,265,1600,1627]" captionStart="FIGURE 5" captionStartId="9.[151,250,1632,1656]" captionTargetBox="[169,1422,191,1598]" captionTargetId="figure-19@9.[151,1435,181,1609]" captionTargetPageId="9" captionText="FIGURE 5. Seraphs Montfort, 1810. A. Seraphs minus (Vincent, 1913), length 7.1 mm (Jung 1974, pl. 6, figs. 9, 10). B. Seraphs naricus (Vredenberg, 1925), holotype, GSI 12614, length 38.6 mm (Jung 1974, pl. 6, figs. 20–22). C. Seraphs olivaceus (Cossmann, 1889), MNHN A28577, Collection de Paleontologie, Paris, length 32 mm (Caze et al. 2010, figs. 6H, I). D. Seraphs peterjungi Caze et al., 2010, holotype, MNHN A28578, Collection de Paleontologie, length 16 mm (Caze et al. 2010, figs. 15J, K, N). E. Seraphs placiferus (Bayan, 1870), lectotype, MNHN G2100, length 33.7 mm (Jung 1974, pl. 7, figs. 5–7). F. Seraphs sopita (Brander, 1766), lectotype, NHMUK GG21010, length 26.4 mm (Jung 1974, pl. 1, figs. 1–3). G. Seraphs squamosus (Martin, 1914), Lectotype, RGM, Leiden St10344, length 37.1 mm (Jung 1974, pl. 6, figs. 17–19). H. Seraphs striatus (von Koenen, 1889), holotype, length 38.5 mm (von Koenen 1889, pl. I, figs. 7A–C). I. Seraphs subconvolutus (d’Orbigny, 1852), lectotype, MNHN UBT65-2-120, length 28 mm (Caze et al. 2010, figs. 14A, B). J. Seraphs volutatus (Brander, 1766), MNHN A28776, Collection de Paleontologie, Paris, length 17 mm (Caze et al. 2010, figs. 6A–C)." figureDoi="http://doi.org/10.5281/zenodo.5026789" httpUri="https://zenodo.org/record/5026789/files/figure.png" pageId="14" pageNumber="415">Figure 5D</figureCitation>
</paragraph>
<paragraph id="8B8D36958754FFFEFF1EF6BF11D2F90F" blockId="14.[151,764,1668,1693]" box="[151,764,1668,1693]" pageId="14" pageNumber="415">
2010
<taxonomicName id="4C324D168754FFFEFF59F6BE112DF90E" authority="Caze et al." authorityName="Caze" box="[208,515,1668,1693]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="415" phylum="Mollusca" rank="species" species="peterjungi">
<emphasis id="B946EA878754FFFEFF59F6BE12BEF90E" box="[208,400,1668,1692]" italics="true" pageId="14" pageNumber="415">Seraphs peterjungi</emphasis>
Caze
<emphasis id="B946EA878754FFFEFE59F6BF112DF90E" box="[464,515,1668,1692]" italics="true" pageId="14" pageNumber="415">et al.</emphasis>
</taxonomicName>
, p. 445, figs. 15J–N, 19.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8754FFFEFF1EF6F111BEF8BF" pageId="14" pageNumber="415" type="materials_examined">
<paragraph id="8B8D36958754FFFEFF1EF6F113CEF89B" blockId="14.[151,1437,1738,2017]" pageId="14" pageNumber="415">
<emphasis id="B946EA878754FFFEFF1EF6F11268F976" bold="true" box="[151,326,1738,1765]" pageId="14" pageNumber="415">Type material.</emphasis>
<materialsCitation id="3B5A3CC88754FFFEFEC4F6F113CEF89B" ID-GBIF-Occurrence="3320081317" accessionNumber="A28578" location="MNHN" pageId="14" pageNumber="415" specimenCount="1" stateProvince="Monaghan" typeStatus="holotype">
<typeStatus id="548988378754FFFEFEC4F6F1129AF977" box="[333,436,1739,1765]" pageId="14" pageNumber="415" type="holotype">Holotype</typeStatus>
—
<collectingRegion id="49F6F8778754FFFEFE58F6F11109F977" box="[465,551,1739,1765]" country="Ireland" name="Monaghan" pageId="14" pageNumber="415">MNHN</collectingRegion>
<accessionNumber id="9461AB768754FFFEFDA4F6F011A5F977" box="[557,651,1738,1765]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28578" pageId="14" pageNumber="415" type="EnaNcbi">A28578</accessionNumber>
. The type measures
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length, with a width of
<quantity id="4CCA9B708754FFFEFB58F6F1163DF977" box="[1233,1299,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="415" unit="mm" value="3.0">3 mm</quantity>
(
<bibRefCitation id="EFA34B648754FFFEFAABF6F113FAF89B" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." pageId="14" pageNumber="415" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878754FFFEFAEAF6F116B3F977" box="[1379,1437,1739,1765]" italics="true" pageId="14" pageNumber="415">et al.</emphasis>
2010
</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8D36958754FFFEFF4EF72911BEF8BF" blockId="14.[151,1437,1738,2017]" box="[199,656,1810,1837]" pageId="14" pageNumber="415">
<materialsCitation id="3B5A3CC88754FFFEFF4EF72911A2F8BF" ID-GBIF-Occurrence="3320081312" box="[199,652,1810,1837]" country="France" location="Chaussy" municipality="Lutetian" pageId="14" pageNumber="415" specimenCount="1" typeStatus="holotype">
<emphasis id="B946EA878754FFFEFF4EF7291248F8BE" bold="true" box="[199,358,1810,1837]" pageId="14" pageNumber="415">
<typeStatus id="548988378754FFFEFF4EF729122CF8BF" box="[199,258,1811,1837]" pageId="14" pageNumber="415">Type</typeStatus>
locality.
</emphasis>
<location id="8EED604E8754FFFEFEE5F72912E2F8BF" LSID="urn:lsid:plazi:treatment:039B87838754FFFFFF1EF62112F0FED3:8EED604E8754FFFEFEE5F72912E2F8BF" box="[364,460,1811,1837]" country="France" municipality="Lutetian" name="Chaussy" pageId="14" pageNumber="415">Chaussy</location>
,
<collectingCountry id="F32576058754FFFEFE5EF729110DF8BF" box="[471,547,1811,1837]" name="France" pageId="14" pageNumber="415">France</collectingCountry>
<collectingMunicipality id="6BE9ACEF8754FFFEFDA2F72911A2F8BF" box="[555,652,1811,1837]" pageId="14" pageNumber="415">Lutetian</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8754FFFEFF4EF70C1041F873" pageId="14" pageNumber="415" type="diagnosis">
<paragraph id="8B8D36958754FFFEFF4EF70C1041F873" blockId="14.[151,1437,1738,2017]" pageId="14" pageNumber="415">
<emphasis id="B946EA878754FFFEFF4EF70C126CF8C2" bold="true" box="[199,322,1846,1872]" pageId="14" pageNumber="415">Diagnosis.</emphasis>
The shell is moderately ovate and elongated, with the widest part being just below the adapical end of the aperture. The apex is moderately blunt. The surface of the shell is smooth with faint sinuous growth lines. The labial callus is well-developed and conspicuous. The labrum is not thickened and has a moderately quadrate profile. The columella is recurved backward near the base. The base of the labrum does not extend as far as the end of the columella. The anterior of the body whorl has fine axial grooves.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8755FFFFFF4EF0AC1153FF6B" pageId="15" pageNumber="416" type="distribution">
<paragraph id="8B8D36958755FFFFFF4EF0AC1153FF6B" blockId="15.[151,1436,150,321]" pageId="15" pageNumber="416">
<emphasis id="B946EA878755FFFFFF4EF0AC124FFF22" bold="true" box="[199,353,150,176]" pageId="15" pageNumber="416">Distribution.</emphasis>
Lutetian—
<emphasis id="B946EA878755FFFFFE68F0AD111CFF22" box="[481,562,151,176]" italics="true" pageId="15" pageNumber="416">
<collectingCountry id="F32576058755FFFFFE68F0AD111CFF22" box="[481,562,151,176]" name="France" pageId="15" pageNumber="416">France</collectingCountry>
</emphasis>
Chaussy, Val d’Oise (
<bibRefCitation id="EFA34B648755FFFFFCA0F0AD10C8FF23" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[809,998,151,177]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFCE3F0AD108CFF23" box="[874,930,151,177]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
</bibRefCitation>
); Grignon, Yvelines (
<bibRefCitation id="EFA34B648755FFFFFB5AF0AD16BEFF23" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[1235,1424,151,177]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFA9DF0AD1662FF23" box="[1300,1356,151,177]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
</bibRefCitation>
); Beynes, Yvelines (
<bibRefCitation id="EFA34B648755FFFFFEEDF0811130FF47" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[356,542,187,213]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFE2DF08112F5FF47" box="[420,475,187,213]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
</bibRefCitation>
); Saulx-Marchais, Yvelines (
<bibRefCitation id="EFA34B648755FFFFFCD4F0811739FF47" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[861,1047,187,213]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFC15F08110FAFF47" box="[924,980,187,213]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
</bibRefCitation>
); Fercourt, Oise (
<bibRefCitation id="EFA34B648755FFFFFB5FF08116BEFF47" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[1238,1424,187,213]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFA9FF0811663FF47" box="[1302,1357,187,213]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
</bibRefCitation>
); Parnes, Oise (
<bibRefCitation id="EFA34B648755FFFFFEBBF0E51284FF6B" author="Jung, P." box="[306,426,223,249]" pageId="15" pageNumber="416" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
;
<bibRefCitation id="EFA34B648755FFFFFE3FF0E5115FFF6B" author="Caze, B. &amp; Merle, D. &amp; Pacaud, J. - M. &amp; Martin, J. - P." box="[438,625,223,249]" pageId="15" pageNumber="416" pagination="417 - 477" refId="ref33015" refString="Caze, B., Merle, D., Pacaud, J. - M. &amp; Martin, J. - P. (2010) First systematic study using the variability of the residual colour patterns: the case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea). Geodiversitas, 32, 417 - 477. https: // doi. org / 10.5252 / g 2010 n 3 a 4" type="journal article" year="2010">
Caze
<emphasis id="B946EA878755FFFFFE7CF0E51103FF6B" box="[501,557,223,249]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
2010
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).
</paragraph>
</subSubSection>
<subSubSection id="C328651E8755FFFFFF4EF13912F0FED3" pageId="15" pageNumber="416" type="discussion">
<paragraph id="8B8D36958755FFFFFF4EF13912F0FED3" blockId="15.[151,1436,150,321]" pageId="15" pageNumber="416">
<emphasis id="B946EA878755FFFFFF4EF1391215FE8F" bold="true" box="[199,315,259,285]" pageId="15" pageNumber="416">Remarks.</emphasis>
This shell is only distinguishable from
<taxonomicName id="4C324D168755FFFFFD7BF13910BDFE8F" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[754,915,259,285]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="416" phylum="Mollusca" rank="species" species="chilophorus">
<emphasis id="B946EA878755FFFFFD7BF13910BDFE8F" box="[754,915,259,285]" italics="true" pageId="15" pageNumber="416">S. chilophorus</emphasis>
</taxonomicName>
by the residual colour under UV; for compara- tive remarks see that species.
</paragraph>
</subSubSection>
</treatment>
</document>
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