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<document id="2EC6E4C1CB98784E0C5EB13368A59C96" ID-DOI="10.11646/zootaxa.3884.1.5" ID-GBIF-Dataset="1b9a7949-14c3-4367-a7b9-b125a92597c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="287450" ID-ZooBank="0DFB8B35-C265-4E96-876A-3B9BB051DF57" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460964658797" checkinUser="plazi" docAuthor="Meyer, Kirstin S., Cumberlidge, Neil &amp; Koppin, Jennifer C." docDate="2014" docId="A9132D0CFFF6A11225B8DA58FE5CFEF2" docLanguage="en" docName="zt03884p072.pdf" docOrigin="Zootaxa 3884 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glabrithelphusa Meyer, Cumberlidge &amp; Koppin, 2014, n. gen." docType="treatment" docVersion="6" lastPageNumber="67" masterDocId="552A5574FFF7A110252FD86FFFA9FF97" masterDocTitle="A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae)" masterLastPageNumber="72" masterPageNumber="65" pageNumber="66" updateTime="1698557657870" updateUser="plazi">
<mods:mods id="F0B8DFBB4DF4027EC6AC53512DD8B985" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="13A2F05088ECEAD2C298147C70C8A10B">
<mods:title id="FE218C76416B88EB3FF5F789C4114816">A new genus and species of freshwater crab from Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae)</mods:title>
</mods:titleInfo>
<mods:name id="2585F7FC6D4861E7F3C538A26C55DDE9" type="personal">
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<mods:namePart id="F7BD7B8BBB50E189993E4A5738F8031B">Meyer, Kirstin S.</mods:namePart>
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<mods:namePart id="325B4D2B77624989C392F47E8053EFB6">Cumberlidge, Neil</mods:namePart>
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<mods:namePart id="9F13F1370B297C7A2C341F30A78E93CF">Koppin, Jennifer C.</mods:namePart>
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<mods:typeOfResource id="3323D8D95EA51545FBD0C10AACA960C0">text</mods:typeOfResource>
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<mods:title id="8ACDBAAA375F41CD6F534E85B79B334A">Zootaxa</mods:title>
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<mods:part id="CA770FB26D26E367BA4168021E1811BE">
<mods:date id="141DEDF69E8E59EECEC92E65BE240F78">2014</mods:date>
<mods:detail id="19E63CFDB9DDFF9A6B2532B463AF92C9" type="volume">
<mods:number id="B0E70DF533AAC70EAB5753FAC2475B5C">3884</mods:number>
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<mods:detail id="CB045EC13477ADE375E6560772A434D2" type="issue">
<mods:number id="04CD75C5785DCE696ACEE82F14A4225A">1</mods:number>
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<mods:classification id="70BC91D6086C8BD1C7660DDC9F3B9531">journal article</mods:classification>
<mods:identifier id="E2D3AEEAE65A91E2789F999D881E210C" type="DOI">10.11646/zootaxa.3884.1.5</mods:identifier>
<mods:identifier id="FD05C088D7196A23816CB84DFE262395" type="GBIF-Dataset">1b9a7949-14c3-4367-a7b9-b125a92597c6</mods:identifier>
<mods:identifier id="6489B4E12B7FDC6B591582E2A425C298" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="666B3849B0CFF31667324373A31B8092" type="Zenodo-Dep">287450</mods:identifier>
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<treatment id="A9132D0CFFF6A11225B8DA58FE5CFEF2" ID-DOI="http://doi.org/10.5281/zenodo.6139263" ID-GBIF-Taxon="127670770" ID-Zenodo-Dep="6139263" LSID="urn:lsid:plazi:treatment:A9132D0CFFF6A11225B8DA58FE5CFEF2" httpUri="http://treatment.plazi.org/id/A9132D0CFFF6A11225B8DA58FE5CFEF2" lastPageId="2" lastPageNumber="67" pageId="1" pageNumber="66">
<subSubSection id="69A0CF91FFF6A11125B8DA58FEBBFDE4" pageId="1" pageNumber="66" type="nomenclature">
<paragraph id="21059C1AFFF6A11125B8DA58FE1CFDC5" blockId="1.[151,437,567,627]" box="[151,437,567,594]" pageId="1" pageNumber="66">
<heading id="7A4D2B76FFF6A11125B8DA58FE1CFDC5" bold="true" box="[151,437,567,594]" fontSize="11" level="1" pageId="1" pageNumber="66" reason="1">
<emphasis id="13CE4008FFF6A11125B8DA58FE1CFDC5" bold="true" box="[151,437,567,594]" pageId="1" pageNumber="66">
<taxonomicName id="E6BAE799FFF6A11125B8DA58FEF5FDC6" box="[151,348,567,593]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A11125B8DA58FEF5FDC6" bold="true" box="[151,348,567,593]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08FDFD73FFF6A111244CDA57FE1CFDC5" box="[355,437,568,594]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="21059C1AFFF6A11125B8DA35FEBBFDE4" blockId="1.[151,437,567,627]" box="[151,274,602,627]" pageId="1" pageNumber="66">
(
<figureCitation id="B981809FFFF6A11125B0DA35FEA0FDE4" box="[159,265,602,627]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1862,1884]" captionStartId-1="3.[151,250,1469,1491]" captionStartId-2="4.[151,250,1877,1899]" captionStartId-3="5.[151,250,664,686]" captionTargetBox-0="[179,1416,430,1790]" captionTargetBox-1="[311,1265,206,1428]" captionTargetBox-2="[226,1344,230,1804]" captionTargetBox-3="[152,1432,222,617]" captionTargetId-0="figure@2.[162,1435,411,1801]" captionTargetId-1="figure@3.[305,1282,195,1447]" captionTargetId-2="figure@4.[151,1436,193,1856]" captionTargetId-3="figure@5.[151,1436,221,620]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." captionText-1="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." captionText-2="FIGURE 3. Glabrithelphusa angene n. gen n. sp.. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). A, dorsal view; B, sternal view; C, frontal view. Scale bar: 14 mm." captionText-3="FIGURE 4. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). A, left G 1, dorsal view; B, left G 1, ventral view; C, terminal article of left G 1, close-up, dorsal view; D, terminal article of left G 1, closeup, ventral view; E, left G 2, ventral view. Scale bar: 5 mm." httpUri-0="https://zenodo.org/record/287451/files/figure.png" httpUri-1="https://zenodo.org/record/287452/files/figure.png" httpUri-2="https://zenodo.org/record/287453/files/figure.png" httpUri-3="https://zenodo.org/record/287454/files/figure.png" pageId="1" pageNumber="66">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="69A0CF91FFF6A11125B8DACDFC9EFD2C" box="[151,823,674,699]" pageId="1" pageNumber="66" type="materials_examined">
<paragraph id="21059C1AFFF6A11125B8DACDFC9EFD2C" blockId="1.[151,1437,674,2031]" box="[151,823,674,699]" pageId="1" pageNumber="66">
<emphasis id="13CE4008FFF6A11125B8DACDFE9BFD2C" bold="true" box="[151,306,674,699]" pageId="1" pageNumber="66">
<typeStatus id="FE0122B8FFF6A11125B8DACDFF7BFD2C" box="[151,210,674,699]" pageId="1" pageNumber="66">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="E6BAE799FFF6A1112416DACDFDE1FD2C" box="[313,584,674,699]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="angene" status="sp. nov.">
<emphasis id="13CE4008FFF6A1112416DACDFDE1FD2C" box="[313,584,674,699]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa angene</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A1112760DACDFD27FD2C" bold="true" box="[591,654,674,699]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A1112760DACDFD27FD2C" box="[591,654,674,699]" pageId="1" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="69A0CF91FFF6A11125E8DAA9FD8BFCB0" pageId="1" pageNumber="66" type="etymology">
<paragraph id="21059C1AFFF6A11125E8DAA9FD8BFCB0" blockId="1.[151,1437,674,2031]" pageId="1" pageNumber="66">
<emphasis id="13CE4008FFF6A11125E8DAA9FEE7FD48" bold="true" box="[199,334,710,735]" pageId="1" pageNumber="66">Etymology.</emphasis>
The name
<taxonomicName id="E6BAE799FFF6A11124E1DAA9FD2CFD48" box="[462,645,710,735]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A11124E1DAA9FD2CFD48" box="[462,645,710,735]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
refers to the characteristic smooth (
<emphasis id="13CE4008FFF6A1112130DAA9FB28FD48" box="[1055,1153,710,735]" italics="true" pageId="1" pageNumber="66">glabrous</emphasis>
in Latin) and untextured nature of the dorsal carapace of this taxon, in suite with the traditional freshwater crab genus name
<taxonomicName id="E6BAE799FFF6A11121DEDA85FACCFC94" box="[1265,1381,746,771]" class="Malacostraca" family="Potamonautidae" genus="Thelphusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11121DEDA85FACCFC94" box="[1265,1381,746,771]" italics="true" pageId="1" pageNumber="66">Thelphusa</emphasis>
</taxonomicName>
, and is a masculine noun in the singular.
</paragraph>
</subSubSection>
<subSubSection id="69A0CF91FFF6A11125E8DB5DFE76FBFC" pageId="1" pageNumber="66" type="diagnosis">
<paragraph id="21059C1AFFF6A11125E8DB5DFE76FBFC" blockId="1.[151,1437,674,2031]" pageId="1" pageNumber="66">
<emphasis id="13CE4008FFF6A11125E8DB5DFEE8FCDC" bold="true" box="[199,321,818,843]" pageId="1" pageNumber="66">Diagnosis.</emphasis>
Mandibular palp 2-segmented with small, hard, anterior lobe at junction between segments, 0.25x length of terminal segment (
<figureCitation id="B981809FFFF6A11124FFDB39FDBAFCF8" box="[464,531,854,879]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
D,E). Entire dorsal surface of carapace smooth; cervical grooves short; suborbital region smooth; postfrontal crest faint, incomplete, epibranchial, exorbital teeth weak, low; space or cleft between epibranchial tooth and exorbital tooth reduced to small but distinct shallow notch (
<figureCitation id="B981809FFFF6A111216BDBF1FB2DFC20" box="[1092,1156,926,951]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A, 3A); third maxilliped with deep vertical sulcus on ischium (
<figureCitation id="B981809FFFF6A111277DDBADFD3DFC4C" box="[594,660,962,987]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
C). Sternal sulcus s3/s4, faint, reduced to 2 side-notches (
<figureCitation id="B981809FFFF6A1112018DBADFAD4FC4C" box="[1335,1405,962,987]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
F). Major cheliped with 3 unfused molars (
<figureCitation id="B981809FFFF6A111274FDB89FD0BFC68" box="[608,674,998,1023]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
F). G1 terminal article with conspicuously raised lobe on superior margin, distal third of G1 subterminal segment curved outward (
<figureCitation id="B981809FFFF6A11126BCDC65FC72FBB4" box="[915,987,1034,1059]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
A,B, 4A,B,C,D); G2 terminal segment flagellum-like, straight, no distal curve (
<figureCitation id="B981809FFFF6A111275FDC41FD1EFBD0" box="[624,695,1070,1095]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
C, 4E). Walking legs (p2‒p5) normal length, not elongated (p5 merus/CW = 0.28) (
<figureCitation id="B981809FFFF6A1112458DC3DFE13FBFC" box="[375,442,1106,1131]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[226,1344,230,1804]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Glabrithelphusa angene n. gen n. sp.. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). A, dorsal view; B, sternal view; C, frontal view. Scale bar: 14 mm." httpUri="https://zenodo.org/record/287453/files/figure.png" pageId="1" pageNumber="66">Fig. 3</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="69A0CF91FFF6A11225E8DC19FDC7FED7" lastPageId="2" lastPageNumber="67" pageId="1" pageNumber="66" type="discussion">
<paragraph id="21059C1AFFF6A11125E8DC19FDB9F93C" blockId="1.[151,1437,674,2031]" pageId="1" pageNumber="66">
<emphasis id="13CE4008FFF6A11125E8DC19FE92FB18" bold="true" box="[199,315,1142,1167]" pageId="1" pageNumber="66">Remarks.</emphasis>
<taxonomicName id="E6BAE799FFF6A111246ADC19FE54FB18" box="[325,509,1142,1167]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A111246ADC19FE54FB18" box="[325,509,1142,1167]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A1112727DC19FDF3FB18" bold="true" box="[520,602,1142,1167]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A1112727DC19FDF3FB18" box="[520,602,1142,1167]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
is assigned to the
<taxonomicName id="E6BAE799FFF6A1112613DC19FBD1FB18" authority="Bott, 1970" authorityName="Bott" authorityYear="1970" box="[828,1144,1142,1167]" class="Malacostraca" family="Potamonautidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="family">
Potamonautidae
<bibRefCitation id="452BE1EBFFF6A11126D4DC18FBD1FB18" author="Bott" box="[1019,1144,1143,1167]" pageId="1" pageNumber="66" refString="Bott, R. (1970) Betrachtungen uber die Entwicklungsgeschichte und Verbreitung der Sußwasser-Krabben nach der Sammlung des Naturhistorischen Museums in Genf / Schweiz. Revue suisse de Zoologie, 77 (2), 327 - 344." type="journal article" year="1970">Bott, 1970</bibRefCitation>
</taxonomicName>
, because of the common possession of a suite of diagnostic familial characters, including a 2-segmented mandibular palp (
<figureCitation id="B981809FFFF6A111202CDCF5FAE3FB24" box="[1283,1354,1178,1203]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
D,E), a triangular abdomen (
<figureCitation id="B981809FFFF6A11124A9DCD1FE65FB40" box="[390,460,1214,1239]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
F), and a G1 with a medium-length terminal article (
<figureCitation id="B981809FFFF6A1112101DCD1FBDDFB40" box="[1070,1140,1214,1239]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
A,B, 4A,B) (
<bibRefCitation id="452BE1EBFFF6A1112027DCD1FF7AFB6C" author="Cumberlidge" pageId="1" pageNumber="66" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. Faune et Flore tropicales 35. IRD, Paris, 382 pp." type="book" year="1999">Cumberlidge 1999</bibRefCitation>
).
<taxonomicName id="E6BAE799FFF6A11125C7DC8DFE09FB6C" box="[232,416,1250,1275]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A11125C7DC8DFE09FB6C" box="[232,416,1250,1275]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
can be distinguished from the other
<collectingCountry id="59ADDC8AFFF6A1112667DC8DFC1CFB6C" box="[840,949,1250,1275]" name="Madagascar" pageId="1" pageNumber="66">Malagasy</collectingCountry>
freshwater crab genera by the form of the mandibles. Although all potamonautid crabs possess a 2-segmented mandibular palp within this family, there is a great deal of variation of mandible characters, especially among the crabs found in
<collectingCountry id="59ADDC8AFFF6A1112141DD45FB5CFAD4" box="[1134,1269,1322,1347]" name="Madagascar" pageId="1" pageNumber="66">Madagascar</collectingCountry>
(
<bibRefCitation id="452BE1EBFFF6A1112027DD45FF7AFAF0" author="Cumberlidge" pageId="1" pageNumber="66" refString="Cumberlidge, N. (1999) The Freshwater Crabs of West Africa, Family Potamonautidae. Faune et Flore tropicales 35. IRD, Paris, 382 pp." type="book" year="1999">Cumberlidge 1999</bibRefCitation>
;
<bibRefCitation id="452BE1EBFFF6A11125F1DD21FDEFFAF0" author="Cumberlidge" box="[222,582,1358,1383]" pageId="1" pageNumber="66" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">Cumberlidge &amp; Sternberg 2002</bibRefCitation>
).
<bibRefCitation id="452BE1EBFFF6A1112776DD21FBEEFAF0" author="Cumberlidge" box="[601,1095,1358,1383]" pageId="1" pageNumber="66" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">
Cumberlidge &amp; Sternberg (2002
<figureCitation id="B981809FFFF6A11126E2DD21FBA7FAF0" box="[973,1038,1358,1383]" captionStart="FIGURE 4" captionStartId="5.[151,250,664,686]" captionTargetBox="[152,1432,222,617]" captionTargetId="figure@5.[151,1436,221,620]" captionTargetPageId="5" captionText="FIGURE 4. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). A, left G 1, dorsal view; B, left G 1, ventral view; C, terminal article of left G 1, close-up, dorsal view; D, terminal article of left G 1, closeup, ventral view; E, left G 2, ventral view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/287454/files/figure.png" pageId="1" pageNumber="66">Fig. 4</figureCitation>
A-L)
</bibRefCitation>
grouped
<collectingCountry id="59ADDC8AFFF6A111219EDD21FAB7FAF0" box="[1201,1310,1358,1383]" name="Madagascar" pageId="1" pageNumber="66">Malagasy</collectingCountry>
freshwater crabs based on mandibular palp characters as follows. Taxa with a simple mandible, completely lacking a lobe of any kind at the junction between the segments (
<taxonomicName id="E6BAE799FFF6A1112790DDF9FBADFA38" authority="Bott, 1965" authorityName="Bott" authorityYear="1965" box="[703,1028,1430,1455]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A1112790DDF9FCD7FA38" box="[703,894,1430,1455]" italics="true" pageId="1" pageNumber="66">Madagapotamon</emphasis>
<bibRefCitation id="452BE1EBFFF6A11126A7DDF8FBADFA38" author="Bott" box="[904,1028,1430,1455]" pageId="1" pageNumber="66" refString="Bott, R. (1965) Die Sußwasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle, Paris, 37 (2), 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
</taxonomicName>
); taxa with a small, hard ledge-like anterior lobe on the mandible about 0.1 times as long as the terminal segment (
<taxonomicName id="E6BAE799FFF6A1112139DDD5FA31FA44" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[1046,1432,1466,1491]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A1112139DDD5FB06FA44" box="[1046,1199,1466,1491]" italics="true" pageId="1" pageNumber="66">Skelosophusa</emphasis>
Ng &amp;
<bibRefCitation id="452BE1EBFFF6A11121D1DDD5FA31FA44" author="Takeda" box="[1278,1432,1466,1491]" pageId="1" pageNumber="66" refString="Ng, P. K. L. &amp; Takeda, M. (1994) Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Series A (Zoology), 20 (4), 161 - 172." type="journal article" year="1994">Takeda, 1994</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="E6BAE799FFF6A11125E4DDB1FDC0FA60" authority="Cumberlidge, 2010" authorityName="Cumberlidge" authorityYear="2010" box="[203,617,1502,1527]" class="Malacostraca" family="Potamonautidae" genus="Boreathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11125E4DDB1FED2FA60" box="[203,379,1502,1527]" italics="true" pageId="1" pageNumber="66">Boreathelphusa</emphasis>
<bibRefCitation id="452BE1EBFFF6A11124A8DDB1FDC0FA60" author="Cumberlidge" box="[391,617,1502,1527]" pageId="1" pageNumber="66" refString="Cumberlidge, N. (2010) Boreathelphusa, a replacement name for Boreas Cumberlidge and Sternberg, 2002 (Crustacea: Brachyura: Potamonautidae), preoccupied by Boreas Morris, 1980 (Bryozoa: Hippothoidae). Zootaxa, 2450, 68." type="journal article" year="2010">Cumberlidge, 2010</bibRefCitation>
</taxonomicName>
); and taxa with a clearly bilobed terminal segment with a substantial medium-sized anterior lobe about 0.5 times as long as the terminal segment (
<taxonomicName id="E6BAE799FFF6A1112128DE6DFF7BF9A8" authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A1112128DE6DFB12F98C" box="[1031,1211,1538,1563]" italics="true" pageId="1" pageNumber="66">Hydrothelphusa</emphasis>
<bibRefCitation id="452BE1EBFFF6A11121ECDE6CFF7BF9A8" author="Milne-Edwards" pageId="1" pageNumber="66" refString="Milne-Edwards, A. (1872) Note sur les crabes d'eau douce de Madagascar. Bibliographie Ecole Hautes Etudes (Section Sciences naturelles) Paris, 5 (8), 1 - 3." type="journal article" year="1872">A. Milne-Edwards, 1872</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E6BAE799FFF6A11125F3DE48FD5CF9A8" authority="Cumberlidge, Boyko" authorityName="Cumberlidge, Boyko" authorityYear="2002" box="[220,757,1574,1599]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11125F3DE48FEE8F9A9" box="[220,321,1575,1598]" italics="true" pageId="1" pageNumber="66">Marojejy</emphasis>
<bibRefCitation id="452BE1EBFFF6A1112467DE49FD5CF9A8" author="Cumberlidge" box="[328,757,1574,1599]" pageId="1" pageNumber="66" refString="Cumberlidge, N., Boyko, C. B. &amp; Harvey, A. W. (2002) A new genus and species of freshwater crab (Decapoda, Crustacea, Potamoidea) from northern Madagascar, and a second new species associated with Pandanus leaf axils. Journal of Natural History, 36 (1), 65 - 77. http: // dx. doi. org / 10.1080 / 00222930010003800" type="journal article" year="2002">Cumberlidge, Boyko, &amp; Harvey, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E6BAE799FFF6A11127D0DE49FB42F9A8" authority="Cumberlidge &amp; Sternberg, 2002" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[767,1259,1574,1599]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11127D0DE49FCD5F9A8" box="[767,892,1574,1599]" italics="true" pageId="1" pageNumber="66">Malagasya</emphasis>
<bibRefCitation id="452BE1EBFFF6A11126ADDE49FB42F9A8" author="Cumberlidge" box="[898,1259,1574,1599]" pageId="1" pageNumber="66" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">Cumberlidge &amp; Sternberg, 2002</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="E6BAE799FFF6A111200BDE48FE3AF9F4" authority="Reed &amp; Cumberlidge, 2006" authorityName="Reed &amp; Cumberlidge" authorityYear="2006" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A111200BDE48FAF5F9A9" box="[1316,1372,1575,1598]" italics="true" pageId="1" pageNumber="66">Foza</emphasis>
<bibRefCitation id="452BE1EBFFF6A111204DDE49FE3AF9F4" author="Reed" pageId="1" pageNumber="66" refString="Reed, S. K. &amp; Cumberlidge, N. (2006) Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proceedings of the Biological Society of Washington, 119 (1), 58 - 66. http: // dx. doi. org / 10.2988 / 0006 - 324 x (2006) 119 [58: franga] 2.0. co; 2" type="journal article" year="2006">Reed &amp; Cumberlidge, 2006</bibRefCitation>
</taxonomicName>
). Here we recognize a fourth kind of mandibular structure in Madagascan freshwater crabs for
<taxonomicName id="E6BAE799FFF6A1112591DE01FEDCF910" box="[190,373,1646,1671]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A1112591DE01FEDCF910" box="[190,373,1646,1671]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A1112452DE01FE65F910" bold="true" box="[381,460,1646,1671]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A1112452DE01FE65F910" box="[381,460,1646,1671]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
in which the mandible has a small, hard ledge-like anterior lobe about 0.25x as long as the terminal segment (
<figureCitation id="B981809FFFF6A11124BEDEFDFE7CF93C" box="[401,469,1682,1707]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
D,E).
</paragraph>
<paragraph id="21059C1AFFF6A11225E8DED9FDC7FED7" blockId="1.[151,1437,674,2031]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="67" pageId="1" pageNumber="66">
Besides the mandibles, other important taxonomic characters that set
<taxonomicName id="E6BAE799FFF6A11126CFDED9FB31F958" box="[992,1176,1718,1743]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A11126CFDED9FB31F958" box="[992,1176,1718,1743]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A111218EDED9FB58F958" bold="true" box="[1185,1265,1718,1743]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A111218EDED9FB58F958" box="[1185,1265,1718,1743]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
apart from the seven other
<collectingCountry id="59ADDC8AFFF6A111240CDEB5FE03F964" box="[291,426,1754,1779]" name="Madagascar" pageId="1" pageNumber="66">Madagascar</collectingCountry>
freshwater crab genera include its completely smooth dorsal carapace surface (
<figureCitation id="B981809FFFF6A1112013DEB5FAD4F964" box="[1340,1405,1754,1779]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A, 3A;
<bibRefCitation id="452BE1EBFFF6A11125E6DE91FD85F880" author="Cumberlidge" box="[201,556,1790,1815]" pageId="1" pageNumber="66" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">Cumberlidge &amp; Sternberg 2002</bibRefCitation>
<figureCitation id="B981809FFFF6A111271BDE91FDDFF880" box="[564,630,1790,1815]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="1" pageNumber="66">Fig. 2</figureCitation>
A-F).
<taxonomicName id="E6BAE799FFF6A1112793DE91FCDDF880" box="[700,884,1790,1815]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A1112793DE91FCDDF880" box="[700,884,1790,1815]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A1112654DE91FC63F880" bold="true" box="[891,970,1790,1815]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A1112654DE91FC63F880" box="[891,970,1790,1815]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
can also be distinguished from
<taxonomicName id="E6BAE799FFF6A111201CDE90FAC2F881" box="[1331,1387,1791,1814]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A111201CDE90FAC2F881" box="[1331,1387,1791,1814]" italics="true" pageId="1" pageNumber="66">Foza</emphasis>
</taxonomicName>
and
<taxonomicName id="E6BAE799FFF6A11125B8DF4DFEBDF8AC" box="[151,276,1826,1851]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11125B8DF4DFEBDF8AC" box="[151,276,1826,1851]" italics="true" pageId="1" pageNumber="66">Malagasya</emphasis>
</taxonomicName>
by the smooth anterolateral margin of the carapace (
<figureCitation id="B981809FFFF6A111264BDF4DFC0BF8AC" box="[868,930,1826,1851]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A, 3A), which has small teeth in
<taxonomicName id="E6BAE799FFF6A1112035DF4BFAFBF8AC" box="[1306,1362,1828,1851]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A1112035DF4BFAFBF8AC" box="[1306,1362,1828,1851]" italics="true" pageId="1" pageNumber="66">Foza</emphasis>
</taxonomicName>
(Reed &amp; Cumberlidge
<figureCitation id="B981809FFFF6A1112471DF29FE0FF8C8" box="[350,422,1862,1887]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
), and conspicuous teeth in
<emphasis id="13CE4008FFF6A111262BDF29FC21F8C8" box="[772,904,1862,1887]" italics="true" pageId="1" pageNumber="66">
<taxonomicName id="E6BAE799FFF6A111262BDF29FC2DF8C8" box="[772,900,1862,1887]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">Malagasya</taxonomicName>
.
</emphasis>
The new genus can be distinguished from
<taxonomicName id="E6BAE799FFF6A11125B8DF05FEE2F814" box="[151,331,1898,1923]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11125B8DF05FEE2F814" box="[151,331,1898,1923]" italics="true" pageId="1" pageNumber="66">Hydrothelphusa</emphasis>
</taxonomicName>
and
<taxonomicName id="E6BAE799FFF6A11124A2DF03FE5BF814" box="[397,498,1900,1923]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11124A2DF03FE5BF814" box="[397,498,1900,1923]" italics="true" pageId="1" pageNumber="66">Marojejy</emphasis>
</taxonomicName>
by the small notch-like space between the exorbital and epibranchial teeth in
<taxonomicName id="E6BAE799FFF6A11125B8DFE1FEE6F830" box="[151,335,1934,1959]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A11125B8DFE1FEE6F830" box="[151,335,1934,1959]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A111247BDFE1FE0BF830" bold="true" box="[340,418,1934,1959]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A111247BDFE1FE0BF830" box="[340,418,1934,1959]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B981809FFFF6A111249EDFE1FE46F830" box="[433,495,1934,1959]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A, 3A), versus a wide space or cleft between these teeth in both
<taxonomicName id="E6BAE799FFF6A1112195DFE1FAC7F830" box="[1210,1390,1934,1959]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A1112195DFE1FAC7F830" box="[1210,1390,1934,1959]" italics="true" pageId="1" pageNumber="66">Hydrothelphusa</emphasis>
</taxonomicName>
and
<taxonomicName id="E6BAE799FFF6A11125B8DFDBFF54F85C" box="[151,253,1972,1995]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF6A11125B8DFDBFF54F85C" box="[151,253,1972,1995]" italics="true" pageId="1" pageNumber="66">Marojejy</emphasis>
</taxonomicName>
(
<bibRefCitation id="452BE1EBFFF6A1112423DFDDFCB5F85C" author="Cumberlidge" box="[268,796,1970,1995]" pageId="1" pageNumber="66" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">
Cumberlidge &amp; Sternberg 2002
<figureCitation id="B981809FFFF6A1112756DFDDFD1FF85C" box="[633,694,1970,1995]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A-C, 2F
</bibRefCitation>
). The walking legs (not elongated or slim) and eyestalks and corneas (normal length and size) of
<taxonomicName id="E6BAE799FFF6A1112771DFB9FCBCF878" box="[606,789,2006,2031]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF6A1112771DFB9FCBCF878" box="[606,789,2006,2031]" italics="true" pageId="1" pageNumber="66">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF6A1112632DFB9FCC4F878" bold="true" box="[797,877,2006,2031]" pageId="1" pageNumber="66">
<taxonomicNameLabel id="08FDFD73FFF6A1112632DFB9FCC4F878" box="[797,877,2006,2031]" pageId="1" pageNumber="66" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B981809FFFF6A1112652DFB9FC15F878" box="[893,956,2006,2031]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
A, 3A) further distinguish this genus from
<taxonomicName id="E6BAE799FFF5A11225B8D8F6FF54FF27" box="[151,253,153,176]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF5A11225B8D8F6FF54FF27" box="[151,253,153,176]" italics="true" pageId="2" pageNumber="67">Marojejy</emphasis>
</taxonomicName>
whose walking legs are long, slender, and elongated, and whose eyestalks taper distally and have reduced corneas (
<bibRefCitation id="452BE1EBFFF5A1122428D8D3FC92FF43" author="Cumberlidge" box="[263,827,188,213]" pageId="2" pageNumber="67" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">
Cumberlidge &amp; Sternberg 2002
<figureCitation id="B981809FFFF5A112278BD8D3FD5BFF43" box="[676,754,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1469,1491]" captionTargetBox="[311,1265,206,1428]" captionTargetId="figure@3.[305,1282,195,1447]" captionTargetPageId="3" captionText="FIGURE 2. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) right G 1, ventral view; (B) right G 1, dorsal view; (C) right G 2, ventral view; (D) mandibular palp, frontal view; (E) mandibular palp, superior view; (F) sternum. Scale bar: 2.9 mm (A, B, C), 5.0 mm (D, E), 7.1 mm (F)." httpUri="https://zenodo.org/record/287452/files/figure.png" pageId="2" pageNumber="67">Fig. 2</figureCitation>
F, 8F
</bibRefCitation>
).
<taxonomicName id="E6BAE799FFF5A1122673D8D3FBBDFF42" box="[860,1044,188,213]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF5A1122673D8D3FBBDFF42" box="[860,1044,188,213]" italics="true" pageId="2" pageNumber="67">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF5A1122108D8D3FB2BFF42" bold="true" box="[1063,1154,188,213]" pageId="2" pageNumber="67">
<taxonomicNameLabel id="08FDFD73FFF5A1122108D8D3FB2BFF42" box="[1063,1154,188,213]" pageId="2" pageNumber="67" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="E6BAE799FFF5A11225B8D8B0FEFFFF6F" box="[151,342,223,248]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF5A11225B8D8B0FEFFFF6F" box="[151,342,223,248]" italics="true" pageId="2" pageNumber="67">Madagapotamon</emphasis>
</taxonomicName>
by the length of the flagellum of the exopod of the third maxilliped, which is long and well developed in
<taxonomicName id="E6BAE799FFF5A1122401D96BFE4CFE8A" box="[302,485,260,285]" class="Malacostraca" family="Potamonautidae" genus="Glabrithelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="13CE4008FFF5A1122401D96BFE4CFE8A" box="[302,485,260,285]" italics="true" pageId="2" pageNumber="67">Glabrithelphusa</emphasis>
</taxonomicName>
<emphasis id="13CE4008FFF5A11224C2D96BFD92FE8A" bold="true" box="[493,571,260,285]" pageId="2" pageNumber="67">
<taxonomicNameLabel id="08FDFD73FFF5A11224C2D96BFD92FE8A" box="[493,571,260,285]" pageId="2" pageNumber="67" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B981809FFFF5A1122764D96BFD23FE8B" box="[587,650,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1862,1884]" captionTargetBox="[179,1416,430,1790]" captionTargetId="figure@2.[162,1435,411,1801]" captionTargetPageId="2" captionText="FIGURE 1. Glabrithelphusa angene n. gen n. sp. Holotype, adult male from Madagascar (cw 41.5, cl 29.3 mm). (A) carapace, dorsal view; (B) frontal view; (C) third maxilliped; (D) carpus and merus of right cheliped, dorsal view; (E) carpus and merus of right cheliped, ventral view; (F) right cheliped, frontal view; (G) left cheliped, frontal view. Scale bar: 9.0 mm (A, B, D, E, F, G), 3.1 mm (C)." httpUri="https://zenodo.org/record/287451/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
C), but almost absent, and reduced to a short stub in
<taxonomicName id="E6BAE799FFF5A11221F3D96BFA32FE8A" box="[1244,1435,260,285]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="13CE4008FFF5A11221F3D96BFA32FE8A" box="[1244,1435,260,285]" italics="true" pageId="2" pageNumber="67">Madagapotamon</emphasis>
</taxonomicName>
(
<bibRefCitation id="452BE1EBFFF5A11225B1D948FDC8FED7" author="Cumberlidge" box="[158,609,295,320]" pageId="2" pageNumber="67" refString="Cumberlidge, N. &amp; Sternberg, R. (2002) The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema, 24 (1), 41 - 79." type="journal article" year="2002">Cumberlidge &amp; Sternberg 2002 Fig. 5H</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="69A0CF91FFF5A11225E8D923FE5CFEF2" box="[199,501,332,357]" pageId="2" pageNumber="67" type="distribution">
<paragraph id="21059C1AFFF5A11225E8D923FE5CFEF2" blockId="2.[151,1437,151,357]" box="[199,501,332,357]" pageId="2" pageNumber="67">
<emphasis id="13CE4008FFF5A11225E8D923FEC9FEF2" bold="true" box="[199,352,332,357]" pageId="2" pageNumber="67">Distribution.</emphasis>
<collectingCountry id="59ADDC8AFFF5A1122448D923FE58FEF2" box="[359,497,332,357]" name="Madagascar" pageId="2" pageNumber="67">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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