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<document id="261A29C2CCC23875C589E45FCCD8BBCC" ID-DOI="10.5852/ejt.2023.897.2293" ID-GBIF-Dataset="6ce49ff3-917a-45e5-964c-7e88d414455b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8405868" ID-ZooBank="A340BD2E-094A-4C84-88B3-5B4ED220F476" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696412047706" checkinUser="plazi" docAuthor="Valdez-Mondragón, Alejandro &amp; Cabrera-Espinosa, Luis A." docDate="2023" docId="FF24F9045A766523068BFECC28B8FBE9" docLanguage="en" docName="EJT.2023.897.1-56.pdf" docOrigin="European Journal of Taxonomy 897" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2293/9923" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Latrodectus occidentalis Valdez-Mondragon 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="031D817C5A6C6509076AFFC42C59FFD9" masterDocTitle="Phylogenetic analyses and description of a new species of black widow spider of the genus Latrodectus Walckenaer (Araneae, Theridiidae) from Mexico; one or more species?" masterLastPageNumber="56" masterPageNumber="1" pageNumber="27" updateTime="1701999844797" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="69DF4A7892BF31193CC4565634AAB4AF">Phylogenetic analyses and description of a new species of black widow spider of the genus Latrodectus Walckenaer (Araneae, Theridiidae) from Mexico; one or more species?</mods:title>
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<mods:namePart id="EA5B08989236ECFD0BCB256A079CDD5B">Valdez-Mondragón, Alejandro</mods:namePart>
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<mods:affiliation id="F38F020A7075657995105C26157153C8">Colección de Aracnología (CARCIB), Programa Académico de Planeación Ambiental y Conservación (PLAYCO), Centro de Investigaciones Biológicas del Noroeste (CIBNOR), S. C. Km 1 Carretera a San Juan de La Costa ́ El Comitán », La Paz, Baja California Sur, C. P. 23205, Mexico</mods:affiliation>
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<mods:namePart id="A95D611C171753512970FB3B25054716">Cabrera-Espinosa, Luis A.</mods:namePart>
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<mods:affiliation id="6A17842FD7428E76123A5BEB50FF7AD4">Colección de Aracnología (CARCIB), Programa Académico de Planeación Ambiental y Conservación (PLAYCO), Centro de Investigaciones Biológicas del Noroeste (CIBNOR), S. C. Km 1 Carretera a San Juan de La Costa ́ El Comitán », La Paz, Baja California Sur, C. P. 23205, Mexico</mods:affiliation>
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<taxonomicName id="B08D33915A766513068BFECC2FA9FEFA" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[481,1008,264,291]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A766513068BFECC2F5EFEFA" bold="true" box="[481,775,264,291]" italics="true" pageId="26" pageNumber="27">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
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<figureCitation id="EFB654975A76651305B2FE8B2F02FEB0" box="[728,859,335,361]" captionStart-0="Figs 1318" captionStart-1="Figs 1924" captionStart-2="Figs 2528" captionStart-3="Figs 2931" captionStart-4="Figs 3243" captionStart-5="Figs 4448" captionStart-6="Figs 4960" captionStart-7="Figs 6169" captionStart-8="Figs 7075" captionStart-9="Figs 7681" captionStartId-0="30.[189,242,1748,1774]" captionStartId-1="31.[189,242,1775,1801]" captionStartId-2="33.[189,242,1928,1954]" captionStartId-3="34.[185,238,1926,1952]" captionStartId-4="35.[187,240,1822,1848]" captionStartId-5="36.[189,242,1801,1827]" captionStartId-6="38.[188,241,1857,1883]" captionStartId-7="39.[189,242,1741,1767]" captionStartId-8="40.[188,241,1857,1883]" captionStartId-9="41.[189,242,1693,1719]" captionTargetBox-0="[189,1398,268,1700]" captionTargetBox-1="[189,1398,268,1701]" captionTargetBox-2="[189,1398,512,1882]" captionTargetBox-3="[189,1398,958,1879]" captionTargetBox-4="[189,1398,708,1771]" captionTargetBox-5="[189,1398,265,1746]" captionTargetBox-6="[189,1398,745,1807]" captionTargetBox-7="[189,1398,268,1683]" captionTargetBox-8="[189,1398,442,1810]" captionTargetBox-9="[189,1398,267,1639]" captionTargetId-0="figure-14@30.[189,1398,268,1700]" captionTargetId-1="figure-11@31.[189,1398,268,1701]" captionTargetId-2="figure-97@33.[189,1398,512,1882]" captionTargetId-3="figure-253@34.[189,1398,958,1884]" captionTargetId-4="figure-157@35.[189,1398,708,1771]" captionTargetId-5="figure-13@36.[189,1398,265,1746]" captionTargetId-6="figure-193@38.[189,1398,745,1807]" captionTargetId-7="figure-11@39.[189,1398,268,1683]" captionTargetId-8="figure-64@40.[189,1398,442,1810]" captionTargetId-9="figure-11@41.[189,1398,267,1639]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="36" captionTargetPageId-6="38" captionTargetPageId-7="39" captionTargetPageId-8="40" captionTargetPageId-9="41" captionText-0="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" captionText-1="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." captionText-2="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." captionText-3="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." captionText-4="Figs 3243. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the seminal receptacles of females, dorsal view. 3233. Cocula, Jalisco, Mexico (type locality). 3435. Jilotitlán de los Dolores, Jalisco, Mexico. 3637. Minatitlán, Colima, Mexico. 3839. Coahuayana de Hidalgo, Michoacán, Mexico. 4041. Quechultenango, Guerrero, Mexico. 4243. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm." captionText-5="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." captionText-6="Figs 4960. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the male left palps, prolateral views. 4952. Quechultenango, Guerrero, Mexico. 5356. Minatitlán, Colima, Mexico. 57 60. Cocula, Jalisco, Mexico (type locality). Ventral view (49, 53, 57); retrolateral-frontal view (50, 54, 59); retrolateral view (51, 55, 59); posterior view (52, 56, 60). 4956. Postcopulatory palps. Scale bars = 0.2 mm." captionText-7="Figs 6169. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the dorsal and ventral pattern coloration of the abdomen of the females of Latrodectus occidentalis sp. nov. Dorsal, posterior, and ventral views respectively. 6163. Chilpancingo, Guerrero, Mexico. 6466. Minatitlán, Colima, Mexico. 6769. Cocula, Jalisco, Mexico (type locality). Red coloration pattern gets lost under ethanol. Scale bars = 1 mm." captionText-8="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." captionText-9="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi-0="http://doi.org/10.5281/zenodo.8405894" figureDoi-1="http://doi.org/10.5281/zenodo.8405896" figureDoi-2="http://doi.org/10.5281/zenodo.8405898" figureDoi-3="http://doi.org/10.5281/zenodo.8405900" figureDoi-4="http://doi.org/10.5281/zenodo.8405902" figureDoi-5="http://doi.org/10.5281/zenodo.8405904" figureDoi-6="http://doi.org/10.5281/zenodo.8405906" figureDoi-7="http://doi.org/10.5281/zenodo.8405908" figureDoi-8="http://doi.org/10.5281/zenodo.8405910" figureDoi-9="http://doi.org/10.5281/zenodo.8405912" httpUri-0="https://zenodo.org/record/8405894/files/figure.png" httpUri-1="https://zenodo.org/record/8405896/files/figure.png" httpUri-2="https://zenodo.org/record/8405898/files/figure.png" httpUri-3="https://zenodo.org/record/8405900/files/figure.png" httpUri-4="https://zenodo.org/record/8405902/files/figure.png" httpUri-5="https://zenodo.org/record/8405904/files/figure.png" httpUri-6="https://zenodo.org/record/8405906/files/figure.png" httpUri-7="https://zenodo.org/record/8405908/files/figure.png" httpUri-8="https://zenodo.org/record/8405910/files/figure.png" httpUri-9="https://zenodo.org/record/8405912/files/figure.png" pageId="26" pageNumber="27">Figs 1381</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3F971B995A76652907D7FE542939FE2F" lastPageId="32" lastPageNumber="33" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="773248125A76651307D7FE542D90FE73" blockId="26.[189,457,400,426]" box="[189,457,400,426]" pageId="26" pageNumber="27">
<heading id="2C7AFF7E5A76651307D7FE542D90FE73" bold="true" box="[189,457,400,426]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="45F994005A76651307D7FE542D90FE73" bold="true" box="[189,457,400,426]" pageId="26" pageNumber="27">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A76652907D7FE7B2939FE2F" blockId="26.[187,1399,447,615]" lastBlockId="32.[189,1401,265,502]" lastPageId="32" lastPageNumber="33" pageId="26" pageNumber="27">
Females of
<taxonomicName id="B08D33915A7665130628FE7B2E3BFE00" authority="Valdez-Mondragón &amp; Cabrera-Espinosa, 2023" authorityName="Valdez-Mondragón &amp; Cabrera-Espinosa" authorityYear="2023" box="[322,610,447,473]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A7665130628FE7B2E3BFE00" box="[322,610,447,473]" italics="true" pageId="26" pageNumber="27">Latrodectus occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A766513050DFE042E9BFE03" box="[615,706,448,474]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
can be identified from those of other species of
<taxonomicName id="B08D33915A7665130381FE7B292FFE00" authorityName="Walckenaer" authorityYear="1805" box="[1259,1398,447,473]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="45F994005A7665130381FE7B292FFE00" box="[1259,1398,447,473]" italics="true" pageId="26" pageNumber="27">Latrodectus</emphasis>
</taxonomicName>
from
<collectingRegion id="B54986F05A7665130794FE262D25FE25" box="[254,380,482,508]" country="Mexico" name="Distrito Federal" pageId="26" pageNumber="27">Mexico by</collectingRegion>
the combination of several features. Dorsal coloration: distinct red stripes and dots on dorsal surface of the abdomen: 1) one V-shaped stripe on anterior part (
<figureCitation id="EFB654975A766513036EFDC22802FDF8" box="[1028,1115,518,545]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="26" pageNumber="27">Fig. 25</figureCitation>
, red arrow), 2) long and sigmoid stripe dorsally along the abdomen (
<figureCitation id="EFB654975A76651305D8FDED2F5FFD9D" box="[690,774,553,580]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="26" pageNumber="27">Fig. 25</figureCitation>
, blue arrow), and 4) two pairs of stripes towards lateral part of abdomen (
<figureCitation id="EFB654975A76651306F9FD882DB4FDBE" box="[403,493,588,615]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="26" pageNumber="27">Fig. 25</figureCitation>
, green arrows). Ventral coloration: abdomen with hourglass pattern always complete, wide, and never separated as in
<taxonomicName id="B08D33915A77651205D6FECE2F1CFEFA" authorityName="Chamberlin &amp; Ivie" authorityYear="1935" box="[700,837,265,291]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="hesperus">
<emphasis id="45F994005A77651205D6FECE2F1CFEFA" box="[700,837,265,291]" italics="true" pageId="27" pageNumber="28">L. hesperus</emphasis>
</taxonomicName>
and
<taxonomicName id="B08D33915A77651204E9FECE285CFEFA" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[899,1029,266,291]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mactans">
<emphasis id="45F994005A77651204E9FECE285CFEFA" box="[899,1029,266,291]" italics="true" pageId="27" pageNumber="28">L. mactans</emphasis>
</taxonomicName>
(
<figureCitation id="EFB654975A776512037FFECD2836FEFA" box="[1045,1135,265,291]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="27" pageNumber="28">Fig. 26</figureCitation>
). Body measurements (
<figureCitation id="EFB654975A77651207ADFEE82D6DFE9E" box="[199,308,300,327]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="23.[189,232,1928,1954]" captionStartId-1="24.[189,232,1928,1954]" captionTargetBox-0="[196,1398,717,1873]" captionTargetBox-1="[196,1398,717,1874]" captionTargetId-0="figure-119@23.[189,1398,710,1885]" captionTargetId-1="figure-103@24.[189,1398,710,1885]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="Fig. 5. Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that showed significant differences (p&gt;0.05) in females of three putative species of Latrodectus Walckenaer, 1805 from Mexico." captionText-1="Fig. 6. Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that showed significant differences (p&gt;0.05) in females of three putative species of Latrodectus Walckenaer, 1805 from Mexico." figureDoi-0="http://doi.org/10.5281/zenodo.8405880" figureDoi-1="http://doi.org/10.5281/zenodo.8405882" httpUri-0="https://zenodo.org/record/8405880/files/figure.png" httpUri-1="https://zenodo.org/record/8405882/files/figure.png" pageId="27" pageNumber="28">Figs 56</figureCitation>
): sternum coefficient (length/width) longer than other species analyzed herein, tibia-patella I longer than other species, tibia-patella coefficient longer than other analyzed species, femur I longer than other species, femur I and IV coefficient longer than other species, TT1 and TT4 coefficients slightly longer than other species. Angle between the spermathecae position in dorsal view longer than other species. Genital opening of epigynum wider than other species (
<figureCitation id="EFB654975A7765120493FE7D28F5FE0A" box="[1017,1196,441,467]" captionStart-0="Figs 2931" captionStart-1="Figs 3243" captionStartId-0="34.[185,238,1926,1952]" captionStartId-1="35.[187,240,1822,1848]" captionTargetBox-0="[189,1398,958,1879]" captionTargetBox-1="[189,1398,708,1771]" captionTargetId-0="figure-253@34.[189,1398,958,1884]" captionTargetId-1="figure-157@35.[189,1398,708,1771]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." captionText-1="Figs 3243. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the seminal receptacles of females, dorsal view. 3233. Cocula, Jalisco, Mexico (type locality). 3435. Jilotitlán de los Dolores, Jalisco, Mexico. 3637. Minatitlán, Colima, Mexico. 3839. Coahuayana de Hidalgo, Michoacán, Mexico. 4041. Quechultenango, Guerrero, Mexico. 4243. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8405900" figureDoi-1="http://doi.org/10.5281/zenodo.8405902" httpUri-0="https://zenodo.org/record/8405900/files/figure.png" httpUri-1="https://zenodo.org/record/8405902/files/figure.png" pageId="27" pageNumber="28">Figs 29, 3142</figureCitation>
(ventral views)). Copulatory ducts forming four or five loops around spermathecae (
<figureCitation id="EFB654975A776512049CFE18280CFE2F" box="[1014,1109,476,502]" captionStart="Figs 2931" captionStartId="34.[185,238,1926,1952]" captionTargetBox="[189,1398,958,1879]" captionTargetId="figure-253@34.[189,1398,958,1884]" captionTargetPageId="34" captionText="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="27" pageNumber="28">Figs 30</figureCitation>
,
<figureCitation id="EFB654975A776512030FFE1828F7FE2F" box="[1125,1198,476,502]" captionStart="Figs 3243" captionStartId="35.[187,240,1822,1848]" captionTargetBox="[189,1398,708,1771]" captionTargetId="figure-157@35.[189,1398,708,1771]" captionTargetPageId="35" captionText="Figs 3243. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the seminal receptacles of females, dorsal view. 3233. Cocula, Jalisco, Mexico (type locality). 3435. Jilotitlán de los Dolores, Jalisco, Mexico. 3637. Minatitlán, Colima, Mexico. 3839. Coahuayana de Hidalgo, Michoacán, Mexico. 4041. Quechultenango, Guerrero, Mexico. 4243. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405902" httpUri="https://zenodo.org/record/8405902/files/figure.png" pageId="27" pageNumber="28">3343</figureCitation>
(ventral views), 72). Males: similar to females, but abdomen oval (
<figureCitation id="EFB654975A7765120446FE3B2FD3FDC3" box="[812,906,511,538]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="27" pageNumber="28">Figs 17</figureCitation>
,
<figureCitation id="EFB654975A77651204F1FDC42FB3FDC3" box="[923,1002,512,538]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="27" pageNumber="28">2728</figureCitation>
). Coloration: ventrally, abdomen with hourglass pattern always complete, thinner than females (
<figureCitation id="EFB654975A77651204FCFDE62FB4FDE4" box="[918,1005,546,573]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="27" pageNumber="28">Fig. 28</figureCitation>
). Body measurements (
<figureCitation id="EFB654975A7765120394FDE62930FDE4" box="[1278,1385,546,573]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="26.[189,232,1928,1954]" captionStartId-1="27.[189,232,1928,1954]" captionTargetBox-0="[198,1398,713,1871]" captionTargetBox-1="[199,1369,716,1873]" captionTargetId-0="figure-106@26.[189,1398,707,1882]" captionTargetId-1="figure-155@27.[189,1398,709,1884]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="Fig. 7. Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that show significant differences (p&gt;0.05) in males of three putative species of Latrodectus Walckenaer, 1805 from Mexico." captionText-1="Fig. 8. Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that show significant differences (p&gt;0.05) in males of three putative species of Latrodectus Walckenaer, 1805 from Mexico." figureDoi-0="http://doi.org/10.5281/zenodo.8405884" figureDoi-1="http://doi.org/10.5281/zenodo.8405886" httpUri-0="https://zenodo.org/record/8405884/files/figure.png" httpUri-1="https://zenodo.org/record/8405886/files/figure.png" pageId="27" pageNumber="28">Figs 78</figureCitation>
): sternum coefficient (length/width) longer than other species analyzed herein, tibia-patella 1 length and tibia-patella coefficient longer than other species, tibia-patella 1 coefficient longer than other species, tibia-patella 1 coefficient longer than other species, length and coefficient of femur I longer than other species, TT1 coefficient longer than other species, tibia-patella IV longer than other species, femur IV longer than on other species. Embolus with 3 coils, located distally on cymbium (
<figureCitation id="EFB654975A4C652903D0FEB6291AFE54" box="[1210,1347,370,397]" captionStart="Figs 4448" captionStartId="36.[189,242,1801,1827]" captionTargetBox="[189,1398,265,1746]" captionTargetId="figure-13@36.[189,1398,265,1746]" captionTargetPageId="36" captionText="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="32" pageNumber="33">Figs 4448</figureCitation>
, red arrows), last coil extending medially, curving downwards along retrolateral part of palp, continuing ventrally, ending in a long thin tip (
<figureCitation id="EFB654975A4C65290501FE7D2EA1FE0A" box="[619,760,441,467]" captionStart-0="Figs 4448" captionStart-1="Figs 4960" captionStartId-0="36.[189,242,1801,1827]" captionStartId-1="38.[188,241,1857,1883]" captionTargetBox-0="[189,1398,265,1746]" captionTargetBox-1="[189,1398,745,1807]" captionTargetId-0="figure-13@36.[189,1398,265,1746]" captionTargetId-1="figure-193@38.[189,1398,745,1807]" captionTargetPageId-0="36" captionTargetPageId-1="38" captionText-0="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." captionText-1="Figs 4960. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the male left palps, prolateral views. 4952. Quechultenango, Guerrero, Mexico. 5356. Minatitlán, Colima, Mexico. 57 60. Cocula, Jalisco, Mexico (type locality). Ventral view (49, 53, 57); retrolateral-frontal view (50, 54, 59); retrolateral view (51, 55, 59); posterior view (52, 56, 60). 4956. Postcopulatory palps. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8405904" figureDoi-1="http://doi.org/10.5281/zenodo.8405906" httpUri-0="https://zenodo.org/record/8405904/files/figure.png" httpUri-1="https://zenodo.org/record/8405906/files/figure.png" pageId="32" pageNumber="33">Figs 4460</figureCitation>
). In
<taxonomicName id="B08D33915A4C65290458FE7E2FECFE0A" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[818,949,442,467]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mactans">
<emphasis id="45F994005A4C65290458FE7E2FECFE0A" box="[818,949,442,467]" italics="true" pageId="32" pageNumber="33">L. mactans</emphasis>
</taxonomicName>
and
<taxonomicName id="B08D33915A4C6529049AFE7E2823FE0A" authorityName="Chamberlin &amp; Ivie" authorityYear="1935" box="[1008,1146,441,467]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hesperus">
<emphasis id="45F994005A4C6529049AFE7E2823FE0A" box="[1008,1146,441,467]" italics="true" pageId="32" pageNumber="33">L. hesperus</emphasis>
</taxonomicName>
(native species from North America), the embolus has 2 coils (
<bibRefCitation id="131C35E35A4C652905C2FE182891FE2F" author="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A." box="[680,1224,476,502]" pageId="32" pageNumber="33" pagination="923665" refId="ref21933" refString="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A. 2021. Distribucion y modelaje de nicho ecologico, comentarios biogeograficos y taxonomicos del genero de aranas Latrodectus Walckenaer (Araneae, Theridiidae) de Mexico. Revista Mexicana de Biodiversidad 92: e 923665. https: // doi. org / 10.22201 / ib. 20078706 e. 2021.92.3665" type="journal article" year="2021">Cabrera-Espinosa &amp; Valdez-Mondragón 2021</bibRefCitation>
: figs 3037).
</paragraph>
</subSubSection>
<caption id="23F2189A5A76651307D7F84C2EB3F831" ID-DOI="http://doi.org/10.5281/zenodo.8405884" ID-Zenodo-Dep="8405884" httpUri="https://zenodo.org/record/8405884/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1928,1954]" targetBox="[198,1398,713,1871]" targetPageId="26">
<paragraph id="773248125A76651307D7F84C2EB3F831" blockId="26.[187,1399,1928,2025]" pageId="26" pageNumber="27">
<emphasis id="45F994005A76651307D7F84C2D50F87B" bold="true" box="[189,265,1928,1954]" pageId="26" pageNumber="27">Fig. 7.</emphasis>
Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that show significant differences (p&gt;0.05) in males of three putative species of
<taxonomicName id="B08D33915A76651307B7F80A2E18F830" authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" box="[221,577,1998,2025]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="45F994005A76651307B7F80A2D31F831" box="[221,360,1998,2024]" italics="true" pageId="26" pageNumber="27">Latrodectus</emphasis>
Walckenaer, 1805
</taxonomicName>
from Mexico.
</paragraph>
</caption>
<caption id="23F2189A5A77651207D7F84C2EB3F831" ID-DOI="http://doi.org/10.5281/zenodo.8405886" ID-Zenodo-Dep="8405886" httpUri="https://zenodo.org/record/8405886/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1928,1954]" targetBox="[199,1369,716,1873]" targetPageId="27">
<paragraph id="773248125A77651207D7F84C2EB3F831" blockId="27.[187,1399,1928,2025]" pageId="27" pageNumber="28">
<emphasis id="45F994005A77651207D7F84C2D50F87B" bold="true" box="[189,265,1928,1954]" pageId="27" pageNumber="28">Fig. 8.</emphasis>
Comparison of variances (ANOVA) and medians (Kruskal-Wallis) of different structures analyzed under linear morphometry that show significant differences (p&gt;0.05) in males of three putative species of
<taxonomicName id="B08D33915A77651207B7F80A2E18F830" authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" box="[221,577,1998,2025]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="45F994005A77651207B7F80A2D31F831" box="[221,360,1998,2024]" italics="true" pageId="27" pageNumber="28">Latrodectus</emphasis>
Walckenaer, 1805
</taxonomicName>
from Mexico.
</paragraph>
</caption>
<caption id="23F2189A5A70651507D7F8852E14F831" ID-DOI="http://doi.org/10.5281/zenodo.8405890" ID-Zenodo-Dep="8405890" httpUri="https://zenodo.org/record/8405890/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,242,1857,1883]" targetBox="[244,1346,283,1804]" targetPageId="28">
<paragraph id="773248125A70651507D7F8852E14F831" blockId="28.[189,1399,1857,2025]" pageId="28" pageNumber="29">
<emphasis id="45F994005A70651507D7F8852D03F882" bold="true" box="[189,346,1857,1883]" pageId="28" pageNumber="29">Figs 910. 9.</emphasis>
Records and Species Distribution Model (SDM) (red) of
<taxonomicName id="B08D33915A706515036FF886290EF885" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1029,1367,1857,1884]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mactans">
<emphasis id="45F994005A706515036FF88628DFF882" box="[1029,1158,1858,1883]" italics="true" pageId="28" pageNumber="29">L. mactans</emphasis>
(Fabricius, 1775)
</taxonomicName>
in Mexico and the USA.
<emphasis id="45F994005A70651506ACF8A12DB2F8A6" bold="true" box="[454,491,1893,1919]" pageId="28" pageNumber="29">10.</emphasis>
Records and SDM (yellow) of
<taxonomicName id="B08D33915A7065150435F8A228AFF8A6" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[863,1270,1893,1919]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A7065150435F8A22855F8A6" box="[863,1036,1893,1919]" italics="true" pageId="28" pageNumber="29">L. occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A7065150397F8A12901F8A6" box="[1277,1368,1893,1919]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
in Mexico. Abbreviations: CHIS = Chiapas; COL = Colima; EMX = Edo. Mex; GRO = Guerrero; GTO = Guanajuato; JAL = Jalisco; MICH= Michoacán; MOR = Morelos; NAY = Nayarit; OAX = Oaxaca; PUE = Puebla; QRO = Querétaro.
</paragraph>
</caption>
<caption id="23F2189A5A71651407D7F91C2D10F830" ID-DOI="http://doi.org/10.5281/zenodo.8405892" ID-Zenodo-Dep="8405892" httpUri="https://zenodo.org/record/8405892/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,242,1752,1778]" targetBox="[279,1310,281,1706]" targetPageId="29">
<paragraph id="773248125A71651407D7F91C2D10F830" blockId="29.[189,1399,1752,2025]" pageId="29" pageNumber="30">
<emphasis id="45F994005A71651407D7F91C2D23F92B" bold="true" box="[189,378,1752,1778]" pageId="29" pageNumber="30">Figs 1112. 11.</emphasis>
Records and Species Distribution Model (SDM) of
<taxonomicName id="B08D33915A716514049DF91D2CA1F8CF" authority="Chamberlin &amp; Ivie, 1935" authorityName="Chamberlin &amp; Ivie" authorityYear="1935" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hesperus">
<emphasis id="45F994005A716514049DF91D28D8F92B" box="[1015,1153,1752,1778]" italics="true" pageId="29" pageNumber="30">L. hesperus</emphasis>
<bibRefCitation id="131C35E35A71651403E1F91C2CA1F8CF" author="Chamberlin R. V. &amp; Ivie W." pageId="29" pageNumber="30" pagination="1 - 29" refId="ref22088" refString="Chamberlin R. V. &amp; Ivie W. 1935. The black widow spider and its varieties in the United States. Bulletin of the University of Utah 25 (8): 1 - 29." type="journal article" year="1935">Chamberlin &amp; Ivie, 1935</bibRefCitation>
</taxonomicName>
for Mexico and the USA.
<emphasis id="45F994005A7165140558F93F2E0EF8CC" bold="true" box="[562,599,1787,1813]" pageId="29" pageNumber="30">12.</emphasis>
SDM of
<taxonomicName id="B08D33915A71651405ACF9382841F8CF" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[710,1048,1787,1814]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mactans">
<emphasis id="45F994005A71651405ACF9382F1EF8CC" box="[710,839,1788,1813]" italics="true" pageId="29" pageNumber="30">L. mactans</emphasis>
(Fabricius, 1775)
</taxonomicName>
(red),
<taxonomicName id="B08D33915A7165140302F9382D1FF8E1" authority="ValdezMondragon" authorityName="Valdez-Mondragon" authorityYear="2023" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A7165140302F938294CF8CC" box="[1128,1301,1787,1813]" italics="true" pageId="29" pageNumber="30">L. occidentalis</emphasis>
ValdezMondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A716514063AF8DB2DF7F8E0" box="[336,430,1823,1849]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
(yellow), and
<taxonomicName id="B08D33915A7165140509F8DB2EB4F8E1" authorityName="Chamberlin &amp; Ivie" authorityYear="1935" box="[611,749,1822,1848]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hesperus">
<emphasis id="45F994005A7165140509F8DB2EB4F8E1" box="[611,749,1822,1848]" italics="true" pageId="29" pageNumber="30">L. hesperus</emphasis>
</taxonomicName>
(purple) confined only to Mexico, showing a lack of overlap in their distributions, using the biogeographical provinces proposed by
<bibRefCitation id="131C35E35A716514031AF886297EF885" author="Morrone J." box="[1136,1319,1858,1884]" pageId="29" pageNumber="30" pagination="149 - 162" refId="ref24464" refString="Morrone J. 2004. Panbiogeografia, componentes bioticos y zonas de transicion. Revista Brasileira de Entomologia 48 (2): 149 - 162. https: // doi. org / 10.1590 / S 0085 - 56262004000200001" type="journal article" year="2004">Morrone (2004</bibRefCitation>
,
<bibRefCitation id="131C35E35A716514025EF885292BF885" author="Morrone J." box="[1332,1394,1857,1884]" pageId="29" pageNumber="30" pagination="207 - 252" refId="ref24504" refString="Morrone J. 2005. Hacia una sintesis biogeografica de Mexico. Revista Mexicana de Biodiversidad 76 (2): 207 - 252. https: // doi. org / 10.22201 / ib. 20078706 e. 2005.002.303" type="journal article" year="2005">2005</bibRefCitation>
,
<bibRefCitation id="131C35E35A71651407D5F8A12CA6F8A6" author="Morrone J." box="[191,255,1893,1919]" pageId="29" pageNumber="30" pagination="277 - 279" refId="ref24544" refString="Morrone J. 2017. Mexican biogeographic provinces: map and shapefiles. Zootaxa 4277 (2): 277 - 279. https: // doi. org / 10.11646 / zootaxa. 4277.2.8" type="journal article" year="2017">2017</bibRefCitation>
). Abbreviations: BB = Balsas Basin; BC = Baja Californian; Ca = Californian; CD = Chihuahuan Desert; PL = Pacific Lowlands; SMO = Sierra Madre Oriental; SMOc = Sierra Madre Occidental: SMS = Sierra Madre del Sur; So = Sonoran; Ta = Tamaulipas; TVB = Transmexican Volcanic Belt; Ve = Veracruzan.
</paragraph>
</caption>
<caption id="23F2189A5A72651707D7F91028FAF846" ID-DOI="http://doi.org/10.5281/zenodo.8405894" ID-Zenodo-Dep="8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,242,1748,1774]" targetBox="[189,1398,268,1700]" targetPageId="30">
<paragraph id="773248125A72651707D7F91028FAF846" blockId="30.[186,1399,1748,1951]" pageId="30" pageNumber="31">
<emphasis id="45F994005A72651707D7F9102D14F937" bold="true" box="[189,333,1748,1774]" pageId="30" pageNumber="31">Figs 1318.</emphasis>
Live females (1315) and males (1618) of
<taxonomicName id="B08D33915A726517040BF910292EF937" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[865,1399,1748,1774]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A726517040BF91028D3F937" box="[865,1162,1748,1774]" italics="true" pageId="30" pageNumber="31">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A72651707D7F93C2D47F8CB" box="[189,286,1784,1810]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="45F994005A7265170643F9332D21F8C8" bold="true" box="[297,376,1783,1809]" pageId="30" pageNumber="31">13, 16</emphasis>
. Salvatierra, Guanajuato, Mexico.
<emphasis id="45F994005A726517045AF9332F17F8C8" bold="true" box="[816,846,1783,1809]" pageId="30" pageNumber="31">14</emphasis>
. Camichines, Cocula, Jalisco, Mexico (type locality).
<emphasis id="45F994005A726517065FF8DF2D0AF8EC" bold="true" box="[309,339,1819,1845]" pageId="30" pageNumber="31">15</emphasis>
. Hostotipaquillo, Jalisco, Mexico.
<emphasis id="45F994005A7265170460F8DF2F71F8EC" bold="true" box="[778,808,1819,1845]" pageId="30" pageNumber="31">17</emphasis>
. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico.
<emphasis id="45F994005A726517064FF8FA2D1AF881" bold="true" box="[293,323,1854,1880]" pageId="30" pageNumber="31">18</emphasis>
. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by
<bibRefCitation id="131C35E35A72651703F6F8FA2D57F8A2" author="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A." pageId="30" pageNumber="31" pagination="923665" refId="ref21933" refString="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A. 2021. Distribucion y modelaje de nicho ecologico, comentarios biogeograficos y taxonomicos del genero de aranas Latrodectus Walckenaer (Araneae, Theridiidae) de Mexico. Revista Mexicana de Biodiversidad 92: e 923665. https: // doi. org / 10.22201 / ib. 20078706 e. 2021.92.3665" type="journal article" year="2021">Cabrera-Espinosa (2021)</bibRefCitation>
. Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294
</paragraph>
</caption>
<caption id="23F2189A5A73651607D7F92B2D29F860" ID-DOI="http://doi.org/10.5281/zenodo.8405896" ID-Zenodo-Dep="8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,242,1775,1801]" targetBox="[189,1398,268,1701]" targetPageId="31">
<paragraph id="773248125A73651607D7F92B2D29F860" blockId="31.[187,1399,1775,1977]" pageId="31" pageNumber="32">
<emphasis id="45F994005A73651607D7F92B2D15F8D0" bold="true" box="[189,332,1775,1801]" pageId="31" pageNumber="32">Figs 1924.</emphasis>
Habitats and microhabitats of
<taxonomicName id="B08D33915A73651605AEF92B2881F8D0" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[708,1240,1775,1801]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A73651605AEF92B2FB0F8D0" box="[708,1001,1775,1801]" italics="true" pageId="31" pageNumber="32">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A736516038BF92B2967F8D0" box="[1249,1342,1775,1801]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
Red arrow indicates the microhabitat where the specimens were collected.
<emphasis id="45F994005A736516036DF8D62809F8F5" bold="true" box="[1031,1104,1810,1836]" pageId="31" pageNumber="32">19, 22</emphasis>
. “Las Letras”, Pénjamo, Guanajuato, Mexico.
<emphasis id="45F994005A73651606DCF8F12D8AF896" bold="true" box="[438,467,1845,1871]" pageId="31" pageNumber="32">20</emphasis>
. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.
<emphasis id="45F994005A73651603AEF8F128B8F896" bold="true" box="[1220,1249,1845,1871]" pageId="31" pageNumber="32">21</emphasis>
. Salvatierra, Guanajuato, Mexico.
<emphasis id="45F994005A73651606D5F89C2E5EF8AB" bold="true" box="[447,519,1880,1906]" pageId="31" pageNumber="32">2324</emphasis>
. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 19
<emphasis id="45F994005A736516044AF8B82F76F84F" bold="true" box="[800,815,1916,1942]" pageId="31" pageNumber="32"></emphasis>
22 by
<bibRefCitation id="131C35E35A7365160415F8B828F5F84F" author="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A." box="[895,1196,1916,1942]" pageId="31" pageNumber="32" pagination="923665" refId="ref21933" refString="Cabrera-Espinosa L. A. &amp; Valdez-Mondragon A. 2021. Distribucion y modelaje de nicho ecologico, comentarios biogeograficos y taxonomicos del genero de aranas Latrodectus Walckenaer (Araneae, Theridiidae) de Mexico. Revista Mexicana de Biodiversidad 92: e 923665. https: // doi. org / 10.22201 / ib. 20078706 e. 2021.92.3665" type="journal article" year="2021">Cabrera-Espinosa (2021)</bibRefCitation>
; 2324 by Jared Lacayo (2021).
</paragraph>
</caption>
<subSubSection id="3F971B995A4C652907D7FDDA2EB0FD75" pageId="32" pageNumber="33" type="etymology">
<paragraph id="773248125A4C652907D7FDDA2D1DFDE1" blockId="32.[189,324,542,568]" box="[189,324,542,568]" pageId="32" pageNumber="33">
<heading id="2C7AFF7E5A4C652907D7FDDA2D1DFDE1" bold="true" box="[189,324,542,568]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="45F994005A4C652907D7FDDA2D1DFDE1" bold="true" box="[189,324,542,568]" pageId="32" pageNumber="33">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A4C652907D7FD882EB0FD75" blockId="32.[188,1399,588,685]" pageId="32" pageNumber="33">
This species is a name in apposition and refers to the distribution of the species in the Spanish language:
<emphasis id="45F994005A4C652907ADFDAB2D88FD53" box="[199,465,623,650]" italics="true" pageId="32" pageNumber="33">
Occidente de
<collectingRegion id="B54986F05A4C65290611FDB42D88FD53" box="[379,465,624,650]" country="Mexico" name="Mexico" pageId="32" pageNumber="33">México</collectingRegion>
</emphasis>
(western Mexico),
<taxonomicName id="B08D33915A4C6529059DFDAB2FDCFD50" authorityName="Valdez-Mondragon" authorityYear="2023" box="[759,901,623,649]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="45F994005A4C6529059DFDAB2FDCFD50" box="[759,901,623,649]" italics="true" pageId="32" pageNumber="33">occidentalis</emphasis>
</taxonomicName>
, that includes the states of
<collectingRegion id="B54986F05A4C6529026FFDAB292BFD50" box="[1285,1394,623,649]" country="Mexico" name="Guerrero" pageId="32" pageNumber="33">Guerrero</collectingRegion>
,
<collectingRegion id="B54986F05A4C652907D7FD562D1BFD75" box="[189,322,658,684]" country="Mexico" name="Michoacan de Ocampo" pageId="32" pageNumber="33">Michoacán</collectingRegion>
,
<collectingRegion id="B54986F05A4C65290624FD562DFEFD75" box="[334,423,658,684]" country="Mexico" name="Colima" pageId="32" pageNumber="33">Colima</collectingRegion>
, and
<collectingRegion id="B54986F05A4C6529068FFD562E6CFD75" box="[485,565,658,684]" country="Mexico" name="Jalisco" pageId="32" pageNumber="33">Jalisco</collectingRegion>
(
<typeStatus id="A836F6B05A4C6529052EFD572E21FD74" box="[580,632,659,685]" pageId="32" pageNumber="33">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="3F971B995A4C652B07D7FD102ED2FDE4" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="773248125A4C652907D7FD102DF3FD37" blockId="32.[189,426,724,750]" box="[189,426,724,750]" pageId="32" pageNumber="33">
<heading id="2C7AFF7E5A4C652907D7FD102DF3FD37" bold="true" box="[189,426,724,750]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="45F994005A4C652907D7FD102DF3FD37" bold="true" box="[189,426,724,750]" pageId="32" pageNumber="33">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A4C65290780FCC7286CFCBD" blockId="32.[189,1398,771,868]" pageId="32" pageNumber="33">
<materialsCitation id="C7E5424F5A4C65290780FCC7286CFCBD" ID-GBIF-Occurrence="4421595302" collectingDate="2020-06-01" collectingMethod="night collecting" collectionCode="CNAN-T" collectorName="C. I. Navarro-Rodriguez" country="Mexico" elevation="1268" latitude="20.4991" location="Camichines" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCode="CNAN-T01589" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="45F994005A4C65290780FCC72D05FCC4" bold="true" box="[234,348,771,797]" pageId="32" pageNumber="33">
<typeStatus id="A836F6B05A4C65290780FCC72D05FCC4" box="[234,348,771,797]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="0F9A08825A4C652907D7FCE22D68FC99" box="[189,305,806,832]" name="Mexico" pageId="32" pageNumber="33">MEXICO</collectingCountry>
<emphasis id="45F994005A4C65290624FCE12DFAFCE6" bold="true" box="[334,419,805,831]" pageId="32" pageNumber="33">
<collectingRegion id="B54986F05A4C65290624FCE12DFAFCE6" box="[334,419,805,831]" country="Mexico" name="Jalisco" pageId="32" pageNumber="33">Jalisco</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4C652906D6FCE22D8AFC99" box="[444,467,806,832]" pageId="32" pageNumber="33" type="female"></specimenCount>
;
<collectingMunicipality id="9756D2685A4C65290688FCE22EAEFC99" box="[482,759,806,832]" pageId="32" pageNumber="33">Municipality of Cocula</collectingMunicipality>
,
<location id="72521EC95A4C65290469FCE22FC8FC99" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C65290469FCE22FC8FC99" box="[771,913,806,832]" country="Mexico" latitude="20.4991" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" name="Camichines" pageId="32" pageNumber="33" stateProvince="Jalisco">Camichines</location>
;
<geoCoordinate id="12B92ED55A4C652904F7FCE22871FC99" box="[925,1064,806,832]" degrees="20.4991" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="20.4991">20.4991° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C6529035EFCE2288BFC99" box="[1076,1234,806,832]" degrees="103.8020" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-103.802">103.8020° W</geoCoordinate>
;
<quantity id="B075E5F75A4C652903B5FCE22961FC99" box="[1247,1336,806,832]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="32" pageNumber="33" unit="m" value="1268.0">
<elevation id="FCA0AF215A4C652903B5FCE22961FC99" box="[1247,1336,806,832]" metricMagnitude="3" metricUnit="m" metricValue="1.268" pageId="32" pageNumber="33" unit="m" value="1268.0">1268 m</elevation>
</quantity>
a.s.l;
<date id="03336ED25A4C652907D7FC8D2D11FCBA" box="[189,328,841,867]" pageId="32" pageNumber="33" value="2020-06-01">
<collectingDate id="1377973A5A4C652907D7FC8D2D11FCBA" box="[189,328,841,867]" pageId="32" pageNumber="33" value="2020-06-01">1 Jun. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C6529063FFC8D2E35FCBA" box="[341,620,841,867]" pageId="32" pageNumber="33">C.I. Navarro-Rodríguez</collectorName>
leg.;
<collectingMethod id="AECC30055A4C652905C4FC8D2F30FCBA" box="[686,873,841,867]" pageId="32" pageNumber="33">night collecting</collectingMethod>
;
<specimenCode id="272BE0695A4C6529041FFC8D2868FCBD" box="[885,1073,841,868]" collectionCode="CNAN-T" pageId="32" pageNumber="33">CNAN-T01589</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="773248125A4C65290780FC4E2FEEFC35" blockId="32.[189,1398,906,1004]" pageId="32" pageNumber="33">
<materialsCitation id="C7E5424F5A4C65290780FC4E282EFC11" ID-GBIF-Occurrence="4421595317" collectingDate="2020-06-01" collectingMethod="night collecting" collectionCode="CNAN-T" collectorName="C. I. Navarro-Rodriguez" country="Mexico" elevation="1268" latitude="20.4991" location="Jalisco" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCode="CNAN-T01590" specimenCount="1" stateProvince="Jalisco" typeStatus="paratype">
<emphasis id="45F994005A4C65290780FC4E2D3EFC7D" bold="true" box="[234,359,906,932]" pageId="32" pageNumber="33">
<typeStatus id="A836F6B05A4C65290780FC4E2D3EFC7D" box="[234,359,906,932]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="0F9A08825A4C652907D7FC6A2D68FC11" box="[189,305,942,968]" name="Mexico" pageId="32" pageNumber="33">MEXICO</collectingCountry>
<emphasis id="45F994005A4C6529063BFC692DFFFC1E" bold="true" box="[337,422,941,967]" pageId="32" pageNumber="33">
<collectingRegion id="B54986F05A4C6529063BFC692DFFFC1E" box="[337,422,941,967]" country="Mexico" name="Jalisco" pageId="32" pageNumber="33">Jalisco</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4C652906ABFC6A2DA9FC1E" box="[449,496,939,968]" pageId="32" pageNumber="33">1 ♁</specimenCount>
; same collection data as for holotype;
<specimenCode id="272BE0695A4C652904D4FC69282EFC11" box="[958,1143,941,968]" collectionCode="CNAN-T" pageId="32" pageNumber="33">CNAN-T01590</specimenCode>
</materialsCitation>
<materialsCitation id="C7E5424F5A4C652903F8FC6A2FEAFC35" ID-GBIF-Occurrence="4421595314" collectingDate="2020-06-01" collectingMethod="night collecting" collectionCode="CNAN-T" collectorName="C. I. Navarro-Rodriguez" country="Mexico" elevation="1268" latitude="20.4991" location="Camichines" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCode="CNAN-T01591, CNAN-T01593" specimenCount="5" specimenCount-female="3" stateProvince="Jalisco" typeStatus="paratype">
<specimenCount id="618B839B5A4C652903F8FC6A288EFC11" box="[1170,1239,942,968]" pageId="32" pageNumber="33" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="618B839B5A4C6529038CFC6A2973FC1E" box="[1254,1322,939,968]" pageId="32" pageNumber="33">2 ♁♁</specimenCount>
; same collection data as for holotype;
<specimenCode id="272BE0695A4C6529055AFC152EB2FC35" box="[560,747,977,1004]" collectionCode="CNAN-T" pageId="32" pageNumber="33">CNAN-T01591</specimenCode>
,
<specimenCode id="272BE0695A4C6529059CFC152FEAFC35" box="[758,947,977,1004]" collectionCode="CNAN-T" pageId="32" pageNumber="33">CNAN-T01593</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="773248125A4C65290780FBD62DFFFBF5" blockId="32.[187,1400,1042,2019]" box="[234,422,1042,1068]" pageId="32" pageNumber="33">
<emphasis id="45F994005A4C65290780FBD62DFFFBF5" bold="true" box="[234,422,1042,1068]" pageId="32" pageNumber="33">Other material</emphasis>
</paragraph>
<paragraph id="773248125A4C652B07D7FBF12ED2FDE4" blockId="32.[187,1400,1042,2019]" lastBlockId="34.[188,1399,265,573]" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<materialsCitation id="C7E5424F5A4C652907D7FBF12D69FB4F" ID-GBIF-Occurrence="4421595310" collectingDate="2020-11-16" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="296" latitude="19.1676" location="watering place Los Amiales" longLatPrecision="52" longitude="-103.834" municipality="Municipality of Coquimatlan" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Colima">
<collectingCountry id="0F9A08825A4C652907D7FBF12D68FB96" box="[189,305,1077,1103]" name="Mexico" pageId="32" pageNumber="33">MEXICO</collectingCountry>
<emphasis id="45F994005A4C6529063CFBF12DE8FB96" bold="true" box="[342,433,1077,1103]" pageId="32" pageNumber="33">
<collectingRegion id="B54986F05A4C6529063CFBF12DE8FB96" box="[342,433,1077,1103]" country="Mexico" name="Colima" pageId="32" pageNumber="33">Colima</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4C652906BBFBF12E58FB89" box="[465,513,1077,1104]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="9756D2685A4C65290579FBF12F2AFB96" box="[531,883,1077,1103]" pageId="32" pageNumber="33">Municipality of Coquimatlán</collectingMunicipality>
,
<location id="72521EC95A4C652904EBFBF1288FFB96" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C652904EBFBF1288FFB96" box="[897,1238,1077,1104]" country="Mexico" latitude="19.1676" longLatPrecision="52" longitude="-103.834" municipality="Municipality of Coquimatlan" name="watering place Los Amiales" pageId="32" pageNumber="33" stateProvince="Colima">watering place Los Amiales</location>
;
<geoCoordinate id="12B92ED55A4C6529038EFBF1292BFB89" box="[1252,1394,1077,1104]" degrees="19.1676" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="19.1676">19.1676° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C652907D7FB9D2D16FBAA" box="[189,335,1113,1139]" degrees="103.834" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-103.834">103.834° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290637FB9C2DB2FBAB" box="[349,491,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="32" pageNumber="33" unit="m" value="296.0">
<quantity id="B075E5F75A4C65290637FB9C2DF0FBAA" box="[349,425,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="32" pageNumber="33" unit="m" value="296.0">296 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C65290690FB9C2EF9FBAA" box="[506,672,1112,1139]" pageId="32" pageNumber="33" value="2020-11-16">
<collectingDate id="1377973A5A4C65290690FB9C2EF9FBAA" box="[506,672,1112,1139]" pageId="32" pageNumber="33" value="2020-11-16">16 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C652905C7FB9D2F7CFBAB" box="[685,805,1112,1139]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C65290458FB9D2FE9FBAA" box="[818,944,1113,1139]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C652904D6FB9D2869FBAA" box="[956,1072,1113,1139]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C65290357FB9D28E7FBAB" box="[1085,1214,1112,1139]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.; LATLAX Ara-1074
</materialsCitation>
<materialsCitation id="C7E5424F5A4C6529062CFBB82FA3FB05" ID-GBIF-Occurrence="4421595309" collectingDate="2020-11-16" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="195" latitude="19.438" location="Municipality of Minatitlan" longLatPrecision="55" longitude="-104.0019" pageId="32" pageNumber="33" specimenCount="8" specimenCount-female="5" specimenCount-immature="2" stateProvince="Colima">
<specimenCount id="618B839B5A4C6529062CFBB82DD1FB4F" box="[326,392,1148,1175]" pageId="32" pageNumber="33" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="618B839B5A4C652906FFFBB82D99FB4F" box="[405,448,1146,1174]" pageId="32" pageNumber="33">1 ♁</specimenCount>
,
<specimenCount id="618B839B5A4C652906A7FBB82E3BFB4F" box="[461,610,1148,1174]" pageId="32" pageNumber="33" type="immature">2 immatures</specimenCount>
;
<location id="72521EC95A4C65290507FBB82FFFFB4F" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C65290507FBB82FFFFB4F" box="[621,934,1148,1174]" country="Mexico" latitude="19.438" longLatPrecision="55" longitude="-104.0019" name="Municipality of Minatitlan" pageId="32" pageNumber="33" stateProvince="Colima">Municipality of Minatitlán</location>
,
<locationDeviation id="668051305A4C652904DBFBB8292DFB4F" box="[945,1396,1148,1175]" pageId="32" pageNumber="33">
<quantity id="B075E5F75A4C652904DBFBB82858FB4F" box="[945,1025,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="km" value="2.5">2.5 km</quantity>
NW of San Antonio crossroads
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4C652907D7FB5B2D60FB63" box="[189,313,1183,1210]" degrees="19.438" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="55" value="19.438">19.438° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C6529062CFB5B2DBCFB63" box="[326,485,1183,1210]" degrees="104.0019" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-104.0019">104.0019° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290698FB5B2E27FB60" box="[498,638,1183,1210]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="32" pageNumber="33" unit="m" value="195.0">
<quantity id="B075E5F75A4C65290698FB5B2E64FB63" box="[498,573,1183,1210]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="32" pageNumber="33" unit="m" value="195.0">195 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C652905E6FB5B2F76FB60" box="[652,815,1183,1210]" pageId="32" pageNumber="33" value="2020-11-16">
<collectingDate id="1377973A5A4C652905E6FB5B2F76FB60" box="[652,815,1183,1210]" pageId="32" pageNumber="33" value="2020-11-16">16 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290450FB5B2FECFB60" box="[826,949,1183,1209]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C652904D0FB5B286BFB63" box="[954,1074,1183,1210]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C65290356FB5B28F6FB63" box="[1084,1199,1183,1210]" pageId="32" pageNumber="33">A. Juarez</collectorName>
,
<collectorName id="DA782DC45A4C652903D1FB5B2962FB60" box="[1211,1339,1183,1209]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4C652907D4FB062DD7FB05" box="[190,398,1218,1244]" pageId="32" pageNumber="33">diurnal collecting</collectingMethod>
; LATLAX Ara-1073, Ara-1051, Ara-1061, Ara-1066
</materialsCitation>
.
<materialsCitation id="C7E5424F5A4C6529037CFB062D86FAFA" ID-GBIF-Occurrence="4421595303" collectingDate="2018-07-01" collectorName="P. Solis &amp; S. Rodriguez" country="Mexico" elevation="1205" latitude="17.3119" location="Municipality of Chilpancingo de los Bravo" longLatPrecision="7" longitude="-99.2979" municipality="Municipality of Chilpancingo" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero">
<emphasis id="45F994005A4C6529037CFB0628D5FB05" bold="true" box="[1046,1164,1218,1244]" pageId="32" pageNumber="33">
<collectingRegion id="B54986F05A4C6529037CFB0628D5FB05" box="[1046,1164,1218,1244]" country="Mexico" name="Guerrero" pageId="32" pageNumber="33">Guerrero</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4C652903CFFB062888FB04" box="[1189,1233,1218,1245]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<location id="72521EC95A4C652903B5FB062E4CFAD9" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C652903B5FB062E4CFAD9" country="Mexico" latitude="17.3119" longLatPrecision="7" longitude="-99.2979" municipality="Municipality of Chilpancingo" name="Municipality of Chilpancingo de los Bravo" pageId="32" pageNumber="33" stateProvince="Guerrero">Municipality of Chilpancingo de los Bravo</location>
;
<geoCoordinate id="12B92ED55A4C6529054AFB212EF1FAD9" box="[544,680,1253,1280]" degrees="17.3119" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="17.3119">17.3119° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C652905D8FB212F18FAD9" box="[690,833,1253,1280]" degrees="99.2979" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-99.2979">99.2979° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290426FB212FBBFB26" box="[844,994,1253,1280]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="32" pageNumber="33" unit="m" value="1205.0">
<quantity id="B075E5F75A4C65290426FB212FFDFAD9" box="[844,932,1253,1280]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="32" pageNumber="33" unit="m" value="1205.0">1205 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C65290484FB212828FB26" box="[1006,1137,1253,1279]" pageId="32" pageNumber="33" value="2018-07-01">
<collectingDate id="1377973A5A4C65290484FB212828FB26" box="[1006,1137,1253,1279]" pageId="32" pageNumber="33" value="2018-07-01">1 Jul. 2018</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290316FB22288AFB26" box="[1148,1235,1253,1280]" pageId="32" pageNumber="33">P. Solís</collectorName>
,
<collectorName id="DA782DC45A4C652903B4FB21292FFB26" box="[1246,1398,1253,1279]" pageId="32" pageNumber="33">S. Rodríguez</collectorName>
leg.; LATLAX Ara-0771
</materialsCitation>
<materialsCitation id="C7E5424F5A4C65290692FACD2F65FAB0" ID-GBIF-Occurrence="4421595315" collectingDate="2018-10-15" collectorName="A. Valdez &amp; P. Solis &amp; I. Navarro &amp; A. Juarez &amp; L. Cabrera" country="Mexico" county="Municipality of Pilcaya" elevation="1145" latitude="18.6703" location="Botanical garden" longLatPrecision="7" longitude="-99.5134" municipality="Botanical" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Guerrero">
<specimenCount id="618B839B5A4C65290692FACD2E62FAFA" box="[504,571,1289,1315]" pageId="32" pageNumber="33" type="female">2 ♀♀</specimenCount>
;
<collectingCounty id="9E53309E5A4C65290520FACC2F3BFAFB" box="[586,866,1288,1314]" pageId="32" pageNumber="33">Municipality of Pilcaya</collectingCounty>
,
<location id="72521EC95A4C65290404FACC2860FAFB" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C65290404FACC2860FAFB" box="[878,1081,1288,1314]" country="Mexico" county="Municipality of Pilcaya" latitude="18.6703" longLatPrecision="7" longitude="-99.5134" municipality="Botanical" name="Botanical garden" pageId="32" pageNumber="33" stateProvince="Guerrero">Botanical garden</location>
,
<location id="72521EC95A4C6529032EFACD2928FAFB" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C6529032EFACD2928FAFB" box="[1092,1393,1288,1315]" country="Mexico" county="Municipality of Pilcaya" latitude="18.6703" longLatPrecision="7" longitude="-99.5134" municipality="Botanical" name="Grutas de Cacahuamilma" pageId="32" pageNumber="33" stateProvince="Guerrero">Grutas de Cacahuamilma</location>
;
<geoCoordinate id="12B92ED55A4C652907D7FAE82D1CFA9F" box="[189,325,1324,1350]" degrees="18.6703" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="18.6703">18.6703° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C65290625FAE82D85FA9F" box="[335,476,1324,1351]" degrees="99.5134" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-99.5134">99.5134° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C6529068CFAE82E20FA9F" box="[486,633,1324,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="32" pageNumber="33" unit="m" value="1145.0">
<quantity id="B075E5F75A4C6529068CFAE82E65FA9F" box="[486,572,1324,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="32" pageNumber="33" unit="m" value="1145.0">1145 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C652905EEFAE82F42FA9F" box="[644,795,1324,1351]" pageId="32" pageNumber="33" value="2018-10-15">
<collectingDate id="1377973A5A4C652905EEFAE82F42FA9F" box="[644,795,1324,1351]" pageId="32" pageNumber="33" value="2018-10-15">15 Oct. 2018</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C6529044EFAE82FC1FA9F" box="[804,920,1324,1350]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C652904CBFAE82FAEFA9F" box="[929,1015,1324,1350]" pageId="32" pageNumber="33">P. Solís</collectorName>
,
<collectorName id="DA782DC45A4C6529036BFAE82822FA9F" box="[1025,1147,1324,1350]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C652903E8FAE828ABFA9F" box="[1154,1266,1324,1350]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C65290391FAE8292EFA9F" box="[1275,1399,1324,1350]" pageId="32" pageNumber="33">L. Cabrera</collectorName>
leg.; diurnal collecting; LATLAX Ara-0767, Ara-0772
</materialsCitation>
<materialsCitation id="C7E5424F5A4C6529043CFA8B2F22FA69" ID-GBIF-Occurrence="4421595301" collectingDate="2018-10-14" collectorName="A. Valdez &amp; P. Solis &amp; I. Navarro &amp; A. Juarez &amp; L. Cabrera" country="Mexico" elevation="923" latitude="17.4392" location="Tourist Center Grutas de Juxtlahuaca" longLatPrecision="7" longitude="-99.1593" municipality="Municipality of Quechultenango" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero">
<specimenCount id="618B839B5A4C6529043CFA8B2FDAFAB3" box="[854,899,1359,1386]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="9756D2685A4C652904F9FA8B294CFAB0" box="[915,1301,1359,1385]" pageId="32" pageNumber="33">Municipality of Quechultenango</collectingMunicipality>
,
<location id="72521EC95A4C6529024BFA8B2E40FA55" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C6529024BFA8B2E40FA55" country="Mexico" latitude="17.4392" longLatPrecision="7" longitude="-99.1593" municipality="Municipality of Quechultenango" name="Tourist Center Grutas de Juxtlahuaca" pageId="32" pageNumber="33" stateProvince="Guerrero">Tourist Center Grutas de Juxtlahuaca</location>
;
<geoCoordinate id="12B92ED55A4C6529054FFAB62EE9FA54" box="[549,688,1394,1421]" degrees="17.4392" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="17.4392">17.4392° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C652905D6FAB62F15FA54" box="[700,844,1394,1421]" degrees="99.1593" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-99.1593">99.1593° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290432FAB62FBBFA55" box="[856,994,1394,1421]" metricMagnitude="2" metricUnit="m" metricValue="9.23" pageId="32" pageNumber="33" unit="m" value="923.0">
<quantity id="B075E5F75A4C65290432FAB62FFAFA54" box="[856,931,1394,1421]" metricMagnitude="2" metricUnit="m" metricValue="9.23" pageId="32" pageNumber="33" unit="m" value="923.0">923 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C6529049AFAB628D5FA55" box="[1008,1164,1394,1420]" pageId="32" pageNumber="33" value="2018-10-14">
<collectingDate id="1377973A5A4C6529049AFAB628D5FA55" box="[1008,1164,1394,1420]" pageId="32" pageNumber="33" value="2018-10-14">14 Oct. 2018</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C652903FDFAB62954FA55" box="[1175,1293,1394,1420]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C65290273FAB72928FA55" box="[1305,1393,1394,1421]" pageId="32" pageNumber="33">P. Solís</collectorName>
,
<collectorName id="DA782DC45A4C652907D7FA522D62FA69" box="[189,315,1430,1456]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C6529062DFA522DE5FA76" box="[327,444,1429,1456]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C652906A0FA522E13FA76" box="[458,586,1429,1456]" pageId="32" pageNumber="33">L. Cabrera</collectorName>
leg.; LATLAX Ara-1057
</materialsCitation>
<materialsCitation id="C7E5424F5A4C652904F3FA512F79FA2E" ID-GBIF-Occurrence="4421595318" collectingDate="2018-09-21" collectorName="D. Montiel" country="Mexico" elevation="2330" latitude="18.5908" location="Municipality of Taxco de Alarcon" longLatPrecision="7" longitude="-99.6033" municipality="Municipality of Taxco de Alarcon" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero">
<specimenCount id="618B839B5A4C652904F3FA512F91FA69" box="[921,968,1429,1456]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="9756D2685A4C652904B3FA51292AFA76" box="[985,1395,1429,1456]" pageId="32" pageNumber="33">Municipality of Taxco de Alarcón</collectingMunicipality>
,
<locationDeviation id="668051305A4C652907D7FA7D2FD5FA0B" box="[189,908,1464,1491]" pageId="32" pageNumber="33">
<quantity id="B075E5F75A4C652907D7FA7D2D50FA0B" box="[189,265,1464,1491]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="km" value="10.0">10 km</quantity>
SW of Taxco, on the way to Cerro de Huixteco park
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4C652904F0FA7C287FFA0A" box="[922,1062,1464,1491]" degrees="18.5908" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="18.5908">18.5908° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C6529035EFA7C289FFA0A" box="[1076,1222,1464,1491]" degrees="99.6033" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-99.6033">99.6033° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C652903BEFA7D2929FA0B" box="[1236,1392,1464,1491]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="32" pageNumber="33" unit="m" value="2330.0">
<quantity id="B075E5F75A4C652903BEFA7D2976FA0A" box="[1236,1327,1465,1491]" metricMagnitude="3" metricUnit="m" metricValue="2.33" pageId="32" pageNumber="33" unit="m" value="2330.0">2330 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C652907D5FA182D07FA2F" box="[191,350,1500,1526]" pageId="32" pageNumber="33" value="2018-09-21">
<collectingDate id="1377973A5A4C652907D5FA182D07FA2F" box="[191,350,1500,1526]" pageId="32" pageNumber="33" value="2018-09-21">21 Sep. 2018</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290607FA182DB6FA2F" box="[365,495,1500,1526]" pageId="32" pageNumber="33">D. Montiel</collectorName>
leg.; LATLAX Ara-1065
</materialsCitation>
<materialsCitation id="C7E5424F5A4C65290457FA182E09F9E4" ID-GBIF-Occurrence="4421595306" collectingDate="2017-09-19" collectorName="A. Valdez &amp; I. Navarro &amp; P. Solis &amp; J. Valerdi" country="Mexico" elevation="930" latitude="18.4197" location="Miguel Hidalgo Street" longLatPrecision="52" longitude="-99.545" municipality="Municipality of Taxco de Alarcon" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero">
<specimenCount id="618B839B5A4C65290457FA182F32FA21" box="[829,875,1500,1528]" pageId="32" pageNumber="33" type="female">
1
<emphasis id="45F994005A4C6529043FFA182F32FA21" bold="true" box="[853,875,1500,1528]" pageId="32" pageNumber="33"></emphasis>
</specimenCount>
;
<collectingMunicipality id="9756D2685A4C65290416FA18294CFA2F" box="[892,1301,1500,1526]" pageId="32" pageNumber="33">Municipality of Taxco de Alarcón</collectingMunicipality>
,
<location id="72521EC95A4C65290248FA182D35F9C0" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C65290248FA182D35F9C0" country="Mexico" latitude="18.4197" longLatPrecision="52" longitude="-99.545" municipality="Municipality of Taxco de Alarcon" name="Miguel Hidalgo Street" pageId="32" pageNumber="33" stateProvince="Guerrero">Miguel Hidalgo Street</location>
, #23;
<geoCoordinate id="12B92ED55A4C652906D9FA3B2E64F9C3" box="[435,573,1535,1562]" degrees="18.4197" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="18.4197">18.4197° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C65290522FA3B2E9EF9C3" box="[584,711,1535,1562]" degrees="99.545" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-99.545">99.545° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C652905B9FA3B2F04F9C0" box="[723,861,1535,1562]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="32" pageNumber="33" unit="m" value="930.0">
<quantity id="B075E5F75A4C652905B9FA3B2F44F9C3" box="[723,797,1535,1562]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="32" pageNumber="33" unit="m" value="930.0">930 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C65290400FA3B285EF9C0" box="[874,1031,1535,1561]" pageId="32" pageNumber="33" value="2017-09-19">
<collectingDate id="1377973A5A4C65290400FA3B285EF9C0" box="[874,1031,1535,1561]" pageId="32" pageNumber="33" value="2017-09-19">19 Sep. 2017</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C6529037BFA3B28DEF9C0" box="[1041,1159,1535,1561]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C652903F8F9C42954F9C3" box="[1170,1293,1536,1562]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C65290273F9C42928F9C0" box="[1305,1393,1535,1562]" pageId="32" pageNumber="33">P. Solís</collectorName>
,
<collectorName id="DA782DC45A4C652907D7F9E72D73F9E5" box="[189,298,1570,1597]" pageId="32" pageNumber="33">J. Valerdi</collectorName>
leg.; LATLAX Ara-1075
</materialsCitation>
<materialsCitation id="C7E5424F5A4C6529050CF9E62E68F97F" ID-GBIF-Occurrence="4421595322" collectingDate="2020-11-22" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" county="Municipality of Tecpan de Galeana" elevation="79" latitude="17.2918" location="path to microwave repeater El Tamarindo" longLatPrecision="53" longitude="-101.045" municipality="El Tamarindo" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="2" specimenCount-immature="1" stateProvince="Guerrero">
<specimenCount id="618B839B5A4C6529050CF9E62EFEF9E6" box="[614,679,1570,1599]" pageId="32" pageNumber="33" type="female">
2
<emphasis id="45F994005A4C65290511F9E72EFEF9E6" bold="true" box="[635,679,1571,1599]" pageId="32" pageNumber="33">♀♀</emphasis>
</specimenCount>
, 1 ♁,
<specimenCount id="618B839B5A4C65290587F9E62F2CF9E5" box="[749,885,1570,1596]" pageId="32" pageNumber="33" type="immature">1 immature</specimenCount>
;
<collectingCounty id="9E53309E5A4C652904EAF9E62944F9E5" box="[896,1309,1570,1596]" pageId="32" pageNumber="33">Municipality of Técpan de Galeana</collectingCounty>
,
<location id="72521EC95A4C6529024DF9E62E13F986" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C6529024DF9E62E13F986" country="Mexico" county="Municipality of Tecpan de Galeana" latitude="17.2918" longLatPrecision="53" longitude="-101.045" municipality="El Tamarindo" name="path to microwave repeater El Tamarindo" pageId="32" pageNumber="33" stateProvince="Guerrero">path to microwave repeater El Tamarindo</location>
,
<locationDeviation id="668051305A4C6529053EF981281FF9B9" box="[596,1094,1605,1632]" pageId="32" pageNumber="33">
<quantity id="B075E5F75A4C6529053EF9812EFAF986" box="[596,675,1605,1632]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="km" value="1.5">1.5 km</quantity>
SW of viewpoint Bahía de Papanoa
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4C6529033BF9812880F9B9" box="[1105,1241,1605,1632]" degrees="17.2918" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="17.2918">17.2918° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C65290389F981292BF9B9" box="[1251,1394,1605,1632]" degrees="101.045" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-101.045">101.045° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C652907D1F9AD2D6CF95B" box="[187,309,1640,1667]" metricMagnitude="1" metricUnit="m" metricValue="7.9" pageId="32" pageNumber="33" unit="m" value="79.0">
<quantity id="B075E5F75A4C652907D1F9AD2CAFF95A" box="[187,246,1641,1667]" metricMagnitude="1" metricUnit="m" metricValue="7.9" pageId="32" pageNumber="33" unit="m" value="79.0">79 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C6529062BF9AD2DB8F95A" box="[321,481,1641,1667]" pageId="32" pageNumber="33" value="2020-11-22">
<collectingDate id="1377973A5A4C6529062BF9AD2DB8F95A" box="[321,481,1641,1667]" pageId="32" pageNumber="33" value="2020-11-22">22 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290681F9AD2E39F95B" box="[491,608,1640,1667]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C65290500F9AD2EBCF95A" box="[618,741,1641,1667]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C65290584F9AD2F06F95A" box="[750,863,1641,1667]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C65290403F9AD2FBEF95B" box="[873,999,1640,1667]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4C6529034DF9AC28A1F95B" box="[1063,1272,1640,1666]" pageId="32" pageNumber="33">diurnal collecting</collectingMethod>
; LATLAX Ara-1056, Ara-1063, Ara-1064
</materialsCitation>
.
<materialsCitation id="C7E5424F5A4C65290532F94F2F6CF913" ID-GBIF-Occurrence="4421595304" collectingDate="2018-09-21" collectingMethod="diurnal collecting" collectorName="I. Navarro" country="Mexico" elevation="1268" latitude="20.4991" location="Jalisco" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco">
<emphasis id="45F994005A4C65290532F94F2EF4F97C" bold="true" box="[600,685,1675,1701]" pageId="32" pageNumber="33">
<collectingRegion id="B54986F05A4C65290532F94F2EF4F97C" box="[600,685,1675,1701]" country="Mexico" name="Jalisco" pageId="32" pageNumber="33">Jalisco</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4C652905A0F9482EA0F97F" box="[714,761,1676,1702]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<date id="03336ED25A4C652903A5F948292BF97F" box="[1231,1394,1676,1702]" pageId="32" pageNumber="33" value="2018-09-21">
<collectingDate id="1377973A5A4C652903A5F948292BF97F" box="[1231,1394,1676,1702]" pageId="32" pageNumber="33" value="2018-09-21">21 Sep. 2018</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C652907D7F9742D6DF913" box="[189,308,1712,1738]" pageId="32" pageNumber="33">I. Navarro</collectorName>
leg.;
<collectingMethod id="AECC30055A4C6529061EF96B2E1DF910" box="[372,580,1711,1737]" pageId="32" pageNumber="33">diurnal collecting</collectingMethod>
; LATLAX Ara-0765
</materialsCitation>
<materialsCitation id="C7E5424F5A4C65290426F96B2EC5F935" ID-GBIF-Occurrence="4421595308" collectingDate="2019-01-05" collectingMethod="night collecting" collectorName="I. Navarro" country="Mexico" elevation="1268" latitude="20.4991" location="Camichines" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco">
<specimenCount id="618B839B5A4C65290426F96B2F21F913" box="[844,888,1711,1738]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<date id="03336ED25A4C65290259F96B2CA2F935" pageId="32" pageNumber="33" value="2019-01-05">
<collectingDate id="1377973A5A4C65290259F96B2CA2F935" pageId="32" pageNumber="33" value="2019-01-05">5 Jan. 2019</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C6529066EF9172D23F934" box="[260,378,1747,1773]" pageId="32" pageNumber="33">I. Navarro</collectorName>
leg.; LATLAX Ara-0776
</materialsCitation>
<materialsCitation id="C7E5424F5A4C652905C5F91628E2F8D6" ID-GBIF-Occurrence="4421595307" collectingDate="2019-06-21" collectingMethod="night collecting" collectorName="A. Valdez &amp; I. Navarro &amp; L. Cabrera &amp; J. Flores" country="Mexico" elevation="1268" latitude="20.4991" location="Camichines" longLatPrecision="7" longitude="-103.802" municipality="Municipality of Cocula" pageId="32" pageNumber="33" specimenCount="4" specimenCount-female="1" specimenCount-immature="3" stateProvince="Jalisco">
<specimenCount id="618B839B5A4C652905C5F9162E81F934" box="[687,728,1746,1773]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
,
<specimenCount id="618B839B5A4C6529058EF9162F2EF935" box="[740,887,1746,1772]" pageId="32" pageNumber="33" type="immature">3 immatures</specimenCount>
; same collection data as for holotype;
<date id="03336ED25A4C6529024FF9162CA2F8D6" pageId="32" pageNumber="33" value="2019-06-21">
<collectingDate id="1377973A5A4C6529024FF9162CA2F8D6" pageId="32" pageNumber="33" value="2019-06-21">21 Jun. 2019</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C6529066DF9322D26F8D6" box="[263,383,1781,1808]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C652906E7F9322E53F8C9" box="[397,522,1782,1808]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C6529057DF9322EC3F8D6" box="[535,666,1781,1808]" pageId="32" pageNumber="33">L. Cabrera</collectorName>
,
<collectorName id="DA782DC45A4C652905CDF9322F55F8D6" box="[679,780,1781,1808]" pageId="32" pageNumber="33">J. Flores</collectorName>
leg.; LATLAX Ara-1083, Ara-1084
</materialsCitation>
<materialsCitation id="C7E5424F5A4C652903BDF9312DAFF8A3" ID-GBIF-Occurrence="4421595319" collectingDate="2020-11-08" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1336" latitude="21.0314" location="municipality of Hostotipaquillo" longLatPrecision="7" longitude="-104.0668" municipality="municipality of Hostotipaquillo" pageId="32" pageNumber="33" specimenCount="15" specimenCount-female="6" specimenCount-immature="6" stateProvince="Jalisco">
<specimenCount id="618B839B5A4C652903BDF9312945F8C9" box="[1239,1308,1781,1808]" pageId="32" pageNumber="33" type="female">6 ♀♀</specimenCount>
,
<specimenCount id="618B839B5A4C65290241F9312929F8D6" box="[1323,1392,1779,1807]" pageId="32" pageNumber="33">3 ♁♁</specimenCount>
,
<specimenCount id="618B839B5A4C652907D7F8DC2D0AF8EB" box="[189,339,1816,1842]" pageId="32" pageNumber="33" type="immature">6 immatures</specimenCount>
;
<collectingMunicipality id="9756D2685A4C65290635F8DC2E8CF8EB" box="[351,725,1816,1842]" pageId="32" pageNumber="33">municipality of Hostotipaquillo</collectingMunicipality>
,
<locationDeviation id="668051305A4C6529058AF8DD2869F8EB" box="[736,1072,1816,1843]" location="Hostotipaquillo" pageId="32" pageNumber="33">
<quantity id="B075E5F75A4C6529058AF8DD2F42F8EB" box="[736,795,1816,1843]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="km" value="3.0">3 km</quantity>
SW of Hostotipaquillo
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4C65290356F8DD289EF8EA" box="[1084,1223,1817,1843]" degrees="21.0314" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="21.0314">21.0314° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C652903B9F8DC292BF8EA" box="[1235,1394,1816,1843]" degrees="104.0668" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-104.0668">104.0668° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C652907D7F8F82D0DF88F" box="[189,340,1852,1878]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" pageId="32" pageNumber="33" unit="m" value="1336.0">
<quantity id="B075E5F75A4C652907D7F8F82D4CF88F" box="[189,277,1852,1878]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" pageId="32" pageNumber="33" unit="m" value="1336.0">1336 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C6529060AF8F82DABF88F" box="[352,498,1852,1878]" pageId="32" pageNumber="33" value="2020-11-08">
<collectingDate id="1377973A5A4C6529060AF8F82DABF88F" box="[352,498,1852,1878]" pageId="32" pageNumber="33" value="2020-11-08">8 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290691F8F82E29F88F" box="[507,624,1852,1878]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C65290511F8F82EACF88F" box="[635,757,1852,1878]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C65290594F8F82F36F88F" box="[766,879,1852,1878]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C65290410F8F82FA1F88F" box="[890,1016,1852,1878]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.; LATLAX Ara-1043 to Ara-1050, Ara-1058, Ara-1059
</materialsCitation>
<materialsCitation id="C7E5424F5A4C6529057EF89B2EC5F866" ID-GBIF-Occurrence="4421595305" collectingDate="2020-11-05" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1406" latitude="19.4634" location="Municipality of Jilotlan de los Dolores" longLatPrecision="7" longitude="-102.5112" municipality="Municipality of Jilotlan de los Dolores" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="3" stateProvince="Jalisco">
<specimenCount id="618B839B5A4C6529057EF89B2E00F8A3" box="[532,601,1887,1914]" pageId="32" pageNumber="33" type="female">3 ♀♀</specimenCount>
;
<collectingMunicipality id="9756D2685A4C65290503F89B2865F8A0" box="[617,1084,1887,1913]" pageId="32" pageNumber="33">Municipality of Jilotlán de los Dolores</collectingMunicipality>
,
<locationDeviation id="668051305A4C65290320F89B292BF8A0" box="[1098,1394,1887,1913]" location="Agua Bendita" pageId="32" pageNumber="33">
<quantity id="B075E5F75A4C65290320F89B28DEF8A0" box="[1098,1159,1887,1913]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="33" unit="km" value="4.0">4 km</quantity>
S of Agua Bendita
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4C652907D7F8462D1EF844" box="[189,327,1922,1949]" degrees="19.4634" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="19.4634">19.4634° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C65290638F8462DB6F844" box="[338,495,1922,1949]" degrees="102.5112" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-102.5112">102.5112° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290691F8462ECAF845" box="[507,659,1922,1949]" metricMagnitude="3" metricUnit="m" metricValue="1.406" pageId="32" pageNumber="33" unit="m" value="1406.0">
<quantity id="B075E5F75A4C65290691F8462E0AF844" box="[507,595,1922,1949]" metricMagnitude="3" metricUnit="m" metricValue="1.406" pageId="32" pageNumber="33" unit="m" value="1406.0">1406 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C652905CAF8462F6BF845" box="[672,818,1922,1949]" pageId="32" pageNumber="33" value="2020-11-05">
<collectingDate id="1377973A5A4C652905CAF8462F6BF845" box="[672,818,1922,1949]" pageId="32" pageNumber="33" value="2020-11-05">5 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290456F8462FEBF845" box="[828,946,1922,1948]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C652904D7F8472861F844" box="[957,1080,1923,1949]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C6529032BF84628EAF845" box="[1089,1203,1922,1948]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C652903D4F8462965F845" box="[1214,1340,1922,1948]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.; LATLAX Ara-1060, Ara-1071, Ara-1072
</materialsCitation>
<materialsCitation id="C7E5424F5A4C652805D8F8612F0CFEFD" ID-GBIF-Occurrence="4421595312" collectingDate="2020-11-05" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" county="Municipality of Jilotlan" elevation="1163" lastPageId="33" lastPageNumber="34" latitude="19.4689" location="Agua Bendita" longLatPrecision="52" longitude="-102.546" municipality="Municipality of Jilotlan de los Dolores" pageId="32" pageNumber="33" specimenCount="3" stateProvince="Jalisco">
<specimenCount id="618B839B5A4C652905D8F8612EADF866" box="[690,756,1955,1983]" pageId="32" pageNumber="33">3 ♁♁</specimenCount>
;
<collectingMunicipality id="9756D2685A4C6529046BF861289DF866" box="[769,1220,1957,1983]" pageId="32" pageNumber="33">Municipality of Jilotlán de los Dolores</collectingMunicipality>
,
<location id="72521EC95A4C652903A7F862292BF866" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4C652903A7F862292BF866" box="[1229,1394,1957,1984]" country="Mexico" county="Municipality of Jilotlan" latitude="19.4689" longLatPrecision="52" longitude="-102.546" municipality="Municipality of Jilotlan de los Dolores" name="Agua Bendita" pageId="32" pageNumber="33" stateProvince="Jalisco">Agua Bendita</location>
;
<geoCoordinate id="12B92ED55A4C652907D7F80C2D12F83A" box="[189,331,1992,2019]" degrees="19.4689" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="19.4689">19.4689° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4C65290631F80C2DB7F83A" box="[347,494,1992,2019]" degrees="102.546" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-102.546">102.546° W</geoCoordinate>
;
<elevation id="FCA0AF215A4C65290694F80C2EC7F83B" box="[510,670,1992,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="32" pageNumber="33" unit="m" value="1163.0">
<quantity id="B075E5F75A4C65290694F80C2E03F83A" box="[510,602,1992,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="32" pageNumber="33" unit="m" value="1163.0">1163 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4C652905C5F80C2F13F83A" box="[687,842,1992,2019]" pageId="32" pageNumber="33" value="2020-11-05">
<collectingDate id="1377973A5A4C652905C5F80C2F13F83A" box="[687,842,1992,2019]" pageId="32" pageNumber="33" value="2020-11-05">5 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4C65290432F80D2F8BF83B" box="[856,978,1992,2019]" pageId="32" pageNumber="33">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4C6529048BF80D2838F83A" box="[993,1121,1993,2019]" pageId="32" pageNumber="33">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4C65290305F80D28BDF83A" box="[1135,1252,1993,2019]" pageId="32" pageNumber="33">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4C6529039EF80D292FF83B" box="[1268,1398,1992,2019]" pageId="32" pageNumber="33">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4D65280792FECD2D93FEFA" box="[248,458,265,291]" pageId="33" pageNumber="34">diurnal collecting</collectingMethod>
; LATLAX Ara-1053 to Ara-1055
</materialsCitation>
<materialsCitation id="C7E5424F5A4D65280405FECD2F01FEB0" ID-GBIF-Occurrence="4421595311" collectingDate="2020-11-14" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="109" latitude="19.8327" location="Municipality of Tomatlan" longLatPrecision="52" longitude="-105.304" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco">
<specimenCount id="618B839B5A4D65280405FECD2FC5FEFA" box="[879,924,265,291]" pageId="33" pageNumber="34" type="female">1 ♀</specimenCount>
;
<location id="72521EC95A4D652804C1FECD2885FEFA" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4D652804C1FECD2885FEFA" box="[939,1244,265,291]" country="Mexico" latitude="19.8327" longLatPrecision="52" longitude="-105.304" name="Municipality of Tomatlan" pageId="33" pageNumber="34" stateProvince="Jalisco">Municipality of Tomatlán</location>
,
<locationDeviation id="668051305A4D65280382FECD2D34FE9F" location="Campo Acosta" pageId="33" pageNumber="34">
<quantity id="B075E5F75A4D65280382FECD297AFEFA" box="[1256,1315,265,291]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="km" value="9.0">9 km</quantity>
NO of Campo Acosta
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4D65280610FEE82E5CFE9E" box="[378,517,300,327]" degrees="19.8327" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="19.8327">19.8327° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4D6528057BFEE82EFBFE9E" box="[529,674,300,327]" degrees="105.304" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="55" value="-105.304">105.304° W</geoCoordinate>
;
<elevation id="FCA0AF215A4D652805C4FEE82F63FE9F" box="[686,826,300,327]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="33" pageNumber="34" unit="m" value="109.0">
<quantity id="B075E5F75A4D652805C4FEE82EA0FE9E" box="[686,761,300,327]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="33" pageNumber="34" unit="m" value="109.0">109 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4D65280422FEE82FB2FE9F" box="[840,1003,300,327]" pageId="33" pageNumber="34" value="2020-11-14">
<collectingDate id="1377973A5A4D65280422FEE82FB2FE9F" box="[840,1003,300,327]" pageId="33" pageNumber="34" value="2020-11-14">14 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4D6528049CFEE82834FE9F" box="[1014,1133,300,326]" pageId="33" pageNumber="34">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4D65280313FEE928ACFE9E" box="[1145,1269,301,327]" pageId="33" pageNumber="34">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4D6528026AFEE8292BFE9F" box="[1280,1394,300,326]" pageId="33" pageNumber="34">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4D652807D4FE8B2D19FEB0" box="[190,320,335,361]" pageId="33" pageNumber="34">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4D652806E3FE8B2E04FEB0" box="[393,605,335,361]" pageId="33" pageNumber="34">diurnal collecting</collectingMethod>
; LATLAX Ara-1077
</materialsCitation>
<materialsCitation id="C7E5424F5A4D65280412FE8B2FECFE69" ID-GBIF-Occurrence="4421595313" collectingDate="2020-11-07" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="905" latitude="19.8133" location="Municipality of Tonaya" longLatPrecision="7" longitude="-104.0059" pageId="33" pageNumber="34" specimenCount="2" specimenCount-female="1" specimenCount-immature="1" stateProvince="Jalisco">
<specimenCount id="618B839B5A4D65280412FE8B2FFEFEB3" box="[888,935,335,362]" pageId="33" pageNumber="34" type="female">1 ♀</specimenCount>
,
<specimenCount id="618B839B5A4D652804D3FE8B281FFEB0" box="[953,1094,335,361]" pageId="33" pageNumber="34" type="immature">1 immature</specimenCount>
;
<location id="72521EC95A4D6528033FFE8B292AFEB3" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4D6528033FFE8B292AFEB3" box="[1109,1395,335,362]" country="Mexico" latitude="19.8133" longLatPrecision="7" longitude="-104.0059" name="Municipality of Tonaya" pageId="33" pageNumber="34" stateProvince="Jalisco">Municipality of Tonaya</location>
,
<locationDeviation id="668051305A4D652807D7FEB72DA8FE55" box="[189,497,370,397]" location="Amacuautitlan" pageId="33" pageNumber="34">
<quantity id="B075E5F75A4D652807D7FEB72CA1FE55" box="[189,248,370,397]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="km" value="1.0">1 km</quantity>
W of Amacuautitlán
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4D65280697FEB72EDEFE54" box="[509,647,371,397]" degrees="19.8133" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="19.8133">19.8133° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4D652805F9FEB62F6BFE54" box="[659,818,370,397]" degrees="104.0059" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="-104.0059">104.0059° W</geoCoordinate>
;
<elevation id="FCA0AF215A4D65280454FEB62F90FE55" box="[830,969,370,397]" metricMagnitude="2" metricUnit="m" metricValue="9.05" pageId="33" pageNumber="34" unit="m" value="905.0">
<quantity id="B075E5F75A4D65280454FEB62FD0FE54" box="[830,905,370,397]" metricMagnitude="2" metricUnit="m" metricValue="9.05" pageId="33" pageNumber="34" unit="m" value="905.0">905 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4D652804BCFEB72833FE54" box="[982,1130,371,397]" pageId="33" pageNumber="34" value="2020-11-07">
<collectingDate id="1377973A5A4D652804BCFEB72833FE54" box="[982,1130,371,397]" pageId="33" pageNumber="34" value="2020-11-07">7 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4D6528031EFEB728B2FE55" box="[1140,1259,370,397]" pageId="33" pageNumber="34">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4D6528039CFEB7292BFE54" box="[1270,1394,371,397]" pageId="33" pageNumber="34">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4D652807D7FE522D74FE69" box="[189,301,406,432]" pageId="33" pageNumber="34">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4D6528065DFE522DECFE69" box="[311,437,406,432]" pageId="33" pageNumber="34">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4D6528069EFE522E9DFE69" box="[500,708,406,432]" pageId="33" pageNumber="34">diurnal collecting</collectingMethod>
; LATLAX Ara-1078
</materialsCitation>
<materialsCitation id="C7E5424F5A4D652B04A6FE522FE7FE9F" ID-GBIF-Occurrence="4421595316" collectingDate="2020-11-08" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" county="Municipality of Zapopan" elevation="1605" lastPageId="34" lastPageNumber="35" latitude="20.712" location="La Primavera common" longLatPrecision="522" longitude="-103.57" municipality="Lince Cave" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco">
<specimenCount id="618B839B5A4D652804A6FE522FAEFE69" box="[972,1015,406,432]" pageId="33" pageNumber="34" type="female">1 ♀</specimenCount>
;
<collectingCounty id="9E53309E5A4D6528036EFE522973FE69" box="[1028,1322,406,432]" pageId="33" pageNumber="34">Municipality of Zapopan</collectingCounty>
,
<collectingMunicipality id="9756D2685A4D652B025EFE522CA5FEFA" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">Lince Cave</collectingMunicipality>
,
<location id="72521EC95A4E652B066DFECD2E41FEFA" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4E652B066DFECD2E41FEFA" box="[263,536,265,291]" country="Mexico" county="Municipality of Zapopan" latitude="20.712" longLatPrecision="522" longitude="-103.57" municipality="Lince Cave" name="La Primavera common" pageId="34" pageNumber="35" stateProvince="Jalisco">La Primavera common</location>
;
<geoCoordinate id="12B92ED55A4E652B0548FECD2EC4FEFA" box="[546,669,265,291]" degrees="20.712" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="20.712">20.712° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4E652B05C3FECD2F73FEFA" box="[681,810,265,292]" degrees="103.57" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="555" value="-103.57">103.57° W</geoCoordinate>
;
<elevation id="FCA0AF215A4E652B045FFECD2F94FEFA" box="[821,973,265,292]" metricMagnitude="3" metricUnit="m" metricValue="1.605" pageId="34" pageNumber="35" unit="m" value="1605.0">
<quantity id="B075E5F75A4E652B045FFECD2FD7FEFA" box="[821,910,265,292]" metricMagnitude="3" metricUnit="m" metricValue="1.605" pageId="34" pageNumber="35" unit="m" value="1605.0">1605 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4E652B04B0FECD2835FEFA" box="[986,1132,265,291]" pageId="34" pageNumber="35" value="2020-11-08">
<collectingDate id="1377973A5A4E652B04B0FECD2835FEFA" box="[986,1132,265,291]" pageId="34" pageNumber="35" value="2020-11-08">8 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4E652B031CFECD28B5FEFA" box="[1142,1260,265,291]" pageId="34" pageNumber="35">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4E652B039DFECD292BFEFA" box="[1271,1394,265,291]" pageId="34" pageNumber="35">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4E652B07D6FEE82D77FE9F" box="[188,302,300,326]" pageId="34" pageNumber="35">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4E652B0652FEE82DEFFE9F" box="[312,438,300,326]" pageId="34" pageNumber="35">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4E652B069DFEE82E91FE9F" box="[503,712,300,326]" pageId="34" pageNumber="35">diurnal collecting</collectingMethod>
; LATLAX Ara-1068
</materialsCitation>
.
<materialsCitation id="C7E5424F5A4E652B04B7FEEF2DA4FE69" ID-GBIF-Occurrence="4421595320" collectingDate="2020-11-19" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" county="Municipality of Coahuayana de Hidalgo" elevation="65" latitude="18.6721" location="La Piedra hill" longLatPrecision="7" longitude="-103.6464" municipality="La Piedra" pageId="34" pageNumber="35" specimenCount="18" specimenCount-female="8" specimenCount-immature="10" stateProvince="Michoacan">
<emphasis id="45F994005A4E652B04B7FEEF2831FE9F" bold="true" box="[989,1128,299,326]" pageId="34" pageNumber="35">
<collectingRegion id="B54986F05A4E652B04B7FEEF2831FE9F" box="[989,1128,299,326]" country="Mexico" name="Michoacan de Ocampo" pageId="34" pageNumber="35">Michoacán</collectingRegion>
</emphasis>
<specimenCount id="618B839B5A4E652B03EAFEE8289BFE9E" box="[1152,1218,300,327]" pageId="34" pageNumber="35" type="female">8 ♀♀</specimenCount>
,
<specimenCount id="618B839B5A4E652B03A5FEE8292BFE9F" box="[1231,1394,300,326]" pageId="34" pageNumber="35" type="immature">10 immatures</specimenCount>
;
<collectingCounty id="9E53309E5A4E652B07D7FE8B2ECEFEB0" box="[189,663,335,361]" pageId="34" pageNumber="35">Municipality of Coahuayana de Hidalgo</collectingCounty>
,
<location id="72521EC95A4E652B05C9FE942F1FFEB0" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4E652B05C9FE942F1FFEB0" box="[675,838,335,362]" country="Mexico" county="Municipality of Coahuayana de Hidalgo" latitude="18.6721" longLatPrecision="7" longitude="-103.6464" municipality="La Piedra" name="La Piedra hill" pageId="34" pageNumber="35" stateProvince="Michoacan">La Piedra hill</location>
;
<geoCoordinate id="12B92ED55A4E652B0438FE8B2F85FEB3" box="[850,988,335,362]" degrees="18.6721" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="18.6721">18.6721° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4E652B0482FE8B28DFFEB3" box="[1000,1158,335,362]" degrees="103.6464" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="-103.6464">103.6464° W</geoCoordinate>
;
<elevation id="FCA0AF215A4E652B03F8FE8B2954FEB0" box="[1170,1293,335,362]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="34" pageNumber="35" unit="m" value="65.0">
<quantity id="B075E5F75A4E652B03F8FE8B2894FEB3" box="[1170,1229,335,362]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="34" pageNumber="35" unit="m" value="65.0">65 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4E652B0270FE8B2CA2FE54" pageId="34" pageNumber="35" value="2020-11-19">
<collectingDate id="1377973A5A4E652B0270FE8B2CA2FE54" pageId="34" pageNumber="35" value="2020-11-19">19 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4E652B0662FEB72DD8FE55" box="[264,385,370,397]" pageId="34" pageNumber="35">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4E652B06E4FEB72E55FE54" box="[398,524,371,397]" pageId="34" pageNumber="35">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4E652B0573FEB72ED4FE54" box="[537,653,371,397]" pageId="34" pageNumber="35">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4E652B05F1FEB72F44FE55" box="[667,797,370,397]" pageId="34" pageNumber="35">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4E652B040EFEB62861FE55" box="[868,1080,370,396]" pageId="34" pageNumber="35">diurnal collecting</collectingMethod>
; LATLAX Ara-1062, Ara-1067, Ara-1079, Ara-1080
</materialsCitation>
<materialsCitation id="C7E5424F5A4E652B054BFE522EDFFE2E" ID-GBIF-Occurrence="4421595321" collectingDate="2020-11-06" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" county="Municipality of Cotija de la Paz" elevation="1630" latitude="19.8087" location="Municipality of Cotija de la Paz" longLatPrecision="7" longitude="-102.6884" municipality="Luis" pageId="34" pageNumber="35" specimenCount="1" specimenCount-immature="1" stateProvince="Michoacan de Ocampo">
<specimenCount id="618B839B5A4E652B054BFE522EE9FE69" box="[545,688,406,432]" pageId="34" pageNumber="35" type="immature">1 immature</specimenCount>
;
<collectingCounty id="9E53309E5A4E652B05ABFE52280FFE69" box="[705,1110,406,432]" pageId="34" pageNumber="35">Municipality of Cotija de la Paz</collectingCounty>
,
<locationDeviation id="668051305A4E652B030DFE52292BFE69" box="[1127,1394,406,432]" pageId="34" pageNumber="35">Luis G. Urbina street</locationDeviation>
;
<geoCoordinate id="12B92ED55A4E652B07D7FE7D2D1EFE0A" box="[189,327,441,467]" degrees="19.8087" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="19.8087">19.8087° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4E652B0639FE7D2DA8FE0A" box="[339,497,441,467]" degrees="102.6884" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="-102.6884">102.6884° W</geoCoordinate>
;
<quantity id="B075E5F75A4E652B0697FE7D2E04FE0A" box="[509,605,441,467]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="34" pageNumber="35" unit="m" value="1630.0">
<elevation id="FCA0AF215A4E652B0697FE7D2E04FE0A" box="[509,605,441,467]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="34" pageNumber="35" unit="m" value="1630.0">1630 m</elevation>
</quantity>
.a.s.l;
<date id="03336ED25A4E652B05F6FE7D2F76FE0A" box="[668,815,441,467]" pageId="34" pageNumber="35" value="2020-11-06">
<collectingDate id="1377973A5A4E652B05F6FE7D2F76FE0A" box="[668,815,441,467]" pageId="34" pageNumber="35" value="2020-11-06">6 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4E652B0453FE7D2FF6FE0A" box="[825,943,441,467]" pageId="34" pageNumber="35">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4E652B04D1FE7D286FFE0A" box="[955,1078,441,467]" pageId="34" pageNumber="35">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4E652B032AFE7D28EBFE0A" box="[1088,1202,441,467]" pageId="34" pageNumber="35">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4E652B03D7FE7D2965FE0A" box="[1213,1340,441,467]" pageId="34" pageNumber="35">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4E652B07D7FE182DD6FE2F" box="[189,399,476,502]" pageId="34" pageNumber="35">diurnal collecting</collectingMethod>
; LATLAX Ara-1052
</materialsCitation>
<materialsCitation id="C7E5424F5A4E652B05CBFE182EDEFDE4" ID-GBIF-Occurrence="4421595323" collectingDate="2020-11-03" collectingMethod="diurnal collecting" collectorName="A. Valdez &amp; I. Navarro &amp; A. Juarez &amp; S. Nolasco" country="Mexico" elevation="1466" latitude="19.572" location="Municipality of Tzitzio" longLatPrecision="55" longitude="-100.9235" pageId="34" pageNumber="35" specimenCount="2" specimenCount-female="1" specimenCount-immature="1" stateProvince="Michoacan de Ocampo">
<specimenCount id="618B839B5A4E652B05CBFE182E97FE2E" box="[673,718,476,503]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
,
<specimenCount id="618B839B5A4E652B05B7FE182F31FE2F" box="[733,872,476,502]" pageId="34" pageNumber="35" type="immature">1 immature</specimenCount>
;
<location id="72521EC95A4E652B041FFE1828D5FE2F" LSID="urn:lsid:plazi:treatment:FF24F9045A766523068BFECC28B8FBE9:72521EC95A4E652B041FFE1828D5FE2F" box="[885,1164,476,502]" country="Mexico" latitude="19.572" longLatPrecision="55" longitude="-100.9235" name="Municipality of Tzitzio" pageId="34" pageNumber="35" stateProvince="Michoacan de Ocampo">Municipality of Tzitzio</location>
,
<locationDeviation id="668051305A4E652B03F2FE18292BFE2F" box="[1176,1394,476,502]" location="Tzitzio" pageId="34" pageNumber="35">
<quantity id="B075E5F75A4E652B03F2FE18288CFE2F" box="[1176,1237,476,502]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="35" unit="km" value="2.0">2 km</quantity>
O of Tzitzio
</locationDeviation>
;
<geoCoordinate id="12B92ED55A4E652B07D7FE3B2D60FDC3" box="[189,313,511,538]" degrees="19.572" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="19.572">19.572° N</geoCoordinate>
,
<geoCoordinate id="12B92ED55A4E652B062FFE3B2DBDFDC3" box="[325,484,511,538]" degrees="100.9235" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="-100.9235">100.9235° W</geoCoordinate>
;
<elevation id="FCA0AF215A4E652B069AFE3B2ED3FDC0" box="[496,650,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.466" pageId="34" pageNumber="35" unit="m" value="1466.0">
<quantity id="B075E5F75A4E652B069AFE3B2E10FDC3" box="[496,585,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.466" pageId="34" pageNumber="35" unit="m" value="1466.0">1466 m</quantity>
a.s.l.
</elevation>
;
<date id="03336ED25A4E652B05FDFE3B2F72FDC0" box="[663,811,511,538]" pageId="34" pageNumber="35" value="2020-11-03">
<collectingDate id="1377973A5A4E652B05FDFE3B2F72FDC0" box="[663,811,511,538]" pageId="34" pageNumber="35" value="2020-11-03">3 Nov. 2020</collectingDate>
</date>
;
<collectorName id="DA782DC45A4E652B045CFDC42FF5FDC0" box="[822,940,511,538]" pageId="34" pageNumber="35">A. Valdez</collectorName>
,
<collectorName id="DA782DC45A4E652B04D2FDC4286DFDC3" box="[952,1076,512,538]" pageId="34" pageNumber="35">I. Navarro</collectorName>
,
<collectorName id="DA782DC45A4E652B0354FDC428E9FDC0" box="[1086,1200,511,538]" pageId="34" pageNumber="35">A. Juárez</collectorName>
,
<collectorName id="DA782DC45A4E652B03D6FE3B2962FDC0" box="[1212,1339,511,537]" pageId="34" pageNumber="35">S. Nolasco</collectorName>
leg.;
<collectingMethod id="AECC30055A4E652B07D7FDE62DD6FDE5" box="[189,399,546,572]" pageId="34" pageNumber="35">diurnal collecting</collectingMethod>
; LATLAX Ara-1070
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="23F2189A5A4D652807D7F84C2875F830" ID-DOI="http://doi.org/10.5281/zenodo.8405898" ID-Zenodo-Dep="8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,242,1928,1954]" targetBox="[189,1398,512,1882]" targetPageId="33">
<paragraph id="773248125A4D652807D7F84C2875F830" blockId="33.[189,1398,1928,2025]" pageId="33" pageNumber="34">
<emphasis id="45F994005A4D652807D7F84C2D11F87B" bold="true" box="[189,328,1928,1954]" pageId="33" pageNumber="34">Figs 2528.</emphasis>
<taxonomicName id="B08D33915A4D65280627F84C2F01F87B" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[333,856,1928,1954]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4D65280627F84C2E37F87B" box="[333,622,1928,1954]" italics="true" pageId="33" pageNumber="34">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4D65280437F84C2FECF87B" box="[861,949,1928,1954]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="45F994005A4D652804DDF84C2858F87B" bold="true" box="[951,1025,1928,1954]" pageId="33" pageNumber="34">2526</emphasis>
. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.
<emphasis id="45F994005A4D652805E2F86F2E88F81C" bold="true" box="[648,721,1963,1989]" pageId="33" pageNumber="34">2728</emphasis>
. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="3F971B995A4E652307D7FDA02845FC66" lastPageId="42" lastPageNumber="43" pageId="34" pageNumber="35" type="description">
<paragraph id="773248125A4E652B07D7FDA02D17FDA7" blockId="34.[189,334,612,638]" box="[189,334,612,638]" pageId="34" pageNumber="35">
<heading id="2C7AFF7E5A4E652B07D7FDA02D17FDA7" bold="true" box="[189,334,612,638]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="45F994005A4E652B07D7FDA02D17FDA7" bold="true" box="[189,334,612,638]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A4E652B0780FD572EDEFD74" blockId="34.[185,1399,658,861]" box="[234,647,658,685]" pageId="34" pageNumber="35">
<emphasis id="45F994005A4E652B0780FD572D1CFD74" bold="true" box="[234,325,659,685]" pageId="34" pageNumber="35">Female</emphasis>
(
<typeStatus id="A836F6B05A4E652B063FFD562DE7FD75" box="[341,446,658,684]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
CNAN-T01589)
</paragraph>
<paragraph id="773248125A4E652B07D7FD72293AFC84" blockId="34.[185,1399,658,861]" pageId="34" pageNumber="35">
<smallCapsWord id="71D4DECE5A4E652B07D7FD722D37FD17" baselines="713,713" box="[189,366,694,720]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="34" pageNumber="35">MEASUREMENTS</smallCapsWord>
. Total length 10.70. Carapace 4.30 long, 3.75 wide (ratio l/w: 1.14). Clypeus 0.75 long. Diameter AME 0.25, ALE 0.20, PME 0.22, PLE 0.23. Distance AMEALE 0.14, ALE PLE 0.22, PMEPLE 0.33, PMEPME 0.19. Leg I: total 28.80 (femur 8.10 +patella 2.30 +tibia 7.20 +metatarsus 8.70 +tarsus 2.50); leg II: 18.32 (5.50 +1.80 +4.00 + 5.30 +1.72); leg III: 13.23 (4.20 +1.48 +2.50 + 3.68 +1.37), leg IV: 24.18 (7.20 +2.08 +5.60 + 7.30 +2.00). Leg formula: 1-4-2-3.
</paragraph>
<caption id="23F2189A5A4E652B07D3F842292EF83E" ID-DOI="http://doi.org/10.5281/zenodo.8405900" ID-Zenodo-Dep="8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="34" pageNumber="35" startId="34.[185,238,1926,1952]" targetBox="[189,1398,958,1879]" targetPageId="34">
<paragraph id="773248125A4E652B07D3F842292EF83E" blockId="34.[185,1399,1926,2023]" pageId="34" pageNumber="35">
<emphasis id="45F994005A4E652B07D3F8422D08F879" bold="true" box="[185,337,1926,1952]" pageId="34" pageNumber="35">Figs 2931.</emphasis>
<taxonomicName id="B08D33915A4E652B060AF8422F27F879" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[352,894,1926,1952]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4E652B060AF8422ED3F879" box="[352,650,1926,1952]" italics="true" pageId="34" pageNumber="35">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4E652B04E6F8432FA9F878" box="[908,1008,1927,1953]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="773248125A4F652A07D7FECD2F3DFE69" blockId="35.[188,1399,265,433]" pageId="35" pageNumber="36">
<smallCapsWord id="71D4DECE5A4F652A07D7FECD2D11FEFB" baselines="284,285" box="[189,328,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="35" pageNumber="36">COLORATION</smallCapsWord>
. Live adult specimen dark brown, with distinctive red stripes dorsally on abdomen (
<figureCitation id="EFB654975A4F652A07AFFEE82D17FE9E" box="[197,334,300,327]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="35" pageNumber="36">Figs 1315</figureCitation>
,
<figureCitation id="EFB654975A4F652A0631FEE82D23FE9E" box="[347,378,300,327]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="35" pageNumber="36">25</figureCitation>
) and ventrally with a red hourglass pattern, complete, located between epigynum and spinnerets (red coloration pattern is lost under ethanol) (
<figureCitation id="EFB654975A4F652A0421FE8B2FF8FEB0" box="[843,929,335,361]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
). Chelicerae dark brown, white apically. Sternum, labium, and endites dark brown. Legs dark brown, metatarsi and tarsi paler coloration than other segments (
<figureCitation id="EFB654975A4F652A06EAFE522E5EFE68" box="[384,519,406,433]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="35" pageNumber="36">Figs 1315</figureCitation>
,
<figureCitation id="EFB654975A4F652A0579FE522E38FE68" box="[531,609,406,433]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="35" pageNumber="36">2526</figureCitation>
). Tarsi darker distally.
</paragraph>
<paragraph id="773248125A4F652C07D7FE1C2D03FEB0" blockId="35.[189,1400,471,639]" lastBlockId="37.[189,1398,265,361]" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<smallCapsWord id="71D4DECE5A4F652A07D7FE1C2D7FFE29" baselines="491,491" box="[189,294,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prosoma" pageId="35" pageNumber="36">PROSOMA</smallCapsWord>
. Carapace with small translucid setae scattered across, dark setae on clypeus and posterior part of ocular region. Fovea transverse, M-shaped. In lateral view, clypeus oblique, protruding close to chelicerae. Eyes widely separated, different sizes (
<figureCitation id="EFB654975A4F652A047EFDDA2F37FDE1" box="[788,878,542,568]" captionStart="Figs 7075" captionStartId="40.[188,241,1857,1883]" captionTargetBox="[189,1398,442,1810]" captionTargetId="figure-64@40.[189,1398,442,1810]" captionTargetPageId="40" captionText="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="35" pageNumber="36">Fig. 70</figureCitation>
). In dorsal view of carapace, AER and PER recurved. AME slightly elevating, protruding from carapace. ALE and PLE protruding from carapace (
<figureCitation id="EFB654975A4F652A07ADFDA02D47FDA7" box="[199,286,612,638]" captionStart="Figs 7075" captionStartId="40.[188,241,1857,1883]" captionTargetBox="[189,1398,442,1810]" captionTargetId="figure-64@40.[189,1398,442,1810]" captionTargetPageId="40" captionText="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="35" pageNumber="36">Fig. 70</figureCitation>
). Sternum setose, subtriangular, shield shaped (
<figureCitation id="EFB654975A4F652A0456FDA02FC8FDA7" box="[828,913,612,638]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
). Labium setose, pale distally, trapezoidal, longer setae apically. Endites setose, pale distally, sub square, longer setae apically. Chelicerae with long setae distally, close to the fangs (
<figureCitation id="EFB654975A49652C0511FEE82E8AFE9F" box="[635,723,300,326]" captionStart="Figs 7075" captionStartId="40.[188,241,1857,1883]" captionTargetBox="[189,1398,442,1810]" captionTargetId="figure-64@40.[189,1398,442,1810]" captionTargetPageId="40" captionText="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="37" pageNumber="38">Fig. 71</figureCitation>
). Palps with long setae, more setose in tarsus, with single toothed claw.
</paragraph>
<caption id="23F2189A5A4F652A07D1F8DA2F44F81F" ID-DOI="http://doi.org/10.5281/zenodo.8405902" ID-Zenodo-Dep="8405902" httpUri="https://zenodo.org/record/8405902/files/figure.png" pageId="35" pageNumber="36" startId="35.[187,240,1822,1848]" targetBox="[189,1398,708,1771]" targetPageId="35">
<paragraph id="773248125A4F652A07D1F8DA2F44F81F" blockId="35.[187,1401,1822,1990]" pageId="35" pageNumber="36">
<emphasis id="45F994005A4F652A07D1F8DA2D12F8E1" bold="true" box="[187,331,1822,1848]" pageId="35" pageNumber="36">Figs 3243.</emphasis>
<taxonomicName id="B08D33915A4F652A063EF8DA2F3FF8E1" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[340,870,1822,1848]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4F652A063EF8DA2E21F8E1" box="[340,632,1822,1848]" italics="true" pageId="35" pageNumber="36">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4F652A0404F8DB2F90F8E0" box="[878,969,1823,1849]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
Variation of the seminal receptacles of females, dorsal view.
<emphasis id="45F994005A4F652A06B2F8852E7BF882" bold="true" box="[472,546,1857,1883]" pageId="35" pageNumber="36">3233</emphasis>
. Cocula, Jalisco, Mexico (type locality).
<emphasis id="45F994005A4F652A0368F8852812F882" bold="true" box="[1026,1099,1857,1883]" pageId="35" pageNumber="36">3435</emphasis>
. Jilotitlán de los Dolores, Jalisco, Mexico.
<emphasis id="45F994005A4F652A06E5F8A12D80F8A6" bold="true" box="[399,473,1893,1919]" pageId="35" pageNumber="36">3637</emphasis>
. Minatitlán, Colima, Mexico.
<emphasis id="45F994005A4F652A043FF8A12FC7F8A6" bold="true" box="[853,926,1893,1919]" pageId="35" pageNumber="36">3839</emphasis>
. Coahuayana de Hidalgo, Michoacán, Mexico.
<emphasis id="45F994005A4F652A0642F84C2D28F87B" bold="true" box="[296,369,1928,1954]" pageId="35" pageNumber="36">4041</emphasis>
. Quechultenango, Guerrero, Mexico.
<emphasis id="45F994005A4F652A0451F84C2FDBF87B" bold="true" box="[827,898,1928,1954]" pageId="35" pageNumber="36">4243</emphasis>
. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="23F2189A5A48652D07D7F8CD2D47F868" ID-DOI="http://doi.org/10.5281/zenodo.8405904" ID-Zenodo-Dep="8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="36" pageNumber="37" startId="36.[189,242,1801,1827]" targetBox="[189,1398,265,1746]" targetPageId="36">
<paragraph id="773248125A48652D07D7F8CD2D47F868" blockId="36.[189,1399,1799,1969]" pageId="36" pageNumber="37">
<emphasis id="45F994005A48652D07D7F8CD2D13F8FA" bold="true" box="[189,330,1801,1827]" pageId="36" pageNumber="37">Figs 4448.</emphasis>
<taxonomicName id="B08D33915A48652D0639F8CD2F3AF8FA" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[339,867,1801,1827]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A48652D0639F8CD2E2FF8FA" box="[339,630,1801,1827]" italics="true" pageId="36" pageNumber="37">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A48652D0400F8CE2F91F8FD" box="[874,968,1802,1828]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♁ (CNAN-T01590), right palp.
<emphasis id="45F994005A48652D066AF8E82D44F89F" bold="true" box="[256,285,1836,1862]" pageId="36" pageNumber="37">44</emphasis>
. Anterior view.
<emphasis id="45F994005A48652D06B5F8E82DA5F89F" bold="true" box="[479,508,1836,1862]" pageId="36" pageNumber="37">45</emphasis>
. Dorsal view.
<emphasis id="45F994005A48652D05C7F8E82E90F89F" bold="true" box="[685,713,1836,1862]" pageId="36" pageNumber="37">46</emphasis>
. Ventral view,
<emphasis id="45F994005A48652D0414F8E82FC3F89F" bold="true" box="[894,922,1836,1862]" pageId="36" pageNumber="37">47</emphasis>
. Retrolateral view.
<emphasis id="45F994005A48652D03EDF8E828FAF89F" bold="true" box="[1159,1187,1836,1862]" pageId="36" pageNumber="37">48</emphasis>
). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="773248125A49652C07D7FE552DA8FDCC" blockId="37.[189,1400,401,533]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7FE552CAFFE70" baselines="420,420" box="[189,246,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="37" pageNumber="38">LEGS</smallCapsWord>
. Uniformly moderately setose throughout, tapering in metatarsi and tarsi (
<figureCitation id="EFB654975A49652C0301FE5528AFFE75" box="[1131,1270,401,428]" captionStart="Figs 7075" captionStartId="40.[188,241,1857,1883]" captionTargetBox="[189,1398,442,1810]" captionTargetId="figure-64@40.[189,1398,442,1810]" captionTargetPageId="40" captionText="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="37" pageNumber="38">Figs 7375</figureCitation>
). Femora I wider basally, femora I and IV slightly curved in lateral view. Tibiae I slightly curved distally. Three tarsal claws present, lateral claws with one row of teeth, which become larger distally. Tarsi IV with distinct comb macrosetae.
</paragraph>
<paragraph id="773248125A49652C07D7FDF82958FD8E" blockId="37.[189,1281,572,599]" box="[189,1281,572,599]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7FDF82D75FD8D" baselines="591,591" box="[189,300,572,598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="37" pageNumber="38">ABDOMEN</smallCapsWord>
. Globular, moderately setose, with all setae of uniform length (
<figureCitation id="EFB654975A49652C0379FDF828C0FD8E" box="[1043,1177,572,599]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="37" pageNumber="38">Figs 13, 15</figureCitation>
,
<figureCitation id="EFB654975A49652C03CCFDF828ADFD8E" box="[1190,1268,572,599]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">2526</figureCitation>
).
</paragraph>
<paragraph id="773248125A49652C07D7FDBA2FF5FCFD" blockId="37.[189,1399,637,804]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7FDBA2D69FD4F" baselines="657,657" box="[189,304,638,664]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Epigynum" pageId="37" pageNumber="38">EPIGYNUM</smallCapsWord>
. In ventral view oval, sclerotized and elevated, with oval and continuous opening, which is M-shaped in anterior part, opening 3.4× wider than long (
<figureCitation id="EFB654975A49652C04CFFD642859FD62" box="[933,1024,672,699]" captionStart="Figs 2931" captionStartId="34.[185,238,1926,1952]" captionTargetBox="[189,1398,958,1879]" captionTargetId="figure-253@34.[189,1398,958,1884]" captionTargetPageId="34" captionText="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="37" pageNumber="38">Fig. 29</figureCitation>
). In posterior view, copulatory openings visible, separated by a median septum (
<figureCitation id="EFB654975A49652C0593FD002F09FD07" box="[761,848,708,734]" captionStart="Figs 2931" captionStartId="34.[185,238,1926,1952]" captionTargetBox="[189,1398,958,1879]" captionTargetId="figure-253@34.[189,1398,958,1884]" captionTargetPageId="34" captionText="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="37" pageNumber="38">Fig. 31</figureCitation>
). In dorsal view, four spermathecae, dumbbellshaped, two anterior slightly bigger than others; copulatory ducts thinned-walled, transparent, weakly sclerotized, coiled around duct between spermathecae (
<figureCitation id="EFB654975A49652C042DFCCE2FC6FCFD" box="[839,927,778,804]" captionStart="Figs 2931" captionStartId="34.[185,238,1926,1952]" captionTargetBox="[189,1398,958,1879]" captionTargetId="figure-253@34.[189,1398,958,1884]" captionTargetPageId="34" captionText="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="37" pageNumber="38">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="773248125A49652C0780FC882E30FCBF" blockId="37.[185,1398,843,1046]" box="[234,617,843,870]" pageId="37" pageNumber="38">
<emphasis id="45F994005A49652C0780FC882D73FCBF" bold="true" box="[234,298,844,870]" pageId="37" pageNumber="38">Male</emphasis>
(
<typeStatus id="A836F6B05A49652C0653FC882DF9FCBF" box="[313,416,844,870]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
CNAN-T01590)
</paragraph>
<paragraph id="773248125A49652C07D7FCAB2F65FBCF" blockId="37.[185,1398,843,1046]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7FCAB2D37FC5E" baselines="898,898" box="[189,366,879,905]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="37" pageNumber="38">MEASUREMENTS</smallCapsWord>
. Total length 4.80. Carapace 2.00 long, 1.68 wide (ratio l/w: 1.19). Clypeus 0.40 long. Diameter AME 0.16, ALE 0.13, PME 0.14, PLE 0.13. Distance AMEALE 0.06, ALEPLE 0.06, PMEPLE 0.15, PMEPME 0.10. Leg I: total 17.70 (5.00 +1.06 + 4.50 +5.30 +1.84); leg II: 10.28 (3.00 +0.84 +2.34 + 2.88 +1.22); leg III: 7.25 (2.25 +0.68 +1.48 + 2.00 +0.84), leg IV: 13.53 (4.05 +0.90 +3.16 + 4.00 +1.42). Leg formula: 1-4-2-3.
</paragraph>
<paragraph id="773248125A49652C07D3FBF9292EFAF3" blockId="37.[185,1401,1085,1323]" pageId="37" pageNumber="38">
C OLORATION. Live adult specimens black, paler than female, with distinctive red stripes dorsally on abdomen, white line around stripes (
<figureCitation id="EFB654975A49652C0504FBA42EAFFBA3" box="[622,758,1120,1146]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="37" pageNumber="38">Figs 1618</figureCitation>
,
<figureCitation id="EFB654975A49652C0469FBA42F7DFBA3" box="[771,804,1120,1146]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">27</figureCitation>
). Abdomen ventrally with a red hourglass pattern, complete, thinner than females (red coloration pattern is lost under ethanol) (
<figureCitation id="EFB654975A49652C035DFB4728D7FB44" box="[1079,1166,1155,1181]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">Fig. 28</figureCitation>
). Chelicerae brown, paler than female, white apically. Palp femora and patellae, pale yellow; cymbium dark brown (
<figureCitation id="EFB654975A49652C0265FB622C87FB3D" captionStart="Figs 4448" captionStartId="36.[189,242,1801,1827]" captionTargetBox="[189,1398,265,1746]" captionTargetId="figure-13@36.[189,1398,265,1746]" captionTargetPageId="36" captionText="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="37" pageNumber="38">Figs 44 48</figureCitation>
). Sternum, labium, and endites paler brown than female, sternum with a pale brown line medially (
<figureCitation id="EFB654975A49652C07A2FB292D79FADE" box="[200,288,1261,1287]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">Fig. 28</figureCitation>
). Leg femora dark brown, except femora III, which is pale orange (
<figureCitation id="EFB654975A49652C0353FB292899FADE" box="[1081,1216,1261,1287]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="37" pageNumber="38">Figs 16, 18</figureCitation>
,
<figureCitation id="EFB654975A49652C03A7FB292945FADE" box="[1229,1308,1261,1287]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">2728</figureCitation>
). Tibiae pale orange, becoming dark brown towards joints. Metatarsi and tarsi pale orange (
<figureCitation id="EFB654975A49652C03E3FAD42949FAF3" box="[1161,1296,1296,1322]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="37" pageNumber="38">Figs 1618</figureCitation>
,
<figureCitation id="EFB654975A49652C0276FAD42932FAF3" box="[1308,1387,1296,1322]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">2728</figureCitation>
).
</paragraph>
<paragraph id="773248125A49652C07D7FA96292FF9E6" blockId="37.[187,1399,1361,1599]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7FA962D7FFAB3" baselines="1381,1381" box="[189,294,1362,1388]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prosoma" pageId="37" pageNumber="38">PROSOMA</smallCapsWord>
. Carapace with fewer small translucid setae scattered across than in female, less dark setae on clypeus and posterior part of ocular region than in female. Fovea transverse, M-shaped. In lateral view, clypeus oblique, protruding close to chelicerae (
<figureCitation id="EFB654975A49652C058DFA5C2F67FA6B" box="[743,830,1432,1458]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">Fig. 76</figureCitation>
). Eyes widely separated, different sizes (
<figureCitation id="EFB654975A49652C0279FA5C2930FA6B" box="[1299,1385,1432,1458]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">Fig. 76</figureCitation>
). In dorsal view of carapace, AER and PER recurved (
<figureCitation id="EFB654975A49652C0445FA7F2FD3FA0C" box="[815,906,1467,1493]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">Fig. 76</figureCitation>
). AME slightly elevated, protruding from carapace. ALE and PLE protruding from carapace (
<figureCitation id="EFB654975A49652C044EFA1A2F24FA21" box="[804,893,1502,1528]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">Fig. 76</figureCitation>
). Chelicerae with long setae distally, close to fangs (
<figureCitation id="EFB654975A49652C0644F9C52DDEF9C5" box="[302,391,1537,1564]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">Fig. 77</figureCitation>
). Sternum setose, subtriangular, shield shaped (
<figureCitation id="EFB654975A49652C04D0F9C5284BF9C5" box="[954,1042,1537,1564]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">Fig. 28</figureCitation>
). Labium setose, pale distally, trapezoidal, longer setae apically. Endites setose, pale distally, sub square, longer setae apically (
<figureCitation id="EFB654975A49652C027EF9E12930F9E6" box="[1300,1385,1573,1599]" captionStart="Figs 7075" captionStartId="40.[188,241,1857,1883]" captionTargetBox="[189,1398,442,1810]" captionTargetId="figure-64@40.[189,1398,442,1810]" captionTargetPageId="40" captionText="Figs 7075. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (female). 70. Frontal view of prosoma. 71. Detail of the chelicerae. 72. Dorsal view of epigynum. 7375. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="37" pageNumber="38">Fig. 71</figureCitation>
).
</paragraph>
<paragraph id="773248125A49652C07D1F9A32E75F933" blockId="37.[187,1399,1638,1770]" pageId="37" pageNumber="38">LEGS. Uniformly moderately setose throughout, tapering in metatarsi and tarsi. Femora I slightly wider basally, femora I and IV less curved in lateral view than female. Tibiae I and IV slightly curved distally. Three tarsal claws present, lateral claws with one row of teeth, which become larger distally. Tarsi IV with distinct comb macrosetae.</paragraph>
<paragraph id="773248125A49652C07D7F8D52FD3F896" blockId="37.[189,1398,1809,1871]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7F8D52D75F8F3" baselines="1828,1829" box="[189,300,1809,1835]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="37" pageNumber="38">ABDOMEN</smallCapsWord>
. Oval, longer than wide, and high, moderately setose, with setae all uniform length. ALS longer than PMS and PLS, oval colulus (
<figureCitation id="EFB654975A49652C05F6F8F02EAEF896" box="[668,759,1844,1871]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="37" pageNumber="38">Figs 17</figureCitation>
,
<figureCitation id="EFB654975A49652C046EF8F12F09F896" box="[772,848,1845,1871]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="37" pageNumber="38">2728</figureCitation>
,
<figureCitation id="EFB654975A49652C0436F8F12F25F896" box="[860,892,1845,1871]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">81</figureCitation>
).
</paragraph>
<paragraph id="773248125A49652C07D7F8B22801F80F" blockId="37.[188,1398,1910,2007]" pageId="37" pageNumber="38">
<smallCapsWord id="71D4DECE5A49652C07D7F8B22CADF857" baselines="1929,1929" box="[189,244,1910,1936]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palp" pageId="37" pageNumber="38">PALP</smallCapsWord>
. In retrolateral view, cymbial apophysis (ca) oval, setose: with tegular apophysis (ta), hook-shaped, transparent, weakly sclerotized (
<figureCitation id="EFB654975A49652C052EF85D2E88F86A" box="[580,721,1945,1971]" captionStart="Figs 4448" captionStartId="36.[189,242,1801,1827]" captionTargetBox="[189,1398,265,1746]" captionTargetId="figure-13@36.[189,1398,265,1746]" captionTargetPageId="36" captionText="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="37" pageNumber="38">Figs 4448</figureCitation>
,
<figureCitation id="EFB654975A49652C058BF85D2F76F86A" box="[737,815,1945,1971]" captionStart="Figs 7681" captionStartId="41.[189,242,1693,1719]" captionTargetBox="[189,1398,267,1639]" captionTargetId="figure-11@41.[189,1398,267,1639]" captionTargetPageId="41" captionText="Figs 7681. Scanning electron microscopy of L. occidentalis Valdez-Mondragón sp. nov. (male). 76. Frontal view of carapace. 77. Detail of distal part of the chelicera. 7880. Right palp: prolateral, retrolateral, and dorsal views, respectively. 81. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis." figureDoi="http://doi.org/10.5281/zenodo.8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="37" pageNumber="38">7880</figureCitation>
). In ventral-basal view, median apophysis bent, almost 90° degrees, ending in blunt point, with a notch in the middle (
<figureCitation id="EFB654975A49652C0498F8782812F80F" box="[1010,1099,1980,2006]" captionStart="Figs 4448" captionStartId="36.[189,242,1801,1827]" captionTargetBox="[189,1398,265,1746]" captionTargetId="figure-13@36.[189,1398,265,1746]" captionTargetPageId="36" captionText="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="37" pageNumber="38">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="773248125A4A652F07D7FECC2D6AFEFB" blockId="38.[189,307,264,290]" box="[189,307,264,290]" pageId="38" pageNumber="39">
<heading id="2C7AFF7E5A4A652F07D7FECC2D6AFEFB" bold="true" box="[189,307,264,290]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="45F994005A4A652F07D7FECC2D6AFEFB" bold="true" box="[189,307,264,290]" pageId="38" pageNumber="39">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A4A652307D7FEF328C8FE55" blockId="38.[189,1401,311,654]" lastBlockId="42.[189,1399,265,397]" lastPageId="42" lastPageNumber="43" pageId="38" pageNumber="39">
Females have black carapace and abdomen coloration, black or dark brown legs, never light shades (
<figureCitation id="EFB654975A4A652F07AFFE9E2D09FEAC" box="[197,336,346,373]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="38" pageNumber="39">Figs 2526</figureCitation>
). Globular abdomen always black with distinctive red lines in dorsal region (
<figureCitation id="EFB654975A4A652F0383FE9E292BFEAD" box="[1257,1394,346,372]" captionStart="Figs 6169" captionStartId="39.[189,242,1741,1767]" captionTargetBox="[189,1398,268,1683]" captionTargetId="figure-11@39.[189,1398,268,1683]" captionTargetPageId="39" captionText="Figs 6169. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the dorsal and ventral pattern coloration of the abdomen of the females of Latrodectus occidentalis sp. nov. Dorsal, posterior, and ventral views respectively. 6163. Chilpancingo, Guerrero, Mexico. 6466. Minatitlán, Colima, Mexico. 6769. Cocula, Jalisco, Mexico (type locality). Red coloration pattern gets lost under ethanol. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8405908" httpUri="https://zenodo.org/record/8405908/files/figure.png" pageId="38" pageNumber="39">Figs 6169</figureCitation>
, red coloration is lost under ethanol), lines may be outlined by a thin white line in juveniles. The frontal region of the dorsal sigmoidal line may or may not be separated at an isolated point between the rest of the line and the V-shape. The two pairs of lines laterally may be attenuated or reduced, but always present (
<figureCitation id="EFB654975A4A652F064CFE232DF6FDDB" box="[294,431,487,514]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="38" pageNumber="39">Figs 1315</figureCitation>
,
<figureCitation id="EFB654975A4A652F06D5FE232D86FDDB" box="[447,479,487,514]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="38" pageNumber="39">25</figureCitation>
,
<figureCitation id="EFB654975A4A652F0687FE232E65FDD8" box="[493,572,487,513]" captionStart="Figs 6169" captionStartId="39.[189,242,1741,1767]" captionTargetBox="[189,1398,268,1683]" captionTargetId="figure-11@39.[189,1398,268,1683]" captionTargetPageId="39" captionText="Figs 6169. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the dorsal and ventral pattern coloration of the abdomen of the females of Latrodectus occidentalis sp. nov. Dorsal, posterior, and ventral views respectively. 6163. Chilpancingo, Guerrero, Mexico. 6466. Minatitlán, Colima, Mexico. 6769. Cocula, Jalisco, Mexico (type locality). Red coloration pattern gets lost under ethanol. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8405908" httpUri="https://zenodo.org/record/8405908/files/figure.png" pageId="38" pageNumber="39">6169</figureCitation>
). Males always present a black body coloration (
<figureCitation id="EFB654975A4A652F03E7FE2328BEFDD8" box="[1165,1255,487,513]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="38" pageNumber="39">Fig. 17</figureCitation>
), legs black or dark brown with light brown on the mid tibia, metatarsus, and tarsus (
<figureCitation id="EFB654975A4A652F0342FDCE28EAFDFD" box="[1064,1203,522,548]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="38" pageNumber="39">Figs 1718</figureCitation>
). Abdomen oval with distinctive red lines on the dorsal region, which may be outlined by a thin white line (
<figureCitation id="EFB654975A4A652F0386FDE9292AFD91" box="[1260,1395,557,584]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="38" pageNumber="39">Figs 1718</figureCitation>
,
<figureCitation id="EFB654975A4A652F07D7FD952C87FDB2" box="[189,222,593,619]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="38" pageNumber="39">27</figureCitation>
). As in females, the frontal region of the transverse dorsal line may or may not be separated at an isolated point between the rest of the line and the V-shape (
<figureCitation id="EFB654975A4A652F0405FDB02F9FFD57" box="[879,966,628,654]" captionStart="Figs 2528" captionStartId="33.[189,242,1928,1954]" captionTargetBox="[189,1398,512,1882]" captionTargetId="figure-97@33.[189,1398,512,1882]" captionTargetPageId="33" captionText="Figs 2528. Latrodectus occidentalis Valdez-Mondragón sp. nov.2526. Holotype, ♀ (CNAN-T01589), dorsal and ventral habitus, respectively.2728. Paratype, ♁ (CNAN-T01590), dorsal and ventral habitus, respectively. Red arrow indicates coloration patterns. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405898" httpUri="https://zenodo.org/record/8405898/files/figure.png" pageId="38" pageNumber="39">Fig. 27</figureCitation>
). Males can vary greatly in body size (
<tableCitation id="3A0F7DA95A44652107AFFECD2D46FEFA" box="[197,287,265,291]" captionStart="Table 4" captionStartId="21.[185,255,595,623]" captionTargetPageId="21" captionText="Table 4. Average linear measurements (mm) analyzed of male specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimum and maximum. “T” = length/width of carapace. “TT” = Tibia-Patella length of legs I or IV." httpUri="http://table.plazi.org/id/23F2189A5A79651C07D3FD972CA5FD6F" pageId="40" pageNumber="41" tableUuid="23F2189A5A79651C07D3FD972CA5FD6F">Table 4</tableCitation>
), however sexual dimorphism is always marked, with females being more than twice the size of males, but males with proportionately longer legs (
<tableCitation id="3A0F7DA95A4465210460FEE82FD6FE9E" box="[778,911,300,327]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="21.[185,255,595,623]" captionStartId-1="22.[185,255,536,564]" captionTargetBox-1="[196,1910,663,1744]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Table 4. Average linear measurements (mm) analyzed of male specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimum and maximum. “T” = length/width of carapace. “TT” = Tibia-Patella length of legs I or IV." captionText-1="Table 5. Average linear measurements (mm) analyzed of female specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimums and maximums. “T” = Length/Width of the Cephalothorax. “TT” = Tibia-Patella length of legs I or IV." httpUri-0="http://table.plazi.org/id/23F2189A5A79651C07D3FD972CA5FD6F" httpUri-1="http://table.plazi.org/id/23F2189A5A7A651F07D3FDDC2DEFFDA2" pageId="40" pageNumber="41" tableUuid-0="23F2189A5A79651C07D3FD972CA5FD6F" tableUuid-1="23F2189A5A7A651F07D3FDDC2DEFFDA2">Tables 45</tableCitation>
). The genital opening of females varies in shape, but is always wider than long (
<figureCitation id="EFB654975A4465210506FE8B2E9FFEB0" box="[620,710,335,361]" captionStart="Figs 2931" captionStartId="34.[185,238,1926,1952]" captionTargetBox="[189,1398,958,1879]" captionTargetId="figure-253@34.[189,1398,958,1884]" captionTargetPageId="34" captionText="Figs 2931. Latrodectus occidentalis Valdez-Mondragón sp. nov., holotype, ♀ (CNAN-T01589). Epigynum, ventral, dorsal, and posterior views, respectively. Dotted lines indicate genital opening (29) and copulatory ducts (30). Abbreviations: CD = copulatory duct; S= spermathecae. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405900" httpUri="https://zenodo.org/record/8405900/files/figure.png" pageId="40" pageNumber="41">Figs 29</figureCitation>
,
<figureCitation id="EFB654975A44652105BBFE8B2F40FEB0" box="[721,793,335,361]" captionStart="Figs 3243" captionStartId="35.[187,240,1822,1848]" captionTargetBox="[189,1398,708,1771]" captionTargetId="figure-157@35.[189,1398,708,1771]" captionTargetPageId="35" captionText="Figs 3243. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the seminal receptacles of females, dorsal view. 3233. Cocula, Jalisco, Mexico (type locality). 3435. Jilotitlán de los Dolores, Jalisco, Mexico. 3637. Minatitlán, Colima, Mexico. 3839. Coahuayana de Hidalgo, Michoacán, Mexico. 4041. Quechultenango, Guerrero, Mexico. 4243. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405902" httpUri="https://zenodo.org/record/8405902/files/figure.png" pageId="40" pageNumber="41">3242</figureCitation>
(ventral views)), the spermathecae are arranged in a V-shape surrounded by the copulatory ducts which may have four or five turns around them (
<figureCitation id="EFB654975A4665230399FECD292FFEFA" box="[1267,1398,265,291]" captionStart="Figs 3243" captionStartId="35.[187,240,1822,1848]" captionTargetBox="[189,1398,708,1771]" captionTargetId="figure-157@35.[189,1398,708,1771]" captionTargetPageId="35" captionText="Figs 3243. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the seminal receptacles of females, dorsal view. 3233. Cocula, Jalisco, Mexico (type locality). 3435. Jilotitlán de los Dolores, Jalisco, Mexico. 3637. Minatitlán, Colima, Mexico. 3839. Coahuayana de Hidalgo, Michoacán, Mexico. 4041. Quechultenango, Guerrero, Mexico. 4243. Hostotipaquillo, Jalisco, Mexico. Dotted lines indicate genital opening. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8405902" httpUri="https://zenodo.org/record/8405902/files/figure.png" pageId="42" pageNumber="43">Figs 3343</figureCitation>
(dorsal views)). The angle of the spermathecae may vary, but they are never parallel as in
<taxonomicName id="B08D33915A46652303A2FEE9292CFE9F" authorityName="C. L. Koch" authorityYear="1841" box="[1224,1397,301,326]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="geometricus">
<emphasis id="45F994005A46652303A2FEE9292CFE9F" box="[1224,1397,301,326]" italics="true" pageId="42" pageNumber="43">L. geometricus</emphasis>
</taxonomicName>
(
<tableCitation id="3A0F7DA95A46652307AFFE8B2D79FEB3" box="[197,288,335,362]" captionStart="Table 5" captionStartId="22.[185,255,536,564]" captionTargetBox="[196,1910,663,1744]" captionTargetPageId="22" captionText="Table 5. Average linear measurements (mm) analyzed of female specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimums and maximums. “T” = Length/Width of the Cephalothorax. “TT” = Tibia-Patella length of legs I or IV." httpUri="http://table.plazi.org/id/23F2189A5A7A651F07D3FDDC2DEFFDA2" pageId="42" pageNumber="43" tableUuid="23F2189A5A7A651F07D3FDDC2DEFFDA2">Table 5</tableCitation>
). Palps of males show little variation, sclerites with little variation, embolus always with three complete turns (
<figureCitation id="EFB654975A4665230611FEB62D8AFE55" box="[379,467,370,396]" captionStart="Figs 4448" captionStartId="36.[189,242,1801,1827]" captionTargetBox="[189,1398,265,1746]" captionTargetId="figure-13@36.[189,1398,265,1746]" captionTargetPageId="36" captionText="Figs 4448. Latrodectus occidentalis Valdez-Mondragón sp. nov., paratype, ♁ (CNAN-T01590), right palp. 44. Anterior view. 45. Dorsal view. 46. Ventral view, 47. Retrolateral view. 48). Prolateral view. Red arrows indicate the number of turns of the embolus. Abbreviations: C = conductor; Cb = conductor base; E = embolus; MA = median aphophysis; T = tegulum; TA = tegular aphophysis. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405904" httpUri="https://zenodo.org/record/8405904/files/figure.png" pageId="42" pageNumber="43">Fig. 46</figureCitation>
); however, there is variation in the apical part (
<figureCitation id="EFB654975A4665230491FEB628DCFE55" box="[1019,1157,370,396]" captionStart="Figs 4960" captionStartId="38.[188,241,1857,1883]" captionTargetBox="[189,1398,745,1807]" captionTargetId="figure-193@38.[189,1398,745,1807]" captionTargetPageId="38" captionText="Figs 4960. Latrodectus occidentalis Valdez-Mondragón sp. nov. Variation of the male left palps, prolateral views. 4952. Quechultenango, Guerrero, Mexico. 5356. Minatitlán, Colima, Mexico. 57 60. Cocula, Jalisco, Mexico (type locality). Ventral view (49, 53, 57); retrolateral-frontal view (50, 54, 59); retrolateral view (51, 55, 59); posterior view (52, 56, 60). 4956. Postcopulatory palps. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8405906" httpUri="https://zenodo.org/record/8405906/files/figure.png" pageId="42" pageNumber="43">Figs 4960</figureCitation>
).
</paragraph>
<caption id="23F2189A5A4A652F07D6F8852D6EF830" ID-DOI="http://doi.org/10.5281/zenodo.8405906" ID-Zenodo-Dep="8405906" httpUri="https://zenodo.org/record/8405906/files/figure.png" pageId="38" pageNumber="39" startId="38.[188,241,1857,1883]" targetBox="[189,1398,745,1807]" targetPageId="38">
<paragraph id="773248125A4A652F07D6F8852D6EF830" blockId="38.[188,1400,1857,2025]" pageId="38" pageNumber="39">
<emphasis id="45F994005A4A652F07D6F8852D16F882" bold="true" box="[188,335,1857,1883]" pageId="38" pageNumber="39">Figs 4960.</emphasis>
<taxonomicName id="B08D33915A4A652F0636F8852F2FF882" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[348,886,1857,1883]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4A652F0636F8852EDDF882" box="[348,644,1857,1883]" italics="true" pageId="38" pageNumber="39">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4A652F04E8F8862FB8F885" box="[898,993,1858,1884]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
Variation of the male left palps, prolateral views.
<emphasis id="45F994005A4A652F06E6F8A12D8DF8A6" bold="true" box="[396,468,1893,1919]" pageId="38" pageNumber="39">4952</emphasis>
. Quechultenango, Guerrero, Mexico.
<emphasis id="45F994005A4A652F04F2F8A12FB8F8A6" bold="true" box="[920,993,1893,1919]" pageId="38" pageNumber="39">5356</emphasis>
. Minatitlán, Colima, Mexico.
<emphasis id="45F994005A4A652F0226F8A12C83F87B" bold="true" pageId="38" pageNumber="39">57 60</emphasis>
. Cocula, Jalisco, Mexico (type locality). Ventral view (49, 53, 57); retrolateral-frontal view (50, 54, 59); retrolateral view (51, 55, 59); posterior view (52, 56, 60).
<emphasis id="45F994005A4A652F04CBF86F2FB3F81C" bold="true" box="[929,1002,1963,1989]" pageId="38" pageNumber="39">4956</emphasis>
. Postcopulatory palps. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="23F2189A5A4B652E07D7F9092DF9F8AD" ID-DOI="http://doi.org/10.5281/zenodo.8405908" ID-Zenodo-Dep="8405908" httpUri="https://zenodo.org/record/8405908/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,242,1741,1767]" targetBox="[189,1398,268,1683]" targetPageId="39">
<paragraph id="773248125A4B652E07D7F9092DF9F8AD" blockId="39.[189,1399,1741,1908]" pageId="39" pageNumber="40">
<emphasis id="45F994005A4B652E07D7F9092D15F93E" bold="true" box="[189,332,1741,1767]" pageId="39" pageNumber="40">Figs 6169.</emphasis>
<taxonomicName id="B08D33915A4B652E063DF9092F35F93E" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[343,876,1741,1767]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4B652E063DF9092E25F93E" box="[343,636,1741,1767]" italics="true" pageId="39" pageNumber="40">Latrodectus occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4B652E041FF9092F8BF93E" box="[885,978,1741,1767]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
Variation of the dorsal and ventral pattern coloration of the abdomen of the females of
<taxonomicName id="B08D33915A4B652E044BF934281BF8D3" authority="Valdez-Mondragón &amp; Cabrera-Espinosa, 2023" authorityName="Valdez-Mondragón &amp; Cabrera-Espinosa" authorityYear="2023" box="[801,1090,1776,1802]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4B652E044BF934281BF8D3" box="[801,1090,1776,1802]" italics="true" pageId="39" pageNumber="40">Latrodectus occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A4B652E0323F93528FDF8D2" box="[1097,1188,1777,1803]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
Dorsal, posterior, and ventral views respectively.
<emphasis id="45F994005A4B652E0556F8D72EDCF8F4" bold="true" box="[572,645,1811,1837]" pageId="39" pageNumber="40">6163</emphasis>
. Chilpancingo, Guerrero, Mexico.
<emphasis id="45F994005A4B652E035BF8D72823F8F4" bold="true" box="[1073,1146,1811,1837]" pageId="39" pageNumber="40">6466</emphasis>
. Minatitlán, Colima, Mexico.
<emphasis id="45F994005A4B652E064CF8F22D36F889" bold="true" box="[294,367,1846,1872]" pageId="39" pageNumber="40">6769</emphasis>
. Cocula, Jalisco, Mexico (type locality). Red coloration pattern gets lost under ethanol. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="23F2189A5A44652107D6F8852E86F831" ID-DOI="http://doi.org/10.5281/zenodo.8405910" ID-Zenodo-Dep="8405910" httpUri="https://zenodo.org/record/8405910/files/figure.png" pageId="40" pageNumber="41" startId="40.[188,241,1857,1883]" targetBox="[189,1398,442,1810]" targetPageId="40">
<paragraph id="773248125A44652107D6F8852E86F831" blockId="40.[188,1399,1857,2025]" pageId="40" pageNumber="41">
<emphasis id="45F994005A44652107D6F8852D17F882" bold="true" box="[188,334,1857,1883]" pageId="40" pageNumber="41">Figs 7075.</emphasis>
Scanning electron microscopy of
<taxonomicName id="B08D33915A4465210593F88628CEF882" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[761,1175,1857,1883]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A4465210593F8862FF3F882" box="[761,938,1857,1883]" italics="true" pageId="40" pageNumber="41">L. occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A44652103C8F8862958F885" box="[1186,1281,1858,1884]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
(female).
<emphasis id="45F994005A44652107D7F8A12C83F8A6" bold="true" box="[189,218,1893,1919]" pageId="40" pageNumber="41">70</emphasis>
. Frontal view of prosoma.
<emphasis id="45F994005A4465210573F8A12E6FF8A6" bold="true" box="[537,566,1893,1919]" pageId="40" pageNumber="41">71</emphasis>
. Detail of the chelicerae.
<emphasis id="45F994005A446521040FF8A12FDBF8A6" bold="true" box="[869,898,1893,1919]" pageId="40" pageNumber="41">72</emphasis>
. Dorsal view of epigynum.
<emphasis id="45F994005A44652103A1F8A1294CF8A6" bold="true" box="[1227,1301,1893,1919]" pageId="40" pageNumber="41">7375</emphasis>
. Details of the setae and sockets on the legs. Abbreviations: ALE = anterior lateral eye; AME = anterior median eye; BQ = basichelicera; CD= copulatory duct; F = fang; PLE= posterior lateral eye; PME= posterior median eye; Q= chelicerae; S = spermathecae.
</paragraph>
</caption>
<caption id="23F2189A5A45652007D7F959288DF852" ID-DOI="http://doi.org/10.5281/zenodo.8405912" ID-Zenodo-Dep="8405912" httpUri="https://zenodo.org/record/8405912/files/figure.png" pageId="41" pageNumber="42" startId="41.[189,242,1693,1719]" targetBox="[189,1398,267,1639]" targetPageId="41">
<paragraph id="773248125A45652007D7F959288DF852" blockId="41.[188,1399,1693,1931]" pageId="41" pageNumber="42">
<emphasis id="45F994005A45652007D7F9592D09F96E" bold="true" box="[189,336,1693,1719]" pageId="41" pageNumber="42">Figs 7681.</emphasis>
Scanning electron microscopy of
<taxonomicName id="B08D33915A456520046FF95A28FEF961" authority="Valdez-Mondragon" authorityName="Valdez-Mondragon" authorityYear="2023" box="[773,1191,1694,1720]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="45F994005A456520046FF95A2FE1F961" box="[773,952,1694,1720]" italics="true" pageId="41" pageNumber="42">L. occidentalis</emphasis>
Valdez-Mondragón
</taxonomicName>
<taxonomicNameLabel id="5ECA297B5A45652003DFF95A294FF961" box="[1205,1302,1694,1720]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
(male).
<emphasis id="45F994005A45652007D7F9052CBBF902" bold="true" box="[189,226,1729,1755]" pageId="41" pageNumber="42">76.</emphasis>
Frontal view of carapace.
<emphasis id="45F994005A456520054DF9052E12F902" bold="true" box="[551,587,1729,1755]" pageId="41" pageNumber="42">77.</emphasis>
Detail of distal part of the chelicera.
<emphasis id="45F994005A456520037FF905283CF902" bold="true" box="[1045,1125,1729,1755]" pageId="41" pageNumber="42">7880.</emphasis>
Right palp: prolateral, retrolateral, and dorsal views, respectively.
<emphasis id="45F994005A45652005A6F9202EB0F927" bold="true" box="[716,745,1764,1790]" pageId="41" pageNumber="42">81</emphasis>
. Ventral-apical view of the spinnerets. Abbreviations: ALE = anterior lateral eye; ALS = anterior lateral spinneret; AME = anterior median eye; BQ = basichelicera; C = conductor; Cb = conductor base; Cu = colulus; E = Embolus; F = fang; MA = median aphophysis; PLE= posterior lateral eye; PLS= posterior lateral spinneret; PME= posterior median eye; PMS = posterior median spinneret; Q = chelicera; T = tegulum; TA = tegular aphophysis.
</paragraph>
</caption>
<paragraph id="773248125A46652307D7FE702EF1FDCD" blockId="42.[189,1399,436,532]" pageId="42" pageNumber="43">
Males (N = 12): Tibia-Patella I: 5.316.8 (x = 5.77); T coefficient: 1.081.25 (x = 1.19). Females (N = 30): Tibia-Patella I: 6.49.1 (x = 7.96); T coefficient: 0.981.22 (x = 1.11). See
<tableCitation id="3A0F7DA95A4665230307FE13289CFE28" box="[1133,1221,471,497]" captionStart="Table 4" captionStartId="21.[185,255,595,623]" captionTargetPageId="21" captionText="Table 4. Average linear measurements (mm) analyzed of male specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimum and maximum. “T” = length/width of carapace. “TT” = Tibia-Patella length of legs I or IV." httpUri="http://table.plazi.org/id/23F2189A5A79651C07D3FD972CA5FD6F" pageId="42" pageNumber="43" tableUuid="23F2189A5A79651C07D3FD972CA5FD6F">Table 4</tableCitation>
and
<tableCitation id="3A0F7DA95A4665230395FE132957FE2B" box="[1279,1294,471,498]" captionStart="Table 5" captionStartId="22.[185,255,536,564]" captionTargetBox="[196,1910,663,1744]" captionTargetPageId="22" captionText="Table 5. Average linear measurements (mm) analyzed of female specimens from the putative species of Latrodectus Walckenaer, 1805 in Mexico recorded in this study. Values in parentheses represent minimums and maximums. “T” = Length/Width of the Cephalothorax. “TT” = Tibia-Patella length of legs I or IV." httpUri="http://table.plazi.org/id/23F2189A5A7A651F07D3FDDC2DEFFDA2" pageId="42" pageNumber="43" tableUuid="23F2189A5A7A651F07D3FDDC2DEFFDA2">5</tableCitation>
for intra and interspecific morphological variation.
</paragraph>
<paragraph id="773248125A46652307D7FDFE2D2CFD8D" blockId="42.[189,373,570,596]" box="[189,373,570,596]" pageId="42" pageNumber="43">
<heading id="2C7AFF7E5A46652307D7FDFE2D2CFD8D" bold="true" box="[189,373,570,596]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="45F994005A46652307D7FDFE2D2CFD8D" bold="true" box="[189,373,570,596]" pageId="42" pageNumber="43">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A46652307D7FDAD2845FC66" blockId="42.[188,1400,617,960]" pageId="42" pageNumber="43">
The specimens were collected in their cobwebs in natural and anthropized localities (
<figureCitation id="EFB654975A46652303C1FDAD296CFD5A" box="[1195,1333,617,643]" captionStart="Figs 1924" captionStartId="31.[189,242,1775,1801]" captionTargetBox="[189,1398,268,1701]" captionTargetId="figure-11@31.[189,1398,268,1701]" captionTargetPageId="31" captionText="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." figureDoi="http://doi.org/10.5281/zenodo.8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="42" pageNumber="43">Figs 1924</figureCitation>
). The species is distributed mainly in tropical deciduous forests in the western states of
<collectingCountry id="0F9A08825A46652303D9FD482948FD7F" box="[1203,1297,652,678]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
:
<collectingRegion id="B54986F05A466523024AFD48292AFD7F" box="[1312,1395,652,678]" country="Mexico" name="Jalisco" pageId="42" pageNumber="43">Jalisco</collectingRegion>
,
<collectingRegion id="B54986F05A46652307D7FD6B2D73FD10" box="[189,298,687,713]" country="Mexico" name="Guerrero" pageId="42" pageNumber="43">Guerrero</collectingRegion>
,
<collectingRegion id="B54986F05A4665230653FD6B2DE7FD10" box="[313,446,687,713]" country="Mexico" name="Michoacan de Ocampo" pageId="42" pageNumber="43">Michoacán</collectingRegion>
,
<collectingRegion id="B54986F05A46652306A7FD6B2E7FFD10" box="[461,550,687,713]" country="Mexico" name="Colima" pageId="42" pageNumber="43">Colima</collectingRegion>
,
<collectingRegion id="B54986F05A466523055FFD6B2EE6FD10" box="[565,703,687,713]" country="Mexico" name="Guanajuato" pageId="42" pageNumber="43">Guanajuato</collectingRegion>
,
<collectingRegion id="B54986F05A46652305A7FD6B2F6BFD10" box="[717,818,687,713]" country="Mexico" name="Morelos" pageId="42" pageNumber="43">Morelos</collectingRegion>
, and
<collectingRegion id="B54986F05A466523041CFD6B2F97FD10" box="[886,974,687,713]" country="Mexico" name="Oaxaca" pageId="42" pageNumber="43">Oaxaca</collectingRegion>
(
<figureCitation id="EFB654975A466523048BFD6B2862FD10" box="[993,1083,687,713]" captionStart="Figs 1924" captionStartId="31.[189,242,1775,1801]" captionTargetBox="[189,1398,268,1701]" captionTargetId="figure-11@31.[189,1398,268,1701]" captionTargetPageId="31" captionText="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." figureDoi="http://doi.org/10.5281/zenodo.8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="42" pageNumber="43">Fig. 21</figureCitation>
). The specimens from the
<typeStatus id="A836F6B05A46652307D7FD172CA9FD34" box="[189,240,723,749]" pageId="42" pageNumber="43">type</typeStatus>
locality were collected inside the same house, under chairs, on window frames, under stowed dishes, and furniture (
<figureCitation id="EFB654975A46652306ACFD312E08FCC9" box="[454,593,757,784]" captionStart="Figs 1924" captionStartId="31.[189,242,1775,1801]" captionTargetBox="[189,1398,268,1701]" captionTargetId="figure-11@31.[189,1398,268,1701]" captionTargetPageId="31" captionText="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." figureDoi="http://doi.org/10.5281/zenodo.8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="42" pageNumber="43">Figs 2324</figureCitation>
, red arrows). Specimens collected in nature were found under and between big boulders on the ground (
<figureCitation id="EFB654975A4665230518FCDD2EA0FCEA" box="[626,761,793,819]" captionStart="Figs 1924" captionStartId="31.[189,242,1775,1801]" captionTargetBox="[189,1398,268,1701]" captionTargetId="figure-11@31.[189,1398,268,1701]" captionTargetPageId="31" captionText="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." figureDoi="http://doi.org/10.5281/zenodo.8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="42" pageNumber="43">Figs 19, 22</figureCitation>
, red arrow), under mounds of rocks, inside cavities on walls along road-cuts, at the bases of alive cacti such as “nopales” (
<taxonomicName id="B08D33915A4665230489FCF82824FC8F" box="[995,1149,828,854]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="45F994005A4665230489FCF8281AFC8F" box="[995,1091,828,854]" italics="true" pageId="42" pageNumber="43">Opuntia</emphasis>
spp.
</taxonomicName>
) and columnar cacti, and even under rotten cacti (
<figureCitation id="EFB654975A4665230562FC9B2EC9FCA0" box="[520,656,863,889]" captionStart="Figs 1924" captionStartId="31.[189,242,1775,1801]" captionTargetBox="[189,1398,268,1701]" captionTargetId="figure-11@31.[189,1398,268,1701]" captionTargetPageId="31" captionText="Figs 1924. Habitats and microhabitats of Latrodectus occidentalis Valdez-Mondragón sp. nov. Red arrow indicates the microhabitat where the specimens were collected. 19, 22. “Las Letras”, Pénjamo, Guanajuato, Mexico.20. Zona arqueológica “Plazuelas”, Pénjamo, Guanajuato, Mexico.21. Salvatierra, Guanajuato, Mexico. 2324. Cocula, Jalisco, Mexico (type locality) (red arrow in figure 23 shows the female found below a wasp nest). Photographs 1922 by Cabrera-Espinosa (2021); 2324 by Jared Lacayo (2021)." figureDoi="http://doi.org/10.5281/zenodo.8405896" httpUri="https://zenodo.org/record/8405896/files/figure.png" pageId="42" pageNumber="43">Figs 1921</figureCitation>
). Some females were collected with egg sacs, which are usually big, oval and yellow in coloration (
<figureCitation id="EFB654975A4665230538FC462EFEFC44" box="[594,679,898,925]" captionStart="Figs 1318" captionStartId="30.[189,242,1748,1774]" captionTargetBox="[189,1398,268,1700]" captionTargetId="figure-14@30.[189,1398,268,1700]" captionTargetPageId="30" captionText="Figs 1318. Live females (1315) and males (1618) of Latrodectus occidentalis Valdez-Mondragón sp. nov. 13, 16. Salvatierra, Guanajuato, Mexico. 14. Camichines, Cocula, Jalisco, Mexico (type locality). 15. Hostotipaquillo, Jalisco, Mexico. 17. 1 km North of San Nicolás de Ibarra, Jalisco, Mexico. 18. “Las Letras”, Pénjamo, Guanajuato, Mexico.Photographs 13, 16, 18 by Cabrera-Espinosa (2021). Photograph 14 by Navarro-Rogríguez I. (2021). Photograph 15 by Valdez-Mondragon A. Photograph 17 by Mamole in https://www.naturalista.mx/observations/126985294" figureDoi="http://doi.org/10.5281/zenodo.8405894" httpUri="https://zenodo.org/record/8405894/files/figure.png" pageId="42" pageNumber="43">Fig. 15</figureCitation>
), differing from other Mexican species (e.g.
<taxonomicName id="B08D33915A46652303C4FC46292EFC45" box="[1198,1399,898,925]" class="Arachnida" family="Theridiidae" genus="Latrodectus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="45F994005A46652303C4FC462960FC45" box="[1198,1337,898,924]" italics="true" pageId="42" pageNumber="43">Latrodectus</emphasis>
sp. 1
</taxonomicName>
and sp. 2) that have smaller egg sacs whitish or pale-yellow in coloration.
</paragraph>
</subSubSection>
<subSubSection id="3F971B995A46652307D7FC2328B8FBE9" box="[189,1249,999,1072]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="773248125A46652307D7FC232D0FFBD8" blockId="42.[189,342,999,1025]" box="[189,342,999,1025]" pageId="42" pageNumber="43">
<heading id="2C7AFF7E5A46652307D7FC232D0FFBD8" bold="true" box="[189,342,999,1025]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="45F994005A46652307D7FC232D0FFBD8" bold="true" box="[189,342,999,1025]" pageId="42" pageNumber="43">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="773248125A46652307D7FBD228B8FBE9" blockId="42.[189,1249,1045,1072]" box="[189,1249,1045,1072]" pageId="42" pageNumber="43">
<collectingCountry id="0F9A08825A46652307D7FBD22D6CFBE9" box="[189,309,1046,1072]" name="Mexico" pageId="42" pageNumber="43">MEXICO</collectingCountry>
:
<collectingRegion id="B54986F05A466523062AFBD12DCAFBF6" box="[320,403,1045,1071]" country="Mexico" name="Jalisco" pageId="42" pageNumber="43">Jalisco</collectingRegion>
,
<collectingRegion id="B54986F05A46652306F5FBD12DA1FBF6" box="[415,504,1045,1071]" country="Mexico" name="Colima" pageId="42" pageNumber="43">Colima</collectingRegion>
,
<collectingRegion id="B54986F05A466523056EFBD12ED0FBF6" box="[516,649,1045,1071]" country="Mexico" name="Michoacan de Ocampo" pageId="42" pageNumber="43">Michoacán</collectingRegion>
,
<collectingRegion id="B54986F05A46652305FFFBD22F5BFBE9" box="[661,770,1046,1072]" country="Mexico" name="Guerrero" pageId="42" pageNumber="43">Guerrero</collectingRegion>
,
<collectingRegion id="B54986F05A4665230467FBD22F31FBE9" box="[781,872,1046,1072]" country="Mexico" name="Oaxaca" pageId="42" pageNumber="43">Oaxaca</collectingRegion>
,
<collectingRegion id="B54986F05A466523041EFBD12F80FBF6" box="[884,985,1045,1071]" country="Mexico" name="Morelos" pageId="42" pageNumber="43">Morelos</collectingRegion>
,
<collectingRegion id="B54986F05A466523048EFBD12832FBF6" box="[996,1131,1045,1071]" country="Mexico" name="Guanajuato" pageId="42" pageNumber="43">Guanajuato</collectingRegion>
(
<figureCitation id="EFB654975A4665230316FBD1288AFBE9" box="[1148,1235,1045,1072]" captionStart="Figs 910. 9" captionStartId="28.[189,242,1857,1883]" captionTargetBox="[244,1346,283,1804]" captionTargetId="figure-13@28.[235,1353,265,1809]" captionTargetPageId="28" captionText="Figs 910. 9. Records and Species Distribution Model (SDM) (red) of L. mactans (Fabricius, 1775) in Mexico and the USA. 10. Records and SDM (yellow) of L. occidentalis Valdez-Mondragón sp. nov. in Mexico. Abbreviations: CHIS = Chiapas; COL = Colima; EMX = Edo. Mex; GRO = Guerrero; GTO = Guanajuato; JAL = Jalisco; MICH= Michoacán; MOR = Morelos; NAY = Nayarit; OAX = Oaxaca; PUE = Puebla; QRO = Querétaro." figureDoi="http://doi.org/10.5281/zenodo.8405890" httpUri="https://zenodo.org/record/8405890/files/figure.png" pageId="42" pageNumber="43">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>