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<document id="00739B94C3DC2E2DF1C90F15E5BC1D57" ID-DOI="10.5252/zoosystema2025v47a4" ID-ISSN="1638-9387" ID-Zenodo-Dep="14831210" ID-ZooBank="3749B35A-48E6-43F6-B889-AC340DBD5CF2" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1738917347317" checkinUser="plazi" docAuthor="Tran, Ngat Thi, Phuong, Lien Thi, Anh, Phuong Nguyen, Nguyen, Anh Duc &amp; Engel, Michael S." docDate="2025" docId="03BA0E159464FFFFFF523CBEC1AC4D2C" docLanguage="en" docName="zoosystema.2025.47.4.pdf" docOrigin="Zoosystema 47 (4)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a4.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.10:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="10" docTitle="Maculonomia (Maculonomia) rufomysteria Engel, Tran &amp; A. D. Nguyen, n. sp." docType="treatment" docVersion="1" lastPageNumber="65" masterDocId="FF83766D9468FFEFFF883A2CC32B4B6D" masterDocTitle="New species and records of the nomiine bee genus Maculonomia Wu, 1982 from Vietnam (Hymenoptera, Halictidae)" masterLastPageNumber="73" masterPageNumber="51" pageNumber="61" updateTime="1739212110286" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FFDC1E4496B4C1A3D271F2D3E916348A">New species and records of the nomiine bee genus Maculonomia Wu, 1982 from Vietnam (Hymenoptera, Halictidae)</mods:title>
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<mods:namePart id="6B9EDB1A3D57A21B675A933B7B8E418A">Tran, Ngat Thi</mods:namePart>
<mods:affiliation id="F249CEA149B859B1930D5D7F9D2A73B1">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
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<mods:namePart id="F447AEA6ADC4B18E5407511FC6204D69">Phuong, Lien Thi</mods:namePart>
<mods:affiliation id="07EDD70A0C5DBE5021BCD86146FB92E1">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="B3807166F634148A885D9FA528401EB9" type="email">phuonglientit@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CA04CEBF193308971F98F81728910BC1">Anh, Phuong Nguyen</mods:namePart>
<mods:affiliation id="7586DC826C76D807F32350BB0AFBC438">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
<mods:nameIdentifier id="087CC632E73003741FC46A662D26BB6B" type="email">phuonglientit@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2C50F00351AC616599AA81E11898D97F">Nguyen, Anh Duc</mods:namePart>
<mods:affiliation id="4CEEBAD419B32E34580E4C3D49443A69">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam) and Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi (Vietnam)</mods:affiliation>
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<mods:namePart id="37AFAAF54D99CC5B39AEF893C5E353BA">Engel, Michael S.</mods:namePart>
<mods:affiliation id="D04C5B5DE24C4F1077BC3F19972A79F0">Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, New York 10024 (United States)</mods:affiliation>
<mods:nameIdentifier id="C9920F0B60AE088FBA5C05B6A52C0F1D" type="email">mengel@amnh.org</mods:nameIdentifier>
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<mods:title id="E5CF14ED9DE2AD3EB759E3211E9380B5">Zoosystema</mods:title>
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<mods:date id="6C6329EFC8B58967775F9C1BA863B81C">2025</mods:date>
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<treatment id="03BA0E159464FFFFFF523CBEC1AC4D2C" LSID="urn:lsid:plazi:treatment:03BA0E159464FFFFFF523CBEC1AC4D2C" httpUri="http://treatment.plazi.org/id/03BA0E159464FFFFFF523CBEC1AC4D2C" lastPageId="16" lastPageNumber="65" pageId="12" pageNumber="61">
<subSubSection id="C309EC889464FFE3FF523CBEC1AF4DA0" pageId="12" pageNumber="61" type="nomenclature">
<paragraph id="8BACBF039464FFE3FF523CBEC1AF4DA0" blockId="12.[218,689,1682,1772]" pageId="12" pageNumber="61">
<heading id="D0E4086F9464FFE3FF523CBEC1AF4DA0" centered="true" fontSize="11" level="2" pageId="12" pageNumber="61" reason="3">
<taxonomicName id="4C13C4809464FFE3FF523CBEC1144DA0" authority="Engel, Tran &amp; A.D. Nguyen" authorityName="Engel, Tran &amp; A. D. Nguyen" class="Insecta" family="Halictidae" genus="Maculonomia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufomysteria" subGenus="Maculonomia">
<emphasis id="B96763119464FFE3FF523CBEC25A4DC0" bold="true" box="[218,369,1682,1709]" italics="true" pageId="12" pageNumber="61">Maculonomia</emphasis>
(
<emphasis id="B96763119464FFE3FE093CBEC1334DC0" bold="true" box="[385,536,1682,1709]" italics="true" pageId="12" pageNumber="61">Maculonomia</emphasis>
)
<emphasis id="B96763119464FFE3FDA13CBEC19A4DC1" bold="true" box="[553,689,1682,1708]" italics="true" pageId="12" pageNumber="61">rufomysteria</emphasis>
Engel, Tran &amp; A.D. Nguyen
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,
<taxonomicNameLabel id="A254DE6A9464FFE3FDC13C9FC1AF4DA0" box="[585,644,1715,1741]" pageId="12" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C309EC889464FFE3FE023CFEC1C64C4D" pageId="12" pageNumber="61" type="description">
<paragraph id="8BACBF039464FFE3FE023CFEC12A4D81" blockId="12.[218,689,1682,1772]" box="[394,513,1746,1772]" pageId="12" pageNumber="61">
(
<figureCitation id="1328A3869464FFE3FE193CFEC2D34D81" box="[401,504,1746,1772]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="13.[141,152,2007,2025]" captionStartId-1="14.[132,143,1592,1610]" captionStartId-2="15.[132,143,1466,1484]" captionTargetBox-0="[281,1306,214,1933]" captionTargetBox-1="[231,1356,215,1536]" captionTargetBox-2="[256,1342,216,1391]" captionTargetId-0="figure-16@13.[262,1325,213,1954]" captionTargetId-1="figure-182@14.[231,1356,215,1536]" captionTargetId-2="figure-250@15.[246,1342,216,1406]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIG. 8. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; A.D. Nguyen, n. sp., holotype male: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." captionText-1="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." captionText-2="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14831232" figureDoi-1="http://doi.org/10.5281/zenodo.14831234" figureDoi-2="http://doi.org/10.5281/zenodo.14831236" httpUri-0="https://zenodo.org/record/14831232/files/figure.png" httpUri-1="https://zenodo.org/record/14831234/files/figure.png" httpUri-2="https://zenodo.org/record/14831236/files/figure.png" pageId="12" pageNumber="61">Figs 8-10</figureCitation>
)
</paragraph>
<paragraph id="8BACBF039464FFE3FF173D22C1C64C4D" blockId="12.[159,749,1806,1824]" box="[159,749,1806,1824]" pageId="12" pageNumber="61">
<uri id="FF82B3019464FFE3FF173D22C1C64C4D" box="[159,749,1806,1824]" pageId="12" pageNumber="61">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C309EC889464FFE3FF0C3D67C11D4C11" pageId="12" pageNumber="61" type="materials_examined">
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<collectionCode id="ED0227C69464FFE3FF0C3D67C39E4C0D" box="[132,181,1867,1890]" pageId="12" pageNumber="61">TYPE</collectionCode>
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. —
<materialsCitation id="3B7BB55E9464FFE3FEC13D66C11D4C11" collectingDate="2019-09-24" collectionCode="IEBR" collectorName="Pham N. T." country="Vietnam" location="Bidoup Nui Ba" municipality="Bidoup Nui Ba" pageId="12" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="holotype">
<emphasis id="B96763119464FFE3FEC13D66C2834C0F" bold="true" box="[329,424,1866,1890]" pageId="12" pageNumber="61">
<typeStatus id="54A801A19464FFE3FEC13D66C2834C0F" box="[329,424,1866,1890]" pageId="12" pageNumber="61" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B96763119464FFE3FE3D3D60C1204C09" bold="true" box="[437,523,1868,1892]" pageId="12" pageNumber="61">
<collectingCountry id="F304FF939464FFE3FE3D3D60C1204C09" box="[437,523,1868,1892]" name="Vietnam" pageId="12" pageNumber="61">Vietnam</collectingCountry>
</emphasis>
•
<specimenCount id="9D15748A9464FFE3FDAB3D61C11C4C0C" box="[547,567,1869,1889]" count="1" pageId="12" pageNumber="61" type="male">♂</specimenCount>
;
<collectingRegion id="49D771E19464FFE3FDCD3D67C1984C0E" box="[581,691,1867,1891]" country="Vietnam" name="Lam Dong" pageId="12" pageNumber="61">Lam Dong</collectingRegion>
,
<collectingMunicipality id="6BC825799464FFE3FD363D66C3E64C11" pageId="12" pageNumber="61">Bidoup Nui Ba</collectingMunicipality>
;
<date id="FFAD99C39464FFE3FF5F3D49C2604C10" box="[215,331,1893,1917]" pageId="12" pageNumber="61" value="2019-09-24">
<collectingDate id="EFE9602B9464FFE3FF5F3D49C2604C10" box="[215,331,1893,1917]" pageId="12" pageNumber="61" value="2019-09-24">24.IX.2019</collectingDate>
</date>
;
<collectorName id="26E6DAD59464FFE3FEDD3D49C2E84C11" box="[341,451,1893,1917]" pageId="12" pageNumber="61">Pham N.T.</collectorName>
leg.;
<collectionCode id="ED0227C69464FFE3FE703D49C1194C11" box="[504,562,1893,1916]" pageId="12" pageNumber="61">IEBR</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309EC889464FFE3FF0C3DB2C2224C87" pageId="12" pageNumber="61" type="etymology">
<paragraph id="8BACBF039464FFE3FF0C3DB2C2224C87" blockId="12.[132,775,1949,2026]" pageId="12" pageNumber="61">
<smallCapsWord id="8D4A29DF9464FFE3FF0C3DB2C3D54CD9" baselines="1967,1968" box="[132,254,1950,1973]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="12" pageNumber="61">ETYMOLOGY</smallCapsWord>
. — The specific epithet is a combination of the Latin adjectives
<emphasis id="B96763119464FFE3FF663D94C2344CBD" box="[238,287,1976,2000]" italics="true" pageId="12" pageNumber="61">rūfus</emphasis>
, meaning, a “reddish”, and
<emphasis id="B96763119464FFE3FDC93D95C1B24CBD" box="[577,665,1977,2000]" italics="true" pageId="12" pageNumber="61">mystērius</emphasis>
, meaning, “mysterious”.
</paragraph>
</subSubSection>
<subSubSection id="C309EC889464FFE1FCA53F95C2C44C86" lastPageId="14" lastPageNumber="63" pageId="12" pageNumber="61" type="diagnosis">
<paragraph id="8BACBF039464FFE1FCA53F95C2C44C86" blockId="12.[811,1457,1465,2027]" lastBlockId="14.[131,777,1712,2027]" lastPageId="14" lastPageNumber="63" pageId="12" pageNumber="61">
<smallCapsWord id="8D4A29DF9464FFE3FCA53F95C0B14EA2" baselines="1482,1483" box="[813,922,1465,1488]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="12" pageNumber="61">DIAGNOSIS</smallCapsWord>
. — This species superficially resembles
<taxonomicName id="4C13C4809464FFE3FA933F96C69B4EBC" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[1307,1456,1465,1489]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FA933F96C69B4EBC" box="[1307,1456,1465,1489]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
owing to the overall black coloration combined with abundant reddish orange setae on the apical metasomal terga, as well as the presence of enamel apical bands. Unlike
<taxonomicName id="4C13C4809464FFE3FB3E3C26C6604D4C" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[1206,1355,1545,1569]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FB3E3C26C6604D4C" box="[1206,1355,1545,1569]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
, however, the reddish orange setae are restricted to terga
<collectionCode id="ED0227C69464FFE3FA803C09C6324D51" box="[1288,1305,1573,1596]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="61" type="Museum">V</collectionCode>
and VI, while in
<taxonomicName id="4C13C4809464FFE3FCCF3C6CC0F44D3A" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[839,991,1599,1623]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FCCF3C6CC0F44D3A" box="[839,991,1599,1623]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
the color is extensive across terga III-V. Additionally, in the new species the orange, apical, enamel bands are complete on terga
<collectionCode id="ED0227C69464FFE3FC633C5AC0D74DE0" box="[1003,1020,1654,1677]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="61" type="Museum">V</collectionCode>
and VI, and only present as small spots at the extreme lateral portions of tergum IV, rather than the complete bands found in
<taxonomicName id="4C13C4809464FFE3FB843C80C78C4DAE" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[1036,1191,1707,1731]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FB843C80C78C4DAE" box="[1036,1191,1707,1731]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
on terga III-V. In
<taxonomicName id="4C13C4809464FFE3FAE33C80C0A74DB3" baseAuthorityName="Wu" baseAuthorityYear="1988" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FAE33C80C0A74DB3" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
the wing is uniformly yellowed to yellowish brown, often a bit lighter apically beyond the veins. In the new species the forewing is yellow, then paler just beyond the veins before forming a large, apical, infumate spot, resembling somewhat the apical spot of
<taxonomicName id="4C13C4809464FFE3FC363D1EC7684C24" authorityName="Smith" authorityYear="1875" box="[958,1091,1842,1865]" class="Insecta" family="Halictidae" genus="Nomia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="B96763119464FFE3FC363D1EC7684C24" box="[958,1091,1842,1865]" italics="true" pageId="12" pageNumber="61">M. terminata</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C13C4809464FFE3FBF83D1EC6224C24" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[1136,1289,1841,1865]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FBF83D1EC6224C24" box="[1136,1289,1841,1865]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
the mesoscutellum is unmodified or with weak paramedial callosities, while in the new species there are prominent elevated tubercles on either side of the midline. Naturally, given that the male of
<taxonomicName id="4C13C4809464FFE3FAD13DAFC0514CD8" baseAuthorityName="Wu" baseAuthorityYear="1988" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FAD13DAFC0514CD8" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
is currently unknown it was tempting to consider this male as conspecific with the aforementioned females attributed to
<taxonomicName id="4C13C4809464FFE3FCA33DF8C0E84C86" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[811,963,2003,2027]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119464FFE3FCA33DF8C0E84C86" box="[811,963,2003,2027]" italics="true" pageId="12" pageNumber="61">M. rufocaudata</emphasis>
</taxonomicName>
. We have erred on the side of splitting, however, given the rather prominent habitat differences:
<taxonomicName id="4C13C4809466FFE1FDE53C9DC02D4DA5" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[621,774,1712,1736]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119466FFE1FDE53C9DC02D4DA5" box="[621,774,1712,1736]" italics="true" pageId="14" pageNumber="63">M. rufocaudata</emphasis>
</taxonomicName>
from mountains in the North of
<collectingCountry id="F304FF939466FFE1FE5C3CE6C1004D8F" box="[468,555,1738,1762]" name="Vietnam" pageId="14" pageNumber="63">Vietnam</collectingCountry>
and Tibet, versus the more isolated mountain of Bidoup National Park in Central Highlands
<collectingCountry id="F304FF939466FFE1FF7D3CD3C27B4C7A" box="[245,336,1791,1815]" name="Vietnam" pageId="14" pageNumber="63">Vietnam</collectingCountry>
. Moreover, the pattern of wing coloration is quite divergent between the two. Additionally, the punctures of tergum II are smaller and separated by about a puncture width in
<taxonomicName id="4C13C4809466FFE1FF283D63C2124C0B" baseAuthorityName="Wu" baseAuthorityYear="1988" box="[160,313,1870,1894]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="rufocaudata">
<emphasis id="B96763119466FFE1FF283D63C2124C0B" box="[160,313,1870,1894]" italics="true" pageId="14" pageNumber="63">M. rufocaudata</emphasis>
</taxonomicName>
versus the slightly larger punctures distinctly separated by much less than a puncture width in the male described here. Thus, pending the discovery of a male from Tibet or the mountainous populations of northern
<collectingCountry id="F304FF939466FFE1FDCA3DB2C1B24CDB" box="[578,665,1950,1974]" name="Vietnam" pageId="14" pageNumber="63">Vietnam</collectingCountry>
that might indicate otherwise, we consider this Central Highlands male as distinct but obviously closely allied.
</paragraph>
</subSubSection>
<caption id="DF6CEF8B9465FFE2FF053DFBC6834C84" ID-DOI="http://doi.org/10.5281/zenodo.14831232" ID-Zenodo-Dep="14831232" box="[141,1448,2007,2025]" httpUri="https://zenodo.org/record/14831232/files/figure.png" pageId="13" pageNumber="62" startId="13.[141,152,2007,2025]" targetBox="[281,1306,214,1933]" targetPageId="13" targetType="figure">
<paragraph id="8BACBF039465FFE2FF053DFBC6834C84" blockId="13.[141,1448,2007,2025]" box="[141,1448,2007,2025]" pageId="13" pageNumber="62">
<smallCapsWord id="8D4A29DF9465FFE2FF053DFBC38D4C85" baselines="2020,2021" box="[141,166,2007,2025]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="13" pageNumber="62">FIG</smallCapsWord>
. 8. —
<taxonomicName id="4C13C4809465FFE2FF553DFBC1444C84" authority="Engel" authorityName="Engel" box="[221,623,2007,2025]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="rufomysteria" subGenus="Maculonomia">
<emphasis id="B96763119465FFE2FF553DFBC2664C84" box="[221,333,2007,2025]" italics="true" pageId="13" pageNumber="62">Maculonomia</emphasis>
(
<emphasis id="B96763119465FFE2FEDF3DFBC2EC4C84" box="[343,455,2007,2025]" italics="true" pageId="13" pageNumber="62">Maculonomia</emphasis>
)
<emphasis id="B96763119465FFE2FE593DFBC1134C84" box="[465,568,2007,2025]" italics="true" pageId="13" pageNumber="62">rufomysteria</emphasis>
Engel
</taxonomicName>
, Tran, &amp; A.D. Nguyen,
<taxonomicNameLabel id="A254DE6A9465FFE2FCA03DFBC0704C84" box="[808,859,2007,2025]" pageId="13" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, holotype male:
<emphasis id="B96763119465FFE2FC693DFBC0C54C84" bold="true" box="[993,1006,2007,2025]" pageId="13" pageNumber="62">A</emphasis>
, lateral habitus;
<emphasis id="B96763119465FFE2FBFF3DFBC7AF4C84" bold="true" box="[1143,1156,2007,2025]" pageId="13" pageNumber="62">B</emphasis>
, dorsal habitus. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF6CEF8B9466FFE1FF0C3C14C7234D0C" ID-DOI="http://doi.org/10.5281/zenodo.14831234" ID-Zenodo-Dep="14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="14" pageNumber="63" startId="14.[132,143,1592,1610]" targetBox="[231,1356,215,1536]" targetPageId="14" targetType="figure">
<paragraph id="8BACBF039466FFE1FF0C3C14C7234D0C" blockId="14.[132,1455,1592,1633]" pageId="14" pageNumber="63">
<smallCapsWord id="8D4A29DF9466FFE1FF0C3C14C3B64D24" baselines="1605,1606" box="[132,157,1592,1610]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="14" pageNumber="63">FIG</smallCapsWord>
. 9. —
<taxonomicName id="4C13C4809466FFE1FF5E3C14C1424D27" authority="Engel" authorityName="Engel" box="[214,617,1592,1610]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="rufomysteria" subGenus="Maculonomia">
<emphasis id="B96763119466FFE1FF5E3C14C26D4D27" box="[214,326,1592,1610]" italics="true" pageId="14" pageNumber="63">Maculonomia</emphasis>
(
<emphasis id="B96763119466FFE1FED83C14C2EB4D27" box="[336,448,1592,1610]" italics="true" pageId="14" pageNumber="63">Maculonomia</emphasis>
)
<emphasis id="B96763119466FFE1FE433C14C1194D27" box="[459,562,1592,1610]" italics="true" pageId="14" pageNumber="63">rufomysteria</emphasis>
Engel
</taxonomicName>
, Tran, &amp; AD Nguyen,
<taxonomicNameLabel id="A254DE6A9466FFE1FC943C14C0654D27" box="[796,846,1592,1610]" pageId="14" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
, holotype male:
<emphasis id="B96763119466FFE1FC5F3C14C0CF4D27" bold="true" box="[983,996,1592,1610]" pageId="14" pageNumber="63">A</emphasis>
, facial view;
<emphasis id="B96763119466FFE1FBD93C14C7754D27" bold="true" box="[1105,1118,1592,1610]" pageId="14" pageNumber="63">B</emphasis>
, mesosoma in dorsal view;
<emphasis id="B96763119466FFE1FAC13C14C67C4D27" bold="true" box="[1353,1367,1592,1610]" pageId="14" pageNumber="63">C</emphasis>
, metatibia showing the basal plate;
<emphasis id="B96763119466FFE1FEDC3C63C2494D0C" bold="true" box="[340,354,1615,1633]" pageId="14" pageNumber="63">D</emphasis>
, metasoma in dorsal view;
<emphasis id="B96763119466FFE1FDCE3C63C1794D0C" bold="true" box="[582,594,1615,1633]" pageId="14" pageNumber="63">E</emphasis>
, metasomal T5-T7 in dorsal view. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C309EC889466FFFFFCA53C9FC1AC4D2C" lastPageId="16" lastPageNumber="65" pageId="14" pageNumber="63" type="description">
<paragraph id="8BACBF039466FFE1FCA53C9FC0964DA6" blockId="14.[811,1455,1715,1836]" box="[813,957,1715,1741]" pageId="14" pageNumber="63">
<smallCapsWord id="8D4A29DF9466FFE1FCA53C9FC0964DA6" baselines="1734,1734" box="[813,957,1715,1741]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="14" pageNumber="63">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BACBF039466FFE1FCA53CFEC0494D81" blockId="14.[811,1455,1715,1836]" box="[813,866,1746,1772]" pageId="14" pageNumber="63">
<emphasis id="B96763119466FFE1FCA53CFEC0494D81" box="[813,866,1746,1772]" italics="true" pageId="14" pageNumber="63">Male</emphasis>
</paragraph>
<paragraph id="8BACBF039466FFE1FCA53CDEC0BD4C41" blockId="14.[811,1455,1715,1836]" pageId="14" pageNumber="63">
<emphasis id="B96763119466FFE1FCA53CDEC0FF4C61" bold="true" box="[813,980,1778,1804]" pageId="14" pageNumber="63">Measurements.</emphasis>
Total body length
<quantity id="4CEB12E69466FFE1FB113CDEC7D54C61" box="[1177,1278,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="14" pageNumber="63" unit="mm" value="13.5">13.5 mm</quantity>
, forewing length
<quantity id="4CEB12E69466FFE1FCA33D3DC0B84C41" box="[811,915,1809,1836]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="63" unit="mm" value="11.5">11.5 mm</quantity>
.
</paragraph>
<paragraph id="8BACBF039466FFE0FCA53D7DC7674C86" blockId="14.[811,1458,1873,2028]" lastBlockId="15.[811,1457,1586,2028]" lastPageId="15" lastPageNumber="64" pageId="14" pageNumber="63">
<emphasis id="B96763119466FFE1FCA53D7DC0B54C06" bold="true" box="[813,926,1873,1899]" pageId="14" pageNumber="63">Structure.</emphasis>
Head broader than long, approximately 1.2× as broad as long (
<figureCitation id="1328A3869466FFE1FC5F3D5DC7144CE6" box="[983,1087,1905,1931]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="14" pageNumber="63">Fig. 10A</figureCitation>
); compound eyes approximately 1.4× genal width; mandible simple; clypeus slightly convex laterally, approximately 1.3× as broad as long; supraclypeal area slightly convex (
<figureCitation id="1328A3869466FFE1FB8C3DFDC77C4C86" box="[1028,1111,2001,2027]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="14" pageNumber="63">Fig. 9A</figureCitation>
). Mesosoma approximately 1.2× as long as broad; mesoscutellum with mediolongitudinal depression apically and strongly bigibbous (
<figureCitation id="1328A3869467FFE0FDB53C7EC1BB4D01" box="[573,656,1618,1644]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="15" pageNumber="64">Fig. 9B</figureCitation>
). Forewing with three submarginal cells, first and third submarginal cells broader than second submarginal cell, 1m-cu entering near apex of second marginal cell (
<figureCitation id="1328A3869467FFE0FE403C9EC1354DA1" box="[456,542,1714,1740]" captionStart="FIG" captionStartId="13.[141,152,2007,2025]" captionTargetBox="[281,1306,214,1933]" captionTargetId="figure-16@13.[262,1325,213,1954]" captionTargetPageId="13" captionText="FIG. 8. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; A.D. Nguyen, n. sp., holotype male: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831232" httpUri="https://zenodo.org/record/14831232/files/figure.png" pageId="15" pageNumber="64">Fig. 8A</figureCitation>
). Metafemur slightly swollen (
<figureCitation id="1328A3869467FFE0FF613CFEC2684D81" box="[233,323,1746,1772]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="15" pageNumber="64">Fig. 9C</figureCitation>
); metatibia gradually expanding toward apex and inner surface with slightly raised knob at one-third length from base. Metasomal S4 with deeply, narrowly emarginate medioapically, margin relatively straight and sloping to middle on either side of emargination (
<figureCitation id="1328A3869467FFE0FD123D7DC1D64C01" box="[666,765,1873,1900]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10A</figureCitation>
). S5 with an isosceles-triangle-shaped process medio-apically, and long thin processes on either side of midline bearing abundant setae (
<figureCitation id="1328A3869467FFE0FECA3D9DC2814CA6" box="[322,426,1969,1995]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10B</figureCitation>
). S6 apical margin emarginate medially, emargination shallow and comparatively narrow, apical margin as in
<figureCitation id="1328A3869467FFE0FB803C1EC78D4D20" box="[1032,1190,1586,1613]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Figure 10C. S</figureCitation>
<figureCitation id="1328A3869467FFE0FB2D3C1FC7984D20" box="[1189,1203,1587,1613]" captionStart="FIG" captionStartId="12.[132,143,1362,1380]" captionTargetBox="[246,1342,215,1317]" captionTargetId="figure-361@12.[246,1342,214,1317]" captionTargetPageId="12" captionText="FIG. 7. — Maculonomia (Maculonomia) rufocaudata (Wu, 1988), female from Lao Cai province: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831230" httpUri="https://zenodo.org/record/14831230/files/figure.png" pageId="15" pageNumber="64">7</figureCitation>
apical margin broadly concave with apicolateral lobes on either side of concavity bearing elongate setae, smaller lobe mesad apicolateral lobes and also with many long setae (
<figureCitation id="1328A3869467FFE0FB5E3CBEC66A4DC1" box="[1238,1345,1682,1708]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10D</figureCitation>
). S8 with broad spiculum, apical margin slightly concave with broad medial plate (
<figureCitation id="1328A3869467FFE0FC4C3CFEC7034D81" box="[964,1064,1746,1772]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10E</figureCitation>
). Dorsal gonostylar lobe expanded, flattened, and lamelliform mesally, apex bifid, with ventral portion long, thin and mesally directed, ventral apical margin with smaller lamella, then narrowed toward apex and round apically with many long setae (
<figureCitation id="1328A3869467FFE0FA8F3D7DC6424C01" box="[1287,1385,1873,1900]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10F</figureCitation>
), with elbowed mesal process proximally with thin, flattened plate apically (
<figureCitation id="1328A3869467FFE0FC193DBDC7334CC6" box="[913,1048,1937,1963]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10F, G</figureCitation>
). Ventral gonostylar process narrow with rounded apex, inner margin curved, outer margin strongly wavy (
<figureCitation id="1328A3869467FFE0FC5F3DFDC76B4C86" box="[983,1088,2001,2027]" captionStart="FIG" captionStartId="15.[132,143,1466,1484]" captionTargetBox="[256,1342,216,1391]" captionTargetId="figure-250@15.[246,1342,216,1406]" captionTargetPageId="15" captionText="FIG. 10. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, S4 in frontal view; B, S5 in frontal view; C, S6 in frontal view; D, S7 in frontal view; E, S8 in frontal view; F, male genitalia in dorsal view; G, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64">Fig. 10G</figureCitation>
).
</paragraph>
<caption id="DF6CEF8B9467FFE0FF0C3F96C6BE4E8F" ID-DOI="http://doi.org/10.5281/zenodo.14831236" ID-Zenodo-Dep="14831236" httpUri="https://zenodo.org/record/14831236/files/figure.png" pageId="15" pageNumber="64" startId="15.[132,143,1466,1484]" targetBox="[256,1342,216,1391]" targetPageId="15" targetType="figure">
<paragraph id="8BACBF039467FFE0FF0C3F96C6BE4E8F" blockId="15.[132,1456,1466,1506]" pageId="15" pageNumber="64">
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. 10. —
<taxonomicName id="4C13C4809467FFE0FF563F96C1444EA1" authority="Engel" authorityName="Engel" box="[222,623,1466,1484]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="rufomysteria" subGenus="Maculonomia">
<emphasis id="B96763119467FFE0FF563F96C2654EA1" box="[222,334,1466,1484]" italics="true" pageId="15" pageNumber="64">Maculonomia</emphasis>
(
<emphasis id="B96763119467FFE0FED03F96C2E34EA1" box="[344,456,1466,1484]" italics="true" pageId="15" pageNumber="64">Maculonomia</emphasis>
)
<emphasis id="B96763119467FFE0FE5A3F96C1124EA1" box="[466,569,1466,1484]" italics="true" pageId="15" pageNumber="64">rufomysteria</emphasis>
Engel
</taxonomicName>
, Tran, &amp; AD Nguyen,
<taxonomicNameLabel id="A254DE6A9467FFE0FC953F96C0644EA1" box="[797,847,1466,1484]" pageId="15" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
, holotype male:
<emphasis id="B96763119467FFE0FC5D3F96C0C94EA1" bold="true" box="[981,994,1466,1484]" pageId="15" pageNumber="64">A</emphasis>
, S4 in frontal view;
<emphasis id="B96763119467FFE0FB0C3F96C7BA4EA1" bold="true" box="[1156,1169,1466,1484]" pageId="15" pageNumber="64">B</emphasis>
, S5 in frontal view;
<emphasis id="B96763119467FFE0FABC3F96C6694EA1" bold="true" box="[1332,1346,1466,1484]" pageId="15" pageNumber="64">C</emphasis>
, S6 in frontal view;
<emphasis id="B96763119467FFE0FF3C3FFCC3E94E8F" bold="true" box="[180,194,1488,1506]" pageId="15" pageNumber="64">D</emphasis>
, S7 in frontal view;
<emphasis id="B96763119467FFE0FEEE3FFCC2594E8F" bold="true" box="[358,370,1488,1506]" pageId="15" pageNumber="64">E</emphasis>
, S8 in frontal view;
<emphasis id="B96763119467FFE0FD9D3FFCC10B4E8F" bold="true" box="[533,544,1488,1506]" pageId="15" pageNumber="64">F</emphasis>
, male genitalia in dorsal view;
<emphasis id="B96763119467FFE0FC973FFCC0064E8F" bold="true" box="[799,813,1488,1506]" pageId="15" pageNumber="64">G</emphasis>
, male genitalia in ventral view. Scale bars: A-C, F, G, 1 mm; D, E, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BACBF039478FFFFFF0C3AFBC297499D" blockId="16.[130,777,215,752]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFF0C3AFBC2394B9C" bold="true" box="[132,274,215,241]" pageId="16" pageNumber="65">Sculpturing.</emphasis>
Clypeus with dense punctures on basal half, sparser, and somewhat wrinkled punctures on remainder; punctures on supraclypeal area denser than those on clypeus; paraocular area with dense punctures on basal half and sparser punctures on remainder; scape with small, sparse punctures (
<figureCitation id="1328A3869478FFFFFE8E3B5BC2754AFC" box="[262,350,375,401]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">Fig. 9A</figureCitation>
); frons with dense, nearly contiguous punctures; vertex with shiny, shallow punctures, punctures sparser than those on frons, except outer margin of median ocellus with shiny, smooth, impunctate area. Mesoscutum and mesoscutellum with dense (not contiguous) round punctures (
<figureCitation id="1328A3869478FFFFFE81383AC24F495D" box="[265,356,534,560]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">Fig. 9B</figureCitation>
). Disc of metasomal T1 with round punctures separated by about a puncture width, apical marginal zone broad, with small, dense punctures basally but otherwise impunctate; punctures on discs of T2-T6 smaller and denser than those of T1, with apical margin zones impunctate except small, dense punctures in basal quarters to thirds (
<figureCitation id="1328A3869478FFFFFEDD38F9C284499D" box="[341,431,725,752]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3939C2AD48E2" blockId="16.[132,776,789,911]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFF0C3939C3E04842" bold="true" box="[132,203,789,815]" pageId="16" pageNumber="65">Color.</emphasis>
Body black except forewing yellow, then paler just beyond veins before forming a large, apical, darkly infumate spot (
<figureCitation id="1328A3869478FFFFFF493979C23C4802" box="[193,279,853,879]" captionStart="FIG" captionStartId="13.[141,152,2007,2025]" captionTargetBox="[281,1306,214,1933]" captionTargetId="figure-16@13.[262,1325,213,1954]" captionTargetPageId="13" captionText="FIG. 8. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; A.D. Nguyen, n. sp., holotype male: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831232" httpUri="https://zenodo.org/record/14831232/files/figure.png" pageId="16" pageNumber="65">Fig. 8A</figureCitation>
). Metasomal T5-T6 with shiny, orange bands apically (
<figureCitation id="1328A3869478FFFFFF6C3959C20D48E2" box="[228,294,885,911]" captionStart="FIG" captionStartId="13.[141,152,2007,2025]" captionTargetBox="[281,1306,214,1933]" captionTargetId="figure-16@13.[262,1325,213,1954]" captionTargetPageId="13" captionText="FIG. 8. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; A.D. Nguyen, n. sp., holotype male: A, lateral habitus; B, dorsal habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831232" httpUri="https://zenodo.org/record/14831232/files/figure.png" pageId="16" pageNumber="65">Figs 8</figureCitation>
;
<figureCitation id="1328A3869478FFFFFEBB3959C25248E2" box="[307,377,885,911]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">9D, E</figureCitation>
).
</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3999C24C4EE3" blockId="16.[130,777,949,1422]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFF0C3999C22248A2" bold="true" box="[132,265,949,975]" pageId="16" pageNumber="65">Pubescence.</emphasis>
Clypeus and supraclypeal area with some blackish, plumose setae; paraocular area from base to lower torular tangent with long, dense, blackish, plumose setae; scape with some blackish setae; face above antennal torulus with tufts of long, yellowish-black to blackish, plumose setae (
<figureCitation id="1328A3869478FFFFFF063E78C3CE4F03" box="[142,229,1108,1134]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">Fig. 9A</figureCitation>
). Mesoscutum covered with dense, soft, blackish, plumose setae; setae of mesoscutellum longer than those on mesoscutum (
<figureCitation id="1328A3869478FFFFFEA83EB8C2534FC3" box="[288,376,1172,1198]" captionStart="FIG" captionStartId="14.[132,143,1592,1610]" captionTargetBox="[231,1356,215,1536]" captionTargetId="figure-182@14.[231,1356,215,1536]" captionTargetPageId="14" captionText="FIG. 9. — Maculonomia (Maculonomia) rufomysteria Engel, Tran, &amp; AD Nguyen, n. sp., holotype male: A, facial view; B, mesosoma in dorsal view; C, metatibia showing the basal plate; D, metasoma in dorsal view; E, metasomal T5-T7 in dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14831234" httpUri="https://zenodo.org/record/14831234/files/figure.png" pageId="16" pageNumber="65">Fig. 9B</figureCitation>
). Propodeum with tuft of blackish, plumose setae laterally. Ventral surfaces of coxae, trochanters, pro- and mesofemora with tufts of soft, blackish, plumose setae. Outer surfaces of tibiae and basitarsi with bristle-like and blackish setae, inner surfaces of metatibia and metabasitarsus with bristle-like and ferruginous setae. Metasomal T1-T4 covered with dense, soft, blackish setae; T5-T6 with reddish-orange setae.
</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3F9FC3E64EA0" blockId="16.[132,249,1459,1517]" box="[132,205,1459,1485]" pageId="16" pageNumber="65">
<emphasis id="B96763119478FFFFFF0C3F9FC3E64EA0" box="[132,205,1459,1485]" italics="true" pageId="16" pageNumber="65">Female</emphasis>
</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3FFFC3D24E80" blockId="16.[132,249,1459,1517]" box="[132,249,1491,1517]" pageId="16" pageNumber="65">Unknown.</paragraph>
<paragraph id="8BACBF039478FFFFFF0C3C04C1AC4D2C" blockId="16.[132,647,1575,1602]" box="[132,647,1575,1602]" pageId="16" pageNumber="65">
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(
<taxonomicName id="4C13C4809478FFFFFE503C01C1054D2D" baseAuthorityName="Smith" baseAuthorityYear="1857" box="[472,558,1581,1600]" class="Insecta" family="Halictidae" genus="Maculonomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="apicalis">
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<emphasis id="B96763119478FFFFFE503C01C1054D2D" box="[472,558,1581,1600]" italics="true" pageId="16" pageNumber="65">APICALIS</emphasis>
</smallCapsWord>
</taxonomicName>
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)
</paragraph>
</subSubSection>
</treatment>
</document>
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