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<document id="374CCD0F49730B9EC2EE0101F888D3EE" ID-CLB-Dataset="271660" ID-DOI="10.5852/ejt.2023.900.2299" ID-GBIF-Dataset="8b15583e-a9c6-4c32-8438-f02b6e4490d6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8420708" ID-ZooBank="0B4BF469-77A8-4CF9-A006-3B98FF4187D5" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696845197903" checkinUser="plazi" docAuthor="Cole, Mary L. &amp; Herbert, David G." docDate="2023" docId="E60087E6FFABFFCDFDD45246FBBBFDBF" docLanguage="en" docName="EJT.2023.900.1-31.pdf" docOrigin="European Journal of Taxonomy 900 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2299/9967" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella kevincolei Cole &amp; Herbert 2023, sp. nov." docType="treatment" docUuid="57C359E6-21D6-4E56-98A5-B56C9FFE3215" docUuidSource="ZooBank" docVersion="6" lastPageNumber="13" masterDocId="1A39FF9EFFA2FFC1FFAF547EFF99FFD9" masterDocTitle="Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another" masterLastPageNumber="31" masterPageNumber="1" pageNumber="10" updateTime="1699694286568" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="86FACE3413D1A4A20F5DF3A7EE4624E6">
<mods:title id="B59997B06330EF4B2748794C7065B219">Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another</mods:title>
</mods:titleInfo>
<mods:name id="8154310446805719CF56049090FFE1D1" type="personal">
<mods:role id="B6949E973D10387FACD1C58022FC0217">
<mods:roleTerm id="A19C1139C9CAD8CF78252032B8C17012">Author</mods:roleTerm>
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<mods:namePart id="E00AAAE111308C501103CA28025E736B">Cole, Mary L.</mods:namePart>
<mods:nameIdentifier id="389EBB37BC58CB735E0BC22AE26743B5" type="LSID">FDA80F4A-4E08-401E-8FF6-778E033BBBA7</mods:nameIdentifier>
<mods:affiliation id="76D4970FD5B4474BFCB20E581A5AD1BF">East London Museum, 319 Oxford St, East London, 5201, South Africa. &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa.</mods:affiliation>
<mods:nameIdentifier id="A82AA93A4ED31AD7EA4A1B8DCDF68FDE" type="email">marybursey@elmuseum.za.org</mods:nameIdentifier>
</mods:name>
<mods:name id="411FFE66E081ABC7BB6D305102E6FD8E" type="personal">
<mods:role id="A04458E900DDAAC1AB35A4E5B09D2C09">
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<mods:namePart id="5722FE6C607D6157C74570E72B5D8B1E">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="AB9F65A5EEA2F98E980E1425D49AD710" type="LSID">0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="F4EA59EBACE6DB179C2CD4DB7753E15D">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="575C4A081AA15347913D1D02A0F3173B" type="email">phasianella@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="AC343489FDA1556892F83BB678A5D615">text</mods:typeOfResource>
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<mods:title id="0185357EF5E4201CA108C9B7F85D1B29">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BA56A28015C30B5AB6E02FFE0B359CCD">
<mods:date id="D16A1067F1E0FC8AF405997CA506D43E">2023</mods:date>
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<mods:number id="6A1E62425EEE1001FD6206C27CE6096C">2023-10-09</mods:number>
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<mods:number id="39A913FCBC74BD65E3BC528B36448EAA">900</mods:number>
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</mods:location>
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<mods:identifier id="85766CF35A16A3152A216A5406241210" type="DOI">10.5852/ejt.2023.900.2299</mods:identifier>
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<treatment id="E60087E6FFABFFCDFDD45246FBBBFDBF" ID-DOI="http://doi.org/10.5281/zenodo.8423112" ID-GBIF-Taxon="213740317" ID-Zenodo-Dep="8423112" LSID="urn:lsid:zoobank.org:act:57C359E6-21D6-4E56-98A5-B56C9FFE3215" httpUri="http://treatment.plazi.org/id/E60087E6FFABFFCDFDD45246FBBBFDBF" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="26B3657BFFABFFC8FDD45246FB26F9AE" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="6E1636F0FFABFFC8FDD45246FC21F98A" blockId="9.[372,1215,1592,1689]" box="[635,952,1592,1619]" pageId="9" pageNumber="10">
<heading id="355E819CFFABFFC8FDD45246FC21F98A" box="[635,952,1592,1619]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="A9A94D73FFABFFC8FDD45246FCCFF98A" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[635,854,1592,1619]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="kevincolei" status="sp. nov.">
<emphasis id="5CDDEAE2FFABFFC8FDD45246FCCFF98A" bold="true" box="[635,854,1592,1619]" italics="true" pageId="9" pageNumber="10">Gulella kevincolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFABFFC8FCF15247FC21F98A" box="[862,952,1593,1619]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="6E1636F0FFABFFC8FEDB5222FB26F9AE" blockId="9.[372,1215,1592,1689]" box="[372,1215,1628,1655]" pageId="9" pageNumber="10">
<uri id="1A383AF2FFABFFC8FEDB5222FB26F9AE" box="[372,1215,1628,1655]" pageId="9" pageNumber="10">
urn:lsid:zoobank.org:act:
<uuid id="1A0F0C25FFABFFC8FD385222FB26F9AE" box="[663,1215,1628,1655]" pageId="9" pageNumber="10">57C359E6-21D6-4E56-98A5-B56C9FFE3215</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFABFFC8FD485201FCD4F940" box="[743,845,1663,1689]" pageId="9" pageNumber="10" type="description">
<paragraph id="6E1636F0FFABFFC8FD485201FCD4F940" blockId="9.[372,1215,1592,1689]" box="[743,845,1663,1689]" pageId="9" pageNumber="10">
<figureCitation id="F6922A75FFABFFC8FD485201FCA8F940" box="[743,817,1663,1689]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[189,1398,901,1886]" captionTargetId="figure-219@5.[189,1398,901,1886]" captionTargetPageId="5" captionText="Fig. 3. Distribution map of Gulella benthodon van Bruggen, 1980 (blue triangles), G. judithmastersae sp. nov. (green squares), G. kevincolei sp. nov. (yellow circle) and G. mcmasteri sp. nov. (red diamonds) in the Amathole region of the Eastern Cape province. Contours at 1000 m and 2000 m." figureDoi="http://doi.org/10.5281/zenodo.8420714" httpUri="https://zenodo.org/record/8420714/files/figure.png" pageId="9" pageNumber="10">Figs 3</figureCitation>
,
<figureCitation id="F6922A75FFABFFC8FC915201FCD4F940" box="[830,845,1663,1689]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="9" pageNumber="10">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFABFFC8FF1252BEFBACF84E" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="6E1636F0FFABFFC8FF1252BEFEAFF903" blockId="9.[189,310,1728,1754]" box="[189,310,1728,1754]" pageId="9" pageNumber="10">
<heading id="355E819CFFABFFC8FF1252BEFEAFF903" bold="true" box="[189,310,1728,1754]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="5CDDEAE2FFABFFC8FF1252BEFEAFF903" bold="true" box="[189,310,1728,1754]" pageId="9" pageNumber="10">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFABFFC8FF125291FBACF84E" blockId="9.[189,1399,1775,1943]" pageId="9" pageNumber="10">Shell minute, sub-cylindrical; sculptured by well-developed axial ribs, extending from suture to suture, aperture narrowing towards base on right; apertural dentition four-fold, including an oblique parietal lamella, a large roundly triangular somewhat bicuspid labral slab, a ridge-like basal tooth in centre of base, and a large inset elongate columella lamella; the columella lip is also thickened, and has an indistinct swelling at either end in some specimens; umbilicus widely open.</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA8FFCBFF125577FB8DFE4E" pageId="10" pageNumber="11" type="etymology">
<paragraph id="6E1636F0FFA8FFCBFF125577FEDDFEFA" blockId="10.[189,324,265,291]" box="[189,324,265,291]" pageId="10" pageNumber="11">
<heading id="355E819CFFA8FFCBFF125577FEDDFEFA" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="5CDDEAE2FFA8FFCBFF125577FEDDFEFA" bold="true" box="[189,324,265,291]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF125549FB8DFE4E" blockId="10.[189,1398,311,407]" pageId="10" pageNumber="11">Named for the first author’s husband, Kevin Cole, who was a volunteer at Hobbiton-on-Hogsback helping at 32 two-week camps for disadvantaged children between 1979 and 1986, and subsequently Chairman of its board, and who helped collect specimens of this species.</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA8FFCBFF1255C1FC42FBE5" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="6E1636F0FFA8FFCBFF1255C1FEF5FE00" blockId="10.[189,364,447,473]" box="[189,364,447,473]" pageId="10" pageNumber="11">
<heading id="355E819CFFA8FFCBFF1255C1FEF5FE00" bold="true" box="[189,364,447,473]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="5CDDEAE2FFA8FFCBFF1255C1FEF5FE00" bold="true" box="[189,364,447,473]" pageId="10" pageNumber="11">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF455590FD93FDAB" blockId="10.[189,1398,494,626]" pageId="10" pageNumber="11">
<materialsCitation id="DEC13CADFFA8FFCBFF455590FD93FDAB" ID-GBIF-Occurrence="4423011364" collectingDate="2021-09-27" collectorName="M. Cole" country="South Africa" county="Hogsback" elevation="1065" latitude="-32.6063" location="Auckland Nature Reserve" longLatPrecision="6" longitude="26.9625" municipality="Auckland Nature Reserve" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<emphasis id="5CDDEAE2FFA8FFCBFF455590FEC5FDD1" bold="true" box="[234,348,494,520]" pageId="10" pageNumber="11">
<typeStatus id="B1128852FFA8FFCBFF455590FEC5FDD1" box="[234,348,494,520]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFA8FFCBFF12566FFE08FDF2" box="[189,401,529,555]" name="South Africa" pageId="10" pageNumber="11">SOUTH AFRICA</collectingCountry>
–
<emphasis id="5CDDEAE2FFA8FFCBFE1B566FFDECFDF2" bold="true" box="[436,629,528,555]" pageId="10" pageNumber="11">
<collectingRegion id="AC6DF812FFA8FFCBFE1B566FFDF8FDF3" box="[436,609,528,555]" country="South Africa" name="Eastern Cape" pageId="10" pageNumber="11">Eastern Cape</collectingRegion>
•
</emphasis>
<collectingCounty id="87774E7CFFA8FFCBFDD1566FFD6FFDF2" box="[638,758,529,555]" pageId="10" pageNumber="11">Hogsback</collectingCounty>
,
<collectingMunicipality id="8E72AC8AFFA8FFCBFCAC566FFBA0FDF2" box="[771,1081,529,555]" pageId="10" pageNumber="11">Auckland Nature Reserve</collectingMunicipality>
,
<locationDeviation id="7FA42FD2FFA8FFCBFBE7566FFDBFFD97" pageId="10" pageNumber="11">base of vertical cliff near Madonna and Child waterfall</locationDeviation>
;
<geoCoordinate id="0B9D5037FFA8FFCBFD9A564AFD27FD97" box="[565,702,564,590]" degrees="32.6063" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-32.6063">32.6063° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFA8FFCBFD60564AFCC2FD97" box="[719,859,564,591]" degrees="26.9625" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="26.9625">26.9625° E</geoCoordinate>
;
<elevation id="E584D1C3FFA8FFCBFCC4564AFB95FD97" box="[875,1036,564,591]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">
<quantity id="A9519B15FFA8FFCBFCC4564AFC51FD97" box="[875,968,564,591]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">1065 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFA8FFCBFBB1564AFB5AFD97" box="[1054,1219,564,590]" pageId="10" pageNumber="11" value="2021-09-27">
<collectingDate id="0A53E9D8FFA8FFCBFBB1564AFB5AFD97" box="[1054,1219,564,590]" pageId="10" pageNumber="11" value="2021-09-27">27 Sep. 2021</collectingDate>
</date>
;
<collectorName id="C35C5326FFA8FFCBFB7C564AFAAEFD97" box="[1235,1335,564,590]" pageId="10" pageNumber="11">M. Cole</collectorName>
leg.; NMSA-Mol 0P2357/T4506.
</materialsCitation>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF4556E7FD2EFCE6" blockId="10.[189,1399,665,832]" pageId="10" pageNumber="11">
<materialsCitation id="DEC13CADFFA8FFCBFF4556E7FE93FCC5" ID-GBIF-Occurrence="4423011350" collectingDate="2021-09-27" collectorName="M. Cole" country="South Africa" county="Hogsback Forest" elevation="1065" latitude="-32.6063" location="Hogsback Forest" longLatPrecision="6" longitude="26.9625" municipality="Madonna" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="5CDDEAE2FFA8FFCBFF4556E7FEFEFD6A" bold="true" box="[234,359,665,691]" pageId="10" pageNumber="11">
<typeStatus id="B1128852FFA8FFCBFF4556E7FEFEFD6A" box="[234,359,665,691]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFA8FFCBFF1256C2FE16FD0F" box="[189,399,700,726]" name="South Africa" pageId="10" pageNumber="11">SOUTH AFRICA</collectingCountry>
–
<emphasis id="5CDDEAE2FFA8FFCBFE0456C2FDCFFD0C" bold="true" box="[427,598,699,726]" pageId="10" pageNumber="11">
<collectingRegion id="AC6DF812FFA8FFCBFE0456C2FDCFFD0C" box="[427,598,699,726]" country="South Africa" name="Eastern Cape" pageId="10" pageNumber="11">Eastern Cape</collectingRegion>
</emphasis>
•
<specimenCount id="78AFFD79FFA8FFCBFDDE56C2FD5FFD0F" box="[625,710,700,726]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="87774E7CFFA8FFCBFD7C56C2FC05FD0F" box="[723,924,700,726]" pageId="10" pageNumber="11">Hogsback Forest</collectingCounty>
,
<locationDeviation id="7FA42FD2FFA8FFCBFC0756C2FEE9FD20" pageId="10" pageNumber="11">base of vertical cliff near Madonna and Child waterfall</locationDeviation>
;
<geoCoordinate id="0B9D5037FFA8FFCBFED556A1FE64FD20" box="[378,509,735,761]" degrees="32.6063" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-32.6063">32.6063° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFA8FFCBFDA656A1FD16FD20" box="[521,655,735,762]" degrees="26.9625" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="26.9625">26.9625° E</geoCoordinate>
;
<elevation id="E584D1C3FFA8FFCBFD3556A1FCA9FD20" box="[666,816,735,762]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">
<quantity id="A9519B15FFA8FFCBFD3556A1FD6BFD20" box="[666,754,735,762]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">1065 m</quantity>
a.s.l.
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;
<date id="1A171030FFA8FFCBFC9356A1FC4FFD20" box="[828,982,735,761]" pageId="10" pageNumber="11" value="2021-09-27">
<collectingDate id="0A53E9D8FFA8FFCBFC9356A1FC4FFD20" box="[828,982,735,761]" pageId="10" pageNumber="11" value="2021-09-27">27 Sep. 2021</collectingDate>
</date>
;
<collectorName id="C35C5326FFA8FFCBFC4E56A1FBD9FD20" box="[993,1088,735,761]" pageId="10" pageNumber="11">M. Cole</collectorName>
leg.; NMSA-Mol 0P2358/ T4607
</materialsCitation>
<emphasis id="5CDDEAE2FFA8FFCBFEBC577CFE84FCC5" bold="true" box="[275,285,770,796]" pageId="10" pageNumber="11">•</emphasis>
<materialsCitation id="DEC13CADFFA8FFCBFE89577CFBA5FCC4" ID-GBIF-Occurrence="4423011337" box="[294,1084,770,797]" collectingDate="2021-09-27" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" county="Hogsback Forest" elevation="1065" latitude="-32.6063" location="Child" longLatPrecision="6" longitude="26.9625" municipality="Madonna" pageId="10" pageNumber="11" specimenCode="ELMD18887" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="78AFFD79FFA8FFCBFE89577CFEE4FCC4" box="[294,381,770,797]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<specimenCode id="3E0F9E8BFFA8FFCBFCF6577CFC6FFCC4" box="[857,1014,770,797]" collectionCode="ELMD" pageId="10" pageNumber="11">ELMD18887</specimenCode>
/T235
</materialsCitation>
<emphasis id="5CDDEAE2FFA8FFCBFBE8577CFBC8FCC5" bold="true" box="[1095,1105,770,796]" pageId="10" pageNumber="11">•</emphasis>
<materialsCitation id="DEC13CADFFA8FFCBFBF6577CFD2AFCE6" ID-GBIF-Occurrence="4423011314" collectingDate="2021-09-27" collectionCode="NHMUK" collectorName="M. Cole" country="South Africa" county="Hogsback Forest" elevation="1065" latitude="-32.6063" location="Child" longLatPrecision="6" longitude="26.9625" municipality="Madonna" pageId="10" pageNumber="11" specimenCode="NHMUK 20230167" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="78AFFD79FFA8FFCBFBF6577CFB36FCC4" box="[1113,1199,770,797]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<specimenCode id="3E0F9E8BFFA8FFCBFE6B5758FD2AFCE6" box="[452,691,805,832]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="11" type="Museum">NHMUK 20230167</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF125718FE60FC59" blockId="10.[189,505,870,896]" box="[189,505,870,896]" pageId="10" pageNumber="11">
<heading id="355E819CFFA8FFCBFF125718FE60FC59" bold="true" box="[189,505,870,896]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="5CDDEAE2FFA8FFCBFF125718FE60FC59" bold="true" box="[189,505,870,896]" pageId="10" pageNumber="11">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF1257EBFC42FBE5" blockId="10.[189,1399,917,1085]" pageId="10" pageNumber="11">
<materialsCitation id="DEC13CADFFA8FFCBFF1257EBFD5FFC2C" ID-GBIF-Occurrence="4423011333" collectingDate="2008-12-31" collectionCode="NMSA" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="10" pageNumber="11" specimenCode="W9361" specimenCount="1" stateProvince="Eastern Cape">
<collectingCountry id="16BE7660FFA8FFCBFF1257EBFE0DFC76" box="[189,404,917,943]" name="South Africa" pageId="10" pageNumber="11">SOUTH AFRICA</collectingCountry>
–
<emphasis id="5CDDEAE2FFA8FFCBFE1157EBFD1CFC76" bold="true" box="[446,645,917,943]" pageId="10" pageNumber="11">
<collectingRegion id="AC6DF812FFA8FFCBFE1157EBFDF7FC76" box="[446,622,917,943]" country="South Africa" name="Eastern Cape" pageId="10" pageNumber="11">Eastern Cape</collectingRegion>
•
</emphasis>
<specimenCount id="78AFFD79FFA8FFCBFD3C57EBFD77FC69" box="[659,750,917,944]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="87774E7CFFA8FFCBFCAF57EBFCE1FC76" box="[768,888,917,943]" pageId="10" pageNumber="11">Hogsback</collectingCounty>
,
<collectingMunicipality id="8E72AC8AFFA8FFCBFC2757EBFB5CFC69" box="[904,1221,917,944]" pageId="10" pageNumber="11">Auckland Nature Reserve</collectingMunicipality>
,
<location id="6B76602BFFA8FFCBFB7857E8FE9FFC0B" LSID="urn:lsid:plazi:treatment:E60087E6FFABFFCDFDD45246FBBBFDBF:6B76602BFFA8FFCBFB7857E8FE9FFC0B" country="South Africa" county="Hogsback" latitude="-32.6055" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" name="Tyume River valley" pageId="10" pageNumber="11" stateProvince="Eastern Cape">Tyume River valley</location>
,
<locationDeviation id="7FA42FD2FFA8FFCBFEBB57C6FCA8FC0B" box="[276,817,952,978]" pageId="10" pageNumber="11">downstream of Madonna and Child waterfall</locationDeviation>
;
<geoCoordinate id="0B9D5037FFA8FFCBFC9057C6FC5CFC0A" box="[831,965,952,979]" degrees="32.6055" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-32.6055">32.6055° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFA8FFCBFC7B57C6FBC7FC0A" box="[980,1118,952,979]" degrees="26.9603" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="26.9603">26.9603° E</geoCoordinate>
;
<elevation id="E584D1C3FFA8FFCBFBC357C7FA93FC0B" box="[1132,1290,952,979]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="11" unit="m" value="1030.0">
<quantity id="A9519B15FFA8FFCBFBC357C7FB51FC0A" box="[1132,1224,953,979]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="11" unit="m" value="1030.0">1030 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFA8FFCBFAB657C7FF64FC2F" pageId="10" pageNumber="11" value="2008-12-31">
<collectingDate id="0A53E9D8FFA8FFCBFAB657C7FF64FC2F" pageId="10" pageNumber="11" value="2008-12-31">31 Dec. 2008</collectingDate>
</date>
;
<collectorName id="C35C5326FFA8FFCBFEA257A2FE50FC2F" box="[269,457,988,1014]" pageId="10" pageNumber="11">M. and K. Cole</collectorName>
leg.;
<collectionCode id="08B8AE35FFA8FFCBFDBD57A2FDFCFC2F" box="[530,613,988,1014]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="10" pageNumber="11" type="Museum">NMSA</collectionCode>
<specimenCode id="3E0F9E8BFFA8FFCBFDC057A5FD5FFC2C" box="[623,710,987,1013]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="10" pageNumber="11" type="Museum">W9361</specimenCode>
</materialsCitation>
•
<materialsCitation id="DEC13CADFFA8FFCBFD4A57A2FE9FFBC0" ID-GBIF-Occurrence="4423011362" collectingDate="2008-12-31" collectionCode="NMSA" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA8FFCBFD4A57A2FCA4FC2F" box="[741,829,988,1014]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
; same collection data as for preceding; ELMD 19089
</materialsCitation>
•
<materialsCitation id="DEC13CADFFA8FFCBFE8F5781FC4EFBE5" ID-GBIF-Occurrence="4423011352" collectingDate="2022-01-04" collectorName="M. Cole" country="South Africa" county="Hogsback Forest" elevation="1065" latitude="-32.6063" location="Hogsback Forest" longLatPrecision="6" longitude="26.9625" municipality="Madonna" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA8FFCBFE8F5781FEECFBC0" box="[288,373,1023,1049]" pageId="10" pageNumber="11" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="87774E7CFFA8FFCBFE2E5781FDD3FBC0" box="[385,586,1023,1049]" pageId="10" pageNumber="11">Hogsback Forest</collectingCounty>
,
<locationDeviation id="7FA42FD2FFA8FFCBFDF95781FB79FBC0" box="[598,1248,1023,1049]" pageId="10" pageNumber="11">base of vertical cliff near Madonna and Child waterfall</locationDeviation>
;
<geoCoordinate id="0B9D5037FFA8FFCBFB435781FAE8FBC0" box="[1260,1393,1023,1049]" degrees="32.6063" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-32.6063">32.6063° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFA8FFCBFF12505CFEDDFBE4" box="[189,324,1058,1085]" degrees="26.9625" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="26.9625">26.9625° E</geoCoordinate>
;
<elevation id="E584D1C3FFA8FFCBFEFE505CFE73FBE5" box="[337,490,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">
<quantity id="A9519B15FFA8FFCBFEFE505CFE33FBE4" box="[337,426,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="10" pageNumber="11" unit="m" value="1065.0">1065 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFA8FFCBFE58505CFD18FBE5" box="[503,641,1058,1085]" pageId="10" pageNumber="11" value="2022-01-04">
<collectingDate id="0A53E9D8FFA8FFCBFE58505CFD18FBE5" box="[503,641,1058,1085]" pageId="10" pageNumber="11" value="2022-01-04">4 Jan. 2022</collectingDate>
</date>
;
<collectorName id="C35C5326FFA8FFCBFD22505DFD74FBE5" box="[653,749,1058,1085]" pageId="10" pageNumber="11">M. Cole</collectorName>
leg.; ELMD 18880
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA8FFCBFF12501DFED8F866" pageId="10" pageNumber="11" type="description">
<paragraph id="6E1636F0FFA8FFCBFF12501DFED7FBA4" blockId="10.[189,334,1123,1149]" box="[189,334,1123,1149]" pageId="10" pageNumber="11">
<heading id="355E819CFFA8FFCBFF12501DFED7FBA4" bold="true" box="[189,334,1123,1149]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="5CDDEAE2FFA8FFCBFF12501DFED7FBA4" bold="true" box="[189,334,1123,1149]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF1250ECFD17F940" blockId="10.[188,1401,1170,1689]" pageId="10" pageNumber="11">
<smallCapsWord id="68F0A02CFFA8FFCBFF1250ECFE9BFB72" baselines="1189,1190" box="[189,258,1170,1196]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="10" pageNumber="11">SHELL</smallCapsWord>
(
<figureCitation id="F6922A75FFA8FFCBFEB950ECFEF9FB75" box="[278,352,1170,1197]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
). Shell minute, sub-cylindrical, length 2.5–3.0 mm, width
<quantity id="A9519B15FFA8FFCBFB8D50ECFB2CFB74" box="[1058,1205,1170,1197]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="10" pageNumber="11" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
, L:W 1.98–2.15 (n = 6). Protoconch approx.
<quantity id="A9519B15FFA8FFCBFDB150CBFD1FFB09" box="[542,646,1205,1232]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="0.9">0. 9 mm</quantity>
in diameter, comprising approx. 2.5 whorls, smooth; junction with teleoconch distinct. Teleoconch comprising approx. 4.5 whorls; whorls convex; sculptured with well-developed axial ribs from suture to suture (approx. 32 on penultimate whorl; rib intervals lacking sculpture (
<figureCitation id="F6922A75FFA8FFCBFEEE5161FE53FAE0" box="[321,458,1311,1337]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="10" pageNumber="11">Fig. 6A–B</figureCitation>
). Aperture narrowing towards base on right (in apertural view); peristome thickened and reflected; base flattened rather than rounded, dentition four-fold (
<figureCitation id="F6922A75FFA8FFCBFBC1513CFB55FA85" box="[1134,1228,1346,1372]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="10" pageNumber="11">Fig. 6C</figureCitation>
): 1) an almost vertical parietal lamella, concave on side facing labral sinus, 2) a large roundly triangular labral slab, somewhat bicuspid in some specimens, leaving only a narrow slit between it and parietal lamella, 3) a ridge-like basal tooth in centre of base, 4) a large, deep-set, elongate columella lamella; in addition, in some specimens the thickened columella may bear a low, broad denticle near its parietal insertion and another near its base, but in other specimens these are scarcely evident. Labral tooth corresponds with a deep pit behind outer lip (
<figureCitation id="F6922A75FFA8FFCBFD92526BFD00F9F6" box="[573,665,1557,1583]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="10" pageNumber="11">Fig. 6B</figureCitation>
); basal tooth corresponds with a pit behind basal lip. Apertural tube behind columella lip collar-like with fine close-set riblets. Umbilicus widely open, elongate-oval, approx.
<quantity id="A9519B15FFA8FFCBFEB25222FE1FF9AF" box="[285,390,1628,1654]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="mm" value="0.12">0.12 mm</quantity>
in length (
<figureCitation id="F6922A75FFA8FFCBFDA95225FDFCF9AC" box="[518,613,1627,1653]" captionStart="Fig" captionStartId="11.[191,234,1954,1980]" captionTargetBox="[193,1392,459,1908]" captionTargetId="figure-43@11.[189,1398,445,1916]" captionTargetPageId="11" captionText="Fig. 6. Gulella kevincolei sp. nov., holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="10" pageNumber="11">Fig. 6D</figureCitation>
). Shell almost transparent when fresh, yellow-orange coloration of dried tissue of animal visible internally.
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF1252BEFE2FF903" blockId="10.[189,438,1728,1754]" box="[189,438,1728,1754]" pageId="10" pageNumber="11">
<emphasis id="5CDDEAE2FFA8FFCBFF1252BEFECFF903" bold="true" box="[189,342,1728,1754]" pageId="10" pageNumber="11">Distribution</emphasis>
(
<figureCitation id="F6922A75FFA8FFCBFEC952BEFE35F903" box="[358,428,1728,1754]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[189,1398,901,1886]" captionTargetId="figure-219@5.[189,1398,901,1886]" captionTargetPageId="5" captionText="Fig. 3. Distribution map of Gulella benthodon van Bruggen, 1980 (blue triangles), G. judithmastersae sp. nov. (green squares), G. kevincolei sp. nov. (yellow circle) and G. mcmasteri sp. nov. (red diamonds) in the Amathole region of the Eastern Cape province. Contours at 1000 m and 2000 m." figureDoi="http://doi.org/10.5281/zenodo.8420714" httpUri="https://zenodo.org/record/8420714/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
)
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF125291FEB0F8F5" blockId="10.[188,1398,1775,1836]" pageId="10" pageNumber="11">
Known only from Hogsback in the
<taxonomicName id="A9A94D73FFA8FFCBFDFC5291FCD7F8D0" authority="Mountains" authorityName="Mountains" box="[595,846,1775,1801]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="amathole">Amathole Mountains</taxonomicName>
; at altitudes between
<quantity id="A9519B15FFA8FFCBFBE95291FB04F8D0" box="[1094,1181,1775,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="1000.0">1000 m</quantity>
and
<quantity id="A9519B15FFA8FFCBFB7B5291FAB3F8D0" box="[1236,1322,1775,1801]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="11" unit="m" value="1100.0">1100 m</quantity>
above sea level.
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF12532DFE84F8B4" blockId="10.[189,285,1875,1901]" box="[189,285,1875,1901]" pageId="10" pageNumber="11">
<heading id="355E819CFFA8FFCBFF12532DFE84F8B4" bold="true" box="[189,285,1875,1901]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="5CDDEAE2FFA8FFCBFF12532DFE84F8B4" bold="true" box="[189,285,1875,1901]" pageId="10" pageNumber="11">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA8FFCBFF1253FCFED8F866" blockId="10.[189,1398,1922,1983]" pageId="10" pageNumber="11">
<taxonomicName id="A9A94D73FFA8FFCBFF1253FCFE75F845" authority="Mistbelt Forest" authorityName="Mistbelt Forest" box="[189,492,1922,1948]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="amathole">Amathole Mistbelt Forest</taxonomicName>
(Southern Mistbelt Forest group) (
<bibRefCitation id="0A384B01FFA8FFCBFC2453FCFB38F845" author="Von Maltitz G. &amp; Mucina L. &amp; Geldenhuys C. &amp; Lawes M. &amp; Eeley H. &amp; Adie H. &amp; Vink D. &amp; Fleming G. &amp; Bailey C." box="[907,1185,1922,1948]" pageId="10" pageNumber="11" refId="ref14670" refString="Von Maltitz G., Mucina L., Geldenhuys C., Lawes M., Eeley H., Adie H., Vink D., Fleming G. &amp; Bailey C. 2003. Classification System for South African Indigenous Forests. CSIR, Pretoria." type="book" year="2003">
von Maltitz
<emphasis id="5CDDEAE2FFA8FFCBFBB053FDFBCAF845" box="[1055,1107,1922,1948]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 2003
</bibRefCitation>
); in leaf-litter and under logs.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA9FFCDFF125577FBBBFDBF" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
<paragraph id="6E1636F0FFA9FFCAFF125577FEB6FEFA" blockId="11.[189,303,265,291]" box="[189,303,265,291]" pageId="11" pageNumber="12">
<heading id="355E819CFFA9FFCAFF125577FEB6FEFA" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="5CDDEAE2FFA9FFCAFF125577FEB6FEFA" bold="true" box="[189,303,265,291]" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA9FFCDFF125549FEDFFE69" blockId="11.[188,1398,311,373]" lastBlockId="12.[189,1400,265,432]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<taxonomicName id="A9A94D73FFA9FFCAFF125549FE0BFE88" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[189,402,311,337]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kevincolei" status="sp. nov.">
<emphasis id="5CDDEAE2FFA9FFCAFF125549FE0BFE88" box="[189,402,311,337]" italics="true" pageId="11" pageNumber="12">Gulella kevincolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA9FFCAFE375546FE6BFE8B" box="[408,498,312,338]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
resembles the variable species
<taxonomicName id="A9A94D73FFA9FFCAFCF45549FAEFFE8B" authority="(Melvill &amp; Ponsonby, 1895)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1895" box="[859,1398,311,338]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="farquhari">
<emphasis id="5CDDEAE2FFA9FFCAFCF45549FBB1FE88" box="[859,1064,311,337]" italics="true" pageId="11" pageNumber="12">Gulella farquhari</emphasis>
(Melvill &amp; Ponsonby, 1895)
</taxonomicName>
but the following differences appear to be consistent. The sculpture of
<taxonomicName id="A9A94D73FFA9FFCAFBA35525FB30FEAD" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[1036,1193,346,373]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kevincolei">
<emphasis id="5CDDEAE2FFA9FFCAFBA35525FB30FEAD" box="[1036,1193,346,373]" italics="true" pageId="11" pageNumber="12">G. kevincolei</emphasis>
</taxonomicName>
is much stronger than that of
<taxonomicName id="A9A94D73FFAEFFCDFEFD5577FE71FEFA" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1895" box="[338,488,265,291]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="farquhari">
<emphasis id="5CDDEAE2FFAEFFCDFEFD5577FE71FEFA" box="[338,488,265,291]" italics="true" pageId="12" pageNumber="13">G. farquhari</emphasis>
</taxonomicName>
which has variable sculpture, usually in the form of milling just below the suture or weak axial ribbing.
<taxonomicName id="A9A94D73FFAEFFCDFD8F5552FD60FE9F" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[544,761,300,326]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="kevincolei">
<emphasis id="5CDDEAE2FFAEFFCDFD8F5552FD60FE9F" box="[544,761,300,326]" italics="true" pageId="12" pageNumber="13">Gulella kevincolei</emphasis>
</taxonomicName>
has a larger, ridge-like basal tooth which reaches the aperture margin, distinct from the inset basal tooth of
<taxonomicName id="A9A94D73FFAEFFCDFCE9552EFC40FEB0" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1895" box="[838,985,335,362]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="farquhari">
<emphasis id="5CDDEAE2FFAEFFCDFCE9552EFC40FEB0" box="[838,985,335,362]" italics="true" pageId="12" pageNumber="13">G. farquhari</emphasis>
</taxonomicName>
. The umbilicus of
<taxonomicName id="A9A94D73FFAEFFCDFB10552EFAC5FEB0" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[1215,1372,335,362]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="kevincolei">
<emphasis id="5CDDEAE2FFAEFFCDFB10552EFAC5FEB0" box="[1215,1372,335,362]" italics="true" pageId="12" pageNumber="13">G. kevincolei</emphasis>
</taxonomicName>
is widely open and it has a large, smooth protoconch (
<quantity id="A9519B15FFAEFFCDFC82550DFC13FE54" box="[813,906,371,397]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="mm" value="0.9">0.9 mm</quantity>
in the
<typeStatus id="B1128852FFAEFFCDFC4F550CFBD0FE55" box="[992,1097,370,396]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
) clearly distinct from the teleoconch.
</paragraph>
<caption id="3AD66678FFA9FFCAFF1053DCFADCF839" ID-DOI="http://doi.org/10.5281/zenodo.8420722" ID-Zenodo-Dep="8420722" httpUri="https://zenodo.org/record/8420722/files/figure.png" pageId="11" pageNumber="12" startId="11.[191,234,1954,1980]" targetBox="[193,1392,459,1908]" targetPageId="11">
<paragraph id="6E1636F0FFA9FFCAFF1053DCFADCF839" blockId="11.[187,1399,1954,2016]" pageId="11" pageNumber="12">
<emphasis id="5CDDEAE2FFA9FFCAFF1053DCFE90F865" bold="true" box="[191,265,1954,1980]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
<taxonomicName id="A9A94D73FFA9FFCAFEA053DCFE7AF865" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[271,483,1954,1980]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kevincolei" status="sp. nov.">
<emphasis id="5CDDEAE2FFA9FFCAFEA053DCFE7AF865" box="[271,483,1954,1980]" italics="true" pageId="11" pageNumber="12">Gulella kevincolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA9FFCAFE4753DDFDDAF864" box="[488,579,1955,1981]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMSA-Mol 0P2357/T4506), length 2.9 mm, width 1.5 mm.
<emphasis id="5CDDEAE2FFA9FFCAFF1453BBFF49F806" bold="true" box="[187,208,1989,2015]" pageId="11" pageNumber="12">A</emphasis>
. Aperture view.
<emphasis id="5CDDEAE2FFA9FFCAFE3C53B8FE3EF839" bold="true" box="[403,423,1990,2016]" pageId="11" pageNumber="12">B</emphasis>
. Side view.
<emphasis id="5CDDEAE2FFA9FFCAFD9753BBFDD4F806" bold="true" box="[568,589,1989,2015]" pageId="11" pageNumber="12">C</emphasis>
. Detail of aperture.
<emphasis id="5CDDEAE2FFA9FFCAFC9653B8FCD7F839" bold="true" box="[825,846,1990,2016]" pageId="11" pageNumber="12">D</emphasis>
. Oblique view of base showing umbilicus.
</paragraph>
</caption>
<paragraph id="6E1636F0FFAEFFCDFF1255A9FEFCFE28" blockId="12.[189,357,471,497]" box="[189,357,471,497]" pageId="12" pageNumber="13">
<heading id="355E819CFFAEFFCDFF1255A9FEFCFE28" bold="true" box="[189,357,471,497]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="5CDDEAE2FFAEFFCDFF1255A9FEFCFE28" bold="true" box="[189,357,471,497]" pageId="12" pageNumber="13">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFAEFFCDFF125678FBBBFDBF" blockId="12.[189,1399,518,615]" pageId="12" pageNumber="13">
<taxonomicName id="A9A94D73FFAEFFCDFF125678FE0DFDF9" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[189,404,518,544]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="kevincolei" status="sp. nov.">
<emphasis id="5CDDEAE2FFAEFFCDFF125678FE0DFDF9" box="[189,404,518,544]" italics="true" pageId="12" pageNumber="13">Gulella kevincolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFAEFFCDFE335678FE6EFDF9" box="[412,503,518,544]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
has been found at only one locality, Hogsback, in the
<taxonomicName id="A9A94D73FFAEFFCDFBD95678FAEAFDF9" authority="Mountains" authorityName="Mountains" box="[1142,1395,518,544]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="amathole">Amathole Mountains</taxonomicName>
, despite fairly extensive collecting efforts throughout the mountain range. It therefore appears to be a very narrow-range endemic, restricted to the main Hogsback forest block.
</paragraph>
</subSubSection>
</treatment>
</document>
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