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<document id="319CA62996F3BD848475A4F90D9A950A" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. &amp; Congdon, Bradley C." docDate="2021" docId="039B87838745FFD0FF1EF5C51172FED2" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoterebellum diversiornatum Maxwell &amp; Rymer &amp; Congdon 2021" docType="treatment" docVersion="9" lastPageNumber="433" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="432" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="513BF9E0777C827AEC531502FDE5724E">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="78633A5AB7E0573CA58BB24B5BE87C07">Maxwell, Stephen J.</mods:namePart>
<mods:nameIdentifier id="42EE78B612F1F51FEC7A05D78877C3B8" type="ORCID">0000-0002-4328-6171</mods:nameIdentifier>
<mods:affiliation id="E387527C14776E76B6D1AB672C5C750D">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="6CF2D680C4F297FA78F2A261D1F815B9">Rymer, Tasmin L.</mods:namePart>
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<mods:affiliation id="B477D1B2CBC927F2244B28484969D6F2">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
<mods:nameIdentifier id="D2E3ABDEFDD1C7FAD5D6085D01FF7C0F" type="email">amsin.rymer@jcu.edu.au</mods:nameIdentifier>
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<mods:namePart id="5238B47B3CA3D38034AB62791AD735F6">Congdon, Bradley C.</mods:namePart>
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<mods:affiliation id="AE29F1A99E7A063A5C5C288C5831D217">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
<mods:nameIdentifier id="8A1F2AFDEE0A7AC70BE0976D666AE985" type="email">brad.congdon@jcu.edu.au</mods:nameIdentifier>
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<mods:title id="98F17491C940046000926CC59367C9AC">Zootaxa</mods:title>
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<mods:date id="809A346F88A8415E894B733ED1EDDF63">2021</mods:date>
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<mods:number id="AB3947D88AC81B091D3F871A8B2B9DBA">2021-06-22</mods:number>
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<treatment id="039B87838745FFD0FF1EF5C51172FED2" ID-DOI="http://doi.org/10.5281/zenodo.5088446" ID-GBIF-Taxon="183411038" ID-Zenodo-Dep="5088446" LSID="urn:lsid:plazi:treatment:039B87838745FFD0FF1EF5C51172FED2" httpUri="http://treatment.plazi.org/id/039B87838745FFD0FF1EF5C51172FED2" lastPageId="32" lastPageNumber="433" pageId="31" pageNumber="432">
<subSubSection id="C328651E8745FFEFFF1EF5C511C9F988" box="[151,743,1535,1562]" pageId="31" pageNumber="432" type="nomenclature">
<paragraph id="8B8D36958745FFEFFF1EF5C511C9F988" blockId="31.[151,743,1535,1598]" box="[151,743,1535,1562]" pageId="31" pageNumber="432">
<heading id="D0C581F98745FFEFFF1EF5C511C9F988" bold="true" box="[151,743,1535,1562]" fontSize="11" level="1" pageId="31" pageNumber="432" reason="1">
<taxonomicName id="4C324D168745FFEFFF1EF5C511C9F988" ID-CoL="8KTGT" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" baseAuthorityName="Eames" baseAuthorityYear="1952" box="[151,743,1535,1562]" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="433" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA878745FFEFFF1EF5C511C9F988" bold="true" box="[151,743,1535,1562]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF1EF5C51101F988" bold="true" box="[151,559,1535,1562]" italics="true" pageId="31" pageNumber="432">Pseudoterebellum diversiornatum</emphasis>
(
<bibRefCitation id="EFA34B648745FFEFFDC8F63A11F0F988" author="Eames, F. E." box="[577,734,1536,1562]" pageId="31" pageNumber="432" pagination="1 - 168" refId="ref34832" refString="Eames, F. E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Piunjab and in the Kohat District. Philosophical Transactions of the Royal Society of London, 236, 1 - 168. https: // doi. org / 10.1098 / rstb. 1952.0001" type="journal article" year="1952">Eames, 1952</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF1EF61E11D4F912" pageId="31" pageNumber="432" type="description">
<paragraph id="8B8D36958745FFEFFF1EF61E1224F9AC" blockId="31.[151,743,1535,1598]" box="[151,266,1572,1598]" pageId="31" pageNumber="432">
<figureCitation id="13092A108745FFEFFF1EF61E1224F9AC" box="[151,266,1572,1598]" captionStart="FIGURE 8" captionStartId="32.[151,250,1577,1601]" captionTargetBox="[204,1373,360,1524]" captionTargetId="figure-88@32.[180,1407,344,1553]" captionTargetPageId="32" captionText="FIGURE 8. Pseudoterebellum new genus and Terebellum Röding, 1798. A. Pseudoterebellum diversiornatum (Eames, 1952), holotype, NHMUK GG68239, length 10.1 mm (Jung 1974, pl. 14, figs. 12–14). B. Pseudoterebellum oympiae (Rolando, 2001), holotype, MGPD 28168, length 20.9 mm (Rolando 2001, fig. 2). C. Pseudoterebellum saudiensis (Abbass, 1972), holotype, ASU S53, length 25 mm (Abbass 1972, pl. V, figs. 1, 2). D. Terebellum cinctum Martin, 1916, holotype, RGM St10337, length 34.7 mm (Jung 1974, pl. 13, figs. 4, 6). E. Terebellum obtusum Sowerby, 1840, holotype, NHMUK GG21000, length 42.2 mm (Jung 1974, pl. 13, figs. 17, 18). F. Terebellum papilliferum Martin, 1916, lectotype, RGM St10339, length 28 mm (Jung 1974, pl. 13, figs. 9, 11)." figureDoi="http://doi.org/10.5281/zenodo.5026799" httpUri="https://zenodo.org/record/5026799/files/figure.png" pageId="31" pageNumber="432">Figure 8A</figureCitation>
</paragraph>
<paragraph id="8B8D36958745FFEFFF1EF65211D4F912" blockId="31.[151,762,1640,1665]" box="[151,762,1640,1665]" pageId="31" pageNumber="432">
1952
<taxonomicName id="4C324D168745FFEFFF59F6521100F912" authority="Eames" authorityName="Eames" authorityYear="1952" box="[208,558,1640,1664]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="diversiornatum">
<emphasis id="B946EA878745FFEFFF59F65212CCF912" box="[208,482,1640,1664]" italics="true" pageId="31" pageNumber="432">Terebellum diversiornatum</emphasis>
Eames
</taxonomicName>
, p. 74, pl. 3, fig. 77.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF1EF6951705F883" pageId="31" pageNumber="432" type="materials_examined">
<paragraph id="8B8D36958745FFEFFF1EF695120BF97F" blockId="31.[151,1437,1710,2025]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF1EF6951266F95A" bold="true" box="[151,328,1710,1737]" pageId="31" pageNumber="432">Type material.</emphasis>
<materialsCitation id="3B5A3CC88745FFEFFED8F695120BF97F" ID-GBIF-Occurrence="3320081354" collectionCode="NHMUK" pageId="31" pageNumber="432" specimenCode="GG68239" specimenCount="1" typeStatus="holotype">
<typeStatus id="548988378745FFEFFED8F6951296F95B" box="[337,440,1711,1737]" pageId="31" pageNumber="432" type="holotype">Holotype</typeStatus>
—
<collectionCode id="ED23AE508745FFEFFE5CF6951111F95B" box="[469,575,1711,1737]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="432" type="Museum">NHMUK</collectionCode>
<specimenCode id="DB949EEE8745FFEFFDC0F6951195F95B" box="[585,699,1711,1737]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="432" type="Museum">GG68239</specimenCode>
. The type measures
<quantity id="4CCA9B708745FFEFFC20F6951721F95B" box="[937,1039,1711,1737]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="31" pageNumber="432" unit="mm" value="10.1">10.1 mm</quantity>
in length with a width of
<quantity id="4CCA9B708745FFEFFACDF69416B3F95B" box="[1348,1437,1710,1737]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="432" unit="mm" value="4.5">4.5 mm</quantity>
(
<bibRefCitation id="EFA34B648745FFEFFF16F6E91236F97F" author="Jung, P." box="[159,280,1747,1773]" pageId="31" pageNumber="432" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8D36958745FFEFFF4EF6CD1705F883" blockId="31.[151,1437,1710,2025]" box="[199,1067,1782,1809]" pageId="31" pageNumber="432">
<materialsCitation id="3B5A3CC88745FFEFFF4EF6CD1708F883" ID-GBIF-Occurrence="3320081352" box="[199,1062,1782,1809]" country="Pakistan" county="Western" location="Zinda Pir" municipality="Ghazij Shales" pageId="31" pageNumber="432" specimenCount="1" stateProvince="Punjab" typeStatus="holotype">
<emphasis id="B946EA878745FFEFFF4EF6CD1248F882" bold="true" box="[199,358,1782,1809]" pageId="31" pageNumber="432">
<typeStatus id="548988378745FFEFFF4EF6CD122CF883" box="[199,258,1783,1809]" pageId="31" pageNumber="432">Type</typeStatus>
locality.
</emphasis>
<location id="8EED604E8745FFEFFEE5F6CD12F9F883" LSID="urn:lsid:plazi:treatment:039B87838745FFD0FF1EF5C51172FED2:8EED604E8745FFEFFEE5F6CD12F9F883" box="[364,471,1783,1809]" country="Pakistan" county="Western" municipality="Ghazij Shales" name="Zinda Pir" pageId="31" pageNumber="432" stateProvince="Punjab">Zinda Pir</location>
,
<collectingMunicipality id="6BE9ACEF8745FFEFFE6BF6CD11AEF883" box="[482,640,1783,1809]" pageId="31" pageNumber="432">Ghazij Shales</collectingMunicipality>
,
<collectingCounty id="62EC4E198745FFEFFD03F6CD11CBF883" box="[650,741,1783,1809]" pageId="31" pageNumber="432">Western</collectingCounty>
<collectingRegion id="49F6F8778745FFEFFD65F6CD1013F883" box="[748,829,1783,1809]" country="Pakistan" name="Punjab" pageId="31" pageNumber="432">Punjab</collectingRegion>
,
<collectingCountry id="F32576058745FFEFFCC1F6CD1089F883" box="[840,935,1783,1809]" name="Pakistan" pageId="31" pageNumber="432">Pakistan</collectingCountry>
[Ypresian]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF4EF7201190F87B" pageId="31" pageNumber="432" type="diagnosis">
<paragraph id="8B8D36958745FFEFFF4EF7201190F87B" blockId="31.[151,1437,1710,2025]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF720126CF8A6" bold="true" box="[199,322,1818,1844]" pageId="31" pageNumber="432">Diagnosis.</emphasis>
The shell is sub-elliptical. It has a short conoidal spire that comprises one-seventh of the length of the shell. The teleoconch whorls are flat with linear sutures. The height of the whorls is proportionally slightly less than the width of the whorl. The teleoconch has fine straight axial costae, numbering ten per mm, that are slightly wider than the interval. They fade on the penultimate whorl, with the only ornamentation being ten spiral bands. The body whorl is oliviform and smooth, with the greatest width above the middle and sides slightly convex. The base of the aperture is narrow with no columnar folds.
</paragraph>
</subSubSection>
<subSubSection id="C328651E877AFFD0FF4EF0AC17F2FF23" box="[199,1244,150,177]" pageId="32" pageNumber="433" type="distribution">
<paragraph id="8B8D3695877AFFD0FF4EF0AC17F2FF23" blockId="32.[151,1436,150,320]" box="[199,1244,150,177]" pageId="32" pageNumber="433">
<emphasis id="B946EA87877AFFD0FF4EF0AC124FFF22" bold="true" box="[199,353,150,176]" pageId="32" pageNumber="433">Distribution.</emphasis>
EOCENE—Ypresian—
<emphasis id="B946EA87877AFFD0FDF9F0AC11FCFF22" box="[624,722,150,176]" italics="true" pageId="32" pageNumber="433">
<collectingCountry id="F3257605877AFFD0FDF9F0AC11FCFF22" box="[624,722,150,176]" name="Pakistan" pageId="32" pageNumber="433">Pakistan</collectingCountry>
</emphasis>
Ghazij Shales, Western
<collectingRegion id="49F6F877877AFFD0FC6DF0AC171CFF22" box="[996,1074,150,176]" country="Pakistan" name="Punjab" pageId="32" pageNumber="433">Punjab</collectingRegion>
(
<bibRefCitation id="EFA34B64877AFFD0FBC9F0AC17E1FF23" author="Eames, F. E." box="[1088,1231,150,177]" pageId="32" pageNumber="433" pagination="1 - 168" refId="ref34832" refString="Eames, F. E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Piunjab and in the Kohat District. Philosophical Transactions of the Royal Society of London, 236, 1 - 168. https: // doi. org / 10.1098 / rstb. 1952.0001" type="journal article" year="1952">Eames 1952</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328651E877AFFD0FF4EF0801172FED2" pageId="32" pageNumber="433" type="discussion">
<paragraph id="8B8D3695877AFFD0FF4EF0801172FED2" blockId="32.[151,1436,150,320]" pageId="32" pageNumber="433">
<emphasis id="B946EA87877AFFD0FF4EF0801215FF46" bold="true" box="[199,315,186,212]" pageId="32" pageNumber="433">Remarks.</emphasis>
<bibRefCitation id="EFA34B64877AFFD0FEC8F08112E9FF46" author="Jung, P." box="[321,455,186,213]" pageId="32" pageNumber="433" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung (1974)</bibRefCitation>
described the
<typeStatus id="54898837877AFFD0FDE1F08011E4FF46" box="[616,714,186,212]" pageId="32" pageNumber="433" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C324D16877AFFD0FD67F081109AFF46" authorityName="Eames" authorityYear="1952" box="[750,948,186,212]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="433" phylum="Mollusca" rank="species" species="diversiornatum">
<emphasis id="B946EA87877AFFD0FD67F081109AFF46" box="[750,948,186,212]" italics="true" pageId="32" pageNumber="433">P. diversiornatum</emphasis>
</taxonomicName>
as an incomplete juvenile internal cast with problematic affinities.Additional weight to the generation of the new clade is provided by
<bibRefCitation id="EFA34B64877AFFD0FB09F0E51633FF6B" author="Eames, F. E." box="[1152,1309,222,249]" pageId="32" pageNumber="433" pagination="1 - 168" refId="ref34832" refString="Eames, F. E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Piunjab and in the Kohat District. Philosophical Transactions of the Royal Society of London, 236, 1 - 168. https: // doi. org / 10.1098 / rstb. 1952.0001" type="journal article" year="1952">Eames (1952)</bibRefCitation>
, who noted that
<taxonomicName id="4C324D16877AFFD0FF4EF13912A2FE8E" authorityName="Eames" authorityYear="1952" box="[199,396,258,284]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="433" phylum="Mollusca" rank="species" species="diversiornatum">
<emphasis id="B946EA87877AFFD0FF4EF13912A2FE8E" box="[199,396,258,284]" italics="true" pageId="32" pageNumber="433">P. diversiornatum</emphasis>
</taxonomicName>
belonged to an as yet undefined subgenus of
<taxonomicName id="4C324D16877AFFD0FC09F13810D2FE8E" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[896,1020,258,284]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="433" phylum="Mollusca" rank="genus">
<emphasis id="B946EA87877AFFD0FC09F13810D2FE8E" box="[896,1020,258,284]" italics="true" pageId="32" pageNumber="433">Terebellum</emphasis>
</taxonomicName>
. A defining feature of
<taxonomicName id="4C324D16877AFFD0FB7DF13813EFFED2" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="433" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA87877AFFD0FB7DF13813EFFED2" italics="true" pageId="32" pageNumber="433">Pseudoterebellum</emphasis>
</taxonomicName>
is the shell being widest posteriorly.
</paragraph>
</subSubSection>
</treatment>
</document>
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