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<document id="2F4D51117A7A968B5B11A978FBE170FF" ID-DOI="10.5281/zenodo.5329843" ID-ISSN="0374-1036" ID-Zenodo-Dep="5329843" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630164576859" checkinUser="felipe" docAuthor="Fikáček, Martin, Leschen, Richard A. B., Newton, Alfred F. &amp; Gunter, Nicole" docDate="2012" docId="03BAB44DFFDFFFAD099BFB39FDB3FAA6" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.129-146.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Horelophus walkeri Orchymont 1913" docType="treatment" docVersion="3" lastPageNumber="143" masterDocId="FF83CC35FFDBFFA30830FFA7FFB6FFB2" masterDocTitle="Horelophus walkeri rediscovered: adult morphology and notes on biology (Coleoptera: Hydrophilidae)" masterLastPageNumber="146" masterPageNumber="129" pageNumber="133" updateTime="1699170717191" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="68FDFAB717BAEB8BBD4D3E66EC8FA99B">Horelophus walkeri rediscovered: adult morphology and notes on biology (Coleoptera: Hydrophilidae)</mods:title>
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<mods:namePart id="629249D06359DD4DCEB34D964FEF6086">Fikáček, Martin</mods:namePart>
<mods:affiliation id="6D32241BD4EEE00252A074E2A6409A4E">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Sciences, Charles University in Prague, Viničná 7, CZ- 128 43 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="CCF115ADEBFCD0F68F93B6B09EF65ECA">Leschen, Richard A. B.</mods:namePart>
<mods:affiliation id="54AD49F9E94D81995CE3B70E9530116D">New Zealand Arthropod Collection, Landcare Research, Private Bag 92170, Auckland, New Zealand; e-mail: leschenr @ landcareresearch. co. nz</mods:affiliation>
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<mods:namePart id="4F61AA8CAA46F58C7EFC5B52543D9988">Newton, Alfred F.</mods:namePart>
<mods:affiliation id="0E4AC00C37BA693C522548155EBAC2E9">Division of Insects, Field Museum of Natural History, Chicago, Illinois 60605, USA; e-mail: anewton @ fieldmuseum. org</mods:affiliation>
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<mods:namePart id="A279D9C45551383E7775E65B03ADF139">Gunter, Nicole</mods:namePart>
<mods:affiliation id="026353B73C5D82E12D73DDF0D3EF0689">Australian National Insect Collection, CSIRO Ecosystem Sciences, GPO Box 1700, Canberra, ACT 2601, Australia; e-mail: Nicole. Gunter @ csiro. au</mods:affiliation>
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<mods:title id="D60514EADDF946E60939B65B3E1A0FE0">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<taxonomicName id="4C137ED8FFDFFFA7099BFB39FCE4FB05" ID-CoL="3MP4N" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[427,850,1182,1207]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFDFFFA7099BFB39FCE4FB05" bold="true" box="[427,850,1182,1207]" pageId="4" pageNumber="133">
<emphasis id="B967D949FFDFFFA7099BFB39FD33FB05" bold="true" box="[427,645,1182,1207]" italics="true" pageId="4" pageNumber="133">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFDFFFA70ABCFB38FCE4FB05" author="ORCHYMONT A." box="[652,850,1183,1207]" pageId="4" pageNumber="133" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
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(
<figureCitation id="132819DEFFDFFFA70A07FB1AFD3EFB63" box="[567,648,1213,1233]" captionStart="Figs" captionStartId="2.[151,190,1538,1557]" captionTargetBox="[152,1125,232,1517]" captionTargetId="figure-10@2.[151,1125,215,1519]" captionTargetPageId="2" captionText="Figs. 1–4. Horelophus walkeri Orchymont, 1913.1–2 – general habitus in dorsal and lateral view (photo M.Fikáček); 3–4 – individuals on the wet rock surfaces at Dead Horse Creek, Marlborough (photo R. Leschen)." figureDoi="http://doi.org/10.5281/zenodo.5329845" httpUri="https://zenodo.org/record/5329845/files/figure.png" pageId="4" pageNumber="133">Figs. 1–4</figureCitation>
, 
<figureCitation id="132819DEFFDFFFA70AA2FB1AFD13FB62" box="[658,677,1213,1232]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="4" pageNumber="133">11</figureCitation>
–51)
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<treatmentCitationGroup id="AB032275FFDFFFA708A7FB46FDB5FB41" box="[151,515,1248,1267]" pageId="4" pageNumber="133">
<treatmentCitation id="0AB2234AFFDFFFA708A7FB46FDB6FB41" author="ORCHYMONT A." box="[151,512,1248,1267]" page="97" pageId="4" pageNumber="133" year="1913">
<taxonomicName id="4C137ED8FFDFFFA708A7FB46FDB6FB41" ID-CoL="3MP4N" authority="Orchymont, 1913: 97" authorityName="Orchymont" authorityPageNumber="97" authorityYear="1913" box="[151,512,1248,1267]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFDFFFA708A7FB46FEF6FB41" box="[151,320,1249,1267]" italics="true" pageId="4" pageNumber="133">Horelophus walkeri</emphasis>
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.
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<emphasis id="B967D949FFDFFFA708A7FAADFEA6FAAF" bold="true" box="[151,272,1289,1309]" pageId="4" pageNumber="133">
<typeStatus id="54A8BBF9FFDFFFA708A7FAADFF72FAAF" box="[151,196,1290,1309]" pageId="4" pageNumber="133">Type</typeStatus>
locality.
</emphasis>
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, South Island, Buller district, Reefton.
</paragraph>
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<emphasis id="B967D949FFDFFFA708A7FA94FECFFAF4" bold="true" box="[151,377,1330,1350]" pageId="4" pageNumber="133">Type material examined.</emphasis>
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: 
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(
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): ‘Coll. R. I. Sc. N. B. / Nouvelle 
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// [label glued on the previous:] Reefton / N.Z. 12 1902 / J. J. Walker // det. d’Orchymont / 
<taxonomicName id="4C137ED8FFDFFFA70ADBFAE9FC20FAD3" authorityName="Orchymont" authorityYear="1913" box="[747,918,1358,1377]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="walkeri">Horelophus Walkeri</taxonomicName>
/ d´Orch. / Cotype’.
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<emphasis id="B967D949FFDFFFA708A7FACFFD7AFAC8" bold="true" box="[151,716,1384,1404]" pageId="4" pageNumber="133">
Additional material examined. 
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: 
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</emphasis>
: 
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(
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, 
<collectionCode id="ED029D9EFFDFFFA70B57FACEFC1EFACE" box="[871,936,1385,1404]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="133">ZMUC</collectionCode>
): 
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SW of Pelorus Bridge, 
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a.s.l., 
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, 
<geoCoordinate id="EE27639CFFDFFFA709F1FA24FD89FA25" box="[449,575,1410,1431]" degrees="173" direction="east" minutes="34" orientation="longitude" pageId="4" pageNumber="133" precision="1" seconds="2.89" value="173.56747">173°34′2.89″E</geoCoordinate>
, 
<date id="FFAD239BFFDFFFA70A78FA24FD1CFA24" box="[584,682,1411,1430]" pageId="4" pageNumber="133" value="1984-12-28">28.xii.1984</date>
, on rocks in spray zone of falls, lgt. A. Newton &amp; M. Thayer; 
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(
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, 
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, 
<collectionCode id="ED029D9EFFDFFFA7098CFA39FE44FA03" box="[444,498,1438,1457]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="133">ANIC</collectionCode>
, 
<collectionCode id="ED029D9EFFDFFFA709CBFA39FD8FFA03" box="[507,569,1438,1457]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="133" type="Museum">SEMC</collectionCode>
): S of Canvastown, Dead Horse Creek, ca. 
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a.s.l., wet stones with algae and moss along exposed stream, 
<geoCoordinate id="EE27639CFFDFFFA70A3AFA1FFDDEFA7E" box="[522,616,1464,1484]" degrees="41" direction="south" minutes="19.59" orientation="latitude" pageId="4" pageNumber="133" precision="9" value="-41.3265">41°19.59′S</geoCoordinate>
, 
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, 
<date id="FFAD239BFFDFFFA70AD5FA1EFCF7FA7E" box="[741,833,1465,1484]" pageId="4" pageNumber="133" value="2010-11-30">30.xi.2010</date>
, lgt. Fikáček &amp; Leschen (RL1511) [
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in pure alcohol in 
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]; 
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(
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): same label data [voucher specimens to DNA isolates SLE132 and SLE133, isolated deposited in 
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(
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): same label data [voucher specimens to DNA isolates COL1799 and COL1834, isolates deposited in 
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<specimenCount id="9D15CED2FFDFFFA70A6FF9AEFD2CF9AE" box="[607,666,1545,1564]" pageId="4" pageNumber="133" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDFFFA70A93F9AEFD54F9AE" box="[675,738,1545,1564]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="4" pageNumber="133" type="Other institutional type">NZAC</collectionCode>
): same label data, dip net (RL1512). 
<emphasis id="B967D949FFDFFFA70C3FF9AEFBEEF9AE" bold="true" box="[1039,1112,1545,1564]" pageId="4" pageNumber="133">
<collectingRegion id="49D7CBB9FFDFFFA70C3FF9AEFBE7F9A8" box="[1039,1105,1545,1564]" country="New Zealand" name="Nelson" pageId="4" pageNumber="133">NELSON</collectingRegion>
:
</emphasis>
<specimenCount id="9D15CED2FFDFFFA70C6CF9AEFF75F985" pageId="4" pageNumber="133" type="generic">6 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDFFFA708FCF984FEBDF984" box="[204,267,1571,1590]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="4" pageNumber="133" type="Other institutional type">NZAC</collectionCode>
): Lyell Walkway, Deepwater Creek, 
<quantity id="4CEBA8BEFFDFFFA70A06F984FDDDF985" box="[566,619,1571,1591]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="133" unit="m" value="130.0">130 m</quantity>
a.s.l., 
<geoCoordinate id="EE27639CFFDFFFA70AAEF984FCACF984" box="[670,794,1570,1590]" degrees="41" direction="south" minutes="47" orientation="latitude" pageId="4" pageNumber="133" precision="1" seconds="39.35" value="-41.794262">41°47′39.35″S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDFFFA70B12F984FC16F985" box="[802,928,1570,1591]" degrees="172" direction="east" minutes="3" orientation="longitude" pageId="4" pageNumber="133" precision="1" seconds="20.37" value="172.05566">172°3′20.37″E</geoCoordinate>
, 
<date id="FFAD239BFFDFFFA70B96F984FC48F984" box="[934,1022,1571,1590]" pageId="4" pageNumber="133" value="2010-12-02">2.xii.2010</date>
, lgt. Fikáček &amp; Leschen (RL1525); 
<specimenCount id="9D15CED2FFDFFFA70960F999FE39F9E3" box="[336,399,1598,1617]" pageId="4" pageNumber="133" type="male">2 males</specimenCount>
(
<collectionCode id="ED029D9EFFDFFFA709A9F999FE6CF9E3" box="[409,474,1598,1617]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="133">NMPC</collectionCode>
):same label data [voucher specimens to DNA isolates COL1790 and COL1823, isolates deposited in 
<collectionCode id="ED029D9EFFDFFFA7090FF9FEFEC1F9DE" box="[319,375,1625,1644]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="133">ANIC</collectionCode>
]; 
<specimenCount id="9D15CED2FFDFFFA709B3F9FEFE0AF9DE" box="[387,444,1625,1644]" pageId="4" pageNumber="133" type="generic">3 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDFFFA709F5F9FEFE4EF9DE" box="[453,504,1625,1644]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="4" pageNumber="133">IZAS</collectionCode>
):same label data [
<specimenCount id="9D15CED2FFDFFFA70ABDF9FEFD70F9DE" box="[653,710,1625,1644]" pageId="4" pageNumber="133" type="generic">1 spec.</specimenCount>
is voucher specimen to DNA isolate A31 deposited in 
<collectionCode id="ED029D9EFFDEFFA608B8FF7BFF0DFF5D" box="[136,187,220,239]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="5" pageNumber="134">IZAS</collectionCode>
]; 
<specimenCount id="9D15CED2FFDEFFA608FAFF7BFEB1FF5D" box="[202,263,220,239]" pageId="5" pageNumber="134" type="generic">4 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60922FF7BFEE7FF5D" box="[274,337,220,239]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA6096BFF7BFE29FF5D" box="[347,415,220,239]" pageId="5" pageNumber="134">BMNH</collectionCode>
): same locality, exposed small stream and waterfall (surface scraping in splash zone), 
<date id="FFAD239BFFDEFFA6089BFF51FEB4FEBB" box="[171,258,246,265]" pageId="5" pageNumber="134" value="2010-12-02">2.xii.2010</date>
, lgt. Fikáček &amp; Leschen (RL1527); 
<specimenCount id="9D15CED2FFDEFFA60A03FF50FDD9FEB8" box="[563,623,247,266]" pageId="5" pageNumber="134" type="generic">7 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60A49FF51FD0EFEBB" box="[633,696,246,265]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA60AF1FF50FCB5FEB8" box="[705,771,247,266]" pageId="5" pageNumber="134">BMNH</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA60B3AFF51FCF4FEBB" box="[778,834,246,265]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="134">ANIC</collectionCode>
): same locality, exposed small stream and waterfall (ex mosses), 
<date id="FFAD239BFFDEFFA609A2FEB6FE5CFE96" box="[402,490,273,292]" pageId="5" pageNumber="134" value="2010-12-02">2.xii.2010</date>
, lgt. Fikáček &amp; Leschen (RL1526) [
<specimenCount id="9D15CED2FFDEFFA60B2DFEB6FCECFE96" box="[797,858,273,292]" pageId="5" pageNumber="134" type="generic">3 spec.</specimenCount>
in pure alcohol in 
<collectionCode id="ED029D9EFFDEFFA60BC8FEB6FB8FFE96" box="[1016,1081,273,292]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
]; 
<specimenCount id="9D15CED2FFDEFFA60841FE8BFF0FFE8D" box="[113,185,300,319]" pageId="5" pageNumber="134" type="generic">14 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA608F4FE8BFEB5FE8D" box="[196,259,300,319]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA6093DFE8BFEFFFE8D" box="[269,329,300,319]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="134" type="Other institutional type">NZAC</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA60962FE8BFE2EFE8D" box="[338,408,300,319]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="134" type="Museum">NHMW</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA60991FE8BFE6DFE8D" box="[417,475,300,319]" pageId="5" pageNumber="134">YMSJ</collectionCode>
): same locality, 
<date id="FFAD239BFFDEFFA60A55FE8CFD0BFE8C" box="[613,701,299,318]" pageId="5" pageNumber="134" value="2010-12-02">2.xii.2010</date>
, exposed hygropetric along small stream and waterfall (pyrethrum spraying rock surface), lgt. Fikáček &amp; Leschen (RL1533); 
<specimenCount id="9D15CED2FFDEFFA60B13FEE1FCD6FEE8" box="[803,864,326,346]" pageId="5" pageNumber="134" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60B5BFEE1FC1AFEEB" box="[875,940,326,345]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
): Lyell Walkway, past cemetery, 
<quantity id="4CEBA8BEFFDEFFA608DDFEC6FE96FEC6" box="[237,288,353,372]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="134" unit="m" value="110.0">110 m</quantity>
a.s.l., 
<geoCoordinate id="EE27639CFFDEFFA60964FEC6FE67FEC6" box="[340,465,352,372]" degrees="41" direction="south" minutes="47" orientation="latitude" pageId="5" pageNumber="134" precision="1" seconds="49.43" value="-41.797062">41°47′49.43″S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA609E9FEC6FDFBFEC6" box="[473,589,352,372]" degrees="172" direction="east" minutes="3" orientation="longitude" pageId="5" pageNumber="134" precision="1" seconds="3.97" value="172.0511">172°3′3.97″E</geoCoordinate>
, rock surface, 
<date id="FFAD239BFFDEFFA60AF3FEC6FCADFEC6" box="[707,795,353,372]" pageId="5" pageNumber="134" value="2010-12-02">2.xii.2010</date>
, lgt. Fikáček &amp; Leschen (RL1531); 
<specimenCount id="9D15CED2FFDEFFA60841FEDBFF18FE3D" box="[113,174,380,399]" pageId="5" pageNumber="134" type="generic">3 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60888FEDBFF41FE3D" box="[184,247,380,399]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
, 
<collectionCode id="ED029D9EFFDEFFA60930FEDBFE8EFE3D" box="[256,312,380,399]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="134">ANIC</collectionCode>
): Lyell Walkway, small creek at track junctions, 
<quantity id="4CEBA8BEFFDEFFA60AE5FEDCFCBDFE3D" box="[725,779,379,399]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="134" unit="m" value="115.0">115 m</quantity>
a.s.l., 
<geoCoordinate id="EE27639CFFDEFFA60B73FEDBFC09FE3D" box="[835,959,379,399]" degrees="41" direction="south" minutes="47" orientation="latitude" pageId="5" pageNumber="134" precision="1" seconds="42.59" value="-41.795162">41°47′42.59″S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60BF9FEDBFB8BFE3D" box="[969,1085,379,399]" degrees="172" direction="east" minutes="3" orientation="longitude" pageId="5" pageNumber="134" precision="1" seconds="20.1" value="172.05559">172°3′20.1″E</geoCoordinate>
, 
<date id="FFAD239BFFDEFFA60841FE31FF7FFE1B" box="[113,201,406,425]" pageId="5" pageNumber="134" value="2010-12-02">2.xii.2010</date>
, ex rock surfaces, lgt. Fikáček &amp; Leschen (RL1530); 
<specimenCount id="9D15CED2FFDEFFA60AB9FE31FD72FE18" box="[649,708,406,426]" pageId="5" pageNumber="134" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60AFEFE31FCB9FE1B" box="[718,783,406,425]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="134">NMPC</collectionCode>
): NE of Owen River, Sunrise Bridge track, Halfway creek, ca. 
<quantity id="4CEBA8BEFFDEFFA6097AFE16FE36FE76" box="[330,384,433,452]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="134" unit="m" value="440.0">440 m</quantity>
a.s.l., 
<geoCoordinate id="EE27639CFFDEFFA60986FE16FDA9FE76" box="[438,543,432,452]" degrees="41" direction="south" minutes="35.993" orientation="latitude" pageId="5" pageNumber="134" precision="1" value="-41.599884">41°35.993′S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60A19FE16FD29FE76" box="[553,671,432,452]" degrees="172" direction="east" minutes="32.236" orientation="longitude" pageId="5" pageNumber="134" precision="1" value="172.53726">172°32.236′E</geoCoordinate>
, 
<date id="FFAD239BFFDEFFA60A97FE16FD48FE76" box="[679,766,433,452]" pageId="5" pageNumber="134" value="2010-12-01">1.xii.2010</date>
, at rock along the stream, lgt. Fikáček &amp; Leschen (RL1519) [voucher specimen to DNA isolate COL1835 deposited in 
<collectionCode id="ED029D9EFFDEFFA60B13FE6BFCEDFE6D" box="[803,859,460,479]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="134">ANIC</collectionCode>
]; 
<specimenCount id="9D15CED2FFDEFFA60B5AFE6BFC11FE6D" box="[874,935,460,479]" pageId="5" pageNumber="134" type="generic">8 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60B82FE6BFC47FE6D" box="[946,1009,460,479]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="134" type="Other institutional type">NZAC</collectionCode>
): Nelson, Cawthron Park, ca. 
<quantity id="4CEBA8BEFFDEFFA6092AFE41FEF3FE48" box="[282,325,486,506]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="134" unit="m" value="30.0">30 m</quantity>
a.s.l. (coordinates ca. 
<geoCoordinate id="EE27639CFFDEFFA60A33FE41FDF1FE4B" box="[515,583,485,505]" degrees="41" direction="south" minutes="18" orientation="latitude" pageId="5" pageNumber="134" precision="925" value="-41.3">41°18′S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60A61FE41FD14FE4B" box="[593,674,485,505]" degrees="173" direction="east" minutes="13" orientation="longitude" pageId="5" pageNumber="134" precision="925" value="173.21666">173°13′E</geoCoordinate>
), 
<date id="FFAD239BFFDEFFA60A82FE41FCBFFE4B" box="[690,777,486,505]" pageId="5" pageNumber="134" value="1924-12-04">4.xii.1924</date>
, lgt. E. S. Gourlay; 
<specimenCount id="9D15CED2FFDEFFA60B82FE41FC59FE48" box="[946,1007,486,506]" pageId="5" pageNumber="134" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60BC9FE41FB8EFE4B" box="[1017,1080,486,505]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="134" type="Other institutional type">NZAC</collectionCode>
): same locality, 
<date id="FFAD239BFFDEFFA608D8FDA6FE89FDA6" box="[232,319,513,532]" pageId="5" pageNumber="134" value="1924-12-09">9.xii.1924</date>
, lgt. E. S. Gourlay; 
<specimenCount id="9D15CED2FFDEFFA609D0FDA6FD93FDA6" box="[480,549,513,532]" pageId="5" pageNumber="134" type="generic">11 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60A1DFDA6FDDDFDA6" box="[557,619,513,532]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="5" pageNumber="134">LUNZ</collectionCode>
): Owen River env., 
<quantity id="4CEBA8BEFFDEFFA60B3DFDA6FCF7FDA6" box="[781,833,513,532]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="134" unit="m" value="460.0">460 m</quantity>
a.s.l. (coordinates ca. 
<geoCoordinate id="EE27639CFFDEFFA60BC8FDA6FB8AFDA6" box="[1016,1084,512,532]" degrees="41" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="134" precision="925" value="-41.683334">41°41′S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60841FDBBFF74FD9D" box="[113,194,539,559]" degrees="172" direction="east" minutes="27" orientation="longitude" pageId="5" pageNumber="134" precision="925" value="172.45">172°27′E</geoCoordinate>
), 
<date id="FFAD239BFFDEFFA608E2FDBCFEAFFD9C" box="[210,281,539,558]" pageId="5" pageNumber="134" value="1984-01-02">2.i.1984</date>
, on tent in 
<taxonomicName id="4C137ED8FFDEFFA60948FDBBFE6FFD9C" box="[376,473,540,558]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="134" phylum="Tracheophyta" rank="genus">
<emphasis id="B967D949FFDEFFA60948FDBBFE6FFD9C" box="[376,473,540,558]" italics="true" pageId="5" pageNumber="134">Nothofagus</emphasis>
</taxonomicName>
forest ca. 
<quantity id="4CEBA8BEFFDEFFA60A03FDBCFDC8FD9D" box="[563,638,539,559]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="134" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
from Owen River, lgt. R. M. Emberson [collected in late afternoon in bright sunshine, sitting or landing on a dome tent with alternating panels of dark green and pale beige color; the tent was pitched in a clearing in 
<taxonomicName id="4C137ED8FFDEFFA609EAFDF5FD8DFDD6" box="[474,571,594,612]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="134" phylum="Tracheophyta" rank="genus">
<emphasis id="B967D949FFDEFFA609EAFDF5FD8DFDD6" box="[474,571,594,612]" italics="true" pageId="5" pageNumber="134">Nothofagus</emphasis>
</taxonomicName>
forest within 
<quantity id="4CEBA8BEFFDEFFA60A81FDF6FD4AFDD6" box="[689,764,593,612]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="134" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
of the bank of the Owen River, which at that point is a small fast-flowing river with lots of protruding rocks and little cascades overhung with beech trees (R. Emberson, in litt. 1986 to A. Newton)]; 
<specimenCount id="9D15CED2FFDEFFA609ECFD20FDA1FD28" box="[476,535,647,666]" pageId="5" pageNumber="134" type="generic">7 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60A10FD21FDE9FD2B" box="[544,607,646,665]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="134" type="Other institutional type">NZAC</collectionCode>
): Courthouse Flat, Nuggety Cr, Lutine Pool, 
<date id="FFAD239BFFDEFFA60BE4FD21FB90FD2B" box="[980,1062,646,665]" pageId="5" pageNumber="134" value="2012-01-25">25.i.2012</date>
, R. Leschen, 
<geoCoordinate id="EE27639CFFDEFFA608F1FD06FE87FD06" box="[193,305,673,692]" degrees="41.473360" direction="south" orientation="latitude" pageId="5" pageNumber="134" precision="1" value="-41.47336">41.473360°S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60908FD06FE02FD06" box="[312,436,673,692]" degrees="172.560224" direction="east" orientation="longitude" pageId="5" pageNumber="134" precision="1" value="172.56023">172.560224°E</geoCoordinate>
, along edges of still water (RL1632); 
<specimenCount id="9D15CED2FFDEFFA60ADCFD06FC90FD06" box="[748,806,673,692]" pageId="5" pageNumber="134" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60B1FFD06FCD8FD06" box="[815,878,673,692]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="134" type="Other institutional type">NZAC</collectionCode>
): same locality, 
<date id="FFAD239BFFDEFFA60BC1FD06FBF6FD06" box="[1009,1088,673,692]" pageId="5" pageNumber="134" value="2012-01-26">26.i.2012</date>
(RL1634). 
<emphasis id="B967D949FFDEFFA608E0FD1BFEAEFD7D" bold="true" box="[208,280,700,719]" pageId="5" pageNumber="134">
<collectingRegion id="49D7CBB9FFDEFFA608E0FD1BFEA7FD7F" box="[208,273,700,719]" country="New Zealand" name="BULLER" pageId="5" pageNumber="134">BULLER</collectingRegion>
:
</emphasis>
<specimenCount id="9D15CED2FFDEFFA6092FFD1BFEEAFD7D" box="[287,348,700,719]" pageId="5" pageNumber="134" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED029D9EFFDEFFA60957FD1BFE1AFD7D" box="[359,428,700,719]" pageId="5" pageNumber="134">BMNH</collectionCode>
): Greymouth, Ten Mile Creek [as ‘ 
<quantity id="4CEBA8BEFFDEFFA60AE5FD1BFCACFD7C" box="[725,794,699,719]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="5" pageNumber="134" unit="mi" value="10.0">10 mile</quantity>
creek’], lgt. Helms [mouth of Ten Mile Creek where it meets the west coast is at ca. 
<geoCoordinate id="EE27639CFFDEFFA60A27FD71FDCFFD5B" box="[535,633,725,745]" degrees="42" direction="south" minutes="20" orientation="latitude" pageId="5" pageNumber="134" precision="15" seconds="05" value="-42.33472">42°20′05″S</geoCoordinate>
, 
<geoCoordinate id="EE27639CFFDEFFA60AB2FD71FD47FD58" box="[642,753,725,746]" degrees="171" direction="east" minutes="15" orientation="longitude" pageId="5" pageNumber="134" precision="15" seconds="38" value="171.26056">171°15′38″E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="C30956D0FFDEFFAE0841FCA6FE3CFDA1" lastPageId="13" lastPageNumber="142" pageId="5" pageNumber="134" type="description">
<paragraph id="8BAC055BFFDEFFA60841FCA6FDF5FC48" blockId="5.[113,1089,769,1594]" pageId="5" pageNumber="134">
<emphasis id="B967D949FFDEFFA60841FCA6FEA1FCAB" bold="true" box="[113,279,769,793]" pageId="5" pageNumber="134">Redescription.</emphasis>
Body elongate, strongly depressed (
<figureCitation id="132819DEFFDEFFA60A9DFCA5FCA1FCA8" box="[685,791,770,794]" captionStart="Figs" captionStartId="2.[151,190,1538,1557]" captionTargetBox="[152,1125,232,1517]" captionTargetId="figure-10@2.[151,1125,215,1519]" captionTargetPageId="2" captionText="Figs. 1–4. Horelophus walkeri Orchymont, 1913.1–2 – general habitus in dorsal and lateral view (photo M.Fikáček); 3–4 – individuals on the wet rock surfaces at Dead Horse Creek, Marlborough (photo R. Leschen)." figureDoi="http://doi.org/10.5281/zenodo.5329845" httpUri="https://zenodo.org/record/5329845/files/figure.png" pageId="5" pageNumber="134">Figs. 1–2</figureCitation>
); total length 
<quantity id="4CEBA8BEFFDEFFA60B86FCA5FB8AFCAB" box="[950,1084,770,794]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.1" metricValueMin="2.1" pageId="5" pageNumber="134" unit="mm" value="2.6" valueMax="3.1" valueMin="2.1">2.1–3.1 mm</quantity>
, maximum width of pronotum 0.8–1.0 mm, maximum width of elytra 1.0– 
<quantity id="4CEBA8BEFFDEFFA60BBDFC86FC54FC8B" box="[909,994,801,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="134" unit="mm" value="1.5">1.5 mm</quantity>
. General coloration of dorsal surface piceous brown to black often with greenish tinge, lateral margins of pronotum and elytra with wide pale reddish stripe, anterolateral margins of clypeus vaguely dark reddish; ventral parts dark brown to black, only gular area and prosternum paler brown and epipleura pale reddish; head appendages dark reddish, maxillary palpomere 4 and antennal club dark brown; legs with brownish coxae, pale reddish femora, tibiae and basal tarsomeres, and distal tarsomeres darkened.
</paragraph>
<paragraph id="8BAC055BFFDEFFA408A0FBA5FCCBFEE1" blockId="5.[113,1089,769,1594]" lastBlockId="7.[113,1088,219,339]" lastPageId="7" lastPageNumber="136" pageId="5" pageNumber="134">
<emphasis id="B967D949FFDEFFA608A0FBA5FE2BFBA8" bold="true" box="[144,413,1025,1050]" pageId="5" pageNumber="134">
Head. 
<emphasis id="B967D949FFDEFFA608E8FBA6FE2BFBA8" bold="true" box="[216,413,1025,1050]" italics="true" pageId="5" pageNumber="134">Clypeus and frons</emphasis>
</emphasis>
(
<figureCitation id="132819DEFFDEFFA60999FBA5FE43FBA8" box="[425,501,1026,1050]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="5" pageNumber="134">Fig. 18</figureCitation>
) with sparse but rather coarse punctation, each puncture bearing a short apically widened seta, frons with several trichobothria, clypeus without trichobothria; frontoclypeal suture only distinct laterally, arising closely before eyes; clypeus slightly expanded laterally, covering bases of antennae, anteromedian margin shallowly concave. 
<emphasis id="B967D949FFDEFFA608E8FB25FEBAFB2B" bold="true" box="[216,268,1154,1177]" italics="true" pageId="5" pageNumber="134">Eyes</emphasis>
small, protruding from outline of head, separated by 6× width of one eye. 
<emphasis id="B967D949FFDEFFA60841FB06FF7CFB08" bold="true" box="[113,202,1185,1210]" italics="true" pageId="5" pageNumber="134">Labrum</emphasis>
(
<figureCitation id="132819DEFFDEFFA608E8FB05FEE3FB08" box="[216,341,1186,1210]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="5" pageNumber="134">Figs. 11–12</figureCitation>
) largely exposed dorsally, only slightly retracted under clypeus, widest subbasally, strongly narrowed basally and arcuately narrowing anteriad, shallowly bisinuate on anterior margin; dorsal surface bearing two pairs of long sublateral setae and the ground punctation similar to that on clypeus; epipharynx with a lateral row of three stout setae on each side, median portion with two vertical rows of long cuticular spines and cone-shaped group of similar spines anteriorly, basal portion with densely pubescent membranous cone. 
<emphasis id="B967D949FFDEFFA60841FAC6FF53FAC8" bold="true" box="[113,229,1377,1402]" italics="true" pageId="5" pageNumber="134">Mandibles</emphasis>
(
<figureCitation id="132819DEFFDEFFA608C6FAC5FEF0FACB" box="[246,326,1377,1402]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="5" pageNumber="134">Fig. 15</figureCitation>
) symmetrical, with distinct mandibular angle, mandibular apex bifid; mediodistal portion with a group of long cuticular projections, medioproximal portion with very fine setae, mola rather small, bearing numerous backwards directed setae on median face. 
<emphasis id="B967D949FFDEFFA6089DFA66FEB5FA68" bold="true" box="[173,259,1473,1498]" italics="true" pageId="5" pageNumber="134">Maxilla</emphasis>
(
<figureCitation id="132819DEFFDEFFA60923FA65FED4FA68" box="[275,354,1474,1498]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="5" pageNumber="134">Fig. 13</figureCitation>
) with a simple subtriangular cardo lacking trichobothria; basistipes triangular, bearing few fine setae only; mediostipes rather vaguely delimited from lacinia, the latter membranous, bearing fine hair-like setae mesally and few stouter and longer setae distally; galea short, with distal setae arranged into well-defined rows; palpifer rather small, with few rather long setae; maxillary palpus with 4 palpomeres, palpomere 1 minute, palpomeres 2 and 4 subequal in length, ca. twice length of palpomere 3; base of palpomere 4 with a group of 10–11 digitiform sensilla on dorsolateral surface. 
<emphasis id="B967D949FFDDFFA50B2CFBF8FCC4FBCA" bold="true" box="[796,882,1119,1144]" italics="true" pageId="6" pageNumber="135">Labium</emphasis>
(
<figureCitation id="132819DEFFDDFFA50BB1FBC7FC6CFBCA" box="[897,986,1120,1144]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="6" pageNumber="135">Figs. 14</figureCitation>
, 
<figureCitation id="132819DEFFDDFFA50BD5FBC7FB9EFBCA" box="[997,1064,1120,1144]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="6" pageNumber="135">20–21</figureCitation>
) with submentum ca. as long and wide as mentum, bearing sparsely arranged setae; mentum transverse, ca. 1.5× wider than long, with continually convex anterior margin, its surface bearing a few larger setae and very weak and sparse mesh-like microsculpture, lateral margins without rows of setae; prementum subdivided into two membranous lobes bearing anteromedian row of fine setae (becoming spoon-like medially) and a dorsal sublateral row of setae, palpifer vaguely sclerotized; labial palpus with three palpomeres, palpomere 1 minute, palpomere 2 ca. half as long as palpomere 3; palpomere 3 with one subbasal digitiform sensillum and few minute apical sensilla. 
<emphasis id="B967D949FFDDFFA509A0FAC7FE59FACA" bold="true" box="[400,495,1376,1400]" italics="true" pageId="6" pageNumber="135">Antenna</emphasis>
(
<figureCitation id="132819DEFFDDFFA509CEFAC7FD36FACA" box="[510,640,1376,1400]" captionStart="Figs" captionStartId="6.[151,190,929,948]" captionTargetBox="[159,1110,228,906]" captionTargetId="figure-249@6.[151,1122,215,921]" captionTargetPageId="6" captionText="Figs. 11–17. Mouthparts and head appendages of Horelophus walkeri Orchymont, 1913. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female)." figureDoi="http://doi.org/10.5281/zenodo.5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="6" pageNumber="135">Figs. 16–17</figureCitation>
) with 9 antennomeres, scapus conical, ca. as long as pedicel, pedicel widest proximally, bearing a few pore-like sensilla and one tiny seta, antennomere 3 ca. as long as antennomeres 4–5 combined, cupula small, antennomeres 7–9 forming a distinct, loosely segmented and densely pubescent antennal club; length of antennae slightly sexually dimorphic, slightly longer (and antennomeres slightly more elongate) in males than in females, proportions of antennomere 9 strongly sexually dimorphic, ca. 3× as long as wide in males and 1.5× as long as wide in females, bearing numerous small conical sensilla on ventral surface (intermixed with trichoid setae on whole surface in male, more accumulated in subapical group in female). 
<emphasis id="B967D949FFDCFFA40A7EFF5DFD33FEA1" bold="true" box="[590,645,250,275]" italics="true" pageId="7" pageNumber="136">Gula</emphasis>
(
<figureCitation id="132819DEFFDCFFA40AA5FF5CFD55FEA1" box="[661,739,251,275]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="7" pageNumber="136">Fig. 19</figureCitation>
) narrow, gular sutures narrowly separated at midlength, slightly diverging at tentorial pits, the latter distinct, elongate. 
<emphasis id="B967D949FFDCFFA40C39FEBCFF19FEE0" bold="true" italics="true" pageId="7" pageNumber="136">Temporae</emphasis>
with short but distinct ridge arising from inner margin of each eye.
</paragraph>
<caption id="DF6C55D3FFDDFFA508A7FC06FEC6FC5B" ID-DOI="http://doi.org/10.5281/zenodo.5329851" ID-Zenodo-Dep="5329851" httpUri="https://zenodo.org/record/5329851/files/figure.png" pageId="6" pageNumber="135" startId="6.[151,190,929,948]" targetBox="[159,1110,228,906]" targetPageId="6">
<paragraph id="8BAC055BFFDDFFA508A7FC06FEC6FC5B" blockId="6.[151,1127,929,1002]" pageId="6" pageNumber="135">
Figs. 11–17. Mouthparts and head appendages of 
<taxonomicName id="4C137ED8FFDDFFA50A0EFC05FC33FC06" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[574,901,929,948]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFDDFFA50A0EFC05FD51FC06" box="[574,743,930,948]" italics="true" pageId="6" pageNumber="135">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFDDFFA50ADCFC06FC33FC06" author="ORCHYMONT A." box="[748,901,929,948]" pageId="6" pageNumber="135" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
</taxonomicName>
. 11 – labrum, dorsal view; 12 – labrum, ventral view (epipharynx); 13 – maxilla; 14 – mentum and prementum; 15 – mandible; 16–17 – antenna (16 – male, 17 – female).
</paragraph>
</caption>
<caption id="DF6C55D3FFDCFFA40841FA40FD50F982" ID-DOI="http://doi.org/10.5281/zenodo.5329853" ID-Zenodo-Dep="5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="7" pageNumber="136" startId="7.[113,152,1511,1530]" targetBox="[117,1086,398,1481]" targetPageId="7">
<paragraph id="8BAC055BFFDCFFA40841FA40FD50F982" blockId="7.[113,1090,1511,1584]" pageId="7" pageNumber="136">
Figs.18–24.Morphology of 
<taxonomicName id="4C137ED8FFDCFFA40951FA4FFD14FA48" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[353,674,1511,1530]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFDCFFA40951FA4FFDBEFA48" box="[353,520,1512,1530]" italics="true" pageId="7" pageNumber="136">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFDCFFA40A3BFA40FD14FA48" author="ORCHYMONT A." box="[523,674,1511,1530]" pageId="7" pageNumber="136" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
</taxonomicName>
.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum.
</paragraph>
</caption>
<paragraph id="8BAC055BFFD3FFAB0885FF7BFE0AFD21" blockId="8.[151,1127,219,1587]" pageId="8" pageNumber="137">
<emphasis id="B967D949FFD3FFAB0885FF7BFE1FFF41" bold="true" box="[181,425,220,244]" pageId="8" pageNumber="137">
Prothorax. 
<emphasis id="B967D949FFD3FFAB0909FF7BFE1FFF41" bold="true" box="[313,425,220,243]" italics="true" pageId="8" pageNumber="137">Pronotum</emphasis>
</emphasis>
(
<figureCitation id="132819DEFFD3FFAB0988FF7CFE4EFF46" box="[440,504,219,244]" captionStart="Figs" captionStartId="2.[151,190,1538,1557]" captionTargetBox="[152,1125,232,1517]" captionTargetId="figure-10@2.[151,1125,215,1519]" captionTargetPageId="2" captionText="Figs. 1–4. Horelophus walkeri Orchymont, 1913.1–2 – general habitus in dorsal and lateral view (photo M.Fikáček); 3–4 – individuals on the wet rock surfaces at Dead Horse Creek, Marlborough (photo R. Leschen)." figureDoi="http://doi.org/10.5281/zenodo.5329845" httpUri="https://zenodo.org/record/5329845/files/figure.png" pageId="8" pageNumber="137">Fig. 1</figureCitation>
, 
<figureCitation id="132819DEFFD3FFAB0A36FF7BFD94FF46" box="[518,546,220,244]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="8" pageNumber="137">24</figureCitation>
) subrectangular, widest in anterior third, with weakly projecting anterior corners, lateral margins not forming continuous curve with lateral margins of elytra; surface with one posteromedian and two sublateral depressions, ground punctation sparse and rather coarse, similar to that of clypeus, trichobothria missing; lateral edges slightly sinuate, with fine marginal bead extending onto anterior edge. 
<emphasis id="B967D949FFD3FFAB0B96FEFBFB9FFEC1" bold="true" box="[934,1065,348,371]" italics="true" pageId="8" pageNumber="137">Hypomeron</emphasis>
(
<figureCitation id="132819DEFFD3FFAB0C0BFEFCFF6DFE26" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="8" pageNumber="137">Fig. 22–23</figureCitation>
) with rather narrow lateral glabrous portion and densely pubescent median portion, portions not divided by a ridge, hypomeral process large, rounded mesally. 
<emphasis id="B967D949FFD3FFAB0BD3FE3BFBD1FE01" bold="true" box="[995,1127,412,435]" italics="true" pageId="8" pageNumber="137">Prosternum</emphasis>
(
<figureCitation id="132819DEFFD3FFAB08AFFE1CFF5BFE66" box="[159,237,443,468]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="8" pageNumber="137">Fig. 23</figureCitation>
) rather long anterior to procoxae, ca. 0.6× as long as procoxa, without longitudinal or transverse ridges or impressions, slightly convex on anterior margin, prosternal process indistinct, concealed between procoxae. 
<emphasis id="B967D949FFD3FFAB0A7EFE5CFD5FFDA1" bold="true" box="[590,745,507,532]" italics="true" pageId="8" pageNumber="137">Coxal cavities</emphasis>
closed internally, open posteriorly, coxal fissure rather long, closed, notopleural suture distinct. Accessory ridge below posterior pronotal margin, laterally obliterated, recognizable as short and indistict ‘transverse fold’. 
<emphasis id="B967D949FFD3FFAB08A7FDFCFF4DFDC6" bold="true" box="[151,251,603,628]" italics="true" pageId="8" pageNumber="137">Profurca</emphasis>
(
<figureCitation id="132819DEFFD3FFAB093AFDFCFEEEFDC6" box="[266,344,603,628]" captionStart="Figs" captionStartId="7.[113,152,1511,1530]" captionTargetBox="[117,1086,398,1481]" captionTargetId="figure-12@7.[114,1086,397,1482]" captionTargetPageId="7" captionText="Figs.18–24.Morphology of Horelophus walkeri Orchymont, 1913.18 – head, dorsal view (inset: detail of punctation of frons); 19 – head, ventral view; 20 – maxillary palpomere 4, dorsal view; 21 – detail of mouthparts; 22 – prothorax, posterior view; 23 – prothorax, ventral view; 24 – right side of pronotum." figureDoi="http://doi.org/10.5281/zenodo.5329853" httpUri="https://zenodo.org/record/5329853/files/figure.png" pageId="8" pageNumber="137">Fig. 22</figureCitation>
) short, profurcal arms widely separated, in the form of slightly asymmetrical plate-like extensions.
</paragraph>
<paragraph id="8BAC055BFFD3FFAB0885FD3BFD2FFA61" blockId="8.[151,1127,219,1587]" pageId="8" pageNumber="137">
<emphasis id="B967D949FFD3FFAB0885FD3BFE28FD01" bold="true" box="[181,414,667,692]" pageId="8" pageNumber="137">
Mesothorax. 
<taxonomicName id="4C137ED8FFD3FFAB097AFD3CFE28FD01" box="[330,414,667,691]" class="Gastropoda" family="Fissurellidae" genus="Scutum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="137" phylum="Mollusca" rank="genus">
<emphasis id="B967D949FFD3FFAB097AFD3CFE28FD01" bold="true" box="[330,414,667,691]" italics="true" pageId="8" pageNumber="137">Scutum</emphasis>
</taxonomicName>
</emphasis>
(
<figureCitation id="132819DEFFD3FFAB099CFD3CFE4EFD01" box="[428,504,667,691]" captionStart="Figs" captionStartId="12.[151,190,1307,1326]" captionTargetBox="[151,1110,224,1288]" captionTargetId="figure-78@12.[148,1123,216,1300]" captionTargetPageId="12" captionText="Figs. 42–50. Morphology of Horelophus walkeri Orchymont, 1913. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing." figureDoi="http://doi.org/10.5281/zenodo.5329861" httpUri="https://zenodo.org/record/5329861/files/figure.png" pageId="8" pageNumber="137">Fig. 46</figureCitation>
) with finely microsculptured median portion, bearing sparsely arranged setae; scutellar shield exposed, triangular, pointed posteriorly, slightly longer than wide, with a few fine setae present on its surface. 
<emphasis id="B967D949FFD3FFAB0AEFFD7CFC84FD46" bold="true" box="[735,818,731,756]" italics="true" pageId="8" pageNumber="137">Elytron</emphasis>
(
<figureCitation id="132819DEFFD3FFAB0B71FD7CFC77FD46" box="[833,961,731,756]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="8" pageNumber="137">Figs. 27–30</figureCitation>
) elongate, with a distinct mesal depression in anterior third; sutural stria present, reaching ca. midlength of elytron; elytral series irregular, formed of the punctures of the same size and morphology as subserially arranged interval punctation, hence elytron seemingly bears ca. 18 more or less irregular series of punctures; scutellary stria absent (not visible even in slide-mounted elytron); alternate elytral intervals each with a few short trichobothria, punctures of elytral series and intervals each bearing a short club-like seta; lateral edge with a narrow bead, finely crenulate; epipleuron moderately wide anteriorly, gradually narrowing from level of metaventrite posteriad, reaching subapically, median pubescent portion not delimited from lateral bare one by a line or ridge; ventral elytral surface without any elevated ridges, only with a narrow longitudinal field of fine spines situated sublaterally between anterior fourth and midlength. 
<emphasis id="B967D949FFD3FFAB08A7FB9CFE92FBE1" bold="true" box="[151,292,1083,1107]" italics="true" pageId="8" pageNumber="137">Mesoventrite</emphasis>
(
<figureCitation id="132819DEFFD3FFAB0901FB9CFECAFBE1" box="[305,380,1083,1107]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="8" pageNumber="137">Fig. 25</figureCitation>
) distinctly divided from mesanepisternum by distinct anapleural suture; mesoventrite subtriangular in shape in anterior two thirds, widely extended laterad in posterior third, lateral extensions bearing distinct coxal lobes; whole mesoventrite nearly flat, without distinct protuberances or ridges, whole surface except for the lateral wings bearing sparse pubescence; mesoventral process narrow. 
<emphasis id="B967D949FFD3FFAB0A69FB1CFCBEFB61" bold="true" box="[601,776,1211,1235]" italics="true" pageId="8" pageNumber="137">Mesanepisterna</emphasis>
not meeting anteromesally, very narrowly divided by anterior portion of mesoventrite; anterior collar well-defined, moderately wide; mesal portion of each mesanepisternum pubescent, large lateral portions bare. 
<emphasis id="B967D949FFD3FFAB0C3FFB5CFF68FA81" bold="true" italics="true" pageId="8" pageNumber="137">Mesepimeron</emphasis>
with large ventral portion, not reaching anterior collar or mesanepisternum anteriorly, forming lateral margin of coxal cavity; its whole surface pubescent. 
<emphasis id="B967D949FFD3FFAB0B54FA9CFC4AFAE1" bold="true" box="[868,1020,1339,1364]" italics="true" pageId="8" pageNumber="137">Coxal cavities</emphasis>
obliquely transverse, ca. 1.5× wider than long, very narrowly separated from each other by mesoventral and metaventral processes; internal postcoxal wall moderately wide mesally and posteriorly. 
<emphasis id="B967D949FFD3FFAB08A7FA3CFEB8FA06" bold="true" box="[151,270,1435,1460]" italics="true" pageId="8" pageNumber="137">Mesofurca</emphasis>
(
<figureCitation id="132819DEFFD3FFAB092CFA3CFE2DFA06" box="[284,411,1435,1460]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="8" pageNumber="137">Figs. 31–32</figureCitation>
) well-developed but short, arising as two widely separate plate-like extensions from posterior wall of coxal cavities.
</paragraph>
<paragraph id="8BAC055BFFD3FFA80885FA7BFEDFFE61" blockId="8.[151,1127,219,1587]" lastBlockId="11.[113,1089,219,1619]" lastPageId="11" lastPageNumber="140" pageId="8" pageNumber="137">
<emphasis id="B967D949FFD3FFAB0885FA7BFE66FA41" bold="true" box="[181,464,1500,1524]" pageId="8" pageNumber="137">
Metathorax. 
<emphasis id="B967D949FFD3FFAB097FFA7BFE66FA41" bold="true" box="[335,464,1500,1523]" italics="true" pageId="8" pageNumber="137">Metanotum</emphasis>
</emphasis>
(
<figureCitation id="132819DEFFD3FFAB09D2FA7CFD82FA41" box="[482,564,1499,1523]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="8" pageNumber="137">Fig. 26</figureCitation>
) weakly sclerotized, ca. 2× wider than long, with rather wide anterior membranous area, alacristae slightly diverging posteriad. 
<emphasis id="B967D949FFD3FFAB0BEBFA5CFBD0F9A1" bold="true" box="[987,1126,1531,1555]" italics="true" pageId="8" pageNumber="137">Metaventrite</emphasis>
(
<figureCitation id="132819DEFFD3FFAB08AFF9BCFF5AF981" box="[159,236,1563,1587]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="8" pageNumber="137">Fig. 25</figureCitation>
) ca. 1.5× longer than mesoventrite, evenly convex, without defined median portion, whole surface (except for a small posteromedian area) bearing dense pubescence; katepisternum narrowly exposed, metacoxal process short but distinctly exposed. Postcoxal ridge very narrow but well-defined. 
<emphasis id="B967D949FFD1FFA909EBFA9EFD2BFAE3" bold="true" box="[475,669,1337,1361]" italics="true" pageId="10" pageNumber="139">Metanepisternum</emphasis>
ca. 6× longer than wide, with an obliquely transverse strengthened ridge anteriorly; whole surface pubescent. 
<emphasis id="B967D949FFD1FFA90B61FAFEFC55FAC3" bold="true" box="[849,995,1369,1393]" italics="true" pageId="10" pageNumber="139">Metepimeron</emphasis>
with minute but distinct ventral portion. 
<emphasis id="B967D949FFD1FFA909F5FADFFD8FFA23" bold="true" box="[453,569,1400,1425]" italics="true" pageId="10" pageNumber="139">Metafurca</emphasis>
(
<figureCitation id="132819DEFFD1FFA90A78FADEFD15FA20" box="[584,675,1401,1426]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="10" pageNumber="139">Figs. 31</figureCitation>
, 
<figureCitation id="132819DEFFD1FFA90A9FFADEFD7DFA23" box="[687,715,1401,1425]" captionStart="Figs" captionStartId="12.[151,190,1307,1326]" captionTargetBox="[151,1110,224,1288]" captionTargetId="figure-78@12.[148,1123,216,1300]" captionTargetPageId="12" captionText="Figs. 42–50. Morphology of Horelophus walkeri Orchymont, 1913. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing." figureDoi="http://doi.org/10.5281/zenodo.5329861" httpUri="https://zenodo.org/record/5329861/files/figure.png" pageId="10" pageNumber="139">45</figureCitation>
) rather large, Y-shaped; stalk grooved medially, without basal extensions; lateral arms rather long, with large anterobasal extensions, apical portions roundly plate-like. 
<emphasis id="B967D949FFD1FFA90A3CFA1FFD37FA63" bold="true" box="[524,641,1464,1489]" italics="true" pageId="10" pageNumber="139">Hind wing</emphasis>
(
<figureCitation id="132819DEFFD1FFA90AA2FA1EFD57FA63" box="[658,737,1465,1489]" captionStart="Figs" captionStartId="12.[151,190,1307,1326]" captionTargetBox="[151,1110,224,1288]" captionTargetId="figure-78@12.[148,1123,216,1300]" captionTargetPageId="12" captionText="Figs. 42–50. Morphology of Horelophus walkeri Orchymont, 1913. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing." figureDoi="http://doi.org/10.5281/zenodo.5329861" httpUri="https://zenodo.org/record/5329861/files/figure.png" pageId="10" pageNumber="139">Fig. 50</figureCitation>
) well developed, ca. 2× longer than elytron, venation well-developed in basal half, absent in distal half; anal lobe rather large, well-defined by anal notch; RA tightly attached to ScA except of subbasally, both reaching to triangular radial cell, RP 
<subScript id="1797071EFFD1FFA90987F98EFE79F98A" attach="left" box="[439,463,1577,1592]" fontSize="6" pageId="10" pageNumber="139">3+4</subScript>
rather long, cross vein r4 arising from its distal portion; MP 
<subScript id="1797071EFFD1FFA90C7EF98EFBD1F98A" attach="left" box="[1102,1127,1577,1592]" fontSize="6" pageId="10" pageNumber="139">1+2</subScript>
strong, forming a loop with distal portion of RP, the latter reaching midlength between the loop and wing base, median spur short but distinct; vein complex of MP 
<subScript id="1797071EFFD0FFA80BAEFEACFC10FEA8" attach="left" box="[926,934,267,282]" fontSize="6" pageId="11" pageNumber="140">4</subScript>
+Cu+AA well developed, not connected to MP 
<subScript id="1797071EFFD0FFA809FBFE8CFE55FE88" attach="left" box="[459,483,299,314]" fontSize="6" pageId="11" pageNumber="140">1+2</subScript>
by cross veins, with well-defined and completely closed basal and wedge cells; AA 
<subScript id="1797071EFFD0FFA809BAFEECFE24FEE8" attach="left" box="[394,402,331,346]" fontSize="6" pageId="11" pageNumber="140">4</subScript>
, CuA 
<subScript id="1797071EFFD0FFA809E1FEECFE6FFEE8" attach="left" box="[465,473,331,346]" fontSize="6" pageId="11" pageNumber="140">2</subScript>
and MP 
<subScript id="1797071EFFD0FFA80A02FEECFD8CFEE8" attach="left" box="[562,570,331,346]" fontSize="6" pageId="11" pageNumber="140">4</subScript>
reaching posterior margin of wing in basal third, branching of CuA 
<subScript id="1797071EFFD0FFA80904FECCFE8AFEC8" attach="left" box="[308,316,363,378]" fontSize="6" pageId="11" pageNumber="140">2</subScript>
and MP 
<subScript id="1797071EFFD0FFA809A8FECCFE16FEC8" attach="left" box="[408,416,363,378]" fontSize="6" pageId="11" pageNumber="140">4</subScript>
slightly variable between specimens or even within the specimen, forming a continuous series between X-shaped and H-shaped branching pattern; AA 
<subScript id="1797071EFFD0FFA80C09FE2CFBF7FE28" attach="right" box="[1081,1089,395,410]" fontSize="6" pageId="11" pageNumber="140">4</subScript>
well developed, nearly reaching posterior wing margin; AP 
<subScript id="1797071EFFD0FFA80ADEFE0CFCB0FE08" attach="left" box="[750,774,427,442]" fontSize="6" pageId="11" pageNumber="140">1+2</subScript>
well developed, reaching ca. midlength of anal lobe.
</paragraph>
<caption id="DF6C55D3FFD2FFAA0841FA68FC2AF980" ID-DOI="http://doi.org/10.5281/zenodo.5329857" ID-Zenodo-Dep="5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="9" pageNumber="138" startId="9.[113,152,1487,1506]" targetBox="[113,1084,219,1443]" targetPageId="9">
<paragraph id="8BAC055BFFD2FFAA0841FA68FC2AF980" blockId="9.[113,1089,1487,1587]" pageId="9" pageNumber="138">
Figs. 25–35. Morphology of 
<taxonomicName id="4C137ED8FFD2FFAA0943FA76FD74FA51" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[371,706,1487,1507]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD2FFAA0943FA76FDA9FA51" box="[371,543,1489,1507]" italics="true" pageId="9" pageNumber="138">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFD2FFAA0A17FA68FD74FA51" author="ORCHYMONT A." box="[551,706,1487,1507]" pageId="9" pageNumber="138" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
</taxonomicName>
. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view
</paragraph>
</caption>
<caption id="DF6C55D3FFD1FFA908A7FB27FBE3FB7A" ID-DOI="http://doi.org/10.5281/zenodo.5329859" ID-Zenodo-Dep="5329859" httpUri="https://zenodo.org/record/5329859/files/figure.png" pageId="10" pageNumber="139" startId="10.[151,190,1152,1171]" targetBox="[152,1122,219,1134]" targetPageId="10">
<paragraph id="8BAC055BFFD1FFA908A7FB27FBE3FB7A" blockId="10.[151,1127,1152,1225]" pageId="10" pageNumber="139">
Figs. 36–41. Legs of 
<taxonomicName id="4C137ED8FFD1FFA90975FB26FD30FB21" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[325,646,1152,1171]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD1FFA90975FB26FE5AFB21" box="[325,492,1153,1171]" italics="true" pageId="10" pageNumber="139">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFD1FFA909C0FB27FD30FB21" author="ORCHYMONT A." box="[496,646,1152,1171]" pageId="10" pageNumber="139" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
</taxonomicName>
. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor.
</paragraph>
</caption>
<paragraph id="8BAC055BFFD0FFA808A0FE7BFC59FB21" blockId="11.[113,1089,219,1619]" pageId="11" pageNumber="140">
<emphasis id="B967D949FFD0FFA808A0FE7BFF73FE46" bold="true" box="[144,197,476,500]" pageId="11" pageNumber="140">Legs</emphasis>
(
<figureCitation id="132819DEFFD0FFA808E1FE7CFEE7FE41" box="[209,337,475,499]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="10.[151,190,1152,1171]" captionStartId-1="12.[151,190,1307,1326]" captionTargetBox-0="[152,1122,219,1134]" captionTargetBox-1="[151,1110,224,1288]" captionTargetId-0="figure-145@10.[151,1122,219,1136]" captionTargetId-1="figure-78@12.[148,1123,216,1300]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="Figs. 36–41. Legs of Horelophus walkeri Orchymont, 1913. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor." captionText-1="Figs. 42–50. Morphology of Horelophus walkeri Orchymont, 1913. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing." figureDoi-0="http://doi.org/10.5281/zenodo.5329859" figureDoi-1="http://doi.org/10.5281/zenodo.5329861" httpUri-0="https://zenodo.org/record/5329859/files/figure.png" httpUri-1="https://zenodo.org/record/5329861/files/figure.png" pageId="11" pageNumber="140">Figs. 36–44</figureCitation>
). 
<emphasis id="B967D949FFD0FFA80951FE7CFE19FE41" bold="true" box="[353,431,475,499]" italics="true" pageId="11" pageNumber="140">Coxae:</emphasis>
procoxae subglobular, narrowly transverse, sparsely pubescent ventrally; mesocoxae transverse, rather robust mesally, narrowly separated, finely pubescent ventrally; metacoxae narrowly transverse, subrectangular in ventral view, sparsely pubescent on whole ventral surface. 
<emphasis id="B967D949FFD0FFA8094AFD9DFE48FDE1" bold="true" box="[378,510,570,595]" italics="true" pageId="11" pageNumber="140">Trochanters</emphasis>
with proximal parts concealed by coxae, distal subtriangular parts exposed ventrally, pubescent. 
<emphasis id="B967D949FFD0FFA80A29FDFBFDD9FDC1" bold="true" box="[537,623,604,627]" italics="true" pageId="11" pageNumber="140">Femora</emphasis>
attached to trochanters by their posteromesal (in meso- and metafemora) or anteromesal (on profemora) portions only, anteromesal (in meso- and metafemora) or posteromesal bases (in profemora) free, angulate; pro- and mesofemora densely pubescent in their basal portion, metafemora bearing such pubescence only on extreme basoanterior portions, most of their surface bearing sparsely arranged spine-like setae; tibial grooves not defined on any femora. 
<emphasis id="B967D949FFD0FFA80A45FD5DFD0CFCA1" bold="true" box="[629,698,762,787]" italics="true" pageId="11" pageNumber="140">Tibiae</emphasis>
slightly longer than femora, slightly widening distad; each tibia with three dorsal and three lateral series of spines, distal portion with a group of enlarged spines and two rather short but stout tibial spurs; protibia with two large closely associated spines subdistally on outer margin, outermost series of spines closely aggregated. 
<emphasis id="B967D949FFD0FFA808C6FCDCFE9BFC21" bold="true" box="[246,301,891,915]" italics="true" pageId="11" pageNumber="140">Tarsi</emphasis>
with 5 tarsomeres, basal tarsomere short, subequal in length to each of tarsomeres 2–4, tarsomere 5 as long as tarsomeres 2–4 (in pro- and mesotarsus) or 3–4 (in metatarsus) combined. Ventral setae of pro- and mesotarsomeres 1–3 sexually dimorphic, in shape of long wide plates in males (
<figureCitation id="132819DEFFD0FFA809EEFC7CFDE4FC41" box="[478,594,987,1011]" captionStart="Figs" captionStartId="10.[151,190,1152,1171]" captionTargetBox="[152,1122,219,1134]" captionTargetId="figure-145@10.[151,1122,219,1136]" captionTargetPageId="10" captionText="Figs. 36–41. Legs of Horelophus walkeri Orchymont, 1913. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor." figureDoi="http://doi.org/10.5281/zenodo.5329859" httpUri="https://zenodo.org/record/5329859/files/figure.png" pageId="11" pageNumber="140">Fig. 36–37</figureCitation>
), shorter, spine-like in females (
<figureCitation id="132819DEFFD0FFA80BAAFC7CFC50FC46" box="[922,998,987,1012]" captionStart="Figs" captionStartId="10.[151,190,1152,1171]" captionTargetBox="[152,1122,219,1134]" captionTargetId="figure-145@10.[151,1122,219,1136]" captionTargetPageId="10" captionText="Figs. 36–41. Legs of Horelophus walkeri Orchymont, 1913. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor." figureDoi="http://doi.org/10.5281/zenodo.5329859" httpUri="https://zenodo.org/record/5329859/files/figure.png" pageId="11" pageNumber="140">Fig. 38</figureCitation>
), ventral setae of pro- and mesotarsomeres 4–5 and metatarsomeres 1–5 of both sexes spine-like (
<figureCitation id="132819DEFFD0FFA80C3BFC5CFF0EFB86" captionStart="Figs" captionStartId="10.[151,190,1152,1171]" captionTargetBox="[152,1122,219,1134]" captionTargetId="figure-145@10.[151,1122,219,1136]" captionTargetPageId="10" captionText="Figs. 36–41. Legs of Horelophus walkeri Orchymont, 1913. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor." figureDoi="http://doi.org/10.5281/zenodo.5329859" httpUri="https://zenodo.org/record/5329859/files/figure.png" pageId="11" pageNumber="140">Figs. 39–41</figureCitation>
); claws rather large, arcuate, bearing a subbasal tooth, shape of claws the same in all three pairs of legs and in both sexes; empodium moderately large, with rather massive and sculptured unguitractor plate concealed within distal part of tarsomere 
<quantity id="4CEBA8BEFFD0FFA80B56FBFCFC39FBC1" box="[870,911,1115,1139]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="11" pageNumber="140" unit="in" value="5.0">5 in</quantity>
relaxed position (
<figureCitation id="132819DEFFD0FFA80849FBDCFF4CFB26" box="[121,250,1147,1172]" captionStart="Figs" captionStartId="10.[151,190,1152,1171]" captionTargetBox="[152,1122,219,1134]" captionTargetId="figure-145@10.[151,1122,219,1136]" captionTargetPageId="10" captionText="Figs. 36–41. Legs of Horelophus walkeri Orchymont, 1913. 36 – male, apex of protibia and basal part of protarsus; 37 – male, mesotarsomeres 1–3; 38 – female, mesotarsomeres 1–3; 39 – male, metatarsomere 5 and pretarsus in relaxed condition; 40 – male, metatarsomeres 1–3; 41 – male, metatarsomere 5 and pretarsus with exposed unguitractor." figureDoi="http://doi.org/10.5281/zenodo.5329859" httpUri="https://zenodo.org/record/5329859/files/figure.png" pageId="11" pageNumber="140">Figs. 39, 41</figureCitation>
), exposed portion of empodium bearing a pair of stout subapical setae.
</paragraph>
<paragraph id="8BAC055BFFD0FFA808A0FB3CFD0DFA61" blockId="11.[113,1089,219,1619]" pageId="11" pageNumber="140">
<emphasis id="B967D949FFD0FFA808A0FB3CFF49FB01" bold="true" box="[144,255,1179,1203]" pageId="11" pageNumber="140">Abdomen</emphasis>
(
<figureCitation id="132819DEFFD0FFA80920FB3CFE22FB01" box="[272,404,1179,1203]" captionStart="Figs" captionStartId="9.[113,152,1487,1506]" captionTargetBox="[113,1084,219,1443]" captionTargetId="figure-12@9.[113,1085,219,1443]" captionTargetPageId="9" captionText="Figs. 25–35. Morphology of Horelophus walkeri Orchymont, 1913. 25 – meso- and metathorax, ventral view; 26 – metanotum; 27 – elytral margin, dorsal view; 29 – elytron, dorsal view; 30 – detail of elytral punctation; 31 – meso- and metafurca, lateral view (part of mesoventrite broken off); 32 – mesofurca, posterior view; 33 – detail of projections of laterotergite 3; 34 – abdominal laterotergites 3–4; 35 – abdomen, ventral view" figureDoi="http://doi.org/10.5281/zenodo.5329857" httpUri="https://zenodo.org/record/5329857/files/figure.png" pageId="11" pageNumber="140">Figs. 33–35</figureCitation>
) with five exposed ventrites; ventrite 1 with moderately large bare coxal grooves, remaining portion densely pubescent, median portion without longitudinal carina; ventrites 2–5 subequal in length, densely pubescence on whole surface, with a few larger setae scattered among ground pubescent in posterior portion of ventrite 3–4 and especially ventrite 5; posterior margin of ventrite 5 without median emargination or group of enlarged setae; laterotergite 3 simple, dorsal portion not divided from ventral one by a ridge, bearing an area of goose-head-shaped cuticular projections, without any kind of organized stridulatory file; laterotergites 4–6 step-like, subdivided into elevated ventral and depressed dorsal portion, ventral portion with cuticular projections similar to laterotergite 3; tergites weakly sclerotized, densely pubescent posterolaterally.
</paragraph>
<paragraph id="8BAC055BFFD0FFAF08A0FA7CFE68F9E1" blockId="11.[113,1089,219,1619]" lastBlockId="12.[151,1127,1435,1619]" lastPageId="12" lastPageNumber="141" pageId="11" pageNumber="140">
<emphasis id="B967D949FFD0FFA808A0FA7CFE1CFA41" bold="true" box="[144,426,1498,1523]" pageId="11" pageNumber="140">
Genitalia. 
<emphasis id="B967D949FFD0FFA80939FA7DFE1CFA41" bold="true" box="[265,426,1498,1523]" italics="true" pageId="11" pageNumber="140">Male genitalia</emphasis>
</emphasis>
(
<figureCitation id="132819DEFFD0FFA8098CFA7CFDF7FA46" box="[444,577,1499,1524]" captionStart="Figs" captionStartId="12.[151,190,1307,1326]" captionTargetBox="[151,1110,224,1288]" captionTargetId="figure-78@12.[148,1123,216,1300]" captionTargetPageId="12" captionText="Figs. 42–50. Morphology of Horelophus walkeri Orchymont, 1913. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing." figureDoi="http://doi.org/10.5281/zenodo.5329861" httpUri="https://zenodo.org/record/5329861/files/figure.png" pageId="11" pageNumber="140">Figs. 47–49</figureCitation>
). Aedeagus of simply trilobed 
<typeStatus id="54A8BBF9FFD0FFA80BA4FA7BFC73FA46" box="[916,965,1500,1524]" pageId="11" pageNumber="140">type</typeStatus>
; parameres short, ca. 0.4× as long as phallobase, wide basally, arcuately narrowing to apical fifth on outer margin, mesal margin nearly straight, apices rounded, bent laterad, whole paramere bearing numerous pore-like sensilla; median lobe ca. 1.5× as long as parameres, subtriangular apically, with rather long straight apodemes reaching into phallobase, apex narrowly rounded, gonoporus apical; phallobase with extremely enlarged symmetrical manubrium ca. 0.8× as long as main portion of phallobase. Sternite 9 widely tongue-like, with very short subbasal lateral struts. Sternite 8 crescent-like, finely serrate on posterior margin, with low and wide anterior projection. 
<emphasis id="B967D949FFD7FFAF09FEF9BDFD31F981" bold="true" box="[462,647,1562,1587]" italics="true" pageId="12" pageNumber="141">Female genitalia</emphasis>
examined only externally, with long peg-like gonoxocites 9 and gonostyli 9.
</paragraph>
<caption id="DF6C55D3FFD7FFAF08A7FABCFE3EFAD1" ID-DOI="http://doi.org/10.5281/zenodo.5329861" ID-Zenodo-Dep="5329861" httpUri="https://zenodo.org/record/5329861/files/figure.png" pageId="12" pageNumber="141" startId="12.[151,190,1307,1326]" targetBox="[151,1110,224,1288]" targetPageId="12">
<paragraph id="8BAC055BFFD7FFAF08A7FABCFE3EFAD1" blockId="12.[151,1127,1307,1379]" pageId="12" pageNumber="141">
Figs. 42–50. Morphology of 
<taxonomicName id="4C137ED8FFD7FFAF09A0FABBFD6FFA9C" authority="Orchymont, 1913" authorityName="Orchymont" authorityYear="1913" box="[400,729,1307,1326]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD7FFAF09A0FABBFD8CFA9C" box="[400,570,1308,1326]" italics="true" pageId="12" pageNumber="141">Horelophus walkeri</emphasis>
<bibRefCitation id="EF8278AAFFD7FFAF0A0FFABCFD6FFA9C" author="ORCHYMONT A." box="[575,729,1307,1326]" pageId="12" pageNumber="141" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">Orchymont, 1913</bibRefCitation>
</taxonomicName>
. 42–44 – legs, ventral view (42 – anterior leg; 43 – middle leg; 44 – posterior leg); 45 – metafurca; 46 – scutellum; 47 – male sternite 8; 48 – male sternite 9; 49 – aedeagus; 50 – hind wing.
</paragraph>
</caption>
<paragraph id="8BAC055BFFD6FFAE0841FF7CFE3CFDA1" blockId="13.[113,1090,219,1588]" pageId="13" pageNumber="142">
<emphasis id="B967D949FFD6FFAE0841FF7CFF52FF41" bold="true" box="[113,228,219,243]" pageId="13" pageNumber="142">Variation.</emphasis>
The species is rather constant in all characters examined, with slight variation observed only in the shape of lateral margins of parameres which may be nearly subangular instead of rounded in some specimens. Specimens from Deepwater Creek and Dead Horse Creek exhibit both subangular and circular lateral portions of parameres (and even intermediate states) and we failed to find any other character by which the specimens would differ from each other, for which reason we consider this variation as intraspecific. Moreover, the variation may be possibly partly an artifact of the preparation of the aedeagus, i.e., the shape of the parameres may partly depend on the precise orientation of the aedeagus on the slide. Slight variation was also found in the precise branching of wing veins CuA 
<subScript id="1797071EFFD6FFAE0B75FE4CFCFBFE48" attach="left" box="[837,845,491,506]" fontSize="6" pageId="13" pageNumber="142">2</subScript>
and MP 
<subScript id="1797071EFFD6FFAE0B96FE4CFC18FE48" attach="left" box="[934,942,491,506]" fontSize="6" pageId="13" pageNumber="142">4</subScript>
as mentioned directly in the description.
</paragraph>
</subSubSection>
<subSubSection id="C30956D0FFD6FFAD0841FDBCFDB3FAA6" lastPageId="14" lastPageNumber="143" pageId="13" pageNumber="142" type="biology_ecology">
<paragraph id="8BAC055BFFD6FFAE0841FDBCFBA4FA81" blockId="13.[113,1090,219,1588]" pageId="13" pageNumber="142">
<emphasis id="B967D949FFD6FFAE0841FDBCFF7AFD81" bold="true" box="[113,204,539,563]" pageId="13" pageNumber="142">Biology.</emphasis>
Longer series of 
<taxonomicName id="4C137ED8FFD6FFAE09BFFDBAFDB7FD86" authorityName="Orchymont" authorityYear="1913" box="[399,513,541,564]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD6FFAE09BFFDBAFDB7FD86" box="[399,513,541,564]" italics="true" pageId="13" pageNumber="142">H. walkeri</emphasis>
</taxonomicName>
were first collected in 1984 at Pelorus Bridge by A. Newton &amp; M. Thayer on wet rock in the spray zone of an exposed waterfall (
<figureCitation id="132819DEFFD6FFAE0B8AFD9CFBBCFDE6" box="[954,1034,571,596]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Fig. 10</figureCitation>
), but no specimens were found later at this site during visits in late 
<date id="FFAD239BFFD6FFAE0B3EFDFCFC0CFDC1" box="[782,954,603,627]" pageId="13" pageNumber="142" value="2005-11">November 2005</date>
(by Newton &amp; Thayer) and in 
<date id="FFAD239BFFD6FFAE091FFDDCFE61FD26" box="[303,471,635,660]" pageId="13" pageNumber="142" value="2010-12">December 2010</date>
(by Leschen &amp; Fikáček), possibly because the waterfall is much less sun-exposed at present than it was in 1984. In 2010, we found multiple specimens on sun-exposed wet rocks at sides of streams at Dead Horse Creek (
<collectingRegion id="49D7CBB9FFD6FFAE0B0DFD1CFC7BFD61" box="[829,973,699,723]" country="New Zealand" name="Marlborough" pageId="13" pageNumber="142">Marlborough</collectingRegion>
; 
<figureCitation id="132819DEFFD6FFAE0BE8FD1CFB8BFD61" box="[984,1085,699,723]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Figs. 5–6</figureCitation>
, rocks with algal film) and three localities at Lyell walkway (
<collectingRegion id="49D7CBB9FFD6FFAE0AD8FD7CFC81FD41" box="[744,823,731,755]" country="New Zealand" name="Nelson" pageId="13" pageNumber="142">Nelson</collectingRegion>
; 
<figureCitation id="132819DEFFD6FFAE0B72FD7CFC10FD46" box="[834,934,731,756]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Figs. 7–9</figureCitation>
, rocks without algae), whereas single specimens were collected on a wet algae-covered stone on side of the Halfway Creek or just beyond the spray zone of an unnamed water fall at Lyell walkway. On two consecutive days in 2012 individuals were observed at Nuggety Creek (
<collectingRegion id="49D7CBB9FFD6FFAE0B49FC9CFC70FCE1" box="[889,966,827,851]" country="New Zealand" name="Nelson" pageId="13" pageNumber="142">Nelson</collectingRegion>
) at mid-day on exposed smooth rock surfaces, much like that shown in 
<figureCitation id="132819DEFFD6FFAE0AE6FCFCFCA2FCC1" box="[726,788,859,883]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Fig. 6</figureCitation>
, but with less moss and in an area of high scouring. Individuals hide in wet moss (
<figureCitation id="132819DEFFD6FFAE0A93FCDCFD52FC26" box="[675,740,891,916]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Fig. 9</figureCitation>
), damp rock cracks and crevices (
<figureCitation id="132819DEFFD6FFAE0849FC3CFF70FC06" box="[121,198,923,948]" captionStart="Figs" captionStartId="3.[113,152,1491,1510]" captionTargetBox="[141,1061,219,1473]" captionTargetId="figure-12@3.[140,1062,219,1473]" captionTargetPageId="3" captionText="Figs.5–10.Habitats of Horelophus walkeri Orchymont, 1913. 5–6 – Marlborough, Dead Horse Creek, 30.xi.2010 (5 – R. Leschen collecting specimens walking on rock steps; 6 – general view of the exposed part of Dead Horse Creek where H. walkeri occurred); 7–9 – Nelson, Deepwater Creek, 2.xii.2010 (7 – general view of the small streamlet on which side the specimens were found, see black arrows; 8–9 – details of the microhabitats where H. walkeri occurred); 10 – Marlborough, Pelorus Bridge, 28.xii.1984, A. Newton collecting H. walkeri at the side of the waterfall. Photos by M. Fikáček (5–9) and M. Thayer (10)." figureDoi="http://doi.org/10.5281/zenodo.5329849" httpUri="https://zenodo.org/record/5329849/files/figure.png" pageId="13" pageNumber="142">Fig. 10</figureCitation>
), or along shaded margins of still water and can be extracted from field-collected moss scrapings or collected on rock surfaces sprayed with pyrethrum. Individuals were usually found walking on the surface of the wet rock (
<figureCitation id="132819DEFFD6FFAE0A29FC7CFDC8FC46" box="[537,638,987,1012]" captionStart="Figs" captionStartId="2.[151,190,1538,1557]" captionTargetBox="[152,1125,232,1517]" captionTargetId="figure-10@2.[151,1125,215,1519]" captionTargetPageId="2" captionText="Figs. 1–4. Horelophus walkeri Orchymont, 1913.1–2 – general habitus in dorsal and lateral view (photo M.Fikáček); 3–4 – individuals on the wet rock surfaces at Dead Horse Creek, Marlborough (photo R. Leschen)." figureDoi="http://doi.org/10.5281/zenodo.5329845" httpUri="https://zenodo.org/record/5329845/files/figure.png" pageId="13" pageNumber="142">Figs. 3–4</figureCitation>
, see also the video mentioned below), and were never seen closely attached to the rock and therefore submerged in the surface water film as is normally the case for other hygropetric hydrophilids (e.g., 
<taxonomicName id="4C137ED8FFD6FFAE0B35FBBAFBA3FB86" authority="Erichson, 1837" authorityName="Erichson" authorityYear="1837" box="[773,1045,1051,1076]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD6FFAE0B35FBBAFCC4FB86" box="[773,882,1053,1076]" italics="true" pageId="13" pageNumber="142">Laccobius</emphasis>
Erichson, 1837
</taxonomicName>
and 
<taxonomicName id="4C137ED8FFD6FFAE0841FB9AFEEAFBE6" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[113,348,1083,1108]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD6FFAE0841FB9AFF65FBE6" box="[113,211,1085,1108]" italics="true" pageId="13" pageNumber="142">Oocyclus</emphasis>
Sharp, 1882
</taxonomicName>
). Individuals observed for a period of 2.5 hours at Nuggety Creek were resting or walking along the edges of still water without entering (see the video at http://www. youtube.com/watch?v=g03QtbXUCMI), and were not generally active or observed to feed. Rowan Emberson collected a series of 
<taxonomicName id="4C137ED8FFD6FFAE0A39FB3AFDC1FB06" authorityName="Orchymont" authorityYear="1913" box="[521,631,1181,1204]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD6FFAE0A39FB3AFDC1FB06" box="[521,631,1181,1204]" italics="true" pageId="13" pageNumber="142">H. walkeri</emphasis>
</taxonomicName>
landing in the bright afternoon sunshine on a tent built in a clearing ca. 
<quantity id="4CEBA8BEFFD6FFAE09A2FB1CFE59FB61" box="[402,495,1211,1235]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="13" pageNumber="142" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
from the Owen river in 1984 (see under Material examined for details). We observed that when submerged (examined by putting live specimens in a dish of water), the specimens bear a thin film of air on their ventral surfaces, but cannot swim. Complete records of each collecting event are listed in Material examined when known.
</paragraph>
<paragraph id="8BAC055BFFD6FFAD08A0FA9CFC29FEE6" blockId="13.[113,1090,219,1588]" lastBlockId="14.[150,1127,220,1300]" lastPageId="14" lastPageNumber="143" pageId="13" pageNumber="142">
Though we did not systematically check all creeks in the 
<collectingRegion id="49D7CBB9FFD6FFAE0AD3FA9CFC87FAE1" box="[739,817,1339,1363]" country="New Zealand" name="Nelson" pageId="13" pageNumber="142">Nelson</collectingRegion>
, 
<collectingRegion id="49D7CBB9FFD6FFAE0B0AFA9CFC7CFAE1" box="[826,970,1339,1363]" country="New Zealand" name="Marlborough" pageId="13" pageNumber="142">Marlborough</collectingRegion>
, and Buller regions, all 
<taxonomicName id="4C137ED8FFD6FFAE08DFFAFAFEE9FAC6" authorityName="Orchymont" authorityYear="1913" box="[239,351,1373,1396]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD6FFAE08DFFAFAFEE9FAC6" box="[239,351,1373,1396]" italics="true" pageId="13" pageNumber="142">H. walkeri</emphasis>
</taxonomicName>
localities we visited were first-order and second-order streams in the Pelorus River (Dead Horse Creek and Pelorus Bridge), Buller River (Owen River and Lyell Creek area), and Motueka River (Nuggety Creek) catchments. Locations were with clear cool waters with high gradients flowing on grey-wackes, fine-clastic, or metasedimentary rocks, often with small pools or seams with still waters. Other streams examined in 
<collectingRegion id="49D7CBB9FFD6FFAE0BC5FA7CFBF7FA41" box="[1013,1089,1499,1523]" country="New Zealand" name="Nelson" pageId="13" pageNumber="142">Nelson</collectingRegion>
with streambeds composed of sandstones, or of moderate to coarse clastics did not have 
<taxonomicName id="4C137ED8FFD6FFAE0C17FA5AFF77F986" authorityName="Orchymont" authorityYear="1913" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD6FFAE0C17FA5AFF77F986" italics="true" pageId="13" pageNumber="142">H. walkeri</emphasis>
</taxonomicName>
. Stream sections that were enclosed by a canopy did not yield specimens, and most localities were dominated by beech trees (
<taxonomicName id="4C137ED8FFD5FFAD0C2CFF7AFD16FEA6" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="143" phylum="Tracheophyta" rank="genus">
<emphasis id="B967D949FFD5FFAD0C2CFF7AFD16FEA6" italics="true" pageId="14" pageNumber="143">Nothofagus</emphasis>
</taxonomicName>
) apart from Dead Horse Creek which was surrounded by rejuvenating forest and adjacent to a 
<taxonomicName id="4C137ED8FFD5FFAD0AA4FE9AFC90FEE6" box="[660,806,317,340]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="143" phylum="Tracheophyta" rank="species" species="radiata">
<emphasis id="B967D949FFD5FFAD0AA4FE9AFC90FEE6" box="[660,806,317,340]" italics="true" pageId="14" pageNumber="143">Pinus radiata</emphasis>
</taxonomicName>
plantation.
</paragraph>
<paragraph id="8BAC055BFFD5FFAD0AB4FEFBFBD1FE06" blockId="14.[150,1127,220,1300]" pageId="14" pageNumber="143">
Summarizing all of these data, we conclude that 
<taxonomicName id="4C137ED8FFD5FFAD0AA2FEDAFCB6FE26" authorityName="Orchymont" authorityYear="1913" box="[658,768,381,404]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD0AA2FEDAFCB6FE26" box="[658,768,381,404]" italics="true" pageId="14" pageNumber="143">H. walkeri</emphasis>
</taxonomicName>
is a hygropetric species inhabiting exposed wet rocks along streams and waterfalls,
</paragraph>
<paragraph id="8BAC055BFFD5FFAD0A56FE1BFBD0FD46" blockId="14.[150,1127,220,1300]" pageId="14" pageNumber="143">
and is most common in areas with extensive smooth surfaces. The specimens seem to hide in moss and rock crevices normally, and move to the rock surface probably for feeding on algae (but guts that we dissected were empty). The polarized light reflected from the wet rock surface may be used for finding suitable habitats (as it is in many other hydrophilids; 
<bibRefCitation id="EF8278AAFFD5FFAD0B73FD3BFBA5FD01" author="KRISKA G. &amp; CSABAI Z. &amp; BODA P. &amp; MALIK P. &amp; HORVATH G." box="[835,1043,667,692]" pageId="14" pageNumber="143" pagination="1667 - 1671" refId="ref8090" refString="KRISKA G., CSABAI Z., BODA P., MALIK P. &amp; HORVATH G. 2006: Why do red and dark-coloured cars lure aquatic insects? The attraction of water insects to car paintwork explained by reflection-polarization signals. Proceedings of the Royal Society, Series B 273: 1667 - 1671." type="journal article" year="2006">KRISKA et al. 2006</bibRefCitation>
) which might explain the finding of the specimens on the tent by R. Emberson. This would also indicate that
</paragraph>
<paragraph id="8BAC055BFFD5FFAD0A56FD5AFC03FC86" blockId="14.[150,1127,220,1300]" pageId="14" pageNumber="143">
<taxonomicName id="4C137ED8FFD5FFAD0A56FD5AFD62FCA6" authorityName="Orchymont" authorityYear="1913" box="[614,724,765,788]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD0A56FD5AFD62FCA6" box="[614,724,765,788]" italics="true" pageId="14" pageNumber="143">H. walkeri</emphasis>
</taxonomicName>
is a good flier, which is congruent with its long, well-developed wings.
</paragraph>
<paragraph id="8BAC055BFFD5FFAD0A56FC9BFDB3FAA6" blockId="14.[150,1127,220,1300]" pageId="14" pageNumber="143">
<emphasis id="B967D949FFD5FFAD0A56FC9BFC9AFCE6" bold="true" box="[614,812,828,852]" pageId="14" pageNumber="143">Immature stages.</emphasis>
Unknown. During the trip in Fig 51. Distribution of 
<taxonomicName id="4C137ED8FFD5FFAD09B6FCFCFDF0FCDF" authorityName="Orchymont" authorityYear="1913" box="[390,582,859,877]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD09B6FCFCFDF0FCDF" box="[390,582,859,877]" italics="true" pageId="14" pageNumber="143">Horelophus walkeri</emphasis>
</taxonomicName>
the beginning of 
<date id="FFAD239BFFD5FFAD0B12FCFBFC67FCC6" box="[802,977,860,884]" pageId="14" pageNumber="143" value="2010-12">December 2010</date>
, we collected 
<bibRefCitation id="EF8278AAFFD5FFAD08A7FCD2FE98FC3A" author="ORCHYMONT A." box="[151,302,885,904]" pageId="14" pageNumber="143" pagination="94 - 100" refId="ref8535" refString="ORCHYMONT A. d' 1913: Un nouvel hydrophilide de la faune antartique Neozelandaise. Annales de la Societe Entomologique de Belgique 57: 94 - 100." type="journal article" year="1913">
Orchymont, 
<quantity id="4CEBA8BEFFD5FFAD0933FCD2FE98FC3A" box="[259,302,885,904]" metricMagnitude="1" metricUnit="m" metricValue="4.85902" pageId="14" pageNumber="143" unit="in" value="1913.0">1913</quantity>
</bibRefCitation>
in 
<collectingCountry id="F30445CBFFD5FFAD0979FCD2FE09FC3A" box="[329,447,885,904]" name="New Zealand" pageId="14" pageNumber="143">New Zealand</collectingCountry>
. mosses and brushed the algal film from the microhabitats where adults were found at Dead Horse Creek (
<collectingRegion id="49D7CBB9FFD5FFAD08D4FC1BFEC2FC66" box="[228,372,956,980]" country="New Zealand" name="Marlborough" pageId="14" pageNumber="143">Marlborough</collectingRegion>
) and Deepwater Creek (
<collectingRegion id="49D7CBB9FFD5FFAD0A44FC1BFD73FC66" box="[628,709,956,980]" country="New Zealand" name="Nelson" pageId="14" pageNumber="143">Nelson</collectingRegion>
), and these samples were carefully examined later that day in the laboratory. However, the only larvae which were obtained by this way belong to 
<taxonomicName id="4C137ED8FFD5FFAD0904FC5AFD8FFBA6" authority="Broun, 1903" authorityName="Broun" authorityYear="1903" box="[308,569,1020,1044]" class="Insecta" family="Hydrophilidae" genus="Cylomissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD5FFAD0904FC5AFE1BFBA6" box="[308,429,1021,1044]" italics="true" pageId="14" pageNumber="143">Cylomissus</emphasis>
Broun, 1903
</taxonomicName>
(associated by cox 
<geoCoordinate id="EE27639CFFD5FFAD0B3BFC5BFCFEFBA1" box="[779,840,1019,1044]" degrees="1, 16" direction="south" orientation="latitude" pageId="14" pageNumber="143" precision="555" value="-1.16">1, 16S</geoCoordinate>
and 28S DNA sequences). Likewise, Newton and Thayer in 
<date id="FFAD239BFFD5FFAD09C4FBBBFD2AFB86" box="[500,668,1052,1076]" pageId="14" pageNumber="143" value="1984-12">December 1984</date>
and 
<date id="FFAD239BFFD5FFAD0AFDFBBBFCC1FB81" box="[717,887,1051,1076]" pageId="14" pageNumber="143" value="2005-11">November 2005</date>
collected and berlesed damp debris and mosses from the waterfall where the series of 
<taxonomicName id="4C137ED8FFD5FFAD0B19FB9AFC2FFBE6" authorityName="Orchymont" authorityYear="1913" box="[809,921,1085,1108]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD0B19FB9AFC2FFBE6" box="[809,921,1085,1108]" italics="true" pageId="14" pageNumber="143">H. walkeri</emphasis>
</taxonomicName>
was found in 1984, but also found only 
<taxonomicName id="4C137ED8FFD5FFAD095EFBFAFE51FBC6" authorityName="Broun" authorityYear="1903" box="[366,487,1117,1140]" class="Insecta" family="Hydrophilidae" genus="Cylomissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD5FFAD095EFBFAFE51FBC6" box="[366,487,1117,1140]" italics="true" pageId="14" pageNumber="143">Cylomissus</emphasis>
</taxonomicName>
larvae and no likely candidates for larvae of 
<taxonomicName id="4C137ED8FFD5FFAD0BFCFBFAFBFFFBC6" authorityName=": SHARP" authorityYear="1915" box="[972,1097,1117,1140]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD5FFAD0BFCFBFAFBFFFBC6" box="[972,1097,1117,1140]" italics="true" pageId="14" pageNumber="143">Horelophus</emphasis>
</taxonomicName>
. A single larva collected at Nuggety Creek also agrees with those of 
<taxonomicName id="4C137ED8FFD5FFAD0B75FBDAFC08FB26" authorityName="Broun" authorityYear="1903" box="[837,958,1149,1172]" class="Insecta" family="Hydrophilidae" genus="Cylomissus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B967D949FFD5FFAD0B75FBDAFC08FB26" box="[837,958,1149,1172]" italics="true" pageId="14" pageNumber="143">Cylomissus</emphasis>
</taxonomicName>
even though no hydrophilids other then 
<taxonomicName id="4C137ED8FFD5FFAD09A7FB3AFDDCFB06" authorityName="Orchymont" authorityYear="1913" box="[407,618,1181,1204]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD09A7FB3AFDDCFB06" box="[407,618,1181,1204]" italics="true" pageId="14" pageNumber="143">Horelophus walkeri</emphasis>
</taxonomicName>
were collected or observed at the locality. 
<emphasis id="B967D949FFD5FFAD08A7FB1CFE97FB61" bold="true" box="[151,289,1211,1235]" pageId="14" pageNumber="143">Distribution</emphasis>
(Fig. 51). 
<taxonomicName id="4C137ED8FFD5FFAD09BDFB1AFDE8FB66" authorityName="Orchymont" authorityYear="1913" box="[397,606,1213,1236]" class="Insecta" family="Hydrophilidae" genus="Horelophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="B967D949FFD5FFAD09BDFB1AFDE8FB66" box="[397,606,1213,1236]" italics="true" pageId="14" pageNumber="143">Horelophus walkeri</emphasis>
</taxonomicName>
is endemic to the northern part of the South Island of 
<collectingCountry id="F30445CBFFD5FFAD0886FB7BFEFCFB46" box="[182,330,1244,1268]" name="New Zealand" pageId="14" pageNumber="143">New Zealand</collectingCountry>
, so far known only from the districts of 
<collectingRegion id="49D7CBB9FFD5FFAD0B35FB7BFC23FB46" box="[773,917,1244,1268]" country="New Zealand" name="Marlborough" pageId="14" pageNumber="143">Marlborough</collectingRegion>
, 
<collectingRegion id="49D7CBB9FFD5FFAD0B91FB7BFC5BFB46" box="[929,1005,1244,1268]" country="New Zealand" name="Nelson" pageId="14" pageNumber="143">Nelson</collectingRegion>
and Buller west and north of the Alpine fault.
</paragraph>
</subSubSection>
</treatment>
</document>