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<document id="344EB1B865934131F84D15CF75D1CC09" ID-DOI="10.5281/zenodo.193519" ID-GBIF-Dataset="1d3efd1e-f123-4214-860c-67d8d297b34f" ID-ISSN="1175-5326" ID-Zenodo-Dep="193519" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460151470905" checkinUser="plazi" docAuthor="Millet, Johann &amp; Nel, Andre" docDate="2010" docId="03934D06FFB4FF8EFF11F87259F6FC22" docLanguage="en" docName="zt02353p054.pdf" docOrigin="Zootaxa 2353" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Araripeleon alphonsei" docType="treatment" docVersion="8" lastPageNumber="53" masterDocId="FFAA357EFFB4FF8AFF86FF6D5C62FFE0" masterDocTitle="A new myrmeleontoid genus from the Crato Formation of Northeast Brazil (Lower Cretaceous) (Insecta: Neuroptera: Palaeoleontidae)" masterLastPageNumber="54" masterPageNumber="49" pageNumber="49" updateTime="1698235497634" updateUser="plazi">
<mods:mods id="CEEF80F51C48CA8C11D6B3F80EB03B8A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F84F2AE679E8D795BEB95571D1A3EFEF">
<mods:title id="3DB6E6A6B5D9D1E2D7402511112962CD">A new myrmeleontoid genus from the Crato Formation of Northeast Brazil (Lower Cretaceous) (Insecta: Neuroptera: Palaeoleontidae)</mods:title>
</mods:titleInfo>
<mods:name id="0EFF583E55C965556B7E37FC8367D80F" type="personal">
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<mods:namePart id="9CE1D556986F4CC254EC8CBD7C105910">Millet, Johann</mods:namePart>
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<mods:namePart id="CAD9EFD8ECCBB3392642EAABF2977CB5">Nel, Andre</mods:namePart>
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<mods:typeOfResource id="538367E4D4C5A71ED65449BC480C597F">text</mods:typeOfResource>
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<mods:title id="AF5304528091F26B17263FB834BE5B82">Zootaxa</mods:title>
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<mods:part id="BEB872F480A9988435C5AA0E1BB604A8">
<mods:date id="C20E64A6FAEC9CF4F2F138D15C6B6608">2010</mods:date>
<mods:detail id="54B18B5854B95C1E2B18780AA1FAB9DE" type="volume">
<mods:number id="843B0E669AC9F77F373E4779C014CB7A">2353</mods:number>
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<mods:classification id="7615598244CCF745E5E62DAB7D3BCA9D">journal article</mods:classification>
<mods:identifier id="472F970EC50F374AE6651F53D4F15333" type="DOI">10.5281/zenodo.193519</mods:identifier>
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<mods:identifier id="E4BE1E1F3A14C20E3CCFC7F10302A4F6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="79503436502F2ED86330E4AAFA7A67AB" type="Zenodo-Dep">193519</mods:identifier>
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<treatment id="03934D06FFB4FF8EFF11F87259F6FC22" ID-DOI="http://doi.org/10.5281/zenodo.6205791" ID-GBIF-Taxon="124955980" ID-Zenodo-Dep="6205791" LSID="urn:lsid:plazi:treatment:03934D06FFB4FF8EFF11F87259F6FC22" httpUri="http://treatment.plazi.org/id/03934D06FFB4FF8EFF11F87259F6FC22" lastPageId="4" lastPageNumber="53" pageId="0" pageNumber="49">
<subSubSection id="C320AF9BFFB4FF8AFF11F8725D7AF8BC" pageId="0" pageNumber="49" type="nomenclature">
<paragraph id="8B85FC10FFB4FF8AFF11F8725E5FF8D9" blockId="0.[151,573,1823,1884]" box="[151,573,1823,1849]" pageId="0" pageNumber="49">
<heading id="D0CD4B7CFFB4FF8AFF11F8725E5FF8D9" bold="true" box="[151,573,1823,1849]" fontSize="11" level="1" pageId="0" pageNumber="49" reason="1">
<emphasis id="B94E2002FFB4FF8AFF11F8725E5FF8D9" bold="true" box="[151,573,1823,1849]" pageId="0" pageNumber="49">
<taxonomicName id="4C3A8793FFB4FF8AFF11F8725DC6F8D9" ID-CoL="G62Z" box="[151,420,1823,1849]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="alphonsei">
<emphasis id="B94E2002FFB4FF8AFF11F8725DC6F8D9" bold="true" box="[151,420,1823,1849]" italics="true" pageId="0" pageNumber="49">Araripeleon alphonsei</emphasis>
</taxonomicName>
gen.
<emphasis id="B94E2002FFB4FF8AFE60F8725D99F8D9" bold="true" box="[486,507,1823,1849]" italics="true" pageId="0" pageNumber="49">et</emphasis>
sp n.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B85FC10FFB4FF8AFF11F82F5D7AF8BC" blockId="0.[151,573,1823,1884]" box="[151,280,1858,1884]" pageId="0" pageNumber="49">
(
<figureCitation id="1301E095FFB4FF8AFF19F82F5D6CF8BC" box="[159,270,1858,1884]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,255,1123,1147]" captionStartId-1="2.[151,255,1917,1941]" captionStartId-2="3.[151,255,770,794]" captionTargetBox-0="[228,1358,197,1099]" captionTargetBox-1="[244,1360,954,1861]" captionTargetBox-2="[255,1316,239,665]" captionTargetId-0="figure@1.[228,1360,197,1099]" captionTargetId-1="figure@2.[213,1373,903,1896]" captionTargetId-2="figure@3.[255,1332,194,749]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Araripeleon alphonsei gen. nov., sp. nov., holotype SMNK 2358 PAL, photograph of general habitus (scale bar represents 10 mm)." captionText-1="FIGURE 2. Araripeleon alphonsei gen. nov., sp. nov., holotype SMNK 2358 PAL, drawing of general habitus, with the forewing details removed (scale bar represents 10 mm)." captionText-2="FIGURE 3. Araripeleon alphonsei gen. nov., sp. nov., holotype SMNK 2358 PAL, drawing of left forewing (scale bar represents 10 mm)." httpUri-0="https://zenodo.org/record/193520/files/figure.png" httpUri-1="https://zenodo.org/record/193521/files/figure.png" httpUri-2="https://zenodo.org/record/193522/files/figure.png" pageId="0" pageNumber="49">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<paragraph id="8B85FC10FFB4FF8AFF11F8E15D68F811" pageId="0" pageNumber="49">
<table id="F93A0EB0FFB40075FF11F8E159FEF811" box="[151,1436,1932,2033]" gridcols="9" gridrows="3" pageId="0" pageNumber="49">
<tr id="350AFE52FFB40075FF11F8E159FEF846" box="[151,1436,1932,1958]" gridrow="0" pageId="0" pageNumber="49" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76DB972EFFB40075FF11F8E15EC7F846" box="[151,677,1932,1958]" gridcol="0" gridrow="0" pageId="0" pageNumber="49">
<emphasis id="B94E2002FFB4FF8AFF11F8E15D50F846" bold="true" box="[151,306,1932,1958]" pageId="0" pageNumber="49">Type species</emphasis>
:
<taxonomicName id="4C3A8793FFB4FF8AFEC7F8E05E2AF846" box="[321,584,1933,1958]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="alphonsei" status="sp. nov.">
<emphasis id="B94E2002FFB4FF8AFEC7F8E05E2AF846" box="[321,584,1933,1958]" italics="true" pageId="0" pageNumber="49">Araripeleon alphonsei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D9D79FFB4FF8AFDC9F8E15EECF846" box="[591,654,1932,1958]" pageId="0" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</th>
</tr>
<tr id="350AFE52FFB40075FF11F8DC59FEF82C" box="[151,1436,1969,1996]" gridrow="1" pageId="0" pageNumber="49">
<th id="76DB972EFFB40075FF11F8DC5EC7F82C" box="[151,677,1969,1996]" gridcol="0" gridrow="1" pageId="0" pageNumber="49">
<emphasis id="B94E2002FFB4FF8AFF40F8DC5D51F82B" bold="true" box="[198,307,1969,1995]" pageId="0" pageNumber="49">Material</emphasis>
: Holotype SMNK 2358 PAL,
</th>
<td id="76DB972EFFB40075FD34F8DC5E98F82C" box="[690,762,1969,1996]" gridcol="1" gridrow="1" pageId="0" pageNumber="49">stored</td>
<td id="76DB972EFFB40075FC83F8DC5F78F82C" box="[773,794,1969,1996]" gridcol="2" gridrow="1" pageId="0" pageNumber="49">at</td>
<td id="76DB972EFFB40075FCA1F8DC5F29F82C" box="[807,843,1969,1996]" gridcol="3" gridrow="1" pageId="0" pageNumber="49">the</td>
<td id="76DB972EFFB40075FCD1F8DC5FB7F82C" box="[855,981,1969,1996]" gridcol="4" gridrow="1" pageId="0" pageNumber="49">Staatliches</td>
<td id="76DB972EFFB40075FC64F8DC582BF82C" box="[994,1097,1969,1996]" gridcol="5" gridrow="1" pageId="0" pageNumber="49">Museum</td>
<td id="76DB972EFFB40075FBD2F8DC5814F82C" box="[1108,1142,1969,1996]" gridcol="6" gridrow="1" pageId="0" pageNumber="49">für</td>
<td id="76DB972EFFB40075FB05F8DC5976F82C" box="[1155,1300,1969,1996]" gridcol="7" gridrow="1" pageId="0" pageNumber="49">Naturkunde,</td>
<td id="76DB972EFFB40075FAA7F8DC59FEF82C" box="[1313,1436,1969,1996]" gridcol="8" gridrow="1" pageId="0" pageNumber="49">Karlsruhe,</td>
</tr>
<tr id="350AFE52FFB40075FF11F8BA59FEF811" box="[151,1436,2007,2033]" gridrow="2" pageId="0" pageNumber="49" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76DB972EFFB40075FF11F8BA5EC7F811" box="[151,677,2007,2033]" gridcol="0" gridrow="2" pageId="0" pageNumber="49">Germany.</th>
</tr>
</table>
</paragraph>
<caption id="DF45AC98FFB5FF8BFF11FB0E5DEFFB7A" httpUri="https://zenodo.org/record/193520/files/figure.png" pageId="1" pageNumber="50" targetBox="[228,1358,197,1099]" targetPageId="1">
<paragraph id="8B85FC10FFB5FF8BFF11FB0E5DEFFB7A" blockId="1.[151,1436,1123,1178]" pageId="1" pageNumber="50">
<emphasis id="B94E2002FFB5FF8BFF11FB0E5D7BFB9B" bold="true" box="[151,281,1123,1147]" pageId="1" pageNumber="50">FIGURE 1.</emphasis>
<taxonomicName id="4C3A8793FFB5FF8BFE99FB0E5E6DFB9A" box="[287,527,1123,1146]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="alphonsei" status="gen. nov.">
<emphasis id="B94E2002FFB5FF8BFE99FB0E5E6DFB9A" box="[287,527,1123,1146]" italics="true" pageId="1" pageNumber="50">Araripeleon alphonsei</emphasis>
</taxonomicName>
<emphasis id="B94E2002FFB5FF8BFD93FB0E5EB9FB9B" bold="true" box="[533,731,1123,1147]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="A27D9D79FFB5FF8BFD93FB0E5E18FB9B" box="[533,634,1123,1147]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, holotype SMNK 2358 PAL, photograph of general habitus (scale bar represents 10 mm).
</paragraph>
</caption>
<subSubSection id="C320AF9BFFB5FF8BFF42FBA55F24FAB3" pageId="1" pageNumber="50" type="diagnosis">
<paragraph id="8B85FC10FFB5FF8BFF42FBA55F24FAB3" blockId="1.[151,1436,1224,2035]" pageId="1" pageNumber="50">
<emphasis id="B94E2002FFB5FF8BFF42FBA55D2FFB02" bold="true" box="[196,333,1224,1250]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
Costal area narrow; dense wing venation: numerous cross-veins throughout wings and numerous branches of Rs (21 to 23); first posterior branch of Rs close to stem of Rs; R1 and Rs relatively straight distally; absence of anterior Banksian fold in radial area; broad area of MP2 with posterior branch of MP2 relatively short and ending in CuA; anal area narrow.
</paragraph>
</subSubSection>
<subSubSection id="C320AF9BFFB5FF88FF42FA335D4BFCE7" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="description">
<paragraph id="8B85FC10FFB5FF8BFF42FA335DA1FA23" blockId="1.[151,1436,1224,2035]" pageId="1" pageNumber="50">
<emphasis id="B94E2002FFB5FF8BFF42FA335D37FA98" bold="true" box="[196,341,1374,1400]" pageId="1" pageNumber="50">Description</emphasis>
. A nearly complete specimen with the four wings and at least the fore legs still connected to the body. The hind and fore wings of both sides are partly overlapping. A small beetle is fossilised in ventral view above the abdomen.
</paragraph>
<paragraph id="8B85FC10FFB5FF88FF42FAA35D4BFCE7" blockId="1.[151,1436,1224,2035]" lastBlockId="2.[151,1437,152,850]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
Head transverse,
<quantity id="4CC251F5FFB5FF8BFE09FAA35D88FA08" box="[399,490,1486,1512]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="50" unit="mm" value="2.3">2.3 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFDB3FAA35EEDFA08" box="[565,655,1486,1512]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="50" unit="mm" value="3.2">3.2 mm</quantity>
wide; dorsal surface missing so it is not possible to know if it had ocelli or not; eyes globular, nearly as long as head,
<quantity id="4CC251F5FFB5FF8BFD6CFA9E5F4FF9ED" box="[746,813,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="50" unit="mm" value="2.0">2 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFCF0FA9E5FADF9ED" box="[886,975,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="50" unit="mm" value="1.1">1.1 mm</quantity>
wide,
<quantity id="4CC251F5FFB5FF8BFB9BFA9E5814F9ED" box="[1053,1142,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="50" unit="mm" value="1.9">1.9 mm</quantity>
apart in anterior part and
<quantity id="4CC251F5FFB5FF8BFF11F9745C93F9D3" box="[151,241,1561,1587]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="50" unit="mm" value="3.5">3.5 mm</quantity>
apart posteriorly; antenna
<quantity id="4CC251F5FFB5FF8BFDACF9745EF0F9D3" box="[554,658,1561,1587]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="1" pageNumber="50" unit="mm" value="10.3">10.3 mm</quantity>
long, with 100 flagellomeres, nearly all identical, about as long as wide, apically thin, not clubbed, scape smaller than pedicel, pedicel very large and broad; mouthparts not visible. Thorax
<quantity id="4CC251F5FFB5FF8BFEDAF90E5DA9F99D" box="[348,459,1635,1661]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="50" unit="mm" value="13.1">13.1 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFD9DF90E5E18F99D" box="[539,634,1635,1661]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="50" unit="mm" value="7.3">7.3 mm</quantity>
wide; pronotum trapezoidal elongate,
<quantity id="4CC251F5FFB5FF8BFBDBF90E58D9F99D" box="[1117,1211,1635,1661]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="50" unit="mm" value="3.7">3.7 mm</quantity>
long, narrower in anterior part (
<quantity id="4CC251F5FFB5FF8BFEB3F9E45DF0F943" box="[309,402,1673,1699]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="50" unit="mm" value="2.5">2.5 mm</quantity>
) than in posterior part (
<quantity id="4CC251F5FFB5FF8BFD27F9E45F62F943" box="[673,768,1673,1699]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="50" unit="mm" value="4.5">4.5 mm</quantity>
); mesothorax
<quantity id="4CC251F5FFB5FF8BFC22F9E45F9FF943" box="[932,1021,1673,1699]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="50" unit="mm" value="3.7">3.7 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFBC3F9E458FDF943" box="[1093,1183,1673,1699]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="50" unit="mm" value="7.3">7.3 mm</quantity>
wide; metathorax
<quantity id="4CC251F5FFB5FF8BFAF1F9E45CA7F928" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="50" unit="mm" value="5.7">5.7 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFE94F9C35D0FF928" box="[274,365,1710,1736]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="50" unit="mm" value="4.6">4.6 mm</quantity>
wide. Abdomen rather poorly preserved, with 78 visible segments,
<quantity id="4CC251F5FFB5FF8BFB2CF9C35976F928" box="[1194,1300,1710,1736]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="50" unit="mm" value="25.4">25.4 mm</quantity>
long; max. width
<quantity id="4CC251F5FFB5FF8BFF67F9BE5D5DF90D" box="[225,319,1747,1773]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="50" unit="mm" value="3.4">3.4 mm</quantity>
; left ectoproct partly preserved, curved elongate,
<quantity id="4CC251F5FFB5FF8BFC01F9BE5F83F90D" box="[903,993,1747,1773]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="50" unit="mm" value="5.1">5.1 mm</quantity>
long; a long oblong median structure is present at abdomen apex,
<quantity id="4CC251F5FFB5FF8BFE61F9945E20F8F3" box="[487,578,1785,1811]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="50" unit="mm" value="2.8">2.8 mm</quantity>
long. Fore femora broad and elongate, anteriorly directed,
<quantity id="4CC251F5FFB5FF8BFB78F994593AF8F3" box="[1278,1368,1785,1811]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="50" unit="mm" value="5.4">5.4 mm</quantity>
long,
<quantity id="4CC251F5FFB5FF8BFF11F8735C94F8D8" box="[151,246,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="50" unit="mm" value="1.7">1.7 mm</quantity>
wide, longer than pronotum; fore tibiae partly visible, preserved under the femora, posteriorly directed,
<quantity id="4CC251F5FFB5FF8BFE80F82E5D12F8BD" box="[262,368,1859,1885]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="50" unit="mm" value="0.35">0.35 mm</quantity>
wide; no spines or spurs visible on fore leg. Fore and hind wings nearly identical in size, shape and venation. It was possible to draw correctly the hind wing venation but that of forewing is more fragmentary.
<collectingCountry id="F32DBC80FFB5FF8BFEB3F8E35D12F848" box="[309,368,1934,1960]" name="India" pageId="1" pageNumber="50">Hind</collectingCountry>
wing
<quantity id="4CC251F5FFB5FF8BFE3DF8E35E47F848" box="[443,549,1934,1960]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="1" pageNumber="50" unit="mm" value="42.5">42.5 mm</quantity>
long, 10.0 mm wide; Sc distally fused with R1,
<quantity id="4CC251F5FFB5FF8BFBEFF8E358B1F848" box="[1129,1235,1934,1960]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="50" unit="mm" value="35.5">35.5 mm</quantity>
from wing base; area between Sc and anterior wing margin as broad as area between Sc and R1; 40 to 45 simple and short cross-veins between C and Sc between wing base and point of fusion between R1 and Sc; area between anterior wing margin and Sc+R1 greatly widened, with about 13 long secondary veins, some of them being dichotomously forked and divided into small veinlets near wing margin (end-twigging); area between Sc and R1 with several cross-veins (at least five are visible); base of Rs
<quantity id="4CC251F5FFB6FF88FC0EFF8F5F80FF1C" box="[904,994,226,252]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="51" unit="mm" value="2.5">2.5 mm</quantity>
from wing base; MA separating from Rs 1.0 mm distally,
<quantity id="4CC251F5FFB6FF88FE07FE655DB9FEC2" box="[385,475,264,290]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="51" unit="mm" value="3.5">3.5 mm</quantity>
from wing base; the area covered by the branches of Rs is the largest of the wing, covering wing surface from apex to mid part of posterior wing margin; Rs divided into numerous (at least 21 to 23) posterior branches, all slightly zigzagged but not curved, with nine to 12 rows of cross-veins between them; all branches of Rs apically forked close to posterior wing margin; MA curved, slightly zigzagged and with two distal subdivisions, covering a small area; 19 cross-veins between MA and MP1; MP divided into MP1 and MP2 at its base; MP1 simple, MP1 and MP2a parallel, curved, with 21 cross-veins between them; a small fork at their respective apices; fork of MP2 into MP2a and MP2b
<quantity id="4CC251F5FFB6FF88FC5FFE855823FDE2" box="[985,1089,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="2" pageNumber="51" unit="mm" value="12.9">12.9 mm</quantity>
from wing base; MP2b short,
<quantity id="4CC251F5FFB6FF88FF11FD605C93FDC7" box="[151,241,525,551]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="51" unit="mm" value="2.1">2.1 mm</quantity>
long, reaching CuA; basal part of area between MP2 and CuA crossed by a row of eight cells; MP2a with three posterior branches reaching CuA and nine posterior branches reaching posterior wing margin; CuA and CuP separating
<quantity id="4CC251F5FFB6FF88FE0CFD355D87FD92" box="[394,485,600,626]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="51" unit="mm" value="3.4">3.4 mm</quantity>
from wing base; CuA and CuP+1A parallel with several narrow cells between them; distal end of CuA vanishing in area of MP2a; apex of CuP+1A
<quantity id="4CC251F5FFB6FF88FC4DFD105856FD77" box="[971,1076,637,663]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="51" unit="mm" value="12.6">12.6 mm</quantity>
from wing base; area between CuP+1A and posterior wing margin with 23 rows of cells; area between 2A, 3A, and posterior wing margin with two rows of three cells. In forewing, oblique vein (posterior branch of M) between M and CuA well pronounced.
</paragraph>
</subSubSection>
<subSubSection id="C320AF9BFFB6FF88FF42FC7F5E05FCB2" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8B85FC10FFB6FF88FF42FC7F5E05FCB2" blockId="2.[151,1437,152,850]" pageId="2" pageNumber="51">
<emphasis id="B94E2002FFB6FF88FF42FC7F5D30FCCC" bold="true" box="[196,338,786,812]" pageId="2" pageNumber="51">Etymology.</emphasis>
Named after Araripe, and the suffix -
<emphasis id="B94E2002FFB6FF88FC9FFC7F5F2EFCCB" box="[793,844,786,811]" italics="true" pageId="2" pageNumber="51">leon</emphasis>
of
<taxonomicName id="4C3A8793FFB6FF88FCF3FC7F5F9EFCCB" box="[885,1020,786,811]" class="Insecta" family="Palaeoleontidae" genus="Palaeoleon" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B94E2002FFB6FF88FCF3FC7F5F9EFCCB" box="[885,1020,786,811]" italics="true" pageId="2" pageNumber="51">Palaeoleon</emphasis>
</taxonomicName>
.The species epithet is named after Alphonse Millet, son of the first author.
</paragraph>
</subSubSection>
<caption id="DF45AC98FFB6FF88FF11F8105E8BF854" httpUri="https://zenodo.org/record/193521/files/figure.png" pageId="2" pageNumber="51" targetBox="[244,1360,954,1861]" targetPageId="2">
<paragraph id="8B85FC10FFB6FF88FF11F8105E8BF854" blockId="2.[151,1436,1917,1972]" pageId="2" pageNumber="51">
<emphasis id="B94E2002FFB6FF88FF11F8105D7BF875" bold="true" box="[151,281,1917,1941]" pageId="2" pageNumber="51">FIGURE 2.</emphasis>
<taxonomicName id="4C3A8793FFB6FF88FEA6F8135E6DF875" box="[288,527,1918,1941]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="alphonsei" status="gen. nov.">
<emphasis id="B94E2002FFB6FF88FEA6F8135E6DF875" box="[288,527,1918,1941]" italics="true" pageId="2" pageNumber="51">Araripeleon alphonsei</emphasis>
</taxonomicName>
<emphasis id="B94E2002FFB6FF88FD90F8105EBEF875" bold="true" box="[534,732,1917,1941]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A27D9D79FFB6FF88FD90F8105E19F875" box="[534,635,1917,1941]" pageId="2" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, holotype SMNK 2358 PAL, drawing of general habitus, with the forewing details removed (scale bar represents 10 mm).
</paragraph>
</caption>
<caption id="DF45AC98FFB7FF89FF11FC6F5D07FCD9" httpUri="https://zenodo.org/record/193522/files/figure.png" pageId="3" pageNumber="52" targetBox="[255,1316,239,665]" targetPageId="3">
<paragraph id="8B85FC10FFB7FF89FF11FC6F5D07FCD9" blockId="3.[151,1436,770,825]" pageId="3" pageNumber="52">
<emphasis id="B94E2002FFB7FF89FF11FC6F5D78FCFA" bold="true" box="[151,282,770,794]" pageId="3" pageNumber="52">FIGURE 3.</emphasis>
<taxonomicName id="4C3A8793FFB7FF89FEA7FC6F5E70FCF9" box="[289,530,770,793]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="alphonsei" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FEA7FC6F5E70FCF9" box="[289,530,770,793]" italics="true" pageId="3" pageNumber="52">Araripeleon alphonsei</emphasis>
</taxonomicName>
<emphasis id="B94E2002FFB7FF89FD9FFC6F5E81FCFA" bold="true" box="[537,739,770,794]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A27D9D79FFB7FF89FD9FFC6F5EE2FCFA" box="[537,640,770,794]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, holotype SMNK 2358 PAL, drawing of left forewing (scale bar represents 10 mm).
</paragraph>
</caption>
<subSubSection id="C320AF9BFFB7FF89FF42FC0A5974FC12" pageId="3" pageNumber="52" type="discussion">
<paragraph id="8B85FC10FFB7FF89FF42FC0A5974FC12" blockId="3.[151,1437,871,2018]" pageId="3" pageNumber="52">
<emphasis id="B94E2002FFB7FF89FF42FC0A5D35FC61" bold="true" box="[196,343,871,897]" pageId="3" pageNumber="52">Discussion.</emphasis>
The wing venation of
<taxonomicName id="4C3A8793FFB7FF89FDF2FC055F6BFC61" box="[628,777,872,897]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FDF2FC055F6BFC61" box="[628,777,872,897]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D9D79FFB7FF89FC95FC0A5FEAFC61" box="[787,904,871,897]" pageId="3" pageNumber="52" rank="genus">
<emphasis id="B94E2002FFB7FF89FC95FC0A5FEAFC61" bold="true" box="[787,904,871,897]" pageId="3" pageNumber="52">gen. nov.</emphasis>
</taxonomicNameLabel>
resembles those of the recent
<taxonomicName id="4C3A8793FFB7FF89FA83FC0559FAFC62" box="[1285,1432,872,898]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FF11FCE05D5BFC47" box="[151,313,909,935]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
, and
<taxonomicName id="4C3A8793FFB7FF89FEF1FCE05E5FFC47" box="[375,573,909,935]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
. In the most recent phylogenetic analyses of the
<taxonomicName id="4C3A8793FFB7FF89FBF8FCE05960FC47" box="[1150,1282,909,935]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
proposed by
<bibRefCitation id="EFAB81E1FFB7FF89FF11FCDF5DE4FC2C" author="Aspock" box="[151,390,946,972]" pageId="3" pageNumber="53" refString="Aspock, U., Plant, J. D. &amp; Nemeschkal, H. L. (2001) Cladistic analysis of Neuroptera and their position within Neuropterida (Insecta: Holometabola: Neuropterida: Neuroptera). Systematic Entomology, 26 (1), 73 - 86." type="journal article" year="2001">
Aspöck
<emphasis id="B94E2002FFB7FF89FF7FFCDF5D4CFC2B" box="[249,302,946,971]" italics="true" pageId="3" pageNumber="52">et al</emphasis>
. (2001
</bibRefCitation>
,
<bibRefCitation id="EFAB81E1FFB7FF89FE15FCDF5DB0FC2C" author="Aspock" box="[403,466,946,972]" pageId="3" pageNumber="53" refString="Aspock, U., Aspock, H. &amp; Haring, E. (2003) Phylogeny of the Neuropterida - morphological evidence and the molecular advocatus diaboli. Entomologische Abhandlungen, 61 (2), 157 - 158." type="journal article" year="2003">2003</bibRefCitation>
),
<bibRefCitation id="EFAB81E1FFB7FF89FE6EFCDF5EFEFC2C" author="Aspock" box="[488,668,946,972]" pageId="3" pageNumber="53" refString="Aspock, U. (2002) Phylogeny of the Neuropterida. Zoologica Scripta, 31 (1), 51 - 56." type="journal article" year="2002">Aspöck (2002)</bibRefCitation>
, and
<bibRefCitation id="EFAB81E1FFB7FF89FD5DFCDF5870FC2C" author="Aspock" box="[731,1042,946,972]" pageId="3" pageNumber="53" refString="Aspock, U. &amp; Aspock, H. (2008) Phylogenetic relevance of the genital sclerites of Neuropterida (Insecta: Holometabola). Systematic Entomology, 33 (1), 97 - 127." type="journal article" year="2008">Aspöck &amp; Aspöck (2008)</bibRefCitation>
, these three families are grouped together in a subclade (
<taxonomicName id="4C3A8793FFB7FF89FE20FCB55E53FC12" box="[422,561,984,1010]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
+ (
<taxonomicName id="4C3A8793FFB7FF89FDD1FCB55F79FC12" box="[599,795,984,1010]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
+
<taxonomicName id="4C3A8793FFB7FF89FCBCFCB55FBBFC12" box="[826,985,984,1010]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
)) of the Myrmeleontoidea.
</paragraph>
</subSubSection>
<subSubSection id="C320AF9BFFB7FF89FF42FC905E80F972" pageId="3" pageNumber="52" type="reference_group">
<paragraph id="8B85FC10FFB7FF89FF42FC9058CEFB82" blockId="3.[151,1437,871,2018]" pageId="3" pageNumber="52">
<taxonomicName id="4C3A8793FFB7FF89FF42FC905D30FBF6" ID-CoL="ZRJ" box="[196,338,1021,1046]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FF42FC905D30FBF6" box="[196,338,1021,1046]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
differs from all the Myrmeleontoidea already described from the Crato Formation in its dense wing venation with very numerous cross-veins and branches of Rs (21 to 23), broad area of MP2 with posterior branch of MP2 ending in CuA (
<bibRefCitation id="EFAB81E1FFB7FF89FDFFFB255F07FB82" author="Martins-Neto" box="[633,869,1096,1122]" pageId="3" pageNumber="53" refString="Martins-Neto, R. G. (2000) Remarks on the Neuropterofauna (Insecta, Neuroptera) from the Brazilian Cretaceous with keys for the identification of the known taxons. Acta Geologica Hispanica, 35 (1 - 2), 97 - 118." type="journal article" year="2000">Martins-Neto, 2000</bibRefCitation>
;
<bibRefCitation id="EFAB81E1FFB7FF89FCF7FB2558FDFB82" author="Martins-Neto" box="[881,1183,1096,1122]" pageId="3" pageNumber="53" refString="Martins-Neto, R. G., Heads, S. W. &amp; Bechly, G. (2007) 11.16 Neuropterida: snakeflies, dobsonflies and lacewings. pp. 328 - 340. In: Martill, D., Bechly, G. and Loveridge, R. (eds). The Crato fossil beds of Brazil: Window into an ancient world. Cambridge University Press, Cambridge, 624 pp." type="book chapter" year="2007">
Martins-Neto
<emphasis id="B94E2002FFB7FF89FB91FB255828FB81" box="[1047,1098,1096,1121]" italics="true" pageId="3" pageNumber="52">et al</emphasis>
., 2007
</bibRefCitation>
).
</paragraph>
<paragraph id="8B85FC10FFB7FF89FF42FB005E80F972" blockId="3.[151,1437,871,2018]" pageId="3" pageNumber="52">
Following the key to recent neuropteran families of
<bibRefCitation id="EFAB81E1FFB7FF89FCA2FB005FD1FB67" author="New" box="[804,947,1133,1159]" pageId="3" pageNumber="53" refString="New, T. R. (1990) Planipennia (lacewings). Handbuch der Zoologie, Berlin, 4 (30), (Arthropoda: Insecta), 1 - 132." type="journal article" year="1990">New (1990)</bibRefCitation>
,
<taxonomicName id="4C3A8793FFB7FF89FC39FB00582EFB66" ID-CoL="ZRJ" box="[959,1100,1133,1158]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FC39FB00582EFB66" box="[959,1100,1133,1158]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
would fall near
<taxonomicName id="4C3A8793FFB7FF89FA8AFB0059FAFB67" ID-CoL="DGD" box="[1292,1432,1133,1159]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
, the two other families being excluded for the long filiform antennae without club or apex broadened. Nevertheless
<taxonomicName id="4C3A8793FFB7FF89FEB0FBD55DA6FB31" ID-CoL="ZRJ" box="[310,452,1208,1233]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FEB0FBD55DA6FB31" box="[310,452,1208,1233]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
falls in the Mesozoic family
<taxonomicName id="4C3A8793FFB7FF89FCA6FBD558A8FB32" ID-CoL="626KP" authority="Martins-Neto, 1992" authorityName="Martins-Neto" authorityYear="1992" box="[800,1226,1208,1234]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Palaeoleontidae Martins-Neto, 1992</taxonomicName>
because it has all the diagnostic characters of this family as redefined recently by
<bibRefCitation id="EFAB81E1FFB7FF89FC3CFBB05972FB17" author="Menon" box="[954,1296,1245,1271]" pageId="3" pageNumber="53" refString="Menon, F. &amp; Makarkin, V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato formation of Brazil. Palaeontology, 51 (1), 149 - 162." type="journal article" year="2008">Menon &amp; Makarkin (2008)</bibRefCitation>
, viz. large myrmeleontoids (forewing
<quantity id="4CC251F5FFB7FF89FE5FFA6F5E3BFAFC" box="[473,601,1282,1308]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="7.0" metricValueMin="3.6" pageId="3" pageNumber="52" unit="mm" value="53.0" valueMax="70.0" valueMin="36.0">3670 mm</quantity>
long) with relatively long antennae, unusually dense cross-venation in the majority of genera, easily distinguished from other myrmeleontoid families by the following combination of character states: (1) antennae long, filiform, neither dilated nor clavate distally [dilated distally in
<taxonomicName id="4C3A8793FFB7FF89FF11FA1F5D3EFA6C" ID-CoL="D3X" box="[151,348,1394,1420]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
, Araripeneuridae; clavate in
<taxonomicName id="4C3A8793FFB7FF89FD34FA1F5F35FA6C" ID-CoL="6RX" box="[690,855,1394,1420]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
]; (2) origin of Rs situated near wing base in both wings [removed from base in
<taxonomicName id="4C3A8793FFB7FF89FDB6FAF55F69FA52" ID-CoL="D3X" box="[560,779,1432,1458]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
, Araripeneuridae,
<taxonomicName id="4C3A8793FFB7FF89FB89FAF558A0FA52" ID-CoL="6RX" box="[1039,1218,1432,1458]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FB51FAF559F0FA52" ID-CoL="D8B" box="[1239,1426,1432,1458]" class="Insecta" family="Nemopteridae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nemopteridae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FF11FAD05D27FA37" ID-CoL="6YY" box="[151,325,1469,1495]" class="Insecta" family="Babinskaiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Babinskaiidae</taxonomicName>
]; (3) presectorial cross-veins absent in both wings [present in
<taxonomicName id="4C3A8793FFB7FF89FBA2FAD05888FA37" ID-CoL="D3X" box="[1060,1258,1469,1495]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Myrmeleontidae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FB70FAD059FAFA37" ID-CoL="6RX" box="[1270,1432,1469,1495]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FF11FA8F5D20FA1C" ID-CoL="D8B" box="[151,322,1506,1532]" class="Insecta" family="Nemopteridae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nemopteridae</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FEC8FA8F5D9FFA1C" ID-CoL="6YY" box="[334,509,1506,1532]" class="Insecta" family="Babinskaiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Babinskaiidae</taxonomicName>
]; (4) oblique vein present in forewing [absent in
<taxonomicName id="4C3A8793FFB7FF89FBD0FA8F588EFA1C" ID-CoL="DGD" box="[1110,1260,1506,1532]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
]. Nevertheless all these characters can also be found in
<taxonomicName id="4C3A8793FFB7FF89FD25F9655F58F9C2" ID-CoL="DGD" box="[675,826,1544,1570]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
, except maybe the oblique vein absent or less pronounced in
<taxonomicName id="4C3A8793FFB7FF89FECEF9405DB3F9A7" ID-CoL="DGD" box="[328,465,1581,1607]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
than in
<taxonomicName id="4C3A8793FFB7FF89FDB4F9405E92F9A7" ID-CoL="626KP" box="[562,752,1581,1607]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
. It would be necessary to make a phylogenetic analysis to confirm the value of the separation between the two families. We provisionally include
<taxonomicName id="4C3A8793FFB7FF89FB38F93F592DF98B" ID-CoL="ZRJ" box="[1214,1359,1618,1643]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FB38F93F592DF98B" box="[1214,1359,1618,1643]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C3A8793FFB7FF89FF11F9155D33F972" ID-CoL="626KP" box="[151,337,1656,1682]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
for its pronounced oblique vein.
</paragraph>
</subSubSection>
<subSubSection id="C320AF9BFFB7FF8EFF42F9F059F6FC22" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="description">
<paragraph id="8B85FC10FFB7FF8EFF42F9F05830FE8C" blockId="3.[151,1437,871,2018]" lastBlockId="4.[151,1437,152,962]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
<bibRefCitation id="EFAB81E1FFB7FF89FF42F9F05E64F957" author="Menon" box="[196,518,1693,1719]" pageId="3" pageNumber="53" refString="Menon, F. &amp; Makarkin, V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato formation of Brazil. Palaeontology, 51 (1), 149 - 162." type="journal article" year="2008">Menon &amp; Makarkin (2008)</bibRefCitation>
gave a list of the genera currently included in this family.
<taxonomicName id="4C3A8793FFB7FF89FB3CF9F0592AF956" box="[1210,1352,1693,1718]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FB3CF9F0592AF956" box="[1210,1352,1693,1718]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
differs from
<emphasis id="B94E2002FFB7FF89FF51F9AF5D08F93B" box="[215,362,1730,1755]" italics="true" pageId="3" pageNumber="52">Neurastenyx</emphasis>
Martins-Neto &amp; Vulcano, 1997,
<taxonomicName id="4C3A8793FFB7FF89FD69F9AF581AF93C" authority="Ponomarenko, 1992" authorityName="Ponomarenko" authorityYear="1992" box="[751,1144,1730,1756]" class="Insecta" family="Palaeoleontidae" genus="Baisopardus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B94E2002FFB7FF89FD69F9AF5FE0F93B" box="[751,898,1730,1755]" italics="true" pageId="3" pageNumber="52">Baisopardus</emphasis>
Ponomarenko, 1992
</taxonomicName>
,
<taxonomicName id="4C3A8793FFB7FF89FB02F9AF5DE3F8E2" authority="Menon &amp; Makarkin, 2008" authorityName="Menon &amp; Makarkin" authorityYear="2008" class="Insecta" family="Palaeoleontidae" genus="Parapalaeoleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B94E2002FFB7FF89FB02F9AF5923F93B" box="[1156,1345,1730,1755]" italics="true" pageId="3" pageNumber="52">Parapalaeoleon</emphasis>
<bibRefCitation id="EFAB81E1FFB7FF89FACFF9AF5DE3F8E2" author="Menon" pageId="3" pageNumber="53" refString="Menon, F. &amp; Makarkin, V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato formation of Brazil. Palaeontology, 51 (1), 149 - 162." type="journal article" year="2008">Menon &amp; Makarkin, 2008</bibRefCitation>
</taxonomicName>
, and from
<emphasis id="B94E2002FFB7FF89FD8FF9855EBFF8E1" box="[521,733,1768,1793]" italics="true" pageId="3" pageNumber="52">Paraneurastenyx</emphasis>
Martin-Neto,
<quantity id="4CC251F5FFB7FF89FC16F9855F8CF8E2" box="[912,1006,1768,1794]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="3" pageNumber="52" unit="in" value="1998.0">1998 in</quantity>
the numerous and strictly parallel branches of Rs, absence of anterior Banksian fold in radial area, and the median vein more posteriorly curved (
<bibRefCitation id="EFAB81E1FFB7FF89FF19F85F5DCBF8AC" author="Ponomarenko" box="[159,425,1842,1868]" lastPageId="3" lastPageNumber="54" pageId="3" pageNumber="53" refString="Ponomarenko, A. G. (1992 a) Neuroptera (Insecta) from the Cretaceous of Transbaikalia. Paleontological Journal, 26 (3), 43 - 50." type="journal article" year="1992" yearSuffix="a">Ponomarenko, 1992a</bibRefCitation>
;
<bibRefCitation id="EFAB81E1FFB7FF89FE30F85F5ECFF8AC" author="Martins-Neto" box="[438,685,1842,1868]" pageId="3" pageNumber="53" refString="Martins-Neto, R. G. (2000) Remarks on the Neuropterofauna (Insecta, Neuroptera) from the Brazilian Cretaceous with keys for the identification of the known taxons. Acta Geologica Hispanica, 35 (1 - 2), 97 - 118." type="journal article" year="2000">Martins-Neto, 2000</bibRefCitation>
;
<bibRefCitation id="EFAB81E1FFB7FF89FD3CF85F5FFCF8AC" author="Heads" box="[698,926,1842,1868]" pageId="3" pageNumber="53" refString="Heads, S. W., Martill, D. M. &amp; Loveridge, R. F. (2005) An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology, 48 (6), 1409 - 1417." type="journal article" year="2005">
Heads
<emphasis id="B94E2002FFB7FF89FC88F85F5F27F8AB" box="[782,837,1842,1867]" italics="true" pageId="3" pageNumber="52">et al</emphasis>
., 2005
</bibRefCitation>
;
<bibRefCitation id="EFAB81E1FFB7FF89FC2DF85F5891F8AC" author="Menon" box="[939,1267,1842,1868]" pageId="3" pageNumber="53" refString="Menon, F. &amp; Makarkin, V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato formation of Brazil. Palaeontology, 51 (1), 149 - 162." type="journal article" year="2008">Menon &amp; Makarkin, 2008</bibRefCitation>
).
<taxonomicName id="4C3A8793FFB7FF89FA8FF85F59FFF8AB" box="[1289,1437,1842,1867]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FA8FF85F59FFF8AB" box="[1289,1437,1842,1867]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
differs from
<emphasis id="B94E2002FFB7FF89FEB3F8355DB0F891" box="[309,466,1880,1905]" italics="true" pageId="3" pageNumber="52">Samsonileon</emphasis>
Ponomarenko in
<bibRefCitation id="EFAB81E1FFB7FF89FD29F8355FB7F892" author="Dobruskina" box="[687,981,1880,1906]" pageId="3" pageNumber="53" refString="Dobruskina, I. A., Ponomarenko, A. G. &amp; Rasnitsyn, A. P. (1997) Fossil Insects from Israel. Paleontological Journal, 31 (5), 528 - 533." type="journal article" year="1997">
Dobruskina
<emphasis id="B94E2002FFB7FF89FCC1F8355F1DF891" box="[839,895,1880,1905]" italics="true" pageId="3" pageNumber="52">et al</emphasis>
.,
<quantity id="4CC251F5FFB7FF89FC1EF8355FB7F892" box="[920,981,1880,1906]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="3" pageNumber="52" unit="in" value="1997.0">1997</quantity>
</bibRefCitation>
in the absence of a Banksian fold, a narrower anal area, and absence of a very strong fork of median vein (
<bibRefCitation id="EFAB81E1FFB7FF89FC5CF810589AF877" author="Dobruskina" box="[986,1272,1917,1943]" pageId="3" pageNumber="53" refString="Dobruskina, I. A., Ponomarenko, A. G. &amp; Rasnitsyn, A. P. (1997) Fossil Insects from Israel. Paleontological Journal, 31 (5), 528 - 533." type="journal article" year="1997">
Dobruskina
<emphasis id="B94E2002FFB7FF89FBE8F81058C0F876" box="[1134,1186,1917,1942]" italics="true" pageId="3" pageNumber="52">et al</emphasis>
., 1997
</bibRefCitation>
).
<taxonomicName id="4C3A8793FFB7FF89FA8BF81059FEF876" box="[1293,1436,1917,1942]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB7FF89FA8BF81059FEF876" box="[1293,1436,1917,1942]" italics="true" pageId="3" pageNumber="52">Araripeleon</emphasis>
</taxonomicName>
shares with
<emphasis id="B94E2002FFB7FF89FEA1F8CF5D8CF85B" box="[295,494,1954,1979]" italics="true" pageId="3" pageNumber="52">Metahemerobius</emphasis>
<bibRefCitation id="EFAB81E1FFB7FF89FE70F8CF5ED4F85C" author="Makarkin" box="[502,694,1954,1980]" pageId="3" pageNumber="53" refString="Makarkin, V. N. (1990) [New Neuroptera from the Upper Cretaceous of Asia.] pp. 63 - 68. In: Akimov, I. A. et al. (eds). Novosti faunistiki i sistematiki [News in faunistics and systematics.] Naukova Dumka (publ.), Kiev. [in Russian.]" type="book chapter" year="1990">Makarkin, 1990</bibRefCitation>
the absence of the Banksian fold and branches of Rs more or less parallel, but differs in the number of cross-veins between R1 and Rs, R1 and Rs less curved in their distal parts for
<taxonomicName id="4C3A8793FFB0FF8EFE85FFF55DF0FF51" box="[259,402,152,177]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB0FF8EFE85FFF55DF0FF51" box="[259,402,152,177]" italics="true" pageId="4" pageNumber="53">Araripeleon</emphasis>
</taxonomicName>
, and presence of anterior branches of median vein in its distal part in
<emphasis id="B94E2002FFB0FF8EFB50FFF559F9FF51" box="[1238,1435,152,177]" italics="true" pageId="4" pageNumber="53">Metahemerobius</emphasis>
(
<bibRefCitation id="EFAB81E1FFB0FF8EFF19FFD05D00FF37" author="Makarkin" box="[159,354,189,215]" pageId="4" pageNumber="53" refString="Makarkin, V. N. (1990) [New Neuroptera from the Upper Cretaceous of Asia.] pp. 63 - 68. In: Akimov, I. A. et al. (eds). Novosti faunistiki i sistematiki [News in faunistics and systematics.] Naukova Dumka (publ.), Kiev. [in Russian.]" type="book chapter" year="1990">Makarkin, 1990</bibRefCitation>
). Several very important structures of the
<taxonomicName id="4C3A8793FFB0FF8EFCD2FFD0586CFF37" box="[852,1038,189,215]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
are not preserved in the
<typeStatus id="548142B2FFB0FF8EFAB3FFD059FEFF37" box="[1333,1436,189,215]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3A8793FFB0FF8EFF31FF8F5DA7FF1C" authority="Rice, 1969" authorityName="Rice" authorityYear="1969" box="[183,453,226,252]" class="Insecta" family="Palaeoleontidae" genus="Palaeoleon" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B94E2002FFB0FF8EFF31FF8F5D5EFF1B" box="[183,316,226,251]" italics="true" pageId="4" pageNumber="53">Palaeoleon</emphasis>
<bibRefCitation id="EFAB81E1FFB0FF8EFEC2FF8F5DA7FF1C" author="Rice" box="[324,453,226,252]" pageId="4" pageNumber="54" refString="Rice, H. M. A. (1969) An antlion (Neuroptera) and a stonefly (Plecoptera) of Cretaceous age from Labrador, Newfoundland. Geological Survey of Canada, Paper, 68 - 65, 1 - 11." type="book chapter" year="1969">Rice, 1969</bibRefCitation>
</taxonomicName>
,
<typeStatus id="548142B2FFB0FF8EFE54FF8F5E66FF1C" box="[466,516,226,252]" pageId="4" pageNumber="53">type</typeStatus>
genus of the family, i.e. the base of Rs and the oblique vein. Nevertheless, the Banksian fold is clearly visible in the photograph of this fossil (
<bibRefCitation id="EFAB81E1FFB0FF8EFC00FE655804FEC2" author="Rice" box="[902,1126,264,290]" pageId="4" pageNumber="54" refString="Rice, H. M. A. (1969) An antlion (Neuroptera) and a stonefly (Plecoptera) of Cretaceous age from Labrador, Newfoundland. Geological Survey of Canada, Paper, 68 - 65, 1 - 11." type="book chapter" year="1969">Rice, 1969: plate 1</bibRefCitation>
). Note that the base of Rs and the oblique vein are not preserved in the
<typeStatus id="548142B2FFB0FF8EFD24FE405F6BFEA7" box="[674,777,301,327]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
of
<emphasis id="B94E2002FFB0FF8EFCA8FE405F90FEA6" box="[814,1010,301,326]" italics="true" pageId="4" pageNumber="53">Metahemerobius</emphasis>
, and that the oblique vein is also not clearly figured for
<taxonomicName id="4C3A8793FFB0FF8EFEF5FE3F5E64FE8B" box="[371,518,338,363]" class="Insecta" family="Palaeoleontidae" genus="Baisopardus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B94E2002FFB0FF8EFEF5FE3F5E64FE8B" box="[371,518,338,363]" italics="true" pageId="4" pageNumber="53">Baisopardus</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFAB81E1FFB0FF8EFDAAFE3F5F6EFE8C" box="[556,780,338,364]" pageId="4" pageNumber="53" refString="Heads, S. W., Martill, D. M. &amp; Loveridge, R. F. (2005) An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology, 48 (6), 1409 - 1417." type="journal article">
Heads
<emphasis id="B94E2002FFB0FF8EFDFAFE3F5ED2FE8B" box="[636,688,338,363]" italics="true" pageId="4" pageNumber="53">et al</emphasis>
. (2005)
</bibRefCitation>
and
<bibRefCitation id="EFAB81E1FFB0FF8EFCC3FE3F582CFE8C" author="Ponomarenko" box="[837,1102,338,364]" lastPageId="4" lastPageNumber="54" pageId="4" pageNumber="53" refString="Ponomarenko, A. G. (1992 a) Neuroptera (Insecta) from the Cretaceous of Transbaikalia. Paleontological Journal, 26 (3), 43 - 50." type="journal article" year="1992" yearSuffix="a">Ponomarenko (1992a)</bibRefCitation>
.
</paragraph>
<paragraph id="8B85FC10FFB0FF8EFF42FE15595EFCE7" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="53">
Lastly
<emphasis id="B94E2002FFB0FF8EFE99FE155DC2FE71" box="[287,416,376,401]" italics="true" pageId="4" pageNumber="53">Cretoleon</emphasis>
Ponomarenko, 1992 was considered possibly to belong to
<taxonomicName id="4C3A8793FFB0FF8EFB23FE155911FE72" box="[1189,1395,376,402]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
by Ponomarenko (in
<bibRefCitation id="EFAB81E1FFB0FF8EFEEDFEF05E1CFE57" author="Dobruskina" box="[363,638,413,439]" pageId="4" pageNumber="53" refString="Dobruskina, I. A., Ponomarenko, A. G. &amp; Rasnitsyn, A. P. (1997) Fossil Insects from Israel. Paleontological Journal, 31 (5), 528 - 533." type="journal article" year="1997">
Dobruskina
<emphasis id="B94E2002FFB0FF8EFE7AFEF05E52FE56" box="[508,560,413,438]" italics="true" pageId="4" pageNumber="53">et al</emphasis>
. 1997
</bibRefCitation>
), listed among the genera of this family by
<bibRefCitation id="EFAB81E1FFB0FF8EFB05FEF05908FE57" box="[1155,1386,413,439]" pageId="4" pageNumber="53" refString="Heads, S. W., Martill, D. M. &amp; Loveridge, R. F. (2005) An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology, 48 (6), 1409 - 1417." type="journal article">
Heads
<emphasis id="B94E2002FFB0FF8EFB53FEF0596BFE56" box="[1237,1289,413,438]" italics="true" pageId="4" pageNumber="53">et al</emphasis>
. (2005)
</bibRefCitation>
, but tentatively removed from this family by
<bibRefCitation id="EFAB81E1FFB0FF8EFD07FEAF5FABFE3C" author="Menon" box="[641,969,450,476]" pageId="4" pageNumber="53" refString="Menon, F. &amp; Makarkin, V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato formation of Brazil. Palaeontology, 51 (1), 149 - 162." type="journal article" year="2008">Menon &amp; Makarkin (2008)</bibRefCitation>
for the following forewing characters: costal space is broad; Rs originates relatively distant from the wing base; the anterior Banksian fold is absent; the subcostal veinlets in the pterostigmal region are very closely spaced and simple; cross-veins are rare in the distal portion of the wing. All these characters, except the position of the origin of Rs, are not listed in the diagnosis of this family of these last authors. The origin of Rs in
<emphasis id="B94E2002FFB0FF8EFC10FD355868FD91" box="[918,1034,600,625]" italics="true" pageId="4" pageNumber="53">Cretoleon</emphasis>
remains very basal if slightly less basal than in other
<taxonomicName id="4C3A8793FFB0FF8EFE05FD105E26FD77" box="[387,580,637,663]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
(unknown in the
<typeStatus id="548142B2FFB0FF8EFC98FD105F30FD77" box="[798,850,637,663]" pageId="4" pageNumber="53">type</typeStatus>
genus of the family). Also the Banksian fold is absent in the palaeoleontid genus
<emphasis id="B94E2002FFB0FF8EFDB2FDCF5E9FFD5B" box="[564,765,674,699]" italics="true" pageId="4" pageNumber="53">Metahemerobius</emphasis>
. Thus the removal of
<emphasis id="B94E2002FFB0FF8EFB8AFDCF58E6FD5B" box="[1036,1156,674,699]" italics="true" pageId="4" pageNumber="53">Cretoleon</emphasis>
from this family looks rather weakly supported.
<taxonomicName id="4C3A8793FFB0FF8EFE39FDA55E2FFD01" box="[447,589,712,737]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB0FF8EFE39FDA55E2FFD01" box="[447,589,712,737]" italics="true" pageId="4" pageNumber="53">Araripeleon</emphasis>
</taxonomicName>
differs from
<emphasis id="B94E2002FFB0FF8EFD63FDA55F3BFD01" box="[741,857,712,737]" italics="true" pageId="4" pageNumber="53">Cretoleon</emphasis>
in the first posterior branch of Rs close to base of Rs, more numerous cross-veins between R1 and Rs, and costal area narrower (
<bibRefCitation id="EFAB81E1FFB0FF8EFBA8FD80594DFCE7" author="Ponomarenko" box="[1070,1327,749,775]" pageId="4" pageNumber="54" refString="Ponomarenko, A. G. (1992 b) [New lacewings (Insecta, Neuroptera) from the Mesozoic of Mongolia.] pp. 101 - 111. In: Grunt, T. A. (ed.) Novye iskopaemye bespozronochnye Mongolii [= New species of fossil invertebrates of Mongolia]. Trudy Sovmestnaia Sovetsko-Mongol'skaia Paleontologicheskaia Ekspeditsiia. [Transactions of the Joint Russian-Mongolian Paleontological Expedition], Akademii nauk SSSR, 41." type="journal article" year="1992" yearSuffix="b">Ponomarenko, 1992b</bibRefCitation>
).
</paragraph>
<paragraph id="8B85FC10FFB0FF8EFF42FC7F59F6FC22" blockId="4.[151,1437,152,962]" pageId="4" pageNumber="53">
In conclusion
<taxonomicName id="4C3A8793FFB0FF8EFEEDFC7F5D9AFCCB" box="[363,504,786,811]" class="Insecta" family="Palaeoleontidae" genus="Araripeleon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94E2002FFB0FF8EFEEDFC7F5D9AFCCB" box="[363,504,786,811]" italics="true" pageId="4" pageNumber="53">Araripeleon</emphasis>
</taxonomicName>
has enough differences from all these genera to attribute it to a new genus. We tentatively attribute it to the
<taxonomicName id="4C3A8793FFB0FF8EFE62FC555EFFFCB2" box="[484,669,824,850]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
because it shares all the diagnostic characters of the family, even if it has not the Banksian fold present in nearly all the palaeoleontid genera. The confirmation of its position will need a phylogenetic analysis that would include all the Mesozoic myrmeleontoid taxa, and precise the limit of the
<taxonomicName id="4C3A8793FFB0FF8EFEA6FCC55DB8FC22" box="[288,474,936,962]" class="Insecta" family="Palaeoleontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="family">Palaeoleontidae</taxonomicName>
and the relationships between the families currently included in this superfamily.
</paragraph>
</subSubSection>
</treatment>
</document>