treatments-xml/data/E6/00/87/E60087E6FFA1FFC7FDAF5672FC41FAFD.xml

<document id="FB6CB95686B1E0CD4DB50D29BAD5456B" ID-CLB-Dataset="271660" ID-DOI="10.5852/ejt.2023.900.2299" ID-GBIF-Dataset="8b15583e-a9c6-4c32-8438-f02b6e4490d6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8420708" ID-ZooBank="0B4BF469-77A8-4CF9-A006-3B98FF4187D5" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696845197903" checkinUser="plazi" docAuthor="Cole, Mary L. &amp; Herbert, David G." docDate="2023" docId="E60087E6FFA1FFC7FDAF5672FC41FAFD" docLanguage="en" docName="EJT.2023.900.1-31.pdf" docOrigin="European Journal of Taxonomy 900 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2299/9967" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella benthodon van Bruggen 1980, stat. rev." docType="treatment" docVersion="6" lastPageNumber="7" masterDocId="1A39FF9EFFA2FFC1FFAF547EFF99FFD9" masterDocTitle="Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another" masterLastPageNumber="31" masterPageNumber="1" pageNumber="4" updateTime="1699694286568" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2FD0298E0DF15F3BF61E4D2801D16269">Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another</mods:title>
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<mods:namePart id="9696378BF49A7525DEF961AA0EAB6DB0">Cole, Mary L.</mods:namePart>
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<mods:affiliation id="6D4FC81A2E41769F052E2C23F613A2FD">East London Museum, 319 Oxford St, East London, 5201, South Africa. &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa.</mods:affiliation>
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<mods:namePart id="3ABE9B54C6B6822748A39969857AE38A">Herbert, David G.</mods:namePart>
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<mods:affiliation id="76A7C0E0ADDB73547B33E20077BEB693">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, United Kingdom.</mods:affiliation>
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<mods:title id="EB97A5269A3A7E16131370312244700E">European Journal of Taxonomy</mods:title>
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<mods:date id="5C7C898C2A848E8CB74D69CC69250889">2023</mods:date>
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<paragraph id="6E1636F0FFA1FFC2FDAF5672FBADFDFE" blockId="3.[512,1076,524,586]" box="[512,1076,524,551]" pageId="3" pageNumber="4">
<heading id="355E819CFFA1FFC2FDAF5672FBADFDFE" box="[512,1076,524,551]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="A9A94D73FFA1FFC2FDAF5672FC53FDFE" authority="van Bruggen, 1980" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[512,970,524,551]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="benthodon" status="stat. nov.">
<emphasis id="5CDDEAE2FFA1FFC2FDAF5672FD7BFDFE" bold="true" box="[512,738,524,551]" italics="true" pageId="3" pageNumber="4">Gulella benthodon</emphasis>
van 
<bibRefCitation id="0A384B01FFA1FFC2FCB45673FC53FDFE" author="Bruggen A. C. van" box="[795,970,525,551]" pageId="3" pageNumber="4" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen, 1980</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA1FFC2FC7E5673FBADFDFE" box="[977,1076,525,551]" pageId="3" pageNumber="4">stat. rev.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="6E1636F0FFA1FFC2FD48564EFCD4FD93" blockId="3.[512,1076,524,586]" box="[743,845,560,586]" pageId="3" pageNumber="4">
<figureCitation id="F6922A75FFA1FFC2FD48564EFCD4FD93" box="[743,845,560,586]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1963,1989]" captionStartId-1="5.[189,232,1928,1954]" captionTargetBox-0="[226,1361,279,1909]" captionTargetBox-1="[189,1398,901,1886]" captionTargetId-0="figure-20@4.[224,1363,265,1917]" captionTargetId-1="figure-219@5.[189,1398,901,1886]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 2.Gulella benthodon van Bruggen, 1980 status revised (NMSA W25), length 3.4 mm, width 1.4 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." captionText-1="Fig. 3. Distribution map of Gulella benthodon van Bruggen, 1980 (blue triangles), G. judithmastersae sp. nov. (green squares), G. kevincolei sp. nov. (yellow circle) and G. mcmasteri sp. nov. (red diamonds) in the Amathole region of the Eastern Cape province. Contours at 1000 m and 2000 m." figureDoi-0="http://doi.org/10.5281/zenodo.8420712" figureDoi-1="http://doi.org/10.5281/zenodo.8420714" httpUri-0="https://zenodo.org/record/8420712/files/figure.png" httpUri-1="https://zenodo.org/record/8420714/files/figure.png" pageId="3" pageNumber="4">Figs 2–3</figureCitation>
</paragraph>
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<subSubSection id="26B3657BFFA1FFC2FF12560FFDD7FD77" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="6E1636F0FFA1FFC2FF12560FFDD7FD77" blockId="3.[189,1399,625,687]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="4EB911DEFFA1FFC2FF12560FFDD7FD77" pageId="3" pageNumber="4">
<treatmentCitation id="EF0810E1FFA1FFC2FF12560FFCFEFD52" author="Bruggen A. C. van" box="[189,871,625,652]" page="19" pageId="3" pageNumber="4" year="1980">
<taxonomicName id="A9A94D73FFA1FFC2FF12560FFCFEFD52" ID-CoL="7GVQF" authority="van Bruggen, 1980: 17 - 19" authorityName="van Bruggen" authorityPageNumber="17 - 19" authorityYear="1980" box="[189,871,625,652]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="subSpecies" species="darglensis" subSpecies="benthodon">
<emphasis id="5CDDEAE2FFA1FFC2FF12560FFD84FD52" box="[189,541,625,651]" italics="true" pageId="3" pageNumber="4">Gulella darglensis benthodon</emphasis>
van 
<bibRefCitation id="0A384B01FFA1FFC2FDF5560CFCFEFD52" author="Bruggen A. C. van" box="[602,871,625,652]" pageId="3" pageNumber="4" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen, 1980: 17–19</bibRefCitation>
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, fig. 6 (
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locality: Pirie Forest near King Williams Town [R. Godfrey]).
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</paragraph>
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<emphasis id="5CDDEAE2FFA1FFC2FF1256A8FEAFFD29" bold="true" box="[189,310,726,752]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA1FFC2FF12577AFE48FC16" blockId="3.[189,1399,772,975]" pageId="3" pageNumber="4">Shell very small, cylindrical; sculptured with strong axial ribs; apertural dentition eight-fold, including a strong in-running parietal lamella, a large mid-labral complex with a ridge-like upper margin running into aperture with a tooth near lip edge and another further into aperture, and a larger tooth on its lower margin further from lip edge, a low, inset transverse basal tooth to right of centre, an in-running ridge-like basal tooth to left of centre and a large inset bicuspid columella lamella with two ridge-like teeth; umbilicus widely open.</paragraph>
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<emphasis id="5CDDEAE2FFA1FFC2FF125788FEF5FBC9" bold="true" box="[189,364,1014,1040]" pageId="3" pageNumber="4">Type material</emphasis>
(not examined)
</paragraph>
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<materialsCitation id="DEC13CADFFA1FFC2FF45505BFD21FB5C" ID-GBIF-Occurrence="4423011325" collectionCode="NHMUK" collectorName="R. Godfrey" country="South Africa" location="Pirie Forest" municipality="Pirie Forest" pageId="3" pageNumber="4" specimenCode="NHMUK 1937.12" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<emphasis id="5CDDEAE2FFA1FFC2FF45505BFEC5FBE6" bold="true" box="[234,348,1061,1087]" pageId="3" pageNumber="4">
<typeStatus id="B1128852FFA1FFC2FF45505BFEC5FBE6" box="[234,348,1061,1087]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFA1FFC2FF125036FE08FBBB" box="[189,401,1096,1122]" name="South Africa" pageId="3" pageNumber="4">SOUTH AFRICA</collectingCountry>
– 
<emphasis id="5CDDEAE2FFA1FFC2FE195036FDFAFBB8" bold="true" box="[438,611,1095,1122]" pageId="3" pageNumber="4">
<collectingRegion id="AC6DF812FFA1FFC2FE195036FDFAFBB8" box="[438,611,1095,1122]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="4">Eastern Cape</collectingRegion>
</emphasis>
• 
<collectingMunicipality id="8E72AC8AFFA1FFC2FD2E5036FC96FBBB" box="[641,783,1096,1122]" pageId="3" pageNumber="4">Pirie Forest</collectingMunicipality>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FCB15036FBCCFBBB" box="[798,1109,1096,1122]" pageId="3" pageNumber="4">near King Williams Town</locationDeviation>
[now Qonce]; ca 1920; 
<collectorName id="C35C5326FFA1FFC2FF125015FED8FB5C" box="[189,321,1131,1157]" pageId="3" pageNumber="4">R. Godfrey</collectorName>
leg.; 
<specimenCode id="3E0F9E8BFFA1FFC2FE2C5015FDC4FB5C" box="[387,605,1131,1157]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="4" type="Museum">NHMUK 1937.12</specimenCode>
.30.781.
</materialsCitation>
</paragraph>
<paragraph id="6E1636F0FFA1FFC2FF4550D2FB6DFB33" blockId="3.[189,1268,1196,1258]" pageId="3" pageNumber="4">
<materialsCitation id="DEC13CADFFA1FFC2FF4550D2FB69FB33" ID-GBIF-Occurrence="4423011338" collectionCode="NHMUK" collectorName="R. Godfrey" country="South Africa" location="SOUTH AFRICA" municipality="Pirie Forest" pageId="3" pageNumber="4" specimenCode="NHMUK 1937.12" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="5CDDEAE2FFA1FFC2FF4550D2FEC5FB1F" bold="true" box="[234,348,1196,1222]" pageId="3" pageNumber="4">
<typeStatus id="B1128852FFA1FFC2FF4550D2FEC5FB1F" box="[234,348,1196,1222]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFA1FFC2FF1250AEFE17FB33" box="[189,398,1232,1258]" name="South Africa" pageId="3" pageNumber="4">SOUTH AFRICA</collectingCountry>
• 
<specimenCount id="78AFFD79FFA1FFC2FE0850AEFE65FB33" box="[423,508,1232,1258]" pageId="3" pageNumber="4" type="generic">1 spec.</specimenCount>
; same collection data as for holotype; 
<specimenCode id="3E0F9E8BFFA1FFC2FC1050AEFB01FB33" box="[959,1176,1232,1258]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="4" type="Museum">NHMUK 1937.12</specimenCode>
.30.782
</materialsCitation>
.
</paragraph>
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<paragraph id="6E1636F0FFA1FFC2FF12516FFE33FAF2" blockId="3.[189,426,1297,1323]" box="[189,426,1297,1323]" pageId="3" pageNumber="4">
<heading id="355E819CFFA1FFC2FF12516FFE33FAF2" bold="true" box="[189,426,1297,1323]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="5CDDEAE2FFA1FFC2FF12516FFE33FAF2" bold="true" box="[189,426,1297,1323]" pageId="3" pageNumber="4">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA1FFC2FF12513EFD39F8FA" blockId="3.[189,1400,1343,1828]" pageId="3" pageNumber="4">
<materialsCitation id="DEC13CADFFA1FFC2FF12513EFD40FA78" ID-GBIF-Occurrence="4423011334" collectingDate="2002-01-25" collectionCode="NMSA" collectorName="D. Herbert &amp; M. Bursey &amp; G. Redman" country="South Africa" county="Hogsback" elevation="1043" latitude="-32.60701" location="Auckland Nature Reserve" longLatPrecision="4" longitude="26.9625" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCode="W25" specimenCount="5" stateProvince="Eastern Cape">
<collectingCountry id="16BE7660FFA1FFC2FF12513EFE0BFA83" box="[189,402,1344,1370]" name="South Africa" pageId="3" pageNumber="4">SOUTH AFRICA</collectingCountry>
– 
<emphasis id="5CDDEAE2FFA1FFC2FE15513EFDE4FA83" bold="true" box="[442,637,1343,1370]" pageId="3" pageNumber="4">
<collectingRegion id="AC6DF812FFA1FFC2FE15513EFDF1FA80" box="[442,616,1343,1370]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="4">Eastern Cape</collectingRegion>
•
</emphasis>
<specimenCount id="78AFFD79FFA1FFC2FD285141FD7DFA83" box="[647,740,1343,1370]" pageId="3" pageNumber="4">5 specs</specimenCount>
; 
<collectingCounty id="87774E7CFFA1FFC2FD5B5141FCF5FA80" box="[756,876,1343,1369]" pageId="3" pageNumber="4">Hogsback</collectingCounty>
, 
<collectingMunicipality id="8E72AC8AFFA1FFC2FCD55141FB2AFA83" box="[890,1203,1343,1370]" pageId="3" pageNumber="4">Auckland Nature Reserve</collectingMunicipality>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FB6C5141FDB0FAA4" pageId="3" pageNumber="4">downstream of Madonna and Child waterfall</locationDeviation>
; 
<geoCoordinate id="0B9D5037FFA1FFC2FD95511DFD4BFAA4" box="[570,722,1379,1405]" degrees="32.60701" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-32.60701">32.60701° S</geoCoordinate>
, 
<geoCoordinate id="0B9D5037FFA1FFC2FD4B511CFCE8FAA4" box="[740,881,1378,1405]" degrees="26.9625" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="26.9625">26.9625° E</geoCoordinate>
; 
<elevation id="E584D1C3FFA1FFC2FC2D511DFBBFFAA4" box="[898,1062,1379,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="3" pageNumber="4" unit="m" value="1043.0">
<quantity id="A9519B15FFA1FFC2FC2D511DFC78FAA4" box="[898,993,1379,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="3" pageNumber="4" unit="m" value="1043.0">1043 m</quantity>
a.s.l.
</elevation>
; 
<date id="1A171030FFA1FFC2FB97511CFB42FAA4" box="[1080,1243,1378,1405]" pageId="3" pageNumber="4" value="2002-01-25">
<collectingDate id="0A53E9D8FFA1FFC2FB97511CFB42FAA4" box="[1080,1243,1378,1405]" pageId="3" pageNumber="4" value="2002-01-25">25 Jan. 2002</collectingDate>
</date>
; 
<collectorName id="C35C5326FFA1FFC2FB43511DFAEBFAA4" box="[1260,1394,1379,1405]" pageId="3" pageNumber="4">D. Herbert</collectorName>
, 
<collectorName id="C35C5326FFA1FFC2FF1251F8FEA2FA79" box="[189,315,1414,1440]" pageId="3" pageNumber="4">M. Bursey</collectorName>
and 
<collectorName id="C35C5326FFA1FFC2FED851F8FE66FA79" box="[375,511,1414,1440]" pageId="3" pageNumber="4">G. Redman</collectorName>
leg.; 
<collectionCode id="08B8AE35FFA1FFC2FDEB51F8FD0EFA79" box="[580,663,1414,1440]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
<specimenCode id="3E0F9E8BFFA1FFC2FD0F51F8FD40FA78" box="[672,729,1414,1441]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">W25</specimenCode>
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FD5951F8FCBEFA3F" ID-GBIF-Occurrence="4423011343" collectingDate="2008-12-31" collectionCode="NMSA" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCode="W6660" specimenCount="2" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FD5951F8FCC8FA79" box="[758,849,1414,1440]" pageId="3" pageNumber="4">2 specs</specimenCount>
; 
<collectingCounty id="87774E7CFFA1FFC2FCF051F8FC4EFA79" box="[863,983,1414,1440]" pageId="3" pageNumber="4">Hogsback</collectingCounty>
, 
<collectingMunicipality id="8E72AC8AFFA1FFC2FC4C51F8FA81FA79" box="[995,1304,1414,1440]" pageId="3" pageNumber="4">Auckland Nature Reserve</collectingMunicipality>
, 
<location id="6B76602BFFA1FFC2FA8A51F8FECAFA1A" LSID="urn:lsid:plazi:treatment:E60087E6FFA1FFC7FDAF5672FC41FAFD:6B76602BFFA1FFC2FA8A51F8FECAFA1A" country="South Africa" county="Hogsback" latitude="-32.6055" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" name="Tyume River valley" pageId="3" pageNumber="4" stateProvince="Eastern Cape">Tyume River valley</location>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FECD51D7FC13FA1A" box="[354,906,1449,1475]" pageId="3" pageNumber="4">downstream of Madonna and Child waterfall</locationDeviation>
; 
<geoCoordinate id="0B9D5037FFA1FFC2FC3551D7FBBBFA1A" box="[922,1058,1449,1476]" degrees="32.6055" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="-32.6055">32.6055° S</geoCoordinate>
, 
<geoCoordinate id="0B9D5037FFA1FFC2FB9C51D7FB26FA1D" box="[1075,1215,1449,1476]" degrees="26.9603" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="26.9603">26.9603° E</geoCoordinate>
; 
<elevation id="E584D1C3FFA1FFC2FB6051D7FAE9FA1A" box="[1231,1392,1449,1476]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="4" unit="m" value="1030.0">
<quantity id="A9519B15FFA1FFC2FB6051D7FAB5FA1D" box="[1231,1324,1449,1476]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="4" unit="m" value="1030.0">1030 m</quantity>
a.s.l.
</elevation>
; 
<date id="1A171030FFA1FFC2FF1251B2FEFBFA3F" box="[189,354,1484,1511]" pageId="3" pageNumber="4" value="2008-12-31">
<collectingDate id="0A53E9D8FFA1FFC2FF1251B2FEFBFA3F" box="[189,354,1484,1511]" pageId="3" pageNumber="4" value="2008-12-31">31 Dec. 2008</collectingDate>
</date>
; 
<collectorName id="C35C5326FFA1FFC2FEDE51B3FDB4FA3F" box="[369,557,1484,1511]" pageId="3" pageNumber="4">M. and K. Cole</collectorName>
leg.; 
<collectionCode id="08B8AE35FFA1FFC2FDDB51B2FD5EFA3F" box="[628,711,1484,1510]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
<specimenCode id="3E0F9E8BFFA1FFC2FD7E51B2FCBEFA3F" box="[721,807,1484,1510]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">W6660</specimenCode>
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FCEA51B2FD89F9D3" ID-GBIF-Occurrence="4423011335" collectingDate="2008-12-31" collectionCode="NMSA" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCount="2" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FCEA51B2FC39FA3E" box="[837,928,1484,1511]" pageId="3" pageNumber="4">2 specs</specimenCount>
; same collection data as for preceding; ELMD 16135, ELMW 3343
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FD83518EFEFDF9F4" ID-GBIF-Occurrence="4423011306" collectingDate="2008-12-31" collectionCode="NHMUK" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCode="NHMUK 20230166" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FD83518EFD1BF9D3" box="[556,642,1520,1546]" pageId="3" pageNumber="4" type="generic">1 spec.</specimenCount>
; same collection data as for preceding; 
<specimenCode id="3E0F9E8BFFA1FFC2FBF2518EFAD4F9D0" box="[1117,1357,1519,1546]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="4" type="Museum">NHMUK 20230166</specimenCode>
, ex ELMD 16135
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FE2B526DFE9FF988" ID-GBIF-Occurrence="4423011303" collectingDate="2008-12-31" collectionCode="NMW" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FE2B526DFE45F9F4" box="[388,476,1555,1581]" pageId="3" pageNumber="4" type="generic">1 spec.</specimenCount>
; same collection data as for preceding; 
<collectionCode id="08B8AE35FFA1FFC2FC6C526DFB88F9F4" box="[963,1041,1555,1581]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="4">NMW</collectionCode>
.Z.2023.001.00001, ex ELMD 16135
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FEB35248FA9DF988" ID-GBIF-Occurrence="4423011359" box="[284,1284,1590,1617]" collectingDate="2008-12-31" collectionCode="RMNH" collectorName="M. and K. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FEB35248FEF6F989" box="[284,367,1590,1616]" pageId="3" pageNumber="4" type="generic">1 spec.</specimenCount>
; same collection data as for preceding; 
<collectionCode id="08B8AE35FFA1FFC2FC9B5248FC09F989" box="[820,912,1590,1616]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">RMNH</collectionCode>
.MOL.346278, ex ELMD 16135
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FAB55248FC60F94F" ID-GBIF-Occurrence="4423011317" collectingDate="2014-04-06" collectionCode="NMSA" collectorName="D. Herbert &amp; L. Davis &amp; M. Cole" country="South Africa" county="Hogsback" elevation="1043" latitude="-32.607" location="Hogsback" longLatPrecision="6" longitude="26.9625" municipality="Madonna" pageId="3" pageNumber="4" specimenCode="W9731" specimenCount="2" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FAB55248FAEBF989" box="[1306,1394,1590,1616]" pageId="3" pageNumber="4">2 specs</specimenCount>
; 
<collectingCounty id="87774E7CFFA1FFC2FF125227FEACF9AA" box="[189,309,1625,1651]" pageId="3" pageNumber="4">Hogsback</collectingCounty>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FEEE5227FCB6F9AA" box="[321,815,1625,1651]" pageId="3" pageNumber="4">environs of Madonna and Child Waterfall</locationDeviation>
, stn 14-02; 
<geoCoordinate id="0B9D5037FFA1FFC2FC175227FBA4F9AA" box="[952,1085,1625,1651]" degrees="32.6070" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="-32.607">32.6070° S</geoCoordinate>
, 
<geoCoordinate id="0B9D5037FFA1FFC2FBE55227FB4BF9AD" box="[1098,1234,1625,1652]" degrees="26.9625" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="26.9625">26.9625° E</geoCoordinate>
; 
<quantity id="A9519B15FFA1FFC2FB715227FAA1F9AD" box="[1246,1336,1625,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="3" pageNumber="4" unit="m" value="1043.0">
<elevation id="E584D1C3FFA1FFC2FB715227FAA1F9AD" box="[1246,1336,1625,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="3" pageNumber="4" unit="m" value="1043.0">1043 m</elevation>
</quantity>
a.s.l; 
<date id="1A171030FFA1FFC2FF125202FEC8F94F" box="[189,337,1660,1686]" pageId="3" pageNumber="4" value="2014-04-06">
<collectingDate id="0A53E9D8FFA1FFC2FF125202FEC8F94F" box="[189,337,1660,1686]" pageId="3" pageNumber="4" value="2014-04-06">6 Apr. 2014</collectingDate>
</date>
; 
<collectorName id="C35C5326FFA1FFC2FECF5203FE7DF94F" box="[352,484,1660,1687]" pageId="3" pageNumber="4">D. Herbert</collectorName>
, 
<collectorName id="C35C5326FFA1FFC2FE5D5203FDC3F94F" box="[498,602,1660,1687]" pageId="3" pageNumber="4">L. Davis</collectorName>
and 
<collectorName id="C35C5326FFA1FFC2FD365203FD65F94F" box="[665,764,1660,1687]" pageId="3" pageNumber="4">M. Cole</collectorName>
leg.; 
<collectionCode id="08B8AE35FFA1FFC2FCEA5202FC01F94F" box="[837,920,1660,1686]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">NMSA</collectionCode>
<specimenCode id="3E0F9E8BFFA1FFC2FC0D5202FC60F94F" box="[930,1017,1660,1686]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="4" type="Museum">W9731</specimenCode>
</materialsCitation>
• 
<materialsCitation id="DEC13CADFFA1FFC2FBB75202FBEEF904" ID-GBIF-Occurrence="4423011342" collectingDate="2022-01-04" collectorName="M. Cole" country="South Africa" county="Hogsback" elevation="1030" latitude="-32.6055" location="Tyume River valley" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FBB75202FBE9F94E" box="[1048,1136,1660,1687]" pageId="3" pageNumber="4" type="generic">1 spec.</specimenCount>
; 
<collectingCounty id="87774E7CFFA1FFC2FBD05202FB6EF94F" box="[1151,1271,1660,1686]" pageId="3" pageNumber="4">Hogsback</collectingCounty>
, 
<collectingMunicipality id="8E72AC8AFFA1FFC2FAAB5202FEECF963" pageId="3" pageNumber="4">Auckland Nature Reserve</collectingMunicipality>
, 
<location id="6B76602BFFA1FFC2FE2E52DEFDF5F960" LSID="urn:lsid:plazi:treatment:E60087E6FFA1FFC7FDAF5672FC41FAFD:6B76602BFFA1FFC2FE2E52DEFDF5F960" box="[385,620,1695,1722]" country="South Africa" county="Hogsback" latitude="-32.6055" longLatPrecision="6" longitude="26.9603" municipality="Auckland Nature Reserve" name="Tyume River valley" pageId="3" pageNumber="4" stateProvince="Eastern Cape">Tyume River valley</location>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FDD852E1FAEBF960" box="[631,1394,1695,1721]" pageId="3" pageNumber="4">downstream of Madonna and Child waterfall, south-facing slope</locationDeviation>
; 
<geoCoordinate id="0B9D5037FFA1FFC2FF1252BCFEDAF904" box="[189,323,1730,1757]" degrees="32.6055" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="-32.6055">32.6055° S</geoCoordinate>
, 
<geoCoordinate id="0B9D5037FFA1FFC2FEFE52BDFE43F904" box="[337,474,1731,1757]" degrees="26.9603" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="26.9603">26.9603° E</geoCoordinate>
; 
<elevation id="E584D1C3FFA1FFC2FE4752BDFD1AF904" box="[488,643,1731,1757]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="4" unit="m" value="1030.0">
<quantity id="A9519B15FFA1FFC2FE4752BDFDDBF904" box="[488,578,1731,1757]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="4" unit="m" value="1030.0">1030 m</quantity>
a.s.l.
</elevation>
; 
<date id="1A171030FFA1FFC2FD3D52BDFC87F904" box="[658,798,1731,1757]" pageId="3" pageNumber="4" value="2022-01-04">
<collectingDate id="0A53E9D8FFA1FFC2FD3D52BDFC87F904" box="[658,798,1731,1757]" pageId="3" pageNumber="4" value="2022-01-04">4 Jan. 2022</collectingDate>
</date>
; 
<collectorName id="C35C5326FFA1FFC2FC8352BDFC14F904" box="[812,909,1731,1757]" pageId="3" pageNumber="4">M. Cole</collectorName>
leg.; ELMD 19100
</materialsCitation>
<emphasis id="5CDDEAE2FFA1FFC2FB2E52BDFB12F904" bold="true" box="[1153,1163,1731,1757]" pageId="3" pageNumber="4">•</emphasis>
<materialsCitation id="DEC13CADFFA1FFC2FB3D52BDFD05F8FA" ID-GBIF-Occurrence="4423011336" collectingDate="2022-01-04" collectorName="M. Cole" country="South Africa" county="Hogsback" elevation="1086" latitude="-32.6495" location="Hogsback" longLatPrecision="6" longitude="27.0025" municipality="Zinguka area" pageId="3" pageNumber="4" specimenCount="2" stateProvince="Eastern Cape">
<specimenCount id="78AFFD79FFA1FFC2FB3D52BDFB75F904" box="[1170,1260,1731,1757]" pageId="3" pageNumber="4">2 specs</specimenCount>
; 
<collectingCounty id="87774E7CFFA1FFC2FB5552BDFAEBF904" box="[1274,1394,1731,1757]" pageId="3" pageNumber="4">Hogsback</collectingCounty>
, 
<locationDeviation id="7FA42FD2FFA1FFC2FF115298FC0DF8D9" box="[190,916,1766,1792]" pageId="3" pageNumber="4">Zinguka area, at base of large boulder near Wolfridge Road</locationDeviation>
; 
<geoCoordinate id="0B9D5037FFA1FFC2FC0D5298FBB3F8D9" box="[930,1066,1766,1793]" degrees="32.6495" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="-32.6495">32.6495° S</geoCoordinate>
, 
<geoCoordinate id="0B9D5037FFA1FFC2FB965298FB5DF8D9" box="[1081,1220,1766,1793]" degrees="27.0025" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="27.0025">27.0025° E</geoCoordinate>
; 
<elevation id="E584D1C3FFA1FFC2FB7C5298FAEBF8D9" box="[1235,1394,1766,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.0859999999999999" pageId="3" pageNumber="4" unit="m" value="1086.0">
<quantity id="A9519B15FFA1FFC2FB7C5298FAB6F8D9" box="[1235,1327,1766,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.0859999999999999" pageId="3" pageNumber="4" unit="m" value="1086.0">1086 m</quantity>
a.s.l.
</elevation>
; 
<date id="1A171030FFA1FFC2FF125377FEDEF8FA" box="[189,327,1801,1828]" pageId="3" pageNumber="4" value="2022-01-04">
<collectingDate id="0A53E9D8FFA1FFC2FF125377FEDEF8FA" box="[189,327,1801,1828]" pageId="3" pageNumber="4" value="2022-01-04">4 Jan. 2022</collectingDate>
</date>
; 
<collectorName id="C35C5326FFA1FFC2FEFC5374FE2AF8FA" box="[339,435,1801,1828]" pageId="3" pageNumber="4">M. Cole</collectorName>
leg.; ELMD 18881
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA1FFC7FF125334FED8FEAA" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="6E1636F0FFA1FFC2FF125334FED7F8BD" blockId="3.[189,334,1866,1892]" box="[189,334,1866,1892]" pageId="3" pageNumber="4">
<heading id="355E819CFFA1FFC2FF125334FED7F8BD" bold="true" box="[189,334,1866,1892]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="5CDDEAE2FFA1FFC2FF125334FED7F8BD" bold="true" box="[189,334,1866,1892]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA1FFC4FF125307FBDBFD5D" blockId="3.[189,1399,1913,2010]" lastBlockId="5.[188,1400,231,644]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<smallCapsWord id="68F0A02CFFA1FFC2FF125307FE9BF84B" baselines="1932,1933" box="[189,258,1913,1939]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="3" pageNumber="4">SHELL</smallCapsWord>
(
<figureCitation id="F6922A75FFA1FFC2FEB85307FEF8F84A" box="[279,353,1913,1940]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[226,1361,279,1909]" captionTargetId="figure-20@4.[224,1363,265,1917]" captionTargetPageId="4" captionText="Fig. 2.Gulella benthodon van Bruggen, 1980 status revised (NMSA W25), length 3.4 mm, width 1.4 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420712" httpUri="https://zenodo.org/record/8420712/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
). Shell very small, cylindrical, length 3.0– 
<quantity id="A9519B15FFA1FFC2FCC95307FC5CF84D" box="[870,965,1913,1940]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="4" unit="mm" value="3.6">3.6 mm</quantity>
, width 
<quantity id="A9519B15FFA1FFC2FB8F5307FB2DF84D" box="[1056,1204,1913,1940]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="4" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
, L:W 2.18–2.49 (n = 10). Protoconch approx. 
<quantity id="A9519B15FFA1FFC2FD8753E2FD1FF86E" box="[552,646,1948,1975]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="4" unit="mm" value="0.9">0.9 mm</quantity>
in diameter, comprising approx. 2.5 whorls, smooth; junction between protoconch and teleoconch distinct. Teleoconch comprising nearly four whorls; first two whorls moderately convex, subsequent two more weakly so; whorls with faint subsutural angle delimiting narrow subsutural ramp; sculptured by well-developed axial ribs, extending from suture to suture (approx. 30 on penultimate whorl); rib intervals lacking sculpture (
<figureCitation id="F6922A75FFA7FFC4FC775553FBC3FE91" box="[984,1114,301,328]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[226,1361,279,1909]" captionTargetId="figure-20@4.[224,1363,265,1917]" captionTargetPageId="4" captionText="Fig. 2.Gulella benthodon van Bruggen, 1980 status revised (NMSA W25), length 3.4 mm, width 1.4 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420712" httpUri="https://zenodo.org/record/8420712/files/figure.png" pageId="5" pageNumber="6">Fig. 2A–B</figureCitation>
). Aperture sub-quadrate with rounded base and shallowly indented outer lip; dentition eight-fold (
<figureCitation id="F6922A75FFA7FFC4FB9F552FFB16FEB2" box="[1072,1167,337,363]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[226,1361,279,1909]" captionTargetId="figure-20@4.[224,1363,265,1917]" captionTargetPageId="4" captionText="Fig. 2.Gulella benthodon van Bruggen, 1980 status revised (NMSA W25), length 3.4 mm, width 1.4 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420712" httpUri="https://zenodo.org/record/8420712/files/figure.png" pageId="5" pageNumber="6">Fig. 2C</figureCitation>
): 1) an oblique, in-running parietal lamella; 2–4) a large mid-labral complex itself with three teeth, its upper margin ridge-like and running into aperture with a tooth near lip edge and another further into aperture, and a larger rounded tooth on its lower margin set back from lip edge; 5) a low, inset transverse basal tooth to right of centre; 6) an in-running ridge-like basal tooth just to left of centre; 7–8) a large inset columella lamella set with two strong ridge-like teeth (there may also be a smaller bump below these, but it is not always obvious). Labral and basal teeth correspond with deep pits behind outer lip. Umbilicus widely open, elongate-oval, with a deep indentation underlying columella lamella (
<figureCitation id="F6922A75FFA7FFC4FC455639FBD1FDB8" box="[1002,1096,583,609]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[226,1361,279,1909]" captionTargetId="figure-20@4.[224,1363,265,1917]" captionTargetPageId="4" captionText="Fig. 2.Gulella benthodon van Bruggen, 1980 status revised (NMSA W25), length 3.4 mm, width 1.4 mm. A. Aperture view. B. Side view. C. Detail of aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420712" httpUri="https://zenodo.org/record/8420712/files/figure.png" pageId="5" pageNumber="6">Fig. 2D</figureCitation>
). Shell almost transparent when fresh, orange-red coloration of dried tissue of animal visible internally.
</paragraph>
<caption id="3AD66678FFA6FFC5FF1253D5FADCF831" ID-DOI="http://doi.org/10.5281/zenodo.8420712" ID-Zenodo-Dep="8420712" httpUri="https://zenodo.org/record/8420712/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1963,1989]" targetBox="[226,1361,279,1909]" targetPageId="4">
<paragraph id="6E1636F0FFA6FFC5FF1253D5FADCF831" blockId="4.[187,1399,1963,2025]" pageId="4" pageNumber="5">
<emphasis id="5CDDEAE2FFA6FFC5FF1253D5FE90F81C" bold="true" box="[189,265,1963,1989]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="A9A94D73FFA6FFC5FEA353D5FD51F81C" authority="van Bruggen, 1980" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[268,712,1963,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA6FFC5FEA353D5FE7CF81C" box="[268,485,1963,1989]" italics="true" pageId="4" pageNumber="5">Gulella benthodon</emphasis>
van 
<bibRefCitation id="0A384B01FFA6FFC5FDB453D2FD51F81C" author="Bruggen A. C. van" box="[539,712,1963,1990]" pageId="4" pageNumber="5" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen, 1980</bibRefCitation>
</taxonomicName>
status revised (NMSA W25), length 3.4 mm, width 1.4 mm. 
<emphasis id="5CDDEAE2FFA6FFC5FF1453B0FF49F831" bold="true" box="[187,208,1998,2024]" pageId="4" pageNumber="5">A</emphasis>
. Aperture view. 
<emphasis id="5CDDEAE2FFA6FFC5FE3C53B0FE3EF831" bold="true" box="[403,423,1998,2024]" pageId="4" pageNumber="5">B</emphasis>
. Side view. 
<emphasis id="5CDDEAE2FFA6FFC5FD9753B0FDD4F831" bold="true" box="[568,589,1998,2024]" pageId="4" pageNumber="5">C</emphasis>
. Detail of aperture. 
<emphasis id="5CDDEAE2FFA6FFC5FC9653B0FCD7F831" bold="true" box="[825,846,1998,2024]" pageId="4" pageNumber="5">D</emphasis>
. Oblique view of base showing umbilicus.
</paragraph>
</caption>
<paragraph id="6E1636F0FFA7FFC4FF1256D5FE2FFD1F" blockId="5.[189,438,683,710]" box="[189,438,683,710]" pageId="5" pageNumber="6">
<emphasis id="5CDDEAE2FFA7FFC4FF1256D5FECFFD1C" bold="true" box="[189,342,683,709]" pageId="5" pageNumber="6">Distribution</emphasis>
(
<figureCitation id="F6922A75FFA7FFC4FEC956D2FE35FD1F" box="[358,428,684,710]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[189,1398,901,1886]" captionTargetId="figure-219@5.[189,1398,901,1886]" captionTargetPageId="5" captionText="Fig. 3. Distribution map of Gulella benthodon van Bruggen, 1980 (blue triangles), G. judithmastersae sp. nov. (green squares), G. kevincolei sp. nov. (yellow circle) and G. mcmasteri sp. nov. (red diamonds) in the Amathole region of the Eastern Cape province. Contours at 1000 m and 2000 m." figureDoi="http://doi.org/10.5281/zenodo.8420714" httpUri="https://zenodo.org/record/8420714/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
)
</paragraph>
<paragraph id="6E1636F0FFA7FFC4FF1256A4FA81FD2D" blockId="5.[189,1304,730,757]" box="[189,1304,730,757]" pageId="5" pageNumber="6">
Endemic to the 
<taxonomicName id="A9A94D73FFA7FFC4FEDB56A4FDE9FD2D" authority="Mountains" authorityName="Mountains" box="[372,624,730,756]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="amathole">Amathole Mountains</taxonomicName>
; at altitudes between 
<quantity id="A9519B15FFA7FFC4FCC256A4FC5FFD2C" box="[877,966,730,757]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="m" value="1000.0">1000 m</quantity>
and 
<quantity id="A9519B15FFA7FFC4FC5156A4FBCFFD2C" box="[1022,1110,730,757]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="m" value="1100.0">1100 m</quantity>
above sea level.
</paragraph>
<caption id="3AD66678FFA7FFC4FF1253F6FB2FF830" ID-DOI="http://doi.org/10.5281/zenodo.8420714" ID-Zenodo-Dep="8420714" httpUri="https://zenodo.org/record/8420714/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1928,1954]" targetBox="[189,1398,901,1886]" targetPageId="5">
<paragraph id="6E1636F0FFA7FFC4FF1253F6FB2FF830" blockId="5.[188,1399,1928,2025]" pageId="5" pageNumber="6">
<emphasis id="5CDDEAE2FFA7FFC4FF1253F6FE95F87B" bold="true" box="[189,268,1928,1954]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
Distribution map of 
<taxonomicName id="A9A94D73FFA7FFC4FDA953F6FC55F87B" authority="van Bruggen, 1980" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[518,972,1928,1954]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA7FFC4FDA953F6FD7AF87B" box="[518,739,1928,1954]" italics="true" pageId="5" pageNumber="6">Gulella benthodon</emphasis>
van 
<bibRefCitation id="0A384B01FFA7FFC4FCB353F6FC55F87B" author="Bruggen A. C. van" box="[796,972,1928,1954]" pageId="5" pageNumber="6" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen, 1980</bibRefCitation>
</taxonomicName>
(blue triangles), 
<taxonomicName id="A9A94D73FFA7FFC4FB3853F6FAEEF87B" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[1175,1399,1928,1954]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="judithmastersae" status="sp. nov.">
<emphasis id="5CDDEAE2FFA7FFC4FB3853F6FAEEF87B" box="[1175,1399,1928,1954]" italics="true" pageId="5" pageNumber="6">G. judithmastersae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA7FFC4FF1253D2FE8EF81F" box="[189,279,1964,1990]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(green squares), 
<taxonomicName id="A9A94D73FFA7FFC4FE7253D2FDEEF81C" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[477,631,1963,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="kevincolei" status="sp. nov.">
<emphasis id="5CDDEAE2FFA7FFC4FE7253D2FDEEF81C" box="[477,631,1963,1990]" italics="true" pageId="5" pageNumber="6">G. kevincolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA7FFC4FDD053D2FD41F81F" box="[639,728,1964,1990]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(yellow circle) and 
<taxonomicName id="A9A94D73FFA7FFC4FC6F53D2FBC4F81C" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[960,1117,1964,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="mcmasteri" status="sp. nov.">
<emphasis id="5CDDEAE2FFA7FFC4FC6F53D2FBC4F81C" box="[960,1117,1964,1990]" italics="true" pageId="5" pageNumber="6">G. mcmasteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFA7FFC4FBCA53D2FB26F81F" box="[1125,1215,1964,1990]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(red diamonds) in the Amathole region of the Eastern Cape province. Contours at 1000 m and 2000 m.
</paragraph>
</caption>
<paragraph id="6E1636F0FFA4FFC7FF125576FE84FEFB" blockId="6.[189,285,264,290]" box="[189,285,264,290]" pageId="6" pageNumber="7">
<heading id="355E819CFFA4FFC7FF125576FE84FEFB" bold="true" box="[189,285,264,290]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5CDDEAE2FFA4FFC7FF125576FE84FEFB" bold="true" box="[189,285,264,290]" pageId="6" pageNumber="7">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA4FFC7FF125548FED8FEAA" blockId="6.[189,1398,310,371]" pageId="6" pageNumber="7">
<taxonomicName id="A9A94D73FFA4FFC7FF125548FE75FE88" authority="Mistbelt Forest" authorityName="Mistbelt Forest" box="[189,492,310,337]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="amathole">Amathole Mistbelt Forest</taxonomicName>
(Southern Mistbelt Forest group) (
<bibRefCitation id="0A384B01FFA4FFC7FC245549FB38FE88" author="Von Maltitz G. &amp; Mucina L. &amp; Geldenhuys C. &amp; Lawes M. &amp; Eeley H. &amp; Adie H. &amp; Vink D. &amp; Fleming G. &amp; Bailey C." box="[907,1185,310,337]" pageId="6" pageNumber="7" refId="ref14670" refString="Von Maltitz G., Mucina L., Geldenhuys C., Lawes M., Eeley H., Adie H., Vink D., Fleming G. &amp; Bailey C. 2003. Classification System for South African Indigenous Forests. CSIR, Pretoria." type="book" year="2003">
von Maltitz 
<emphasis id="5CDDEAE2FFA4FFC7FBB05549FBCAFE89" box="[1055,1107,310,336]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2003
</bibRefCitation>
); in leaf-litter and under logs.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFA4FFC7FF1255E4FC41FAFD" pageId="6" pageNumber="7" type="discussion">
<paragraph id="6E1636F0FFA4FFC7FF1255E4FEB6FE6D" blockId="6.[189,303,410,436]" box="[189,303,410,436]" pageId="6" pageNumber="7">
<heading id="355E819CFFA4FFC7FF1255E4FEB6FE6D" bold="true" box="[189,303,410,436]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5CDDEAE2FFA4FFC7FF1255E4FEB6FE6D" bold="true" box="[189,303,410,436]" pageId="6" pageNumber="7">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA4FFC7FF1255B7FAD5FD0C" blockId="6.[188,1399,456,725]" pageId="6" pageNumber="7">
We treat 
<taxonomicName id="A9A94D73FFA4FFC7FE8555B6FD9DFE3B" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[298,516,456,482]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA4FFC7FE8555B6FD9DFE3B" box="[298,516,456,482]" italics="true" pageId="6" pageNumber="7">Gulella benthodon</emphasis>
</taxonomicName>
as a separate species and not as a subspecies of 
<taxonomicName id="A9A94D73FFA4FFC7FBFF55B6FE19FDDC" authority="(Melvill &amp; Ponsonby, 1908)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="darglensis">
<emphasis id="5CDDEAE2FFA4FFC7FBFF55B6FB77FE3B" box="[1104,1262,456,482]" italics="true" pageId="6" pageNumber="7">G. darglensis</emphasis>
(Melvill &amp; Ponsonby, 1908)
</taxonomicName>
as originally proposed. It bears very little resemblance to the other two subspecies of 
<taxonomicName id="A9A94D73FFA4FFC7FF125670FEC1FDFE" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" box="[189,344,525,552]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="darglensis">
<emphasis id="5CDDEAE2FFA4FFC7FF125670FEC1FDFE" box="[189,344,525,552]" italics="true" pageId="6" pageNumber="7">G. darglensis</emphasis>
</taxonomicName>
recognised by van 
<bibRefCitation id="0A384B01FFA4FFC7FD955670FD6FFDF1" author="Bruggen A. C. van" box="[570,758,526,552]" pageId="6" pageNumber="7" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen (1980)</bibRefCitation>
. It has a large mid-labral complex with three cusps and not a pair of in-running ridge-like labral teeth. The columella lamella also has two strong ridge-like teeth, but both are inset, while in 
<taxonomicName id="A9A94D73FFA4FFC7FE43562DFD1CFDB4" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" box="[492,645,595,621]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="darglensis">
<emphasis id="5CDDEAE2FFA4FFC7FE43562DFD1CFDB4" box="[492,645,595,621]" italics="true" pageId="6" pageNumber="7">G. darglensis</emphasis>
</taxonomicName>
the lower tooth extends almost to the lip edge. The deeply situated columellar teeth were also noted by van 
<bibRefCitation id="0A384B01FFA4FFC7FD0E5608FCC2FD49" author="Bruggen A. C. van" box="[673,859,630,656]" pageId="6" pageNumber="7" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen (1980)</bibRefCitation>
and were the origin of the subspecific name. 
<taxonomicName id="A9A94D73FFA4FFC7FF1256E6FE09FD6B" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[189,400,664,690]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA4FFC7FF1256E6FE09FD6B" box="[189,400,664,690]" italics="true" pageId="6" pageNumber="7">Gulella benthodon</emphasis>
</taxonomicName>
has stronger and coarser sculpture than both subspecies of 
<taxonomicName id="A9A94D73FFA4FFC7FBB056E6FB21FD6B" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" box="[1055,1208,664,690]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="darglensis">
<emphasis id="5CDDEAE2FFA4FFC7FBB056E6FB21FD6B" box="[1055,1208,664,690]" italics="true" pageId="6" pageNumber="7">G. darglensis</emphasis>
</taxonomicName>
; 
<emphasis id="5CDDEAE2FFA4FFC7FB6A56E6FB65FD6B" box="[1221,1276,664,690]" italics="true" pageId="6" pageNumber="7">G. d.</emphasis>
<taxonomicName id="A9A94D73FFA4FFC7FAAF56E6FD99FD0C" authority="(Melvill &amp; Ponsonby, 1908)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="darglensis">
<emphasis id="5CDDEAE2FFA4FFC7FAAF56E6FAEEFD6B" box="[1280,1399,664,690]" italics="true" pageId="6" pageNumber="7">darglensis</emphasis>
(Melvill &amp; Ponsonby, 1908)
</taxonomicName>
has close-set axial riblets and 
<emphasis id="5CDDEAE2FFA4FFC7FCF356C5FB8EFD0C" box="[860,1047,699,725]" italics="true" pageId="6" pageNumber="7">G. d. illovoensis</emphasis>
(Burnup, 1914) is smooth.
</paragraph>
<paragraph id="6E1636F0FFA4FFC7FF125682FCA8FBD1" blockId="6.[188,1399,764,1033]" pageId="6" pageNumber="7">
The apertural dentition of 
<taxonomicName id="A9A94D73FFA4FFC7FE575682FD4CFCCF" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[504,725,764,790]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA4FFC7FE575682FD4CFCCF" box="[504,725,764,790]" italics="true" pageId="6" pageNumber="7">Gulella benthodon</emphasis>
</taxonomicName>
closely resembles that of 
<taxonomicName id="A9A94D73FFA4FFC7FBA15682FF63FCE1" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="kenbrowni">
<emphasis id="5CDDEAE2FFA4FFC7FBA15682FB36FCCF" box="[1038,1199,764,790]" italics="true" pageId="6" pageNumber="7">G. kenbrowni</emphasis>
<bibRefCitation id="0A384B01FFA4FFC7FB185682FF63FCE1" author="Cole M. L. &amp; Herbert D. G." pageId="6" pageNumber="7" pagination="1 - 32" refId="ref13071" refString="Cole M. L. &amp; Herbert D. G. 2022. Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae) European Journal of Taxonomy 813: 1 - 32. https: // doi. org / 10.5852 / ejt. 2022.813.1729" type="journal article" year="2022">Cole &amp; Herbert, 2022</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="A9A94D73FFA4FFC7FE9D5761FD5BFCE1" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[306,706,798,825]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="fordycei">
<emphasis id="5CDDEAE2FFA4FFC7FE9D5761FE2CFCE1" box="[306,437,798,825]" italics="true" pageId="6" pageNumber="7">G. fordycei</emphasis>
<bibRefCitation id="0A384B01FFA4FFC7FE115760FD5BFCE1" author="Cole M. L. &amp; Herbert D. G." box="[446,706,798,824]" pageId="6" pageNumber="7" pagination="1 - 32" refId="ref13071" refString="Cole M. L. &amp; Herbert D. G. 2022. Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae) European Journal of Taxonomy 813: 1 - 32. https: // doi. org / 10.5852 / ejt. 2022.813.1729" type="journal article" year="2022">Cole &amp; Herbert, 2022</bibRefCitation>
</taxonomicName>
, but 
<taxonomicName id="A9A94D73FFA4FFC7FD535761FC05FCE1" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[764,924,798,825]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA4FFC7FD535761FC05FCE1" box="[764,924,798,825]" italics="true" pageId="6" pageNumber="7">G. benthodon</emphasis>
</taxonomicName>
stands out at a glance by being strongly ribbed and considerably larger. The multi-toothed columella lamella is also reminiscent of 
<taxonomicName id="A9A94D73FFA4FFC7FB6C573FFEEAFCA7" authority="Connolly, 1939" authorityName="Connolly" authorityYear="1939" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="aprosdoketa">
<emphasis id="5CDDEAE2FFA4FFC7FB6C573FFAEDFC82" box="[1219,1396,833,859]" italics="true" pageId="6" pageNumber="7">G. aprosdoketa</emphasis>
Connolly, 1939
</taxonomicName>
, 
<taxonomicName id="A9A94D73FFA4FFC7FED1571AFC8AFCA7" authority="Cole &amp; Herbert, 2009" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[382,787,868,894]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="bomvana">
<emphasis id="5CDDEAE2FFA4FFC7FED1571AFD97FCA7" box="[382,526,868,894]" italics="true" pageId="6" pageNumber="7">G. bomvana</emphasis>
<bibRefCitation id="0A384B01FFA4FFC7FDBB571AFC8AFCA7" author="Cole M. L. &amp; Herbert D. G." box="[532,787,868,894]" pageId="6" pageNumber="7" pagination="547 - 564" refId="ref13017" refString="Cole M. L. &amp; Herbert D. G. 2009. Description of four new species of Gulella Pfeiffer, 1856 from Eastern Cape, South Africa, with additional notes on two poorly known species (Mollusca: Eupulmonata: Streptaxidae). Zoologische Mededelingen 83: 547 - 564." type="journal article" year="2009">Cole &amp; Herbert, 2009</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="A9A94D73FFA4FFC7FCE5571AFB5BFCA7" authority="Cole &amp; Herbert, 2009" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[842,1218,868,894]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="tietzae">
<emphasis id="5CDDEAE2FFA4FFC7FCE5571AFC23FCA4" box="[842,954,868,894]" italics="true" pageId="6" pageNumber="7">G. tietzae</emphasis>
<bibRefCitation id="0A384B01FFA4FFC7FC6E571AFB5BFCA7" author="Cole M. L. &amp; Herbert D. G." box="[961,1218,868,894]" pageId="6" pageNumber="7" pagination="547 - 564" refId="ref13017" refString="Cole M. L. &amp; Herbert D. G. 2009. Description of four new species of Gulella Pfeiffer, 1856 from Eastern Cape, South Africa, with additional notes on two poorly known species (Mollusca: Eupulmonata: Streptaxidae). Zoologische Mededelingen 83: 547 - 564." type="journal article" year="2009">Cole &amp; Herbert, 2009</bibRefCitation>
</taxonomicName>
, but these three species are smooth. The labral complex resembles that of several very small, narrow-range species endemic to Zululand, 
<taxonomicName id="A9A94D73FFA4FFC7FE1257D7FC54FC1A" authority="(Melvill &amp; Ponsonby, 1903)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1903" box="[445,973,937,963]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="genialis">
<emphasis id="5CDDEAE2FFA4FFC7FE1257D7FDE0FC1A" box="[445,633,937,963]" italics="true" pageId="6" pageNumber="7">Gulella genialis</emphasis>
(Melvill &amp; Ponsonby, 1903)
</taxonomicName>
, 
<taxonomicName id="A9A94D73FFA4FFC7FC7657D7FADCFC1D" authority="Burnup, 1925" authorityName="Burnup" authorityYear="1925" box="[985,1349,937,964]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="laevorsa">
<emphasis id="5CDDEAE2FFA4FFC7FC7657D7FB04FC1A" box="[985,1181,937,963]" italics="true" pageId="6" pageNumber="7">Gulella laevorsa</emphasis>
Burnup, 1925
</taxonomicName>
and 
<taxonomicName id="A9A94D73FFA4FFC7FF1257B2FD54FC3F" authority="(Melvill &amp; Ponsonby, 1907)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1907" box="[189,717,972,998]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="vallaris">
<emphasis id="5CDDEAE2FFA4FFC7FF1257B2FEEEFC3F" box="[189,375,972,998]" italics="true" pageId="6" pageNumber="7">Gulella vallaris</emphasis>
(Melvill &amp; Ponsonby, 1907)
</taxonomicName>
in Group 9 of 
<bibRefCitation id="0A384B01FFA4FFC7FC2F57B2FB2CFC3F" author="Herbert D. &amp; Kilburn D." box="[896,1205,972,998]" pageId="6" pageNumber="7" refId="ref13724" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg, South Africa." type="book" year="2004">Herbert &amp; Kilburn (2004)</bibRefCitation>
, but none of the latter species have a multi-toothed columella lamella.
</paragraph>
<paragraph id="6E1636F0FFA4FFC7FF125051FEFCFB90" blockId="6.[189,357,1071,1097]" box="[189,357,1071,1097]" pageId="6" pageNumber="7">
<heading id="355E819CFFA4FFC7FF125051FEFCFB90" bold="true" box="[189,357,1071,1097]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5CDDEAE2FFA4FFC7FF125051FEFCFB90" bold="true" box="[189,357,1071,1097]" pageId="6" pageNumber="7">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFA4FFC7FF125023FC41FAFD" blockId="6.[186,1400,1117,1317]" pageId="6" pageNumber="7">
<taxonomicName id="A9A94D73FFA4FFC7FF125023FE0CFBAE" authorityName="van Bruggen" authorityYear="1980" baseAuthorityName="van Bruggen" baseAuthorityYear="1980" box="[189,405,1117,1143]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="benthodon">
<emphasis id="5CDDEAE2FFA4FFC7FF125023FE0CFBAE" box="[189,405,1117,1143]" italics="true" pageId="6" pageNumber="7">Gulella benthodon</emphasis>
</taxonomicName>
is a narrow-range endemic found only in the 
<taxonomicName id="A9A94D73FFA4FFC7FC3D5023FB1EFBAE" authority="Mountains" authorityName="Mountains" box="[914,1159,1117,1143]" class="Gastropoda" family="Cyclophoridae" genus="Chondrocyclus" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="amathole">Amathole Mountains</taxonomicName>
where, despite fairly extensive collecting efforts throughout the mountain range, it has only been found at three localities. Van 
<bibRefCitation id="0A384B01FFA4FFC7FF1350DDFE12FB64" author="Bruggen A. C. van" box="[188,395,1187,1213]" pageId="6" pageNumber="7" pagination="1 - 62" refId="ref12623" refString="Bruggen A. C. van. 1980. Size clines and subspecies in the streptaxid genus Gulella Pfr. (Mollusca, Gastopoda Pulmonata) in southern Africa. Zoologische Verhandelingen 180: 1 - 62. Available from https: // repository. naturalis. nl / pub / 317764 [accessed 31 Aug. 2023]." type="journal article" year="1980">Bruggen’s (1980)</bibRefCitation>
specimens came from the Pirie Forest near King Williams Town, collected in about 1920. Recently collected specimens are known only from Hogsback and the majority were collected in the Auckland Nature Reserve. In spite of these being protected sites, poaching and illegal harvesting of plant products takes place and there is uncontrolled access of cattle.
</paragraph>
</subSubSection>
</treatment>
</document>