treatments-xml/data/25/CA/B9/25CAB9B1D4545BA8B9DEA23523D650DA.xml
2024-06-21 12:31:41 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/vz.74.e110674" ID-Pensoft-Pub="2625-8498-74-23" ID-Pensoft-UUID="41B30B26C7D754A6847973C6FED0710C" ID-ZooBank="335FBFDDE54340CA80140E16A7977586" ModsDocID="2625-8498-74-23" checkinTime="1705718409033" checkinUser="pensoft" docAuthor="Agarwal, Ishan, Thackeray, Tejas &amp; Khandekar, Akshay" docDate="2024" docId="25CAB9B1D4545BA8B9DEA23523D650DA" docLanguage="en" docName="VertZool 74: 23-83" docOrigin="Vertebrate Zoology 74" docPubDate="2024-01-19" docSource="http://dx.doi.org/10.3897/vz.74.e110674" docTitle="Dravidoseps jawadhuensis Agarwal &amp; Thackeray &amp; Khandekar 2024, sp. nov." docType="treatment" docUuid="D9A6A3BD-173D-4D7C-B4C5-2FF1901434AC" docUuidSource="ZooBank" docVersion="1" id="41B30B26C7D754A6847973C6FED0710C" lastPageNumber="23" masterDocId="41B30B26C7D754A6847973C6FED0710C" masterDocTitle="A non-adaptive radiation of viviparous skinks from the seasonal tropics of India: Systematics of Subdoluseps (Squamata: Scincidae), with description of a new genus and five cryptic new species" masterLastPageNumber="83" masterPageNumber="23" pageNumber="23" updateTime="1705718409033" updateUser="pensoft">
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<mods:title id="E40AADAEE4F3EA522B256794F80600CD">A non-adaptive radiation of viviparous skinks from the seasonal tropics of India: Systematics of Subdoluseps (Squamata: Scincidae), with description of a new genus and five cryptic new species</mods:title>
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<mods:namePart id="371A0AFA07CAE68A3244A7A4758AE0EE">Agarwal, Ishan</mods:namePart>
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<mods:affiliation id="25D461C51610926F21FB014CE86B6101">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
<mods:nameIdentifier id="2FB0D6FD2BA405755D046AF74C0C627F" type="email">ishan.agarwal@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E4C8851535555B0A2B4D15AA30BCB26C">Thackeray, Tejas</mods:namePart>
<mods:nameIdentifier id="5E3432106740F4624BFD370A8421C3A4" type="ORCID">https://orcid.org/0000-0002-9981-8763</mods:nameIdentifier>
<mods:affiliation id="0670312AD109BA101D1E2E9DF435DB6C">Thackeray Wildlife Foundation, Mumbai, 400051, India</mods:affiliation>
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<mods:namePart id="0C8BC981DA96D2E5A73F9B5F179085ED">Khandekar, Akshay</mods:namePart>
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<mods:affiliation id="FF7FE2BFCBACE1931D3E43594DEF3A05">Thackeray Wildlife Foundation, Mumbai, 400051, India &amp; Department of Zoology, Shivaji University, Kolhapur, 416004, India</mods:affiliation>
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<mods:date id="F10677D2146C8CC45A671F2CB121C694">2024</mods:date>
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<mods:identifier id="273A8D6431AE1D9E67380B804A92CCDF" type="DOI">http://dx.doi.org/10.3897/vz.74.e110674</mods:identifier>
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<subSubSection id="32906E3BDDB1B191D59BF81D73A3BF9A" pageId="0" pageNumber="23" type="nomenclature">
<paragraph id="3E56000DC69041CC26877A0200C23783" pageId="0" pageNumber="23">
<taxonomicName id="1BC0A31021DAA2D4E9D00ED39E39CACE" LSID="https://zoobank.org/D9A6A3BD-173D-4D7C-B4C5-2FF1901434AC" authority="Agarwal &amp; Thackeray &amp; Khandekar, 2024" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dravidoseps jawadhuensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="jawadhuensis" status="sp. nov.">Dravidoseps jawadhuensis</taxonomicName>
<taxonomicNameLabel id="419378AE66CB2866C20171856AF26547" pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="786201479BFACAC80B783AF53DE0DF40" pageId="0" pageNumber="23" type="description">
<paragraph id="17C20AEBC9E373D030E5B055EFEE03A7" pageId="0" pageNumber="23">
<figureCitation id="AFAD09EBEFD22444C2D7A55A28902BE8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dravidoseps gen. nov. species in life: A D. pruthi comb. nov. (neotype, NRC-AA- 1291); B D. goaensis comb. nov. (neotype, BNHS 2567); C D. nilgiriensis comb. nov. (ZSI-R- 28604); D D. gingeeensis sp. nov. (holotype, NRC-AA- 8273); E D. jawadhuensis sp. nov. (holotype, NRC-AA- 8274); F D. kalakadensis sp. nov. (holotype, NRC-AA- 8275); G D. srivilliputhurensis sp. nov. (holotype, NRC-AA- 8279); and H D. tamilnaduensis sp. nov. (holotype, NRC-AA- 8286). Photos by Akshay Khandekar (B-D, F-H) and Ishan Agarwal (A, E)." figureDoi="10.3897/vz.74.e110674.figure9" httpUri="https://binary.pensoft.net/fig/970068" pageId="0" pageNumber="23">Figures 9E</figureCitation>
<figureCitation id="C4C33EC352C872B7E9C91DA6C33C8F6F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5CE52603EED37494E96532E2F84780EB" pageId="0" pageNumber="23" type="chresonymy">
<paragraph id="B7E96268746ADE42B9156BEB749F9083" pageId="0" pageNumber="23">Chresonymy.</paragraph>
<paragraph id="5088EAB8DA59AD71865F9793475986E6" pageId="0" pageNumber="23">
<taxonomicName id="0B41302CCE3F58C46C9F1474B972F8FA" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygosoma pruthi" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="pruthi">Lygosoma cf. pruthi</taxonomicName>
-
<bibRefCitation id="1A43EC9B2EF33BC757A503256A25DD4C" author="Ganesh, SR" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="23" pagination="7 - 24" refId="B41" refString="Ganesh, SR, Arumugam, M, 2016. Species richness of montane herpetofauna of southern Eastern Ghats, India: A historical resume and a descriptive checklist. Russian Journal of Herpetology 23: 7 - 24" title="Species richness of montane herpetofauna of southern Eastern Ghats, India: A historical resume and a descriptive checklist." volume="23" year="2016">Ganesh and Arumugam (2016)</bibRefCitation>
</paragraph>
<paragraph id="D8A4B0AB49C21B006A68002418A4AE21" pageId="0" pageNumber="23">
<taxonomicName id="F44889762EABD7A81ED0371DB27319AE" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygosoma pruthi" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="pruthi">
<emphasis id="9B86E2D1E03C0B530E622C8AAE2B6834" italics="true" pageId="0" pageNumber="23">Lygosoma pruthi</emphasis>
</taxonomicName>
-
<bibRefCitation id="5F692EE2266B1A2521DD0B65CB5F9A28" author="Ganesh, SR" journalOrPublisher="Asian Journal of Conservation Biology" pageId="0" pageNumber="23" pagination="128 - 132" refId="B42" refString="Ganesh, SR, Aengals, R, 2018. On further specimens of the poorly-known Pruthi's skink Lygosoma pruthi (Sharma, 1977) with an expanded description. Asian Journal of Conservation Biology 7: 128 - 132" title="On further specimens of the poorly-known Pruthi's skink Lygosoma pruthi (Sharma, 1977) with an expanded description." volume="7" year="2018">Ganesh and Aengals (2018)</bibRefCitation>
</paragraph>
<paragraph id="4E9B8B841C9D03E28FD00445C19D1B47" pageId="0" pageNumber="23">
<taxonomicName id="61EDDD2139D015D9DC6E565604374964" baseAuthorityName="Sharma" baseAuthorityYear="1977" class="Reptilia" family="Scincidae" genus="Subdoluseps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Subdoluseps pruthi" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="pruthi">
<emphasis id="CC89E388C4ACC33FF092EC2D438F767D" italics="true" pageId="0" pageNumber="23">Subdoluseps pruthi</emphasis>
</taxonomicName>
-
<bibRefCitation id="A9CEE290F59A8D84B40D61279822D80E" DOI="https://doi.org/10.11646/zootaxa.4950.2.7" author="Ganesh, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="23" pagination="361 - 376" refId="B44" refString="Ganesh, SR, Srikanthan, AN, Ghosh, A, Adhikari, OD, Kumar, SV, Datta-Roy, A, 2021. A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India. Zootaxa 4950: 361 - 376, DOI: https://doi.org/10.11646/zootaxa.4950.2.7" title="A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India." url="https://doi.org/10.11646/zootaxa.4950.2.7" volume="4950" year="2021">Ganesh et al. (2021)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1319E34725AABC355F8F3D775741569" pageId="0" pageNumber="23" type="holotype">
<paragraph id="7B659D519B36EB145F406DDFD907196F" pageId="0" pageNumber="23">Holotype.</paragraph>
<paragraph id="383875DFCE796A5821A31DAAC1DD5FCB" pageId="0" pageNumber="23">
NRC-AA-8274 (CES 09/930), adult male, from near Beeman falls (
<geoCoordinate id="000CA790E5A1A992058EAD1206E5435B" degrees="12.60174" direction="north" orientation="latitude" precision="1" value="12.60174">12.60174°N</geoCoordinate>
,
<geoCoordinate id="FDBAB5E17F121A661A3FDC44F8AB8FCF" degrees="78.84590" direction="east" orientation="longitude" precision="1" value="78.8459">78.84590°E</geoCoordinate>
; elevation ca. 450 m asl.), Jawadhu Hills, Vellore District, Tamil Nadu State, India, collected by Ishan Agarwal and team on 12th July 2009.
</paragraph>
</subSubSection>
<subSubSection id="E8ACF8F91490C41F6D0EC0D27A413BA0" pageId="0" pageNumber="23" type="paratype">
<paragraph id="641C2AABEFEC1DD04282BC9347E915A7" pageId="0" pageNumber="23">Paratype.</paragraph>
<paragraph id="679357534D63CF57CE1FB2E86B0DE802" pageId="0" pageNumber="23">
BNHS 2569 (AK 850), subadult, from Beeman falls car parking (
<geoCoordinate id="245FD1B5444AE80F5788B5F199712974" degrees="12.60513" direction="north" orientation="latitude" precision="1" value="12.60513">12.60513°N</geoCoordinate>
,
<geoCoordinate id="DBE82911D1E9F8E16B2B389DEE333576" degrees="78.86947" direction="east" orientation="longitude" precision="1" value="78.86947">78.86947°E</geoCoordinate>
; elevation ca. 580 m asl.), same data as holotype except collected by Akshay Khandekar, Ishan Agarwal, Swapnil Pawar, Tejas Thackeray and team on 4th June 2019.
</paragraph>
</subSubSection>
<subSubSection id="4597816F6DB23634A603550675FFF5E9" pageId="0" pageNumber="23" type="etymology">
<paragraph id="50EFC3B565C4AAEFDC56B74A46024ED5" pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph id="39B8B0B9EDE4FB06EAF37C39F1C7E756" pageId="0" pageNumber="23">The specific epithet is a toponym for the Jawadhu Hills in Vellore District of Tamil Nadu State, the type and currently only known locality for the new species.</paragraph>
</subSubSection>
<subSubSection id="39D8B4D8529750046EE6037024FD2A7E" pageId="0" pageNumber="23" type="suggested common name">
<paragraph id="BCA99D83DCF6D05228DD910175282DB2" pageId="0" pageNumber="23">Suggested common name.</paragraph>
<paragraph id="B1208B3CD9717831E8AF016025E65A13" pageId="0" pageNumber="23">Jawadhu leaf-litter skink.</paragraph>
</subSubSection>
<subSubSection id="65C17FE7C7A2E9E239EBD2E6F6A9BC63" pageId="0" pageNumber="23" type="diagnosis">
<paragraph id="F77B2B3B790F1B7228AC2A540AE29E7A" pageId="0" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="BAA0386D0CE06BC277F8ED2369BDA48B" pageId="0" pageNumber="23">A medium-sized skink snout to vent length up to 47 mm (n = 2). Seven supralabials and six infralabials up to angle of mouth; fifth supralabial elongate and below eye; two post-supralabials; seven or eight supraciliaries; one or two elongated nuchals on either side, in contact with each other behind parietal; 65 or 66 scales in paravertebral rows; 30-32 scales around mid-body; 66-68 ventral scales; 12 or 13 enlarged precloacal scales; scales on lateral sides of tail base smooth, 22 or 23 scales around the tail. Subdigital lamellae unpaired, smooth on manus and smooth to weakly keeled on pes; six or seven lamellae under digit I of manus and five or six under digit I of pes; 11 lamellae under digit IV of manus and, 16 or 17 under digit IV of pes. Dorsum dark brown with black markings; thick black stripe from rostrum to tail speckled with light spots; males with yellow on lower parts of forebody and flanks; supralabials with white streak; venter glossy off-white with some darker markings.</paragraph>
</subSubSection>
<subSubSection id="EB81E976A6EA73CDAD65E98546F9E7C0" pageId="0" pageNumber="23" type="comparisons">
<paragraph id="626D3D8D7895DC3867DB496442756F09" pageId="0" pageNumber="23">Comparisons.</paragraph>
<paragraph id="FD1D39A132719AAE918F3AD01071F60E" pageId="0" pageNumber="23">
<taxonomicName id="262EE59C2C06589D3C1982F8EF1D604F" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps jawadhuensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="jawadhuensis">
<emphasis id="EDBE2B063D933DAB1FA6CD9688B829A6" italics="true" pageId="0" pageNumber="23">Dravidoseps jawadhuensis</emphasis>
</taxonomicName>
<emphasis id="084CD9292021E4FDDED75B4348825C69" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
can be diagnosed from known congeners based on the following characters: five PoSbO on each side (versus three or four in
<taxonomicName id="03EB7340599058E68CDB695D2F17B026" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps gingeeensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="gingeeensis">
<emphasis id="17555480FC04CF474D17E3C3E563AE0C" italics="true" pageId="0" pageNumber="23">Dravidoseps gingeeensis</emphasis>
</taxonomicName>
<emphasis id="5EF65C2C44236C66333507F24DC1B3E9" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
, four (rarely three or five on one side) in
<taxonomicName id="F032A5C6815D8E0FB10E81EB0AA6B920" lsidName="D. nilgiriensis" pageId="0" pageNumber="23" rank="species" species="nilgiriensis">
<emphasis id="CF84D584939582824EE628E79C9729FF" italics="true" pageId="0" pageNumber="23">D. nilgiriensis</emphasis>
</taxonomicName>
<emphasis id="0250A960D36B9DF3DB219A538B32B14C" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
, four on each side in
<taxonomicName id="688D0875556713DAA7008CCB3D70D4CB" lsidName="D. goaensis" pageId="0" pageNumber="23" rank="species" species="goaensis">
<emphasis id="DD24C26FFCE7D94A067D87A866C0A626" italics="true" pageId="0" pageNumber="23">D. goaensis</emphasis>
</taxonomicName>
<emphasis id="1C43BE66B50EBE78E9DDB6C018FE2E9B" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
and in
<taxonomicName id="88E45E1853A9D17EEC770B432DE03006" lsidName="D. pruthi" pageId="0" pageNumber="23" rank="species" species="pruthi">
<emphasis id="F813824C7108C8FC33A189FB3E88ACD6" italics="true" pageId="0" pageNumber="23">D. pruthi</emphasis>
</taxonomicName>
<emphasis id="74ADD10FF424360C2D8D9097D562A940" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
); 12.5
<normalizedToken id="E06EAAEADBFC2954BE912AE198290C38" originalValue="±">+/-</normalizedToken>
0.71 (12-13) SPCLR (versus 9.2
<normalizedToken id="92DC4D90EF674217CC913308599FF519" originalValue="±">+/-</normalizedToken>
0.60 (8-10) in
<taxonomicName id="2E17CA88254D599FB0ACAC1EF96F6BE2" lsidName="D. nilgiriensis" pageId="0" pageNumber="23" rank="species" species="nilgiriensis">
<emphasis id="EC513393C0A15D87465636AC493E766C" italics="true" pageId="0" pageNumber="23">D. nilgiriensis</emphasis>
</taxonomicName>
<emphasis id="35085E42B3D7F83E4FA4BAD7CB1D0100" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
, 10.0
<normalizedToken id="7BD29D2F618EFEF68AD0D414494A8B39" originalValue="±">+/-</normalizedToken>
0.00 (10) in
<taxonomicName id="20F7E800A685E54B26AE8FD95920ACD5" lsidName="D. pruthi" pageId="0" pageNumber="23" rank="species" species="pruthi">
<emphasis id="245B3BEE8F6CCDCDF106E55C2F063894" italics="true" pageId="0" pageNumber="23">D. pruthi</emphasis>
</taxonomicName>
<emphasis id="99290A21B6C91401CFDCCFDD9F160EC6" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
, and 8.3
<normalizedToken id="DF700D2C13D72D4E8AEB2A2FD18E0F19" originalValue="±">+/-</normalizedToken>
0.76 (8-10) in
<taxonomicName id="C29CB43646C8F148A8FFBFD41E8C73B9" lsidName="D. goaensis" pageId="0" pageNumber="23" rank="species" species="goaensis">
<emphasis id="B29E06ECC067C664F66137FC4BBE4DCC" italics="true" pageId="0" pageNumber="23">D. goaensis</emphasis>
</taxonomicName>
<emphasis id="99D1A63D433C695B9A268045BB53919C" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
); presence of unkeeled scales on tail base (versus keeled scales on tail base in
<taxonomicName id="3197829FB9514869815648EC95F0D365" lsidName="D. goaensis" pageId="0" pageNumber="23" rank="species" species="goaensis">
<emphasis id="E82350A7DF6DCB21C8B515E565FB799A" italics="true" pageId="0" pageNumber="23">D. goaensis</emphasis>
</taxonomicName>
<emphasis id="B447ABE8CDA93E3A326AE615517D1F35" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
); two PoSL on either side (versus single on either in
<taxonomicName id="056FEA5F1E0950BF71BC873B0E2314D3" lsidName="D. pruthi" pageId="0" pageNumber="23" rank="species" species="pruthi">
<emphasis id="24CC3ABBBD197302D2B63A1145916549" italics="true" pageId="0" pageNumber="23">D. pruthi</emphasis>
</taxonomicName>
<emphasis id="B045F9FA5BA3EF27D2F14EA41B8D3CEB" bold="true" pageId="0" pageNumber="23">comb. nov.</emphasis>
); one or two Nu on either side and Sb Nu absent (versus a single Nu on either side and three Sb Nu in
<taxonomicName id="8C6DF7C7E35544CA8E8E55E5235DA325" lsidName="D. gingeeensis" pageId="0" pageNumber="23" rank="species" species="gingeeensis">
<emphasis id="D3B1D30DC221ED68ABF0A31097879012" italics="true" pageId="0" pageNumber="23">D. gingeeensis</emphasis>
</taxonomicName>
<emphasis id="ABD5AE960145439B64C60AF1F5578513" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
); 22 or 23 RTS (versus 21 RTS in
<taxonomicName id="EDAFC261A15D9D179314032D2E3927FF" lsidName="D. gingeeensis" pageId="0" pageNumber="23" rank="species" species="gingeeensis">
<emphasis id="748CA54DB792B2B20D5CF8C1096C5518" italics="true" pageId="0" pageNumber="23">D. gingeeensis</emphasis>
</taxonomicName>
<emphasis id="C2FD86CC3385012CEB863B66060DC405" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
).
<taxonomicName id="AFEEE9EF17C651DA9C9E1E4264671801" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps jawadhuensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="jawadhuensis">
<emphasis id="293B692CE19B7DB114D3C8F46A675894" italics="true" pageId="0" pageNumber="23">Dravidoseps jawadhuensis</emphasis>
</taxonomicName>
<emphasis id="5F4C471CEA1433EED850F92235024743" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
is diagnosed against the new species described below as part of their respective descriptions.
</paragraph>
</subSubSection>
<subSubSection id="7CB7E0AF52CCAE7A485E5170EC80A510" pageId="0" pageNumber="23" type="description">
<paragraph id="EB9465E04703C779227D7174CCDC8E54" pageId="0" pageNumber="23">Description of the holotype.</paragraph>
<paragraph id="9CB75857CDE6756123F5849086FD591D" pageId="0" pageNumber="23">
Adult male (SVL 45.7 mm) in good state of preservation except more than half of the tail missing (Fig.
<figureCitation id="87B0F7B12E68CFAC0C203D9F77C75FD2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14A, B</figureCitation>
). Head short (HL/SVL 0.16), wide (HW/HL 0.77), not strongly depressed (HH/HL 0.55), indistinct from neck. Loreal region not inflated, canthus rostralis indistinct. Snout almost half head length (ES/HL 0.44), two times eye diameter (ES/ED 2.00). Rostral almost twice as wide (1.7 mm) as long (0.9 mm), in broad contact with supranasals posteriorly and supralabial I and nasals on either side; supranasals marginally separated from each other below rostral by frontonasal, in contact with frontonasal posteriorly, nasals and anterior loreals laterally; frontonasal much wider (1.7 mm) than long (1.2 mm), in contact with supranasals anteriorly, prefrontals and frontal posteriorly, anterior loreals laterally; prefrontals relatively small, widely separated on midline, in contact with frontonasal anteriorly, frontal, first supraocular and first supraciliary posteriorly, anterior and posterior loreals laterally. Frontal elongate, roughly bell-shaped, widest anteriorly at the point where prefrontals and first supraocular connect; in contact with frontonasal anteriorly, frontoparietals posteriorly, prefrontals and first two supraoculars on either side; four supraoculars and one small post-supraocular and postocular on either side; frontoparietals in medial contact posterior to frontal, in contact with second, third, and fourth supraoculars anterolaterally and parietals and interparietal posteriorly. Interparietal large, roughly diamond-shaped, slightly projecting posteriorly, eyespot in posterior projection; postinterparietal absent; parietals large, in medial contact posterior to interparietal, in contact anteriorly with frontoparietals, fourth supraocular, and post-supraocular, two nuchal scales posteriorly, first secondary temporal laterally; a single enlarged, elongate, nuchal scale on either side in median contact posterior to parietals (Fig.
<figureCitation id="99A45137B3C745AEB9D9B459112A3095" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14C</figureCitation>
). Nasals small, trapezoidal, widely separated, in contact with rostral anteriorly, supranasal dorsally, anterior loreal posteriorly, first supralabial ventrally; nostril in center of nasal; anterior loreal slightly taller (0.8 mm) than wide (0.5 mm); posterior loreal larger than anterior loreal and slightly wider (0.9 mm) than tall (0.7 mm); a single small supra-preocular, an upper and lower preocular, and a single sub-preocular present on either sides (Fig.
<figureCitation id="FF22831C4DEF029BD48D6AFD54FA00A2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14E</figureCitation>
). Eye small (ED/ HL 0.22) with round pupil; lower eyelid with enlarged, transparent central window; eight supraciliaries on left and seven on right, anterior supraciliary largest, bordered by prefrontal anteriorly, first supraocular dorsally, and pre-supraocular, upper preocular and posterior loreal laterally; posterior superciliary elongate and projecting dorsomedially, bordered by fourth supraocular dorsally, post-supraocular posteriorly, and first post-subocular laterally; five post-suboculars on either side; a single primary temporal, two secondary temporals, and three tertiary temporals on either side; seven supralabials, fifth elongate and in broad contact with pre-subocular, last post-subocular and five small scales on lower eyelid below eye on left and four on right side; two post-supralabials on either side; six infralabials on either side; two scales separating post-supralabial and external ear opening; external ear opening small (EL/HL 0.07), oval, bearing three anterior lobules on either side; tympanum deep (Fig.
<figureCitation id="DCE8841456AA2C1B132CC0C025953DA3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14E</figureCitation>
). Mental twice wide (2.0 mm) as long (1.0 mm); a single large postmental in contact with first and second infralabials on either side; three enlarged pairs of chin shields posterior to postmental; anterior pair large (1.4 mm), roughly rectangular, in medial contact with each other below postmental and bordered by second and third infralabials, middle pair of chin shields, and by a single median gular scale on either side; middle pair largest (1.7 mm), roughly rectangular, separated from each other by two longitudinally arranged gular scales, bordered by third and fourth infralabials, posterior pair, and four gular scales on either side; posterior pair smallest (0.9 mm), roughly square, separated from each other by five transversely arranged gular scales, bordered by fourth and fifth infralabials and three gular scales on either side; rest of the gular scales much smaller than postmentals, cycloid and imbricate, two or three rows bordering infralabials slightly smaller and elongate (Fig.
<figureCitation id="49E041664C9D2C1995386AB0FC708E5F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14D</figureCitation>
).
</paragraph>
<caption id="1EE8B48AF0A168DCE9DA1C5D27337E09" doi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23" start="Figure 14" startId="F14">
<paragraph id="A2E34EC1A56FE1C52CB84A0FAB16B32A" pageId="0" pageNumber="23">
<emphasis id="E0BEA26064F521B2B3690E8406B7896E" bold="true" pageId="0" pageNumber="23">Figure 14.</emphasis>
<taxonomicName id="755647FAA32BAECE1A332F82DFA9E678" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps jawadhuensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="jawadhuensis">
<emphasis id="A182ED059A6AC7A3398731F13D286BEF" italics="true" pageId="0" pageNumber="23">Dravidoseps jawadhuensis</emphasis>
</taxonomicName>
<emphasis id="EBE65204C38DD8E0C39FDA5055FA77F6" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
(holotype, NRC-AA-8274):
<emphasis id="AA5304958FA742C46EF298B60EB26602" bold="true" pageId="0" pageNumber="23">A</emphasis>
dorsal view of body,
<emphasis id="37D37DCEB86D4483F6E7919F10613723" bold="true" pageId="0" pageNumber="23">B</emphasis>
ventral view of body,
<emphasis id="81668B0F2FA9F7181D46F06651827CA0" bold="true" pageId="0" pageNumber="23">C</emphasis>
dorsal view of head,
<emphasis id="DE8F17DFF91E20769F507D039747DEDF" bold="true" pageId="0" pageNumber="23">D</emphasis>
ventral view of head,
<emphasis id="1C6B77EDBC6002B359EEF3058115FCEE" bold="true" pageId="0" pageNumber="23">E</emphasis>
lateral right side view of head,
<emphasis id="D1DC08321DC734F4A9B43E7022F59692" bold="true" pageId="0" pageNumber="23">F</emphasis>
ventral view of left manus, and
<emphasis id="B4C64C0C6B35278DFCD2F70276E40F06" bold="true" pageId="0" pageNumber="23">G</emphasis>
ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar.
</paragraph>
</caption>
<paragraph id="E6AEAC7331DCF8096F52478739374D9B" pageId="0" pageNumber="23">
Body relatively slender (BW/AGL 0.27), elongate (AGL/SVL = 0.59); dorsal scales on body smooth, cycloid, imbricate; ventrals similar to dorsals except subequal from chest to vent, marginally larger on pectoral and precloacal region; 65 scales in paravertebral rows; 30 scales around mid-body; 68 ventral scales; 13 enlarged precloacal scales (Fig.
<figureCitation id="F527E89A223E64EA3456AC4F5882A0CF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14A, B</figureCitation>
). Limbs, robust, short (FL/SVL = 0.07; CL/SVL = 0.09), widely separated when adpressed; dorsal scales wider and slightly larger than ventral scales; palmar scales raised; plantar scales large, raised, coarse granules; all digits short, scales on dorsal surfaces in single row, subdigital lamellae unpaired, smooth on manus and smooth to weakly keeled on pes; lamellae series: 6-8-11-11-8 left manus (Fig.
<figureCitation id="98E51899400E0467811B98E7FDC38FA5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14F</figureCitation>
), 6-10-15-17-12 left pes (Fig.
<figureCitation id="146FDBFEBBB25C81A8FF22A055A37443" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14G</figureCitation>
), 6-8-10-11-8 right manus, 5-11-14-17-12 right pes. Relative length of digits (measurements in mm in parentheses): IV (2.3)&gt; III (2.2)&gt; II (1.6)&gt; V (1.4)&gt; I (0.7) (left manus); IV (4.9)&gt; III (4.1)&gt; V (3.1)&gt; II (2.4)&gt; I (1.5) (left pes).
</paragraph>
<paragraph id="71AAEF54FB831E201BDF4299B0285226" pageId="0" pageNumber="23">
Tail original, cylindrical, more than half broken and lost; dorsal and ventral scales smooth, cycloid, imbricate, similar to those on body dorsum; scales on lateral tail base smooth, 22 scales around the tail (Fig.
<figureCitation id="C3CBED0A4B1D901B026D1F3E40299971" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dravidoseps jawadhuensis sp. nov. (holotype, NRC-AA- 8274): A dorsal view of body, B ventral view of body, C dorsal view of head, D ventral view of head, E lateral right side view of head, F ventral view of left manus, and G ventral view of left pes. Scale bars: A, B = 10 mm; C-E, G = 5 mm; F = 3 mm; photos by Akshay Khandekar." figureDoi="10.3897/vz.74.e110674.figure14" httpUri="https://binary.pensoft.net/fig/970073" pageId="0" pageNumber="23">14A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01706F485F4B2A604665128CC0417022" pageId="0" pageNumber="23" type="colouration in life (fig. 9e)">
<paragraph id="00DD218A5D607C9CEEC3CF71873CA937" pageId="0" pageNumber="23">Colouration in life (Fig. 9E).</paragraph>
<paragraph id="9E9E922A5D1EBF14DA5B1E2B2225C913" pageId="0" pageNumber="23">Dorsal ground colouration of body, head and tail dark brown; head with a few dark blotches on and between supraoculars; dorsal scales of body and tail finely outlined by dark brown, centre of scales with dark markings forming indistinct stripes; limbs darker than body dorsum and with light spots; a thick dark black stripe running from rostrum through orbit and onto flank and tail with scattered light spots; supralabials with a white streak; two or three rows of scales below the dark band between ear opening and mid-body yellow; ventral regions glossy cream with fine dark stripes.</paragraph>
</subSubSection>
<subSubSection id="1D1BBD206C67D78D0D70247A5E6355A1" pageId="0" pageNumber="23" type="variation">
<paragraph id="59D45AA2F45D2F87F93613FECAA8380D" pageId="0" pageNumber="23">Variation and additional information.</paragraph>
<paragraph id="8B49F576FFB4740E5E5A66231997250D" pageId="0" pageNumber="23">
Mensural and meristic data for the subadult paratype (SVL = 32.5 mm) is given in Table
<tableCitation id="D8DCAE0D4A573AF6ADF182657D53D815" captionStart="Table 8" captionStartId="T9" captionText="Table 8. Mensural (mm) and meristic data for Dravidoseps gingeeensis sp. nov. and Dravidoseps jawadhuensis sp. nov.. Abbreviations are listed in Materials and Methods except for: L &amp; R = Left &amp; Right; M = male; F = female; Sa = Subadult; * = tail incomplete." pageId="0" pageNumber="23">8</tableCitation>
. It resembles the holotype male (NRC-AA-8274) in overall morphology and head scalation except for the following variation: supranasals in contact with each other behind rostral; seven SC present on either side; anterior pair of CS separated from each other below postmental; and tail tip collected in molecular grade ethanol for DNA extraction.
</paragraph>
</subSubSection>
<subSubSection id="441A58C932A194441674C8E3076E6CF6" pageId="0" pageNumber="23" type="distribution">
<paragraph id="093F57079CCFC212BCAD806A54E84AAD" pageId="0" pageNumber="23">Distribution and natural history.</paragraph>
<paragraph id="D2F123BF996AAC37FAAD14975233E17D" pageId="0" pageNumber="23">
<taxonomicName id="5BCC0399BDFEDB28A24CA8CCB1194E9E" authorityName="Agarwal &amp; Thackeray &amp; Khandekar" authorityYear="2024" family="Scincidae" genus="Dravidoseps" kingdom="Animalia" lsidName="Dravidoseps jawadhuensis" order="Squamata" pageId="0" pageNumber="23" rank="species" species="jawadhuensis">
<emphasis id="DB720DD927CA287BE13A659CAF67F58D" italics="true" pageId="0" pageNumber="23">Dravidoseps jawadhuensis</emphasis>
</taxonomicName>
<emphasis id="9DA6AD8FE05914E8F8DCE7497E147D0E" bold="true" pageId="0" pageNumber="23">sp. nov.</emphasis>
is known only from the type locality (in and around Beeman falls), Jawadhu Hills in Vellore District, Tamil Nadu, India (Fig.
<figureCitation id="A1FB95644FF39032DE6A7039A7791D25" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Elevation map showing sampling localities from peninsular India. Stars indicate type localities (neotype locality shown for Dravidoseps goaensis comb. nov., original type locality is close to the southernmost point), light blue D. jawadhuensis sp. nov., dark blue D. srivilliputhurensis sp. nov., light green D. kalakadensis sp. nov., dark green D. pruthi comb. nov., red D. gingeeensis sp. nov., yellow D. nilgiriensis comb. nov., pink D. tamilnaduensis sp. nov., and grey, D. sp. Kalrayan (colour scheme representing species is the same as Figs 3 - 6), black squares unsampled previously known localities. The approximate extent of the Central Western Ghats (CWG), Northern Western Ghats (NWG), and Southern Western Ghats (SWG) are shown; major hill ranges and features are marked by bold white text, within the Western Ghats: AG, Agasthyamalai; AN, Anaimalai; DV, Devarmalai; N, Nilgiris; PG, Palghat Gap; PL, Palani; and outside the Western Ghats: BR, Biligirangan; J, Jawadhu; K, Kollimalai; KR, Kalrayan; mm, Male Mahadeshwara; MP, Mysore Plateau; PC, Pachaimalai; PM, Palamalai; S, Sitteri; SR, Sirumalai; T, Tirupati; V, Velikonda; Y, Yercaud; YL, Yelagiri. Inset, global distribution of Dravidoseps gen. nov. (brown) and Subdoluseps sensu stricto (green). Dravidoseps goaensis = Riopa goaensis (see main text)." figureDoi="10.3897/vz.74.e110674.figure1" httpUri="https://binary.pensoft.net/fig/970060" pageId="0" pageNumber="23">1</figureCitation>
). The new species was collected between elevations of 450-580 m from moist deciduous forests with scattered open rocky patches (Fig.
<figureCitation id="1F2E4C6D932C697351330398AFE28ECA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. General habitats of all species of Dravidoseps sp. nov.: A type locality of D. pruthi comb. nov.; B neotype locality of D. goaensis comb. nov.; C habitat photo of D. nilgiriensis comb. nov., from Thamarakarai; D type locality of D. gingeeensis sp. nov.; E type locality of D. jawadhuensis sp. nov.; F type locality of D. kalakadensis sp. nov.; G paratype locality of D. srivilliputhurensis sp. nov., near Atthi Kovil Falls; and H type locality of D. tamilnaduensis sp. nov.. Photos by Akshay Khandekar (A-C, F-H), Ishan Agarwal (D), and Vinod (E)." figureDoi="10.3897/vz.74.e110674.figure11" httpUri="https://binary.pensoft.net/fig/970070" pageId="0" pageNumber="23">11E</figureCitation>
). The holotype was observed active at ~ 1130 hrs at the mouth of an ant burrow in a patch of deciduous forest with leaf litter. The subadult paratype was collected during the day time (1130 hrs) and was found under a rock surrounded by dry leaf-litter along a path leading towards Beeman fall. Sympatric lizards recorded were
<taxonomicName id="A87EEEAA0B7980F79F2C3009808150EC" baseAuthorityName="Beddome" baseAuthorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Calodactylodes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Calodactylodes aureus" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="aureus">
<emphasis id="8B6F191681A60F240DC861C17C5BBCA2" italics="true" pageId="0" pageNumber="23">Calodactylodes aureus</emphasis>
</taxonomicName>
,
<taxonomicName id="C6136827C49F4998D02B3AD7F9AA9C36" baseAuthorityName="Jerdon" baseAuthorityYear="1853" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cnemaspis mysoriensis" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="mysoriensis">Cnemaspis cf. mysoriensis</taxonomicName>
,
<taxonomicName id="74EA037ED31F34B9526849AE54202CFF" lsidName="Cn. otai" pageId="0" pageNumber="23" rank="species" species="otai">
<emphasis id="0B12B6CE27011E7BA441D4E5C22249B7" italics="true" pageId="0" pageNumber="23">Cn. otai</emphasis>
</taxonomicName>
Das &amp; Bauer,
<taxonomicName id="318980AB63EFE317BC08D9C2C433AF07" baseAuthorityName="Beddome" baseAuthorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus (Geckoella) collegalensis" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="collegalensis" subGenus="Geckoella">Cyrtodactylus (Geckoella) cf. collegalensis</taxonomicName>
,
<taxonomicName id="CBAB685A36FD0FCFD9E42A25351CE3AE" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus frenatus" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="73F696329556DCAE805F2813664F50C7" italics="true" pageId="0" pageNumber="23">Hemidactylus frenatus</emphasis>
</taxonomicName>
,
<taxonomicName id="EDBAFAFA4E2AC9340E6AA9255C8EE328" lsidName="H. graniticolus" pageId="0" pageNumber="23" rank="species" species="graniticolus">H. cf. graniticolus</taxonomicName>
,
<taxonomicName id="63838569E0BD7A83514677D438233865" lsidName="H. leschenaultii" pageId="0" pageNumber="23" rank="species" species="leschenaultii">
<emphasis id="3D3826DBEE938BBF59FC194F0EC4D9B1" italics="true" pageId="0" pageNumber="23">H. leschenaultii</emphasis>
</taxonomicName>
,
<taxonomicName id="EFB50566C8362B86E0D4C78FBCB1E8DD" baseAuthorityName="Schneider" baseAuthorityYear="1801" class="Actinopterygii" family="Scincidae" genus="Eutropis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutropis carinata" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="carinata">
<emphasis id="B5DB5F8A8B1BFAE88EF1046CEAB0B507" italics="true" pageId="0" pageNumber="23">Eutropis carinata</emphasis>
</taxonomicName>
,
<taxonomicName id="50EA605679CE615E3A137CB40A7A7122" lsidName="E. macularia" pageId="0" pageNumber="23" rank="species" species="macularia">
<emphasis id="ACDB67077B705C0F5FE4B3469F145CDD" italics="true" pageId="0" pageNumber="23">E. macularia</emphasis>
</taxonomicName>
,
<taxonomicName id="AC4115A569DBA0AB492EB87FFA8159E2" baseAuthorityName="Daudin" baseAuthorityYear="1802" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" lsidName="Calotes versicolor" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="FFC08CF699B9054732D0B891BE24B67D" italics="true" pageId="0" pageNumber="23">Calotes versicolor</emphasis>
</taxonomicName>
, and
<taxonomicName id="19F1999B453C09ECEAA6B85AC1B1EAD7" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Reptilia" family="Agamidae" genus="Psammophilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psammophilus dorsalis" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="03E274FD92E53C3E46B682E880FB4857" italics="true" pageId="0" pageNumber="23">Psammophilus dorsalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5EED121AC801D227523C73C22C0D6A85" pageId="0" pageNumber="23" type="reproduction">
<paragraph id="4C0DA1F9EAE7C8AAFC301211B1952295" pageId="0" pageNumber="23">Reproduction.</paragraph>
<paragraph id="2D2FDB128C964D0E910BFCB88704236B" pageId="0" pageNumber="23">Unknown, there are no females in the type series.</paragraph>
</subSubSection>
<subSubSection id="F2DDB738BFE1401CBED631557A1E847D" pageId="0" pageNumber="23" type="notes">
<paragraph id="486456FE4089FCA51B4EFB729498845B" pageId="0" pageNumber="23">Note.</paragraph>
<paragraph id="BE47C791967D5F9E869C95103A045B4E" pageId="0" pageNumber="23">
We found the following discrepancies in mensural and meristic characters of specimen CES 09/930 as against the data provided by
<bibRefCitation id="C2DF5AE986F5540A888063D6F0FE79FC" DOI="https://doi.org/10.11646/zootaxa.4950.2.7" author="Ganesh, SR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="23" pagination="361 - 376" refId="B44" refString="Ganesh, SR, Srikanthan, AN, Ghosh, A, Adhikari, OD, Kumar, SV, Datta-Roy, A, 2021. A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India. Zootaxa 4950: 361 - 376, DOI: https://doi.org/10.11646/zootaxa.4950.2.7" title="A new species of Asian gracile skink (Scincidae: Lygosominae: Subdoluseps) from the rain-shadow belts of Nilgiri hills, Western Ghats, India." url="https://doi.org/10.11646/zootaxa.4950.2.7" volume="4950" year="2021">Ganesh et al. (2021)</bibRefCitation>
as
<taxonomicName id="B41F1B6A2DB6EBB687C33A1C4716D6EC" baseAuthorityName="Sharma" baseAuthorityYear="1977" class="Reptilia" family="Scincidae" genus="Subdoluseps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Subdoluseps pruthi" order="Squamata" pageId="0" pageNumber="23" phylum="Chordata" rank="species" species="pruthi">
<emphasis id="AC30E62E6350CB07D8F24A555292F92C" italics="true" pageId="0" pageNumber="23">Subdoluseps pruthi</emphasis>
</taxonomicName>
: TL (13.6* mm versus 31.22 mm; * = tail incomplete), HL (7.6 mm versus 6.27 mm), PVS (65 versus 62), RBS (30 versus 34) VS (68 versus 69), and LamT4 (17 versus 16) (Table
<tableCitation id="0D50DC8C241130B7C5972308065666AB" captionStart="Table 8" captionStartId="T9" captionText="Table 8. Mensural (mm) and meristic data for Dravidoseps gingeeensis sp. nov. and Dravidoseps jawadhuensis sp. nov.. Abbreviations are listed in Materials and Methods except for: L &amp; R = Left &amp; Right; M = male; F = female; Sa = Subadult; * = tail incomplete." pageId="0" pageNumber="23">8</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>