391 lines
34 KiB
XML
391 lines
34 KiB
XML
<document id="E0C3B80262D512EC5DBE3C64A6408B22" ID-CLB-Dataset="299803" ID-DOI="10.3897/zookeys.1207.120068" ID-GBIF-Dataset="d7935e1b-a3af-427e-afd7-88ee385dc1e7" ID-ZooBank="8B2E2B5D-CA1D-4B61-8736-ECF013EDC384" ID-publisher-id="120068" URI-arpha="1314A2C0-E9C6-52F9-821B-A77D9D85F5DA" URI-zoobank="http://zoobank.org/8B2E2B5D-CA1D-4B61-8736-ECF013EDC384" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720801545516" checkinUser="pensoft" docAuthor="Porto, Willians, Derkarabetian, Shahan, Giribet, Gonzalo & Pérez-González, Abel" docDate="2024" docId="0FDE685782E9547BAFDCF6658E7A65CB" docLanguage="en" docName="ZooKeys 1207: 1-149" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/120068/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Fresiax fray Porto, Derkarabetian, Giribet & Pérez-González, 2024, sp. nov." docType="treatment" docUuid="0F310737-611E-4AB6-B41D-FD12B585ECA5" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="149" masterDocId="1314A2C0E9C652F9821BA77D9D85F5DA" masterDocTitle="Systematic revision of the South American “ Nuncia ” (Opiliones, Laniatores, Triaenonychidae)" masterLastPageNumber="149" masterPageNumber="1" pageNumber="1" updateTime="1720815513282" updateUser="ExternalLinkService">
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<mods:titleInfo id="0B75ED649F8366EE76C87925039C8F9C">
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<mods:title id="F843FED78EA5DF69932982DB35A248E6">Systematic revision of the South American “ Nuncia ” (Opiliones, Laniatores, Triaenonychidae)</mods:title>
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<mods:name id="BA40BE923057B81F44087611D6AC2FFD" type="personal">
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<mods:namePart id="4B1DECE1D51D3CBBF6222FD2482C394F">Porto, Willians</mods:namePart>
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<mods:nameIdentifier id="3595F68838936EAEE2210EBDF0B979DF" type="ORCID">0000-0002-4209-5664</mods:nameIdentifier>
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<mods:affiliation id="E0EEDA6097A637F6BBBEE90F7F538A23">División Aracnología, Museo Argentino de Ciencias Naturales – CONICET, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina & Sección Aracnología y Miriapodología, Universidad Nacional de La Plata, Facultad de Ciencias Naturales y Museo, Paseo del Bosque s / n, (1900) La Plata, Argentina</mods:affiliation>
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</mods:name>
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<mods:name id="D002AE4F0D2859C1ADC32FD185481C27" type="personal">
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<mods:roleTerm id="AD5632A56C10430B7D0BDB654CECF8E6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="00E51C6EBC875859B05C8049332F667E">Derkarabetian, Shahan</mods:namePart>
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<mods:nameIdentifier id="C7E52AC9BB11F7B38AF0924A89F37E01" type="ORCID">0000-0002-9163-9277</mods:nameIdentifier>
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<mods:affiliation id="E959AC513144488A45BF333DBF9D2DF4">Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA & San Diego Natural History Museum, Department of Entomology, San Diego, CA 92101, USA</mods:affiliation>
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</mods:name>
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<mods:name id="B398A68A658070C707BF2BFA564B1CE0" type="personal">
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<mods:roleTerm id="1AAEBDDDDAF0D8F360552E3E1483F603">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4E67D499D48CD7B916DAD7FC4DF391E5">Giribet, Gonzalo</mods:namePart>
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<mods:nameIdentifier id="3CAD71BFD6A3A5A605BBA77571E0DB4F" type="ORCID">0000-0002-5467-8429</mods:nameIdentifier>
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<mods:affiliation id="FA4CADCEC40729EB5A61CCC2FF424C57">Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA</mods:affiliation>
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</mods:name>
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<mods:name id="4072CB67A8C588AF9628585F4266BE10" type="personal">
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<mods:roleTerm id="6A6276D57A5F3F4B88663A8FBFEDDDAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6262CFB28430D96DADC9BD4542E5171A">Pérez-González, Abel</mods:namePart>
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<mods:nameIdentifier id="87C755B0C51C83A9BA99B05EF53A4C10" type="ORCID">0000-0002-4245-3302</mods:nameIdentifier>
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<mods:affiliation id="563DB5C7466F96469F3B14257DA5DEB0">División Aracnología, Museo Argentino de Ciencias Naturales – CONICET, Av. Ángel Gallardo 470, C 1405 DJR Buenos Aires, Argentina</mods:affiliation>
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<mods:typeOfResource id="2045C77977C8A5E0856FEB805E199CC0">text</mods:typeOfResource>
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<mods:title id="E3B131C45F381EF45CA8A0F0E32E808A">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="E0E22595B63C43BF062F12B5AAA5E8D8">
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<mods:date id="DA4B6A7C75636F4BC36069848198E69D">2024</mods:date>
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<mods:number id="0C5B1E90CCEE21CCD1EEBADD0DBE6BD8">2024-07-12</mods:number>
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<mods:detail id="740B1961A26D9A1EB00E788C08A15F26" type="volume">
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<mods:number id="4497999454AEEB13DB324BA337145205">1207</mods:number>
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<mods:start id="7F334CB87646971AC8BA7DF40A07C0E9">1</mods:start>
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<mods:classification id="95A6A4804708C08AC2D66995C80EE6D7">journal article</mods:classification>
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<mods:identifier id="520EF38A7537D32DC8737E7D979FF9ED" type="CLB-Dataset">299803</mods:identifier>
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<mods:identifier id="A811F54EE9112CF54C9616383046996C" type="DOI">10.3897/zookeys.1207.120068</mods:identifier>
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<mods:identifier id="F01B5A3386FA2E76C25346A2BD417A18" type="GBIF-Dataset">d7935e1b-a3af-427e-afd7-88ee385dc1e7</mods:identifier>
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<mods:identifier id="F550DBBCB5B1F30794A712D86191EB8B" type="ZooBank">8B2E2B5D-CA1D-4B61-8736-ECF013EDC384</mods:identifier>
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</mods:mods>
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<treatment id="0FDE685782E9547BAFDCF6658E7A65CB" ID-DOI="http://doi.org/10.5281/zenodo.12734474" ID-GBIF-Taxon="235091469" ID-Zenodo-Dep="12734474" ID-arpha="0FDE6857-82E9-547B-AFDC-F6658E7A65CB" ID-zoobank="https://zoobank.org/0F310737-611E-4AB6-B41D-FD12B585ECA5" LSID="urn:lsid:zoobank.org:act:0F310737-611E-4AB6-B41D-FD12B585ECA5" LSID-ZBK="urn:lsid:zoobank.org:act:0F310737-611E-4AB6-B41D-FD12B585ECA5" httpUri="http://treatment.plazi.org/id/0FDE685782E9547BAFDCF6658E7A65CB">
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||
<subSubSection id="3B0B95F99EB578744329EA67FEB6BE84" type="nomenclature">
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||
<paragraph id="1B8BE4AD01331A6FB8C303FA4DD50390">
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||
<taxonomicName id="F934E6C9F6924B4E9EB656BDAA390812" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
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<emphasis id="F5AE57A140B589F52156B16A11BB65BA" italics="true">Fresiax fray</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3FE50E860869C460AEF8EAF6EAB640FD" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="1C830BBF7E4BBCD2EAD2D377900D13B6" type="description">
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||
<paragraph id="AC1D4EC84CA79C3E0E2CD9F614A61098">
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||
<figureCitation id="EE05154EB9DCD2006C43F008FB4D9AA4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Fresiax fray sp. nov. habitus, male A dorsal view C lateral view E ventral view. Female B dorsal view D lateral view F ventral view. Scale bars: 500 µm. Species of Clade A, see Fig. 3." figureDoi="10.3897/zookeys.1207.120068.figure11" httpUri="https://binary.pensoft.net/fig/1090617">Figs 11</figureCitation>
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||
,
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<figureCitation id="55985202BB41D21B56F42AD54F2FF12B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Fresiax fray sp. nov. male, SEM images of habitus A dorsal view B lateral view C, D ventral view. Scale bars: 500 µm (A, B, C); 200 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure12" httpUri="https://binary.pensoft.net/fig/1090618">12</figureCitation>
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||
,
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<figureCitation id="A4381D490C4E21D0C8EA4011AD6C25B4" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Fresiax fray sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1207.120068.figure13" httpUri="https://binary.pensoft.net/fig/1090619">13</figureCitation>
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||
,
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||
<figureCitation id="61D960C9ADB385AD9A90843B0DC11BFB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Fresiax fray sp. nov. legs I A II B III C IV D. Scale bars: 200 µm (A, C); 500 µm (B, D)." figureDoi="10.3897/zookeys.1207.120068.figure14" httpUri="https://binary.pensoft.net/fig/1090620">14</figureCitation>
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||
,
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<figureCitation id="03AD9336709F69913662A8E105EECCE6" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Fresiax fray sp. nov. penis: ventral A, B apical C lateral D, E dorsal F, G. Scale bars: 100 µm (A, D, F); 20 µm (B, C, E, G)." figureDoi="10.3897/zookeys.1207.120068.figure15" httpUri="https://binary.pensoft.net/fig/1090621">15</figureCitation>
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||
,
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||
<figureCitation id="140C730BA14643D7F1D4E5E23AF363BB" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Fresiax fray sp. nov. penis: ventral A apical B lateral C dorsal D. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 20 µm." figureDoi="10.3897/zookeys.1207.120068.figure16" httpUri="https://binary.pensoft.net/fig/1090622">16</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EKMBG" type="material">
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||
<paragraph id="15D234159BB17D806C565D34E7C53642">
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||
<heading id="C0C2E0E9C7E6C8653F8D4A2906CB1E97" reason="title">Material examined.</heading>
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||
</paragraph>
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||
<paragraph id="5F72D537DB2CC903141296E5F1833461">
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||
<materialsCitation id="CF7A29191DF6B6CAFEC95F63C4CAF0C2" ID-GBIF-Occurrence="4905322335" collectingDate="1988-11-03" collectionCode="MNHNCL" collectorName="E. Maury" country="Chile" location="Bosque de Fray Jorge" specimenCount="1" specimenCount-male="1" stateProvince="Coquimbo" typeStatus="holotype">
|
||
<emphasis id="3E03EFEBBB84628055AE5C1B7EA28618" bold="true">
|
||
<emphasis id="D06E95EB0694B011DB03F134FA5F0150" bold="true" italics="true">
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||
<typeStatus id="6597B3C99A0DFD36BBBBDFF8424814D1" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
.
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||
</emphasis>
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||
<specimenCount id="AE88A2ADA8F48C02B7D70E41BD937EDB" count="1" type="male">♂</specimenCount>
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||
<emphasis id="A57EDAE4F47EA3E1187A2E57C9A0AF9A" bold="true">
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||
<collectingCountry id="90A51D5C48A419328D79834CE6D8C616" name="Chile">Chile</collectingCountry>
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||
.
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||
</emphasis>
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||
<collectingRegion id="EF5189A4ECBE627B37F4AC6AAA85EFAE" country="Chile" name="Coquimbo">Coquimbo</collectingRegion>
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||
:
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||
<location id="21E67F9B2B606F92555455291F2D574B" LSID="urn:lsid:plazi:treatment:0FDE685782E9547BAFDCF6658E7A65CB:21E67F9B2B606F92555455291F2D574B" country="Chile" name="Bosque de Fray Jorge" stateProvince="Coquimbo">Bosque de Fray Jorge</location>
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,
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<collectorName id="27250B6C35F23A587F5B51A5D7FBD689">E. Maury</collectorName>
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||
coll.
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<date id="FDCDC489F8988C43EB82090BE437530E" value="1988-11-03">
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<collectingDate id="76571E9A693EB2527F003FA96504AB7C" value="1988-11-03">03. XI. 1988</collectingDate>
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||
</date>
|
||
(
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||
<named-content id="95682934B2F50BD0BEDF8752815E74FA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-historia-natural-santiago" xlink_title="Museo Nacional de Historia Natural, Santiago de Chile">
|
||
<collectionCode id="A8C46CA9BBAD30471E5723988D3F3A7F">MNHNCL</collectionCode>
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||
</named-content>
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||
)
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||
</materialsCitation>
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||
.
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<materialsCitation id="435A788851B1E5F83944E3D67F2118F3" ID-GBIF-Occurrence="4905322360" collectingDate="1988-11-03" collectionCode="MACN" collectorName="P. N. Fray Jorge & E. Maury" country="Chile" location="Bosque Fray Jorge" municipality="Limari" specimenCount="62" specimenCount-female="24" specimenCount-male="38" stateProvince="Coquimbo" typeStatus="paratype">
|
||
<emphasis id="E050A081E238F22D274CDB6B98E3C91D" bold="true">
|
||
<emphasis id="36E25C32FF211FB188950AFE6809B8C9" bold="true" italics="true">
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||
<typeStatus id="4B3029A80616229B266E55434F84B215" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
.
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<collectingCountry id="1DF55161F7F912B1A9CAD575A2B6E0A0" name="Chile">Chile</collectingCountry>
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||
.
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||
</emphasis>
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||
<collectingRegion id="29CCF36BC40CAFE258FDDF9B575BB8E9" country="Chile" name="Coquimbo">Coquimbo</collectingRegion>
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||
:
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<collectingMunicipality id="32C2102E87D0285C037550FAB0867EF5">Limarí</collectingMunicipality>
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||
,
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||
<location id="33EE6E254D75EEE6FFEE87FB592F65B5" LSID="urn:lsid:plazi:treatment:0FDE685782E9547BAFDCF6658E7A65CB:33EE6E254D75EEE6FFEE87FB592F65B5" country="Chile" municipality="Limari" name="Bosque Fray Jorge" stateProvince="Coquimbo">Bosque Fray Jorge</location>
|
||
,
|
||
<collectorName id="708153982CD07E96D98972B7775CCDC1">P. N. Fray Jorge</collectorName>
|
||
,
|
||
<collectorName id="0B3EF97E29C0782F9056682D04975912">E. Maury</collectorName>
|
||
coll.
|
||
<date id="A94D0939240F05738E4530136B14FA42" value="1988-11-03">
|
||
<collectingDate id="B2D693DBD7D6C08DE13E6E310E7A10BE" value="1988-11-03">03. XI. 1988</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="967818009B439192A2B9FC0F071F15BB" count="38" type="male">38 ♂</specimenCount>
|
||
<specimenCount id="81CDB3CF3BF8D49969BFACBE3F7210F6" count="24" type="female">24 ♀</specimenCount>
|
||
, amm. (
|
||
<named-content id="2359E1AE5958661500694AD05201D3E6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales">
|
||
<collectionCode id="766E6D0340828D47BAC8A0E5111B8E83" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
|
||
</named-content>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EFNBG" type="material">
|
||
<paragraph id="EF12DF42FF8EE3CD886A7276B233D85C">
|
||
<heading id="BCA80200B61A42C453EDD4CC508B0E26" reason="title">Additional material.</heading>
|
||
</paragraph>
|
||
<paragraph id="6294CDE8DEA907A65F82ED3255E16A39">
|
||
<collectingCountry id="6AEE48900B9ED84D43CB4ED5B0FBF6F1" name="Chile">Chile</collectingCountry>
|
||
:
|
||
<collectingRegion id="B139460CC98BB7C4F8BCC87A1239DCB9" country="Chile" name="Coquimbo">Coquimbo</collectingRegion>
|
||
: Limarí, Bosque Talinay, P. N. Fray Jorge, relict Valdivian fog forest,
|
||
<collectorName id="989FA3869BA550B43D0828AE674F93A7">R. Schuh</collectorName>
|
||
,
|
||
<collectorName id="8B7557CAF403CCB1DEBD45BC0FE25BA7">N. Platnick</collectorName>
|
||
coll.,
|
||
<date id="3B8F1DC9B478BDCBA5EA78F2FC00AEE8" value="1986-02-08">08. II. 1986</date>
|
||
,
|
||
<specimenCount id="B1910EE25A1A636EE4AC06DD5A9CE845" count="66" type="generic">66 specimens</specimenCount>
|
||
(
|
||
<named-content id="069548D5C819DF70E6E96FA4F3E172DF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History">
|
||
<collectionCode id="839743D8904770D2AA1593FA5FEC5997" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" type="">AMNH</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQNBG" type="etymology">
|
||
<paragraph id="5A7C7D4E8A256F60539DBD955289AA1B">
|
||
<heading id="CD20A99BC452E5BCDD182ED77323B122" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="5BB815F0BD39BE30310C172C19D358F1">
|
||
The epithet
|
||
<taxonomicName id="26171D62295E22755C747CF0855585D5" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray">
|
||
<emphasis id="53E1973E3B21AB549A85933BBDCFEC2A" italics="true">fray</emphasis>
|
||
</taxonomicName>
|
||
, a noun in apposition, refers to the type locality of the species, Bosque Fray Jorge National Park.
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAOBG" type="diagnosis">
|
||
<paragraph id="B2469951BB393B73A248A52644EA3FB7">
|
||
<heading id="5D012D486BED8C7D5200E777DF9012CF" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="CC153FDFEC53B47AE7B48C9DDFD464EF">
|
||
This species can be easily distinguished from the other species of the genus by its small size (<
|
||
<quantity id="FBB4CC44660CD17BFD99307AC6E1730C" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
|
||
long), conical ocularium, with an apophysis at an angle of 45 °.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EFOBG" type="distribution">
|
||
<paragraph id="40888F78CBB6131F7E1E72F2D16A98F9">
|
||
<heading id="6B3659D41F7476FFF49CDAF02CD8F5FF" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="BD4C32040D630FF5F6A1776134A46E48">
|
||
<collectingCountry id="F0E9DE39F43FE10C30ED11A3F5BBC85B" name="Chile">Chile</collectingCountry>
|
||
,
|
||
<collectingRegion id="74EE95F1812C26EE98C650230759F967" country="Chile" name="Coquimbo">Coquimbo Region</collectingRegion>
|
||
(Fig.
|
||
<figureCitation id="7A15F42079D490C5B04AF5D3BC088FD2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maps showing the distribution of the species used in this work. South America (Chile and western Argentina inset). Chile in green, Argentina / South America in grey. Terrestrial ecoregions (following Olson et al. 2001) highlighted in yellow (Chilean Matorral) and orange (Valdivian temperate forests)." figureDoi="10.3897/zookeys.1207.120068.figure4" httpUri="https://binary.pensoft.net/fig/1090610">4 A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EPOBG" type="description">
|
||
<paragraph id="531E03ED4625A17FCE36409C5CE3931C">
|
||
<heading id="7E95D47FCB6C47CB08079EFF32FED494" reason="title">Description of male.</heading>
|
||
</paragraph>
|
||
<paragraph id="34C33EDC5E1B8FAF900BC7573AAE4BA4">Measurements: Total length 1.81, carapace length 0.69, dorsal scutum length 1.47, carapace max. width 1.14, mesotergum max. length 1.49. Appendage measurements: Pedipalps. Length of trochanter 0.13, length of femora 0.80, length of patella 0.46, length of tibia 0.53, length of tarsus 0.62. Leg I: trochanter (tr) 0.16, femora (fe) 0.74, patella (pa) 0.38, tibia (ti) 0.51, metatarsus (mt) 0.62, tarsus (ta) 0.46. II: tr 0.22, fe 1.03, pa 0.41, ti 0.80, mt 0.98, ta 0.93. III: tr 0.19, fe 0.68, pa 0.36, ti 0.60, mt 0.67, ta 0.62. IV: tr 0.26, fe 1.04, pa 0.46, ti 0.80, mt 1.11, ta 0.57.</paragraph>
|
||
<paragraph id="D14E2F9D044EC63DFB1CCB218690B413">
|
||
Dorsum (Fig.
|
||
<figureCitation id="81EB607B47A6B445D17BA3C018B90E80" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Fresiax fray sp. nov. habitus, male A dorsal view C lateral view E ventral view. Female B dorsal view D lateral view F ventral view. Scale bars: 500 µm. Species of Clade A, see Fig. 3." figureDoi="10.3897/zookeys.1207.120068.figure11" httpUri="https://binary.pensoft.net/fig/1090617">11</figureCitation>
|
||
,
|
||
<figureCitation id="AEA6A60F3729FB17FDA73D1623F15B4C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Fresiax fray sp. nov. male, SEM images of habitus A dorsal view B lateral view C, D ventral view. Scale bars: 500 µm (A, B, C); 200 µm (D)." figureDoi="10.3897/zookeys.1207.120068.figure12" httpUri="https://binary.pensoft.net/fig/1090618">12</figureCitation>
|
||
). Eta (η) hourglass-shaped dorsal scutum. Ocularium conical, with a small, forward-pointing apical spine and small setae. Eyes located on the distal region of the ocularium. Dorsal scutum microgranulate with no clear area delimitation. Areas I – III with two small setiferous granules, IV with three, posterior margin with a row of small setiferous granules. All free tergites with a row of small setiferous granules.
|
||
</paragraph>
|
||
<caption id="D6A4524A682AA400F2EA1C85763668E4" ID-DOI="10.3897/zookeys.1207.120068.figure11" ID-arpha="BA96C1EB-DD10-55E3-92CC-960F38FB2262" httpUri="https://binary.pensoft.net/fig/1090617" startId="F11">
|
||
<paragraph id="26D082C365197F52069D7AC22526CC39">
|
||
<label id="BDCCA1D8696E003B0610D31B959180A6">Figure 11.</label>
|
||
</paragraph>
|
||
<paragraph id="55C2A85E89466DA57B0B0BCBA3751739">
|
||
<taxonomicName id="19B75EC87052E2541C3B6E61FAD6FD19" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="A33A9D15F0A0CA01F4306C74D0A65CD5" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B87B30267328F75EF9D3D6C5EF6F84D3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
habitus, male
|
||
<emphasis id="394F6B08BCC89FD0EECD9C1A382B6247" bold="true">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="721C7714D3941C7F353C35E66A0418A2" bold="true">C</emphasis>
|
||
lateral view
|
||
<emphasis id="FA50AB3D308DECEAC858F1921F4D0458" bold="true">E</emphasis>
|
||
ventral view. Female
|
||
<emphasis id="118FEE07443D1B7FE2AD5E620B0A30F8" bold="true">B</emphasis>
|
||
dorsal view
|
||
<emphasis id="A0E99B4D3334CFEB33CEE05D5005D5C5" bold="true">D</emphasis>
|
||
lateral view
|
||
<emphasis id="677CC134DAFCCB064C9AC6D8C2CC271A" bold="true">F</emphasis>
|
||
ventral view. Scale bars: 500 µm. Species of Clade A, see Fig.
|
||
<figureCitation id="E6BDD4D9D277E219A315465DE9D7CB68" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phylogenetics relationships of Triaenonychidae, with clade names as discussed in the main text. Phylogeny is derived from the IQ-TREE analysis. Nodes without circles have < 95 % bootstrap support to AH-aLR and UFBoot." figureDoi="10.3897/zookeys.1207.120068.figure3" httpUri="https://binary.pensoft.net/fig/1090609">3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7BE5D32C940111FFD42EF2E73A5771B0" ID-DOI="10.3897/zookeys.1207.120068.figure12" ID-arpha="C37C395E-4226-5251-BD1F-434085820523" httpUri="https://binary.pensoft.net/fig/1090618" startId="F12">
|
||
<paragraph id="D368652B7EC9D119CDB9DB6C73A688B7">
|
||
<label id="BA524ADBC43227AB8CAB2F2A11E61111">Figure 12.</label>
|
||
</paragraph>
|
||
<paragraph id="013DB9BFD87BC1C979021034E9FA9CA5">
|
||
<taxonomicName id="D91867769657829392A9584CD919AEC3" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="A66BEB90BF7715F847BFD6F424CC549E" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C303A8F810D4CDEBA9C7D25E8D08300C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
male,
|
||
<abbrev content-type="institution" id="ABBRID0EZQBG" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||
images of habitus
|
||
<emphasis id="2CC8D88050FDF70B9B612CC13F25CE5B" bold="true">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="B76CB294473540C43CAE25D317138B90" bold="true">B</emphasis>
|
||
lateral view
|
||
<emphasis id="B0D2EF99110997F7CB2CA83D4A0E4BAF" bold="true">C, D</emphasis>
|
||
ventral view. Scale bars: 500 µm (
|
||
<emphasis id="B3D5DEB104F54F2703D54D769F3C815B" bold="true">A, B, C</emphasis>
|
||
); 200 µm (
|
||
<emphasis id="407583CBBCA114E64B15B63086654F40" bold="true">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="51173CAF526553FBC13797E583B75475">
|
||
Chelicerae (Fig.
|
||
<figureCitation id="856971A219472E2365A66AD36762E60A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Fresiax fray sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1207.120068.figure13" httpUri="https://binary.pensoft.net/fig/1090619">13 A, B</figureCitation>
|
||
). Segment I characterized by a smooth surface without any prominent tubercles or setae. In contrast, segment II with a small frontal tubercle and covered in setae. Segment II with more pronounced texture and setal coverage compared to the smooth surface of segment I.
|
||
</paragraph>
|
||
<caption id="C0F61BC4BC082E8DFF687770672C2BA8" ID-DOI="10.3897/zookeys.1207.120068.figure13" ID-arpha="F2C7108F-2CAB-5C17-8427-21DD36C65EE6" httpUri="https://binary.pensoft.net/fig/1090619" startId="F13">
|
||
<paragraph id="58A64AD95C364C4988AC52D55D62C03C">
|
||
<label id="6CFABECAFD32C52F10F2ED6DE1B19FBC">Figure 13.</label>
|
||
</paragraph>
|
||
<paragraph id="312CBCA2C71C07764B533535BEF55D9A">
|
||
<taxonomicName id="04A5C1212D291C7EDF9AB2269D788801" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="2131C1102568E424B9AA18D173B2DF04" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="38868A7B14C7980875492C63EB929539" rank="species">sp. nov.</taxonomicNameLabel>
|
||
chelicerae: mesal
|
||
<emphasis id="8849C897000542AB0019C8C24F049D66" bold="true">A</emphasis>
|
||
ectal
|
||
<emphasis id="C6D90E44E1AD8065534CE9FBBD21DC7F" bold="true">B</emphasis>
|
||
pedipalps: mesal
|
||
<emphasis id="3DF2922A58DF3DD46721AD3F3DA28EAD" bold="true">C</emphasis>
|
||
ectal
|
||
<emphasis id="2E2F80FDEF2844D06886C065C6DDEA75" bold="true">D</emphasis>
|
||
. Scale bars: 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="84310EA55CFE1F9EE8D190699E2D14D7">
|
||
Pedipalps (Fig.
|
||
<figureCitation id="6CFBE0232616FFCC43E3B078716DD17B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Fresiax fray sp. nov. chelicerae: mesal A ectal B pedipalps: mesal C ectal D. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1207.120068.figure13" httpUri="https://binary.pensoft.net/fig/1090619">13 C, D</figureCitation>
|
||
). Trochanter with a dorsal and a ventral spine, both with setae. Femora with a row of four dorsal spines with subdistal setae, a row of five ventral spines with setae, the proximal one with a subtriangular shape; two dorsal and distal tubercles with setae, a row of five small mesal tubercles with setae, and three small distal tubercles with setae. Patella with two rows of three tubercles with setae, two small mesal tubercles with setae, and few ventral granules. Tibia with four ectal and mesal spines with subdistal setae; three ventral tubercles with setae and two rows of 4–5 granules with setae on the dorsal surface. Tarsus with three mesal and ectal spines with subdistal setae, as well as few setae and granules.
|
||
</paragraph>
|
||
<paragraph id="F21A87C31E0D161AB0A38E111DBA7972">
|
||
Legs (Fig.
|
||
<figureCitation id="E2A262A1EDF547B312AAA0DDF03C17C8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Fresiax fray sp. nov. legs I A II B III C IV D. Scale bars: 200 µm (A, C); 500 µm (B, D)." figureDoi="10.3897/zookeys.1207.120068.figure14" httpUri="https://binary.pensoft.net/fig/1090620">14</figureCitation>
|
||
). Coxa I bearing small setiferous tubercles and a row of three long tubercles with subdistal setae; II – IV with microgranulation, bearing 4–6 bridges between legs II and III, five or six between III and IV, six or seven between leg IV and the opisthosoma. Spiracles not visible. Smooth area occupies 1 / 3 of the leg II, almost ½ of III, and only a small proximal portion of the leg IV. Smooth area of leg II with five small setiferous tubercles, III with a row of four and an anterior process directed to the sternum. Sternum arrow-shaped, with a triangular posterior area. Leg I trochanter with three small ventral setiferous tubercles and one dorsal tubercle. Femora of leg I with a row of seven setiferous tubercles and a dorsal row of small setiferous tubercles. Tibia of leg I with 3–4 ventral tubercles with setae. Legs II – IV covered in setae, with the tarsus and calcaneus area densely covered in setae. Calcaneus smaller than astragalus, ≥ 4 × smaller in leg I, 5 × smaller in leg II, 8 × smaller in leg III, and 7 × smaller in leg IV. These differences in size ratios between the calcaneus and astragalus serve as distinguishing characteristics among the legs of this species. Tarsal count 3–6 – 4 – 4.
|
||
</paragraph>
|
||
<caption id="DC0F04328EB2ACB9CFD24DDD69ED66B9" ID-DOI="10.3897/zookeys.1207.120068.figure14" ID-arpha="934BB51A-AB74-569B-80CD-9B400342E244" httpUri="https://binary.pensoft.net/fig/1090620" startId="F14">
|
||
<paragraph id="C650021A1747FF001E2DD7F47A00E5FC">
|
||
<label id="C3B412A3BA03FF145710238BC73643C5">Figure 14.</label>
|
||
</paragraph>
|
||
<paragraph id="C6F0738D86A6CE2DE0A7B24886A48969">
|
||
<taxonomicName id="B893F24DECEE29A5404CF27D31C00211" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="2A143444C27B7D488777E6E80307F48D" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0B4C1824AA912115E54574C99896A3F0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
legs I
|
||
<emphasis id="B664728D0EFE3E879622EE5231D8FF20" bold="true">A</emphasis>
|
||
II
|
||
<emphasis id="BEB2CA44AE760262576B267C24F90604" bold="true">B</emphasis>
|
||
III
|
||
<emphasis id="9BD251F9567DB2733B77C3765749BC91" bold="true">C</emphasis>
|
||
IV
|
||
<emphasis id="A21FD3FA15280CDF346E4D2CBC858B87" bold="true">D</emphasis>
|
||
. Scale bars: 200 µm (
|
||
<emphasis id="336F9DC7AA94602D6D343DB5AEDCF753" bold="true">A, C</emphasis>
|
||
); 500 µm (
|
||
<emphasis id="0DBA4A3A4A6719AD8B8424FEEC7C1C1C" bold="true">B, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="95904A5182D80BECD0D84E4EA8AC3710">
|
||
Penis (Figs
|
||
<figureCitation id="D1406E553B1519A0F1E658ED8BA82551" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Fresiax fray sp. nov. penis: ventral A, B apical C lateral D, E dorsal F, G. Scale bars: 100 µm (A, D, F); 20 µm (B, C, E, G)." figureDoi="10.3897/zookeys.1207.120068.figure15" httpUri="https://binary.pensoft.net/fig/1090621">15</figureCitation>
|
||
,
|
||
<figureCitation id="C764B4A170E38390AF0CB10CCCCC9942" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Fresiax fray sp. nov. penis: ventral A apical B lateral C dorsal D. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 20 µm." figureDoi="10.3897/zookeys.1207.120068.figure16" httpUri="https://binary.pensoft.net/fig/1090622">16</figureCitation>
|
||
). Pars distalis with a ventral plate and a prominent cleft that divides the plate into two lamellae. Each lamella is equipped with three macrosetae on the ventral surface and one macroseta on the dorsal surface. Capsula externa nearly as long as the capsula interna, having a notch that divides the capsula externa into two halves.
|
||
</paragraph>
|
||
<caption id="4FA4A7ABA9FE390B1B74CCF967A39236" ID-DOI="10.3897/zookeys.1207.120068.figure15" ID-arpha="F1E0C0C5-BC45-53DE-A26B-13B5DAA08C6F" httpUri="https://binary.pensoft.net/fig/1090621" startId="F15">
|
||
<paragraph id="8CAA25472656848E82F90013DE5BBF95">
|
||
<label id="0BCCE8205080B5F3C05E73AFF3C842C3">Figure 15.</label>
|
||
</paragraph>
|
||
<paragraph id="66B6CA2417204EDF07E0AEA76FF2C7D5">
|
||
<taxonomicName id="6A7D28D3A83A6378AF713CB1DB88FAE7" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="14658A240C38718488AC5AAFC9DB6EC8" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0D8A7CB2B988428672A5BE160414F2D7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
penis: ventral
|
||
<emphasis id="1E4EB95E04251EABD902E7ABD4075ED9" bold="true">A, B</emphasis>
|
||
apical
|
||
<emphasis id="426F6B42DB32E0F01F2CFEC74810F4B8" bold="true">C</emphasis>
|
||
lateral
|
||
<emphasis id="FA55ECB9E19E0E99A519EB58171843AD" bold="true">D, E</emphasis>
|
||
dorsal
|
||
<emphasis id="F11ACF3C8C3708322068C06BA15FDFD3" bold="true">F, G</emphasis>
|
||
. Scale bars: 100 µm (
|
||
<emphasis id="3E34B84B8F6C667E0BD696A067413039" bold="true">A, D, F</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="F643CAC1E038054A526CFA3E3191D3F6" bold="true">B, C, E, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="53D142BAEF0BCFA2850F633532CB221E" ID-DOI="10.3897/zookeys.1207.120068.figure16" ID-arpha="4F229E53-98B5-54EC-90BD-CDDB1B1B3A5C" httpUri="https://binary.pensoft.net/fig/1090622" startId="F16">
|
||
<paragraph id="5BCD770D0A5F1665B15D535CE5319B2D">
|
||
<label id="057A3F1B6127A076CFEAC82DC9AE8E17">Figure 16.</label>
|
||
</paragraph>
|
||
<paragraph id="D7542C598A3E34445A8D637B7D7B1D6F">
|
||
<taxonomicName id="22252CF5BC640F10ECEBFB4A48BBACE5" authorityName="Porto & Derkarabetian & Giribet & Pérez-González" authorityYear="2024" class="Arachnida" family="Triaenonychidae" genus="Fresiax" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="fray" status="sp. nov.">
|
||
<emphasis id="A9E004FC5E8893FC868EB9845C439BAD" italics="true">Fresiax fray</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19252E631D1A608AAE7445F01468A5B0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
penis: ventral
|
||
<emphasis id="B0380D03BFC99E5E83067F3B6AB79FAC" bold="true">A</emphasis>
|
||
apical
|
||
<emphasis id="A6DE86F38A79EBC5754B1275BB34425C" bold="true">B</emphasis>
|
||
lateral
|
||
<emphasis id="0B0C3810B9F131AF0F0407CD5A002C32" bold="true">C</emphasis>
|
||
dorsal
|
||
<emphasis id="A5058B90B4E0E0B3D27138DC06C5D5D7" bold="true">D</emphasis>
|
||
. Colors: ventral plate (yellow), capsula externa (blue), capsula interna (red). Scale bars: 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F406941AC390D4D2E3895B2AABABEA3C">
|
||
<emphasis id="A23154087E818723340E94AE63C7CD0B" bold="true">Female.</emphasis>
|
||
Similar to male, with shorter pedipalpal femora.
|
||
</paragraph>
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<paragraph id="7F51D104EC9F809AAF75D58098732AB6">Female measurements. Total length 1.54, length of carapace 0.61, length of dorsal scutum 1.31, max. width of carapace 0.96, max. width of mesotergum 1.35. Appendage measurements: Pedipalps. Length of trochanter 0.11. Length of femora 0.58, length of patella 0.37, length of tibia 0.45, length of tarsus 0.46. Leg I: trochanter (tr) 0.18, femora (fe) 0.62, patella (pa) 0.33, tibia (ti) 0.46, metatarsus (mt) 0.53, tarsus (ta) 0.42. II: tr 0.18, fe 0.88, pa 0.34, ti 0.71, mt 0.84, ta 0.86. III: tr 0.18, fe 0.62, pa 0.26, ti 0.53, mt 0.68, ta 0.49. IV: tr 0.21, fe 0.80, pa 0.37, ti 0.73, mt 0.98, ta 0.57.</paragraph>
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</subSubSection>
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</treatment>
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</document> |