treatments-xml/data/B8/7E/87/B87E87A5FFD0FFA60D48FA815D9F7F28.xml

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<mods:title id="65FCD2F95C1E7B7C04D446180A0246D4">Revision and phylogeny of the European species of the Eurytoma morio species group (Hymenoptera: Eurytomidae), parasitoids of bark and wood boring beetles</mods:title>
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<taxonomicName id="F7D74D30FFD0FFAC0D48FA815AB27C24" ID-CoL="3D7DP" authority="BOHEMAN, 1836" authorityName="Boheman" authorityYear="1836" box="[843,1224,1353,1375]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFD0FFAC0D48FA815A7C7C27" box="[843,1030,1353,1374]" italics="true" pageId="7" pageNumber="377">EURYTOMA AFRA</emphasis>
<bibRefCitation id="54464B42FFD0FFAC0A0CFA815AB27C24" author="Boheman CH" box="[1039,1224,1353,1375]" pageId="7" pageNumber="377" pagination="222 - 259" refId="ref21977" refString="Boheman CH. 1836. Skandinaviska Pteromaliner. Kongliga Vetenskaps-Akademiens Handlingar 1836: 222 - 259." type="journal article" year="1836">BOHEMAN, 1836</bibRefCitation>
</taxonomicName>
(
<figureCitation id="A8EC2A36FFD0FFAC0ADAFA815B1B7C24" box="[1241,1377,1353,1375]" captionStart="Figure 10" captionStartId="16.[164,243,799,818]" captionTargetBox="[323,1283,195,770]" captionTargetId="figure-430@16.[323,1283,195,770]" captionTargetPageId="16" captionText="Figure 10. A–D, Eurytoma afra: A–C, female; D, male. A, head in frontal view; B, head and pronotum in lateral view; C, head and mesosoma in dorsal view; D, antenna." figureDoi="http://doi.org/10.5281/zenodo.5306201" httpUri="https://zenodo.org/record/5306201/files/figure.png" pageId="7" pageNumber="377">FIG. 10A–D</figureCitation>
)
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<treatmentCitation id="B17610A2FFD0FFAC0D25FAB85B607CFF" author="Boheman CH" box="[806,1306,1391,1413]" page="243" pageId="7" pageNumber="377" year="1836">
<taxonomicName id="F7D74D30FFD0FFAC0D25FAB85B607CFF" ID-CoL="3D7DP" authority="Boheman, 1836: 242 - 243" authorityName="Boheman" authorityPageNumber="242 - 243" authorityYear="1836" box="[806,1306,1391,1413]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFD0FFAC0D25FAB85DA27CFF" box="[806,984,1391,1413]" italics="true" pageId="7" pageNumber="377">Eurytoma afra</emphasis>
<bibRefCitation id="54464B42FFD0FFAC0DE6FAA75B607CFF" author="Boheman CH" box="[997,1306,1391,1413]" pageId="7" pageNumber="377" pagination="222 - 259" refId="ref21977" refString="Boheman CH. 1836. Skandinaviska Pteromaliner. Kongliga Vetenskaps-Akademiens Handlingar 1836: 222 - 259." type="journal article" year="1836">Boheman, 1836: 242–243</bibRefCitation>
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. Original description. 
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. 
<treatmentCitation id="B17610A2FFD0FFAC0A17FA465A907CD8" author="Graham MWR de V." box="[1044,1258,1422,1444]" page="147" pageId="7" pageNumber="377" year="1970">
<bibRefCitation id="54464B42FFD0FFAC0A17FA465A907CD8" author="Graham MWR de V." box="[1044,1258,1422,1444]" pageId="7" pageNumber="377" pagination="125 - 152" refId="ref22855" refString="Graham MWR de V. 1970. Taxonomic notes on some western Palaearctic Eurytomidae (Hymenoptera: Chalcidoidea). Proceedings of the Royal Entomological Society of London (B) 39: 125 - 152." type="journal article" year="1970">Graham, 1970: 147</bibRefCitation>
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– revised synonymy, comparison with 
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<emphasis id="02A3EAA1FFD0FFAC0A4AFA655A217CB9" box="[1097,1115,1453,1474]" italics="true" pageId="7" pageNumber="377">E</emphasis>
. 
<emphasis id="02A3EAA1FFD0FFAC0A6FFA655AD47CB9" box="[1132,1198,1453,1474]" italics="true" pageId="7" pageNumber="377">morio</emphasis>
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.
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<paragraph id="306836B3FFD0FFAC0D3CFA035AB37F7B" blockId="7.[806,1422,1391,1903]" pageId="7" pageNumber="377">
<treatmentCitationGroup id="10C7119DFFD0FFAC0D3CFA035AB37F7B" pageId="7" pageNumber="377">
<treatmentCitation id="B17610A2FFD0FFAC0D3CFA035BF07C9B" author="Ashmead WH" box="[831,1418,1483,1505]" page="323" pageId="7" pageNumber="377" year="1894">
<taxonomicName id="F7D74D30FFD0FFAC0D3CFA035BF07C9B" authority="Ashmead, 1894: 322 - 323" authorityName="Ashmead" authorityPageNumber="322 - 323" authorityYear="1894" box="[831,1418,1483,1505]" class="Insecta" family="Eurytomidae" genus="Decatomidea" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFD0FFAC0D3CFA035A2D7C9B" box="[831,1111,1483,1504]" italics="true" pageId="7" pageNumber="377">Decatomidea polygraphi</emphasis>
<bibRefCitation id="54464B42FFD0FFAC0A61FA035BF07C9B" author="Ashmead WH" box="[1122,1418,1483,1505]" pageId="7" pageNumber="377" pagination="318 - 344" refId="ref21954" refString="Ashmead WH. 1894. Descriptions of new parasitic Hymenoptera. Transactions of the American Entomological Society 21: 318 - 344." type="journal article" year="1894">Ashmead, 1894: 322–323</bibRefCitation>
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. Original description. 
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, 
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.
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<treatmentCitationGroup id="10C7119DFFD0FFAC0D3CF9C15DE47F47" pageId="7" pageNumber="377">
= 
<taxonomicName id="F7D74D30FFD0FFAC0D56F9C15A8D7F65" ID-CoL="3D7DS" authority="Boheman, 1836" authorityName="Boheman" authorityYear="1836" box="[853,1271,1544,1566]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="02A3EAA1FFD0FFAC0D56F9C15A417F66" box="[853,1083,1544,1566]" italics="true" pageId="7" pageNumber="377">Eurytoma albimana</emphasis>
<bibRefCitation id="54464B42FFD0FFAC0A40F9C05A8D7F65" author="Boheman CH" box="[1091,1271,1544,1566]" pageId="7" pageNumber="377" pagination="222 - 259" refId="ref21977" refString="Boheman CH. 1836. Skandinaviska Pteromaliner. Kongliga Vetenskaps-Akademiens Handlingar 1836: 222 - 259." type="journal article" year="1836">Boheman, 1836</bibRefCitation>
</taxonomicName>
; 
<treatmentCitation id="B17610A2FFD0FFAC0B07F9C05DE07F47" author="Dalla Torre KW von" page="332" pageId="7" pageNumber="377" year="1898">
<bibRefCitation id="54464B42FFD0FFAC0B07F9C05DE07F47" author="Dalla Torre KW von" pageId="7" pageNumber="377" refId="ref22427" refString="Dalla Torre KW von. 1898. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. V. Chalcididae et Proctotrupidae. Lepzig, 598 pp." type="book" year="1898">Dalla Torre, 1898: 332</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="306836B3FFD0FFAC0D3CF98E5A4B7F01" blockId="7.[806,1422,1391,1903]" pageId="7" pageNumber="377">
<treatmentCitationGroup id="10C7119DFFD0FFAC0D3CF98E5A4B7F01" pageId="7" pageNumber="377">
<taxonomicName id="F7D74D30FFD0FFAC0D3CF98E5AA87F21" ID-CoL="3D82K" authority="Mayr, 1878" authorityName="Mayr" authorityYear="1878" box="[831,1234,1605,1627]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="saliciperdae">
<emphasis id="02A3EAA1FFD0FFAC0D3CF98E5A397F21" box="[831,1091,1605,1627]" italics="true" pageId="7" pageNumber="377">Eurytoma saliciperdae</emphasis>
<bibRefCitation id="54464B42FFD0FFAC0A4EF98E5AA87F21" author="Mayr GL" box="[1101,1234,1605,1627]" pageId="7" pageNumber="377" pagination="297 - 334" refId="ref23395" refString="Mayr GL. 1878. Arten der Chalcidier-Gattung Eurytoma durch Zucht erhalten. Verhandlungen der Kaiserlich Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 28: 297 - 334." type="journal article" year="1878">Mayr, 1878</bibRefCitation>
</taxonomicName>
; 
<treatmentCitation id="B17610A2FFD0FFAC0AE2F98D5D357F02" author="Ferriere C" page="381" pageId="7" pageNumber="377" year="1950">
<bibRefCitation id="54464B42FFD0FFAC0AE2F98D5D357F02" author="Ferriere C" pageId="7" pageNumber="377" pagination="377 - 410" refId="ref22575" refString="Ferriere C. 1950. Notes sur les Eurytoma (Hym., Chalcidoidea). I. les types de Thomson et de Mayr. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 23: 377 - 410." type="journal article" year="1950">Ferrière, 1950: 381</bibRefCitation>
</treatmentCitation>
(misidentification).
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</paragraph>
<paragraph id="306836B3FFD0FFAC0D3CF94A5B3E7FCC" blockId="7.[806,1422,1391,1903]" pageId="7" pageNumber="377">
<treatmentCitationGroup id="10C7119DFFD0FFAC0D3CF94A5B3E7FCC" pageId="7" pageNumber="377">
<taxonomicName id="F7D74D30FFD0FFAC0D3CF94A5AF27FE3" ID-CoL="3PVDD" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[831,1160,1666,1688]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFD0FFAC0D3CF94A5AF27FE3" box="[831,1160,1666,1688]" italics="true" pageId="7" pageNumber="377">Ipideurytoma spessivtsevi</emphasis>
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Boucˇek &amp; 
<treatmentCitation id="B17610A2FFD0FFAC0B20F94A5DAF7FCC" author="Novicky S" page="269" pageId="7" pageNumber="377" year="1954">
<bibRefCitation id="54464B42FFD0FFAC0B20F94A5DAF7FCC" author="Novicky S" pageId="7" pageNumber="377" pagination="266 - 271" refId="ref22101" refString="Boucˇek Z, Novicky S. 1954. Ipideurytoma spessivtsevi n. g., n. sp., ein neuer Borlenkaferparasit. Entomologisk Tidskrift 75: 266 - 271." type="journal article" year="1954">Novicky, 1954: 269–261</bibRefCitation>
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. Original description. 
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.
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</paragraph>
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Host: 
<taxonomicName id="F7D74D30FFD0FFAC0D25F9085B557FAD" authority="(Fabricius, 1787)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[806,1327,1728,1750]" class="Insecta" family="Curculionidae" genus="Trypodendron" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="signatum">
<emphasis id="02A3EAA1FFD0FFAC0D25F9085A347FAE" box="[806,1102,1728,1749]" italics="true" pageId="7" pageNumber="377">Trypodendron signatum</emphasis>
(Fabricius, 1787)
</taxonomicName>
(under 
<taxonomicName id="F7D74D30FFD0FFAC0D25F9165A797F88" authorityName="Erichson" authorityYear="1836" box="[806,1027,1758,1779]" class="Insecta" family="Curculionidae" genus="Xyloterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="02A3EAA1FFD0FFAC0D25F9165A797F88" box="[806,1027,1758,1779]" italics="true" pageId="7" pageNumber="377">Xyloterus lineatus</emphasis>
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). 
<treatmentCitation id="B17610A2FFD0FFAC0A22F9175A8E7F88" author="Zerova MD" box="[1057,1268,1758,1780]" page="32" pageId="7" pageNumber="377" year="1998">
<bibRefCitation id="54464B42FFD0FFAC0A22F9175A8E7F88" author="Zerova MD" box="[1057,1268,1758,1780]" pageId="7" pageNumber="377" pagination="32" refId="ref24564" refString="Zerova MD. 1998. [The identity of Eurytoma spessivtsevi (Boucek et Novicky, 1956) to Decatomoidea polygraphi (Ashmead, 1894) (Hymenoptera, Eurytomidae)]. Vestnik Zoologii, Kiev 32: 32. [in Russian]." type="journal article" year="1998">Zerova, 1998: 32</bibRefCitation>
</treatmentCitation>
: = 
<taxonomicName id="F7D74D30FFD0FFAC0B14F9175A537E68" authority="(Ashmead)" baseAuthorityName="Ashmead" baseAuthorityYear="1894" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFD0FFAC0B14F9175DDB7E69" italics="true" pageId="7" pageNumber="377">Eurytoma polygraphi</emphasis>
(Ashmead)
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.
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</subSubSection>
<subSubSection id="78CD6538FFD0FFAC0D3CF8D35B357E15" pageId="7" pageNumber="377" type="reference_group">
<paragraph id="306836B3FFD0FFAC0D3CF8D35DE47E2B" blockId="7.[806,1422,1391,1903]" pageId="7" pageNumber="377">
<treatmentCitationGroup id="10C7119DFFD0FFAC0D3CF8D35DE47E2B" pageId="7" pageNumber="377">
<taxonomicName id="F7D74D30FFD0FFAC0D3CF8D35B5F7E4A" authority="(Ashmead)" baseAuthorityName="Ashmead" box="[831,1317,1819,1841]" class="Insecta" genus="Ipideurtytoma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFD0FFAC0D3CF8D35D9B7E4B" box="[831,993,1819,1840]" italics="true" pageId="7" pageNumber="377">Ipideurtytoma</emphasis>
[
<emphasis id="02A3EAA1FFD0FFAC0DF0F8D45A6B7E4A" box="[1011,1041,1820,1841]" italics="true" pageId="7" pageNumber="377">sic</emphasis>
] 
<emphasis id="02A3EAA1FFD0FFAC0A20F8D35AE47E4B" box="[1059,1182,1819,1840]" italics="true" pageId="7" pageNumber="377">polygraphi</emphasis>
(Ashmead)
</taxonomicName>
: 
<treatmentCitation id="B17610A2FFD0FFAC0B31F8D35DE07E2B" author="Bugbee RE" page="506" pageId="7" pageNumber="377" year="1956">
<bibRefCitation id="54464B42FFD0FFAC0B31F8D35DE07E2B" author="Bugbee RE" pageId="7" pageNumber="377" pagination="503 - 506" refId="ref22171" refString="Bugbee RE. 1956. Synonymy, new combinations and nomina nuda in the genus Eurytoma Illiger (Chalcidoidea: Hymenoptera). Annals of the Entomological Society of America 49: 503 - 506." type="journal article" year="1956">Bugbee, 1956: 506</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="306836B3FFD0FFAC0D3CF8915B357E15" blockId="7.[806,1422,1391,1903]" box="[831,1359,1881,1903]" pageId="7" pageNumber="377">
= 
<taxonomicName id="F7D74D30FFD0FFAC0D56F8915AA57E14" authority="Ashmead" authorityName="Ashmead" authorityYear="1894" box="[853,1247,1881,1903]" class="Insecta" family="Eurytomidae" genus="Decatomidea" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="377" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFD0FFAC0D56F8915A167E15" box="[853,1132,1881,1902]" italics="true" pageId="7" pageNumber="377">Decatomidea polygraphi</emphasis>
Ashmead
</taxonomicName>
<taxonomicNameLabel id="199057DAFFD0FFAC0AEAF8915B357E15" box="[1257,1359,1881,1902]" pageId="7" pageNumber="377">syn. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="78CD6538FFDFFFA30F6EFF105AA27997" box="[365,1240,216,237]" pageId="8" pageNumber="378" type="nomenclature">
<paragraph id="306836B3FFDFFFA30F6EFF105AA27997" blockId="8.[181,1425,216,1886]" box="[365,1240,216,237]" pageId="8" pageNumber="378">
KEY TO EUROPEAN SPECIES OF 
<taxonomicName id="F7D74D30FFDFFFA30CD6FF105D2B7996" ID-CoL="4H2G" ID-ENA="75202" authority="Illiger, 1807" authorityName="Illiger" authorityYear="1807" box="[725,849,216,237]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="378" phylum="Arthropoda" rank="genus">EURYTOMA</taxonomicName>
BELONGING TO THE 
<emphasis id="02A3EAA1FFDFFFA30A3FFF135AFF7990" box="[1084,1157,219,235]" italics="true" pageId="8" pageNumber="378">MORIO</emphasis>
GROUP
</paragraph>
</subSubSection>
<subSubSection id="78CD6538FFDFFFA30EB6FF365F687868" box="[181,274,254,275]" pageId="8" pageNumber="378" type="multiple">
<paragraph id="306836B3FFDFFFA30EB6FF365F687868" blockId="8.[181,1425,216,1886]" box="[181,274,254,275]" pageId="8" pageNumber="378">
<emphasis id="02A3EAA1FFDFFFA30EB6FF365F687868" box="[181,274,254,275]" italics="true" pageId="8" pageNumber="378">Females</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="78CD6538FFDFFFA20EB6FEED5B017C60" lastPageId="9" lastPageNumber="379" pageId="8" pageNumber="378" type="key">
<key id="44466182FFDFFFA20EB6FEED5B017C60" lastPageId="9" lastPageNumber="379" pageId="8" pageNumber="378">
<keyStep id="8B232E16FFDFFFA30EB6FEED5BEA78BC" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FEED5BEA780A" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FEED5BEA780A" pageId="8" pageNumber="378">
1. Metacoxa densely setose dorsally from base (
<figureCitation id="A8EC2A36FFDFFFA30CE9FEED5D457843" box="[746,831,293,312]" captionStart="Figure 4" captionStartId="11.[144,223,1771,1790]" captionTargetBox="[197,1371,195,1740]" captionTargetId="figure-0@11.[197,1371,195,1740]" captionTargetPageId="11" captionText="Figure 4. A–O, Eurytoma laricis (D, F, H, I, J); Eurytoma morio (G, M, N); Eurytoma striolata (A, B, C, L, O). A, back of head; B, postgena; C, postoccipital region; D–E, clypeus; F–G, lower face; H, male scape; I, scape plate with pores (arrows); J, male antenna; K, mesoscutum; L, female propodeum; M, procoxa; N, metacoxa; O, female petiole. Abbreviations: ATP, anterior tentorial pits; GC, genal carina; LFP, lateral formaminal plate; PGG, postgenal groove, PGL, postgenal lamina; PTP, posterior tentorial pits." figureDoi="http://doi.org/10.5281/zenodo.6987032" httpUri="https://zenodo.org/record/6987032/files/figure.png" pageId="8" pageNumber="378">Figs 4N</figureCitation>
, 22D, 27F). Marginal vein somewhat to evidently thick- ened, 2.90–4.80 times as long as wide (Figs 11G, 22H). Lower surface of costal cell most often bearing numerous short setae, sometimes entirely setose or leaving a narrow posterior bare stripe (Figs 11F, 22G)..............2
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FEB35BEA78BC" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FEB35BEA78BC" pageId="8" pageNumber="378">1′. Metacoxa bare dorsally at base or marginal vein not thickened, at least 6.00 times as long as wide (Figs 12H, 13G, 25J, 30J). Setation of lower surface of costal cell as usual, not especially dense (Figs 16H, 25J, 30J)...............................................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FE185BEA7B42" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FE185BEA7B7B" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FE185BEA7B7B" pageId="8" pageNumber="378">2(1). Notauli appearing as complete impressed lines, not obliterated by the sculpture of the mesoscutum (Fig. 35A, C). Valvulae generally ascending only slightly (Figs 11H, 27H)...........................................................3</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FDC25BEA7B42" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FDC25BEA7B42" pageId="8" pageNumber="378">2′. Notauli hardly impressed and/or obliterated by the sculpture (Figs 14D, 22C). Valvulae evidently ascending (Figs 15A, 22I)...........................................................................................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FD8B5BEA7A79" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FD8B5BF57BD7" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FD8B5BF57BD7" pageId="8" pageNumber="378">
3(2). Body with following parts light: lateral panel of pronotum anteriorly, pro- and mesocoxae, acropleuron, edge of oral fossa. Marginal vein thick, 2.90 times as long as wide (Fig. 11G). Wing membrane with relatively broad diffuse infumation posterior to marginal vein. Median channel of propodeum with complete, mostly regular, median ridge (Fig. 11E)............................................................... 
<emphasis id="02A3EAA1FFDFFFA30A07FD515A827BD7" box="[1028,1272,665,684]" italics="true" pageId="8" pageNumber="378">Eurytoma aloisifilippoi</emphasis>
(Russo, 1938)
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FD7D5BEA7A79" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FD7D5BEA7A79" pageId="8" pageNumber="378">3′. Body with lateral panel of pronotum, pro- and mesocoxae, acropleuron, edge of oral fossa dark. Marginal vein less thick than in alternate, at least 3.50 times as long as wide (Fig. 29D, H). Median channel of propodeum without, or with partial, irregular and sinuous median ridge.............................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FCC35BF57AD6" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FCC35BF57A2C" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FCC35BF57A2C" pageId="8" pageNumber="378">
4(3). Marginal vein wider than in the alternative, 3.50–4.50 times as long as wide and widening distally (Fig. 29H). Lower surface of costal cell completely densely setose (Fig. 29F), the setae mostly white and relatively short (Fig. 29E, G)............................................................................... 
<emphasis id="02A3EAA1FFDFFFA30A0CFC8D5AA27A2C" box="[1039,1240,836,855]" italics="true" pageId="8" pageNumber="378">Eurytoma striolata</emphasis>
Ratzeburg, 1848
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FCA95BF57AD6" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FCA95BF57AD6" pageId="8" pageNumber="378">
4′. Marginal vein slender than in the alternative, about 4.85–5.00 times as long as wide, not widening distally (Fig. 29D). Lower surface of costal cell with a narrow posterior bare stripe (visible only when the wing is tilted; Fig. 29B), setation longer and mostly dark (Fig. 29C). ............. 
<emphasis id="02A3EAA1FFDFFFA30A01FC535A817AD6" box="[1026,1275,922,941]" italics="true" pageId="8" pageNumber="378">Eurytoma melanoneura</emphasis>
Walker, 1871
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FC7F5BF57D5B" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FC7F5BEA7A9C" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FC7F5BEA7A9C" pageId="8" pageNumber="378">
5(2′). Walls of punctures strongly raised on mesonotum, forming transverse crests on pronotum (Fig. 14D–F). Head long and globose, 1.65–1.70 times as wide as long (Fig. 14B) ........................ 
<emphasis id="02A3EAA1FFDFFFA30A5FFC1C5BEA7A9C" bold="true" box="[1116,1424,979,999]" pageId="8" pageNumber="378">
<emphasis id="02A3EAA1FFDFFFA30A5FFC1C5B497A9D" bold="true" box="[1116,1331,979,999]" italics="true" pageId="8" pageNumber="378">Eurytoma cristata</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FC385BF57D5B" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FC385BF57D5B" pageId="8" pageNumber="378">
5′. Walls of punctures not strongly raised, not forming crests on pronotum (Fig. 22C). Head relatively shorter, 1.75–2.0 times as wide as long (Fig. 22A). ............................................ 
<emphasis id="02A3EAA1FFDFFFA30A36FBC55A9B7D64" box="[1077,1249,1037,1055]" italics="true" pageId="8" pageNumber="378">Eurytoma morio</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FBE15BEA7C45" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FBE15BF57DD4" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FBE15BF57DD4" pageId="8" pageNumber="378">
6(1′). Frons forming blunt angle with vertex, which is in the same plan as the pronotal collar (Fig. 10B). Head transverse in frontal view, subtrapezoidal (Fig. 10A). Pronotum with large and well-delimited yellow spots, visible laterally (Fig. 10B). Pronotal collar with small but relatively deep punctures, separated by coriaceous interspaces (Fig. 10C). Axillar grooves narrow and superficial (Fig. 10C). Postgenal carina strongly raised, appearing triangular in lateral view (Fig. 10B) ........................................ 
<emphasis id="02A3EAA1FFDFFFA30A46FB545A987DD4" box="[1093,1250,1180,1199]" italics="true" pageId="8" pageNumber="378">Eurytoma afra</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FB705BEA7C45" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FB705BEA7C45" pageId="8" pageNumber="378">6′. Head rounded in frontal view (Fig. 21A) and/or not so transverse (Fig. 13A). Frons as usual, merging pro- gressively towards vertex (Fig. 18C). Lateral panel of pronotum generally completely black (Fig. 20D), some- times with diffuse yellowish spot. Pronotal collar generally densely punctured, the points separated by narrow interspaces (Figs 13D, 18F), or punctures shallow (Fig. 24D). Axillar grooves deep except sometimes anteri- orly. Postgenal carina not so raised (Fig. 18C).................................................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6FA8F5BF57F58" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6FA8F5BF57CCB" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FA8F5BF57CCB" pageId="8" pageNumber="378">
7(6′). Pronotum finely and sparsely punctured, the points rounded, interspaces distinctly coriaceous (Fig. 5G). Lower face entirely strigose, with radiating crests reaching eyes and antennal toruli (Fig. 21A). Head long and globose, 1.60 times as wide as long when seen in dorsal view, with very convex frons (Fig. 21B). Parasitoids of bark beetles .......................................................................................... 
<emphasis id="02A3EAA1FFDFFFA30A28FA565A987CCB" box="[1067,1250,1438,1456]" italics="true" pageId="8" pageNumber="378">Eurytoma maura</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6FA725BF57F58" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6FA725BF57F58" pageId="8" pageNumber="378">7′. Pronotum densely areolate, punctured with narrow interspaces (Figs 13D, 18F). Lower face most often partly punctured and showing a supraclypeal smooth stripe (Figs 25D, 32B). Head often more transverse than in alternate, 1.70–2.00 times as wide as long (Figs 12A, 13B, 18B, 30B). Hosts sometimes different or species seed eaters................................................................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA30EB6F9E45BF57FB5" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6F9E45BEA7F02" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6F9E45BEA7F02" pageId="8" pageNumber="378">
8(7′). Small size species (2.40–2.90 mm) with elongate gaster (Fig. 24F, 25K). At least procoxa (as in Fig. 26A) and ovipositor reddish, the former sometimes brownish but never black. Species associated with gall midges of the genus 
<emphasis id="02A3EAA1FFDFFFA30F83F9AD5C687F03" box="[384,530,1637,1656]" italics="true" pageId="8" pageNumber="378">Rhabdophaga</emphasis>
on 
<emphasis id="02A3EAA1FFDFFFA30C3DF9AD5C0E7F03" box="[574,628,1637,1656]" italics="true" pageId="8" pageNumber="378">Salix</emphasis>
spp.............................................................................................9
</keyLead>
</paragraph>
<paragraph id="306836B3FFDFFFA30EB6F94A5BF57FB5" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6F94A5BF57FB5" pageId="8" pageNumber="378">8′. Species of larger size (more than 3.0 mm) and/or with relatively less elongate gaster (Figs 12G, 18K, 30K). Procoxa black, ovipositor dark, at most lighter at apex. Parasitoids of bark beetles or other xylophagous Coleoptera within seeds of pines or seed eaters on coniferous trees .................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDFFFA20EB6F9105B067856" lastPageId="9" lastPageNumber="379" pageId="8" pageNumber="378">
<paragraph id="306836B3FFDFFFA30EB6F9105BF57E26" blockId="8.[181,1425,216,1886]" pageId="8" pageNumber="378">
<keyLead id="8B269586FFDFFFA30EB6F9105BF57E26" pageId="8" pageNumber="378">
9(8). Puncturation on pro- and mesonotum superficial with coriaceous interspaces (Fig. 24D). Head long and globose, 1.55 times as wide as long, with very convex frons (Fig. 24B). Lower face almost smooth, showing only ra- diating striolae. Vertex and upper frons coriaceous (Fig. 24C). Lateral prepectus not delimited from sublateral, the reticulate sculpture continuing there (Fig. 24E). Femora and tibiae more extensively fuscous .............. ......................................................................................................... 
<emphasis id="02A3EAA1FFDFFFA30A59F8835B747E26" box="[1114,1294,1866,1885]" italics="true" pageId="8" pageNumber="378">Eurytoma nobbei</emphasis>
Mayr, 1878
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FF0E5B067856" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FF0E5B067856" pageId="9" pageNumber="379">
9′. Puncturation on mesonotum prominent, not superficial (Fig. 25F). Head shorter than in alternate, 1.75 times as wide as long (Fig. 25A). Lower face mostly punctured with short radiating crests (Fig. 25D). Lateral prepectus delimited ventrally by a distinct ridge (as in Fig. 26A). Legs mostly reddish, except metacoxa and small fuscous spots on meso- and metafemora............................................................. 
<emphasis id="02A3EAA1FFDEFFA20A39FED25B067856" bold="true" box="[1082,1404,282,301]" pageId="9" pageNumber="379">
<emphasis id="02A3EAA1FFDEFFA20A39FED25B657856" bold="true" box="[1082,1311,282,301]" italics="true" pageId="9" pageNumber="379">Eurytoma saliphila</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FEFE5B0178E6" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FEFE5B06781E" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FEFE5B06781E" pageId="9" pageNumber="379">10(8′). Gaster relatively short and not laterally compressed, at most as long as mesosoma when excluding the hori- zontal ovipositor (Figs 30K, 16K).................................................................................................11</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FEA65B0178E6" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FEA65B0178E6" pageId="9" pageNumber="379">10′. Gaster distinctly longer than mesosoma and laterally compressed, with ovipositor ascending (Figs 12G, 13H, 18K, 19F, 20G, 31A)..................................................................................................................12</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FE6E5B067B3E" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FE6E5B06788A" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FE6E5B06788A" pageId="9" pageNumber="379">
11(10). Metacoxa sparsely setose dorsally (Fig. 30I). Ovipositor completely reddish (Fig. 30K). Notauli broad and deeply impressed (Fig. 30F). Clypeus distinctly emarginate on ventral margin (Fig. 30A). Head less transverse than in alternate when seen in frontal view, somewhat roundish (Fig. 30A) ............. 
<emphasis id="02A3EAA1FFDEFFA20A56FE165B06788A" bold="true" box="[1109,1404,477,497]" pageId="9" pageNumber="379">
<emphasis id="02A3EAA1FFDEFFA20A56FE165B5A788B" bold="true" box="[1109,1312,477,497]" italics="true" pageId="9" pageNumber="379">Eurytoma sylviae</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FE325B067B3E" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FE325B067B3E" pageId="9" pageNumber="379">
11′. Metacoxa bare dorsally. Ovipositor light only at apex (Fig. 16K). Notauli shallow (Fig. 16F). Clypeus hardly emarginate (Fig. 16A). Head more transverse in frontal view, subtrapezoidal (Fig. 16A). Nearctic species intro- duced in Europe, seed eater on 
<emphasis id="02A3EAA1FFDEFFA20C51FDF95D457B3F" box="[594,831,561,580]" italics="true" pageId="9" pageNumber="379">Pseudotsuga menziesii</emphasis>
.................................... 
<emphasis id="02A3EAA1FFDEFFA20A5CFDFA5B067B3E" bold="true" box="[1119,1404,561,581]" pageId="9" pageNumber="379">
<emphasis id="02A3EAA1FFDEFFA20A5CFDFA5B5A7B3F" bold="true" box="[1119,1312,561,581]" italics="true" pageId="9" pageNumber="379">Eurytoma gatesi</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FD855B067BAB" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FD855B067BE3" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FD855B067BE3" pageId="9" pageNumber="379">
12(10′). Anterior outline of mesepisternum with blunt angle in front of mesocoxae, delimiting anteriorly an apparent ventral shelf when seen in lateral view (Figs 18G, 31B). Lower face entirely strigose, the radiating crests reaching eyes and antennal toruli (Fig. 18A)................................................. 
<emphasis id="02A3EAA1FFDEFFA20A5DFD4D5B067BE3" bold="true" box="[1118,1404,645,664]" pageId="9" pageNumber="379">
<emphasis id="02A3EAA1FFDEFFA20A5DFD4D5B5A7BE3" bold="true" box="[1118,1312,645,664]" italics="true" pageId="9" pageNumber="379">Eurytoma ithma</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FD695B067BAB" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FD695B067BAB" pageId="9" pageNumber="379">12′. Mesepisternum with flat or hardly convex anterior outline (Figs 12F, 13E, 19D, 20D). Lower face with median smooth stripe, crests short (Fig. 13A)........................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FD115B017A03" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FD115B067A5F" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FD115B067A5F" pageId="9" pageNumber="379">13(12′). Gaster elongate, at least 1.90 times as long as third gastral tergite height (Figs 12G, 13H, 19F). Ovipositor moderately ascending. All funicular segments clearly longer than wide (Figs 12B, 13C, 19C). Clypeus and subantennal stripe smooth, lower face otherwise punctured (Fig. 13A) ............................................... 14</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FCE55B017A03" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FCE55B017A03" pageId="9" pageNumber="379">
13′. Gaster less elongate, 1.60 times as long as third gastral tergite height, with ovipositor strongly ascending (Fig. 20G). Funicular segment shorter, F4–F5 hardly longer than wide (Fig. 20B). Lower face punctured strigose (Fig. 32B, C). Larvae phytophagous in seeds of 
<emphasis id="02A3EAA1FFDEFFA20CF8FCAD5D4E7A0C" box="[763,820,869,887]" italics="true" pageId="9" pageNumber="379">Larix</emphasis>
spp.............................. 
<emphasis id="02A3EAA1FFDEFFA20A48FCAD5A847A03" box="[1099,1278,869,888]" italics="true" pageId="9" pageNumber="379">Eurytoma laricis</emphasis>
Yano, 1918
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FC495B067D40" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FC495B017A9C" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FC495B017A9C" pageId="9" pageNumber="379">
14(13). Vertex with coriaceous transverse stripe between lateral ocelli (Fig. 19B). Head globose in dorsal view, 1.77 times as wide as long, the outline of frons strongly convex (Fig. 19A). Outline of genae convex when seen in frontal view. Scape entirely black on holotype. Reared together with xylophagous beetles......................... ................................................................................................... 
<emphasis id="02A3EAA1FFDEFFA20A13FC1D5AA97A9C" box="[1040,1235,980,999]" italics="true" pageId="9" pageNumber="379">Eurytoma kangasi</emphasis>
Hedqvist, 1966
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FC385B067D40" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FC385B067D40" pageId="9" pageNumber="379">14′. Vertex entirely punctured (Figs 12A, 13B). Head more transverse in dorsal view, 1.85–1.95 times as wide as long (Figs 12A, 13B), with outline of frons less convex. Outline of genae straight or almost so (Fig. 13A). Scape at least with base partly reddish–testaceous (Fig. 13C) ................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2FB8C5B017C60" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FB8C5B017DD0" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FB8C5B017DD0" pageId="9" pageNumber="379">
15(14′). Notauli broad but hardly impressed, obliterated by sculpture and appearing crenulate (Fig. 12C, D). Temples angulate with occiput (Fig. 12A). Gaster shorter than in alternate, 1.90 times as long as height of third gastral tergite (Fig. 12G). Cleptoparasite of 
<emphasis id="02A3EAA1FFDEFFA20C83FBB45CA57DF4" box="[640,735,1148,1167]" italics="true" pageId="9" pageNumber="379">Scambus</emphasis>
upon 
<emphasis id="02A3EAA1FFDEFFA20D26FBB45A7C7DF4" box="[805,1030,1148,1167]" italics="true" pageId="9" pageNumber="379">Pissodes validirostris</emphasis>
within cones of 
<emphasis id="02A3EAA1FFDEFFA20AB9FBB55A8C7DF4" box="[1210,1270,1149,1167]" italics="true" pageId="9" pageNumber="379">Pinus</emphasis>
spp........... ................................................................................................... 
<emphasis id="02A3EAA1FFDEFFA20A15FB515AA87DD0" box="[1046,1234,1176,1195]" italics="true" pageId="9" pageNumber="379">Eurytoma annilai</emphasis>
Hedqvist, 1974
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FB7C5B017C60" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FB7C5B017C60" pageId="9" pageNumber="379">
15′. Notauli narrower than in alternate, appearing as impressed lines, not or little obliterated by sculpture (Fig. 13D). Temples merging with occiput (Fig. 13B). Gaster longer than in alternate, 2.20 times as long as height of third gastral tergite (Fig. 13H). Parasitoid of bark beetles on coniferous trees........................................ .................................................................................................... 
<emphasis id="02A3EAA1FFDEFFA20A1AFAC05AAB7C61" box="[1049,1233,1288,1306]" italics="true" pageId="9" pageNumber="379">Eurytoma arctica</emphasis>
Thomson, 1876
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="78CD6538FFDEFFA20EA2FAFB5F477C33" box="[161,317,1331,1352]" pageId="9" pageNumber="379" type="multiple">
<paragraph id="306836B3FFDEFFA20EA2FAFB5F477C33" blockId="9.[161,1405,197,1875]" box="[161,317,1331,1352]" pageId="9" pageNumber="379">
<emphasis id="02A3EAA1FFDEFFA20EA2FAFB5F477C33" box="[161,317,1331,1352]" italics="true" pageId="9" pageNumber="379">Known males</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="78CD6538FFDEFFA10EA2FA925BEA7CDC" lastPageId="10" lastPageNumber="380" pageId="9" pageNumber="379" type="key">
<key id="44466182FFDEFFA10EA2FA925BEA7CDC" lastPageId="10" lastPageNumber="380" pageId="9" pageNumber="379">
<keyStep id="8B232E16FFDEFFA20EA2FA925B017F63" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2FA925B067CB8" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FA925B067CB8" pageId="9" pageNumber="379">
1. Metacoxa densely setose dorsally from base (as in 
<figureCitation id="A8EC2A36FFDEFFA20D15FA925D1C7C16" box="[790,870,1370,1389]" captionStart="Figure 4" captionStartId="11.[144,223,1771,1790]" captionTargetBox="[197,1371,195,1740]" captionTargetId="figure-0@11.[197,1371,195,1740]" captionTargetPageId="11" captionText="Figure 4. A–O, Eurytoma laricis (D, F, H, I, J); Eurytoma morio (G, M, N); Eurytoma striolata (A, B, C, L, O). A, back of head; B, postgena; C, postoccipital region; D–E, clypeus; F–G, lower face; H, male scape; I, scape plate with pores (arrows); J, male antenna; K, mesoscutum; L, female propodeum; M, procoxa; N, metacoxa; O, female petiole. Abbreviations: ATP, anterior tentorial pits; GC, genal carina; LFP, lateral formaminal plate; PGG, postgenal groove, PGL, postgenal lamina; PTP, posterior tentorial pits." figureDoi="http://doi.org/10.5281/zenodo.6987032" httpUri="https://zenodo.org/record/6987032/files/figure.png" pageId="9" pageNumber="379">Fig. 4N</figureCitation>
). Marginal vein somewhat to evidently thickened, 2.90–4.80 times as long as wide (as in Figs 22H, 29D, H). Lower surface of costal cell densely setose (as in Figs 22G, 29A, E). Clypeus emarginate on ventral margin (Fig. 33C, F) or pronotum with cristate sculpture (as in Fig. 14D–F) ...................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2FA045B017F63" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2FA045B017F63" pageId="9" pageNumber="379">1′. Metacoxa bare dorsally at base or marginal vein not thickened, at least 6.0 times as long as wide (as in Figs 12H, 13G, 18J, 25J). Setation of lower surface of costal cell as usual, not especially dense (as in Fig 16H). Clypeus often hardly emarginate (as in Fig. 32C).............................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2F9EA5B067F15" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2F9EA5B017F29" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F9EA5B017F29" pageId="9" pageNumber="379">2(1). Notauli appearing as complete impressed lines, not obliterated by the sculpture of the mesoscutum (as in Fig. 35A)...................................................................................................................................3</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2F9935B067F15" blockId="9.[161,1405,197,1875]" box="[161,1404,1626,1647]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F9935B067F15" box="[161,1404,1626,1647]" pageId="9" pageNumber="379">2′. Notauli hardly impressed and/or obliterated by sculpture (as in Figs 14D, 22C) .................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2F9B05B017F9A" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2F9B05B017FD3" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F9B05B017FD3" pageId="9" pageNumber="379">3(2). Marginal vein shorter and distinctly widening distally, 3.50–4.50 times as long as wide (as in Figs 11G, 29H)...............................................................................................................................................4</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2F9795B017F9A" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F9795B017F9A" pageId="9" pageNumber="379">
3′. Marginal vein less thick than in alternate, 4.85 times as long as wide, and not widened distally (as in Fig. 29D) .................................................................................. 
<emphasis id="02A3EAA1FFDEFFA20DEDF9065A9D7F9A" box="[1006,1255,1742,1761]" italics="true" pageId="9" pageNumber="379">Eurytoma melanoneura</emphasis>
Walker, 1871
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDEFFA20EA2F9225B017E28" pageId="9" pageNumber="379">
<paragraph id="306836B3FFDEFFA20EA2F9225B017E61" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F9225B017E61" pageId="9" pageNumber="379">
4(3). Marginal vein quite wide, 2.90 times as long as wide (as in Fig. 11G). Wing membrane with diffuse infumation below marginal vein ................................................................... 
<emphasis id="02A3EAA1FFDEFFA20DF3F8CF5A9E7E61" box="[1008,1252,1799,1818]" italics="true" pageId="9" pageNumber="379">Eurytoma aloisifilippoi</emphasis>
(Russo, 1938)
</keyLead>
</paragraph>
<paragraph id="306836B3FFDEFFA20EA2F8EC5B017E28" blockId="9.[161,1405,197,1875]" pageId="9" pageNumber="379">
<keyLead id="8B269586FFDEFFA20EA2F8EC5B017E28" pageId="9" pageNumber="379">
4′. Marginal vein less wide, 3.50–4.50 times as long as wide (as in Fig. 29H). Wing membrane without infumation below marginal vein..................................................................... 
<emphasis id="02A3EAA1FFDEFFA20DF8F8895ABE7E28" box="[1019,1220,1856,1875]" italics="true" pageId="9" pageNumber="379">Eurytoma striolata</emphasis>
Ratzeburg, 1848
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FF0E5BF578FA" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FF0E5BEA786A" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FF0E5BEA786A" pageId="10" pageNumber="380">
5(2′). Walls of punctures strongly raised on mesonotum, forming transverse crests on pronotum (as in Fig. 14E, F). Scape with large reticulate surface on inner side (Fig. 14I). Dark species: head, mesosoma, scape, legs, except trochanters and knees, dark to black........................................................ 
<emphasis id="02A3EAA1FFDDFFA10A5FFF365BEA786A" bold="true" box="[1116,1424,254,273]" pageId="10" pageNumber="380">
<emphasis id="02A3EAA1FFDDFFA10A5FFF365B49786A" bold="true" box="[1116,1331,254,273]" italics="true" pageId="10" pageNumber="380">Eurytoma cristata</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FED25BF578FA" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FED25BF578FA" pageId="10" pageNumber="380">
5′. Walls of punctures raised as usual on mesonotum, not forming crests on pronotum (as in Fig. 22C). Scape with a small and smooth ventral swelling (Fig. 23D). Species extremely variable in coloration: sometimes head, mesosoma, petiole, and legs entirely yellow (Fig. 23A, B); in the darkest forms at least part of scape, a spot on lateral panel of pronotum, and procoxae yellow................................. 
<emphasis id="02A3EAA1FFDDFFA10A36FEA65A9B78FB" box="[1077,1249,366,384]" italics="true" pageId="10" pageNumber="380">Eurytoma morio</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FE425BEA7B76" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FE425BEA78AE" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FE425BEA78AE" pageId="10" pageNumber="380">
6(1′). Adscrobal carina raised ventrally on mesepisternum and elbowed, delimiting a ventral shelf when seen in lateral view (Fig. 31B). Scape with long but hardly protruding ventral swelling (Fig. 18L)........................ ............................................................................................................. 
<emphasis id="02A3EAA1FFDDFFA10A71FE0A5BEA78AE" bold="true" box="[1138,1424,449,469]" pageId="10" pageNumber="380">
<emphasis id="02A3EAA1FFDDFFA10A71FE0A5B4978AF" bold="true" box="[1138,1331,449,469]" italics="true" pageId="10" pageNumber="380">Eurytoma ithma</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FE165BEA7B76" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FE165BEA7B76" pageId="10" pageNumber="380">6′. Adscrobal carina not raised and mesepisternum rounded anteriorly in front of mesocoxae (as in Figs 12F, 13E, 19D, 20D). Scape either with distinct ventral swelling or with very short smooth area....................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FDDD5BEA7BE3" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FDDD5BEA7B1B" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FDDD5BEA7B1B" pageId="10" pageNumber="380">7(6′). Scape with a small apico-ventral swelling (Figs 10D, 21H). Pronotum finely and sparsely punctured, the points rounded and interspaces distinctly coriaceous (as in Fig. 5G). Lower face entirely strigose, the radiating crests long, reaching antennal toruli and lower margin of eyes (as in Figs 10A, 21A)......................................8</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FDA25BEA7BE3" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FDA25BEA7BE3" pageId="10" pageNumber="380">7′. Scape with a large ventral swelling, either contrasting in colour with the rest of the surface or strongly promi- nent (Figs 13J, 20I, 34C, 34F) ...................................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FD695BF57A5F" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FD695BF57B97" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FD695BF57B97" pageId="10" pageNumber="380">
8(7). Head transverse in dorsal and frontal views (as in Fig. 10A). Spots on pronotum extended on lateral panels (as in Fig. 10B). Swelling (= sensillar plate bearing pores visible through integument) of scape subcircular, relatively protruding (Fig. 10D)............................................................. 
<emphasis id="02A3EAA1FFDDFFA10A46FD125A987B97" box="[1093,1250,729,748]" italics="true" pageId="10" pageNumber="380">Eurytoma afra</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FD3D5BF57A5F" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FD3D5BF57A5F" pageId="10" pageNumber="380">
8′. Head roundish in frontal view, long and globose in dorsal view (as in Fig. 21A, B). Sensillar plate of scape hardly protruding (Fig. 21H) ............................................................. 
<emphasis id="02A3EAA1FFDDFFA10A28FCD95A987A58" box="[1067,1250,785,803]" italics="true" pageId="10" pageNumber="380">Eurytoma maura</emphasis>
Boheman, 1836
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FCE55BEA7AB7" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FCE55BEA7A03" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FCE55BEA7A03" pageId="10" pageNumber="380">
9(7′). Scape reddish with dark, not very prominent swelling (Fig. 26C). F5 symmetric and without apical peduncle (Fig. 26D). Legs reddish, except meso- and metacoxae and the relevant femora, partly fuscus (Fig. 26A). Species associated with gall midges of the genus 
<emphasis id="02A3EAA1FFDDFFA10CC8FCAD5D277A03" box="[715,861,869,888]" italics="true" pageId="10" pageNumber="380">Rhabdophaga</emphasis>
on 
<emphasis id="02A3EAA1FFDDFFA10D8AFCAD5DC57A03" box="[905,959,869,888]" italics="true" pageId="10" pageNumber="380">Salix</emphasis>
spp. ........... 
<emphasis id="02A3EAA1FFDDFFA10A4DFCAD5BEA7A03" bold="true" box="[1102,1424,868,888]" pageId="10" pageNumber="380">
<emphasis id="02A3EAA1FFDDFFA10A4DFCAD5B497A0C" bold="true" box="[1102,1331,868,888]" italics="true" pageId="10" pageNumber="380">Eurytoma saliphila</emphasis>
sp. nov.
</emphasis>
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FC495BEA7AB7" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FC495BEA7AB7" pageId="10" pageNumber="380">9′. Scape black or nearly so, and quite prominent ventrally (Figs 13J, 20I, 17B, 34C, F). F5 generally strongly asymmetric and with apical peduncle (Figs 13I, 20H, 17A, 34A, 34D). Tibiae and procoxa dark. Parasitoids of seed eaters associated with coniferous trees...............................................................................10</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FC1C5BF57D08" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FC1C5BF57D64" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FC1C5BF57D64" pageId="10" pageNumber="380">10(9′). All funicular segments strongly asymmetric, nodiform, and with an apical peduncle (Figs 17A, 20H, 34A, D). Scape completely black (Figs 17B, 20I). Larvae seed eaters or parasitoids of seed eaters on coniferous cones .............................................................................................................................................11</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FBE05BF57D08" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FBE05BF57D08" pageId="10" pageNumber="380">
10′. F5 symmetric and with very short apical peduncle, other segments elongate and only fairly nodiform (Fig. 13I). Base of scape pale (Fig. 13J). Parasitoid of bark beetles on coniferous trees........................................... .................................................................................................... 
<emphasis id="02A3EAA1FFDDFFA10A2FFBA95A9F7D08" box="[1068,1253,1121,1139]" italics="true" pageId="10" pageNumber="380">Eurytoma arctica</emphasis>
Thomson, 1876
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FBB45BF57D84" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FBB45BF57DBC" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FBB45BF57DBC" pageId="10" pageNumber="380">
11(10). Pilosity of forewing white (Fig. 12H). Propodeum with deep and crenulate median channel, the median crest short and irregular. Cleptoparasite of 
<emphasis id="02A3EAA1FFDDFFA10CABFB505D7D7DD0" box="[680,775,1176,1195]" italics="true" pageId="10" pageNumber="380">Scambus</emphasis>
upon 
<emphasis id="02A3EAA1FFDDFFA10D4EFB505A547DD0" box="[845,1070,1176,1195]" italics="true" pageId="10" pageNumber="380">Pissodes validirostris</emphasis>
within cones of 
<emphasis id="02A3EAA1FFDDFFA10AE1FB505B647DD1" box="[1250,1310,1176,1194]" italics="true" pageId="10" pageNumber="380">Pinus</emphasis>
spp......... ................................................................................................... 
<emphasis id="02A3EAA1FFDDFFA10A2AFB7C5A9C7DBC" box="[1065,1254,1204,1223]" italics="true" pageId="10" pageNumber="380">Eurytoma annilai</emphasis>
Hedqvist, 1974
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FB185BF57D84" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FB185BF57D84" pageId="10" pageNumber="380">11′. Pilosity of forewing entirely dark. Propodeum sometimes with superficial median channel and complete median crest (Fig. 20J). Larvae seed eaters..............................................................................................12</keyLead>
</paragraph>
</keyStep>
<keyStep id="8B232E16FFDDFFA10EB6FAC05BEA7CDC" pageId="10" pageNumber="380">
<paragraph id="306836B3FFDDFFA10EB6FAC05BF57C28" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FAC05BF57C28" pageId="10" pageNumber="380">
12(11′). Median channel of propodeum superficial, but with complete median crest (Fig. 20J). Scape abruptly nar- rowing at apex on ventral margin (Figs 20I, 34C). Larvae phytophagous in seeds of 
<emphasis id="02A3EAA1FFDDFFA10A9AFAEC5AA87C4D" box="[1177,1234,1316,1334]" italics="true" pageId="10" pageNumber="380">Larix</emphasis>
spp. ................. .......................................................................................................... 
<emphasis id="02A3EAA1FFDDFFA10A5CFA885B687C28" box="[1119,1298,1344,1363]" italics="true" pageId="10" pageNumber="380">Eurytoma laricis</emphasis>
Yano, 1918
</keyLead>
</paragraph>
<paragraph id="306836B3FFDDFFA10EB6FA945BEA7CDC" blockId="10.[181,1425,197,1447]" pageId="10" pageNumber="380">
<keyLead id="8B269586FFDDFFA10EB6FA945BEA7CDC" pageId="10" pageNumber="380">
12′. Propodeum with deep and crenulate median channel, the median crest short and irregular (Fig. 17C). Scape progressively narrowing at apex on ventral margin (Fig. 17B). Nearctic species introduced in Europe, phytophagous in seeds of 
<emphasis id="02A3EAA1FFDDFFA10F8AFA5B5C0C7CDD" box="[393,630,1427,1446]" italics="true" pageId="10" pageNumber="380">Pseudotsuga menziesii</emphasis>
............................................................... 
<emphasis id="02A3EAA1FFDDFFA10A70FA5C5B4E7CDD" bold="true" box="[1139,1332,1427,1447]" italics="true" pageId="10" pageNumber="380">Eurytoma gatesi</emphasis>
sp. nov.
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="78CD6538FFDDFFA60EBFFA365F857FCE" lastPageId="13" lastPageNumber="383" pageId="10" pageNumber="380" type="description">
<paragraph id="306836B3FFDDFFA10EBFFA365F7D7F49" blockId="10.[164,778,1533,1586]" pageId="10" pageNumber="380">
= 
<taxonomicName id="F7D74D30FFDDFFA10ED7FA355C697F68" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[212,531,1533,1555]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFDDFFA10ED7FA355C697F68" box="[212,531,1533,1555]" italics="true" pageId="10" pageNumber="380">Ipideurytoma spessivtsevi</emphasis>
</taxonomicName>
Boucˇek &amp; Novicky 
<taxonomicNameLabel id="199057DAFFDDFFA10EA7F9D55F7D7F49" box="[164,263,1565,1586]" pageId="10" pageNumber="380">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="306836B3FFDDFFA10EA7F9AD5F3B7F02" blockId="10.[164,779,1636,1903]" box="[164,321,1636,1658]" pageId="10" pageNumber="380">
<emphasis id="02A3EAA1FFDDFFA10EA7F9AD5F3B7F02" box="[164,321,1636,1658]" italics="true" pageId="10" pageNumber="380">Type material</emphasis>
</paragraph>
<paragraph id="306836B3FFDDFFA10EA7F94B5BDB7F2B" blockId="10.[164,779,1636,1903]" lastBlockId="10.[826,1441,1533,1617]" pageId="10" pageNumber="380">
<taxonomicName id="F7D74D30FFDDFFA10EA7F94B5F317FE3" authorityName="Boheman" authorityYear="1836" box="[164,331,1667,1688]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDDFFA10EA7F94B5F317FE3" box="[164,331,1667,1688]" italics="true" pageId="10" pageNumber="380">Eurytoma afra</emphasis>
</taxonomicName>
. 
<typeStatus id="EF6C8811FFDDFFA10F59F94B5FB27FE3" box="[346,456,1667,1688]" pageId="10" pageNumber="380" type="lectotype">Lectotype</typeStatus>
designated by Claridge and here validated, with the following labelling ‘afra m 
<specimenCount id="26D1FD3AFFDDFFA10CF1F9695D717FCC" box="[754,779,1697,1719]" pageId="10" pageNumber="380" type="male">♂</specimenCount>
flavimana 93’ (manuscript)/(grey label without writing)/ ‘498 Sm’ (manuscript, for Smoland) (red label)/NRM 
<collectingRegion id="F213F851FFDDFFA10EA7F9355F667E68" box="[164,284,1789,1811]" country="Sweden" name="Stockholm" pageId="10" pageNumber="380">Stockholm</collectingRegion>
(in NHRC). 
<taxonomicName id="F7D74D30FFDDFFA10FACF9355CBF7E69" authorityName="Ashmead" authorityYear="1894" box="[431,709,1789,1810]" class="Insecta" family="Eurytomidae" genus="Decatomidea" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFDDFFA10FACF9355CBF7E69" box="[431,709,1789,1810]" italics="true" pageId="10" pageNumber="380">Decatomidea polygraphi</emphasis>
</taxonomicName>
. Type female labelled ‘Morgantown W Va’ (manuscript)/‘type’/ ‘4114’/‘ex 
<taxonomicName id="F7D74D30FFDDFFA10F09F8F25C737E2B" authorityName="Wood &amp; Bright" authorityYear="1992" box="[266,521,1850,1872]" class="Insecta" family="Curculionidae" genus="Polygraphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="rufipennis">Polygraphus rufipennis</taxonomicName>
on spruce’/‘type USNM 2176’ (in USNM). 
<taxonomicName id="F7D74D30FFDDFFA10F72F8915CE97E15" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[369,659,1881,1902]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFDDFFA10F72F8915CE97E15" box="[369,659,1881,1902]" italics="true" pageId="10" pageNumber="380">Ipideurytoma spessivtsevi</emphasis>
</taxonomicName>
. Type ma- terial not examined but the original description leaves no doubt about the identity of the species, which is so distinctive as to be described in a separate genus.
</paragraph>
<paragraph id="306836B3FFDDFFA10D39F94A5DE57FEC" blockId="10.[826,1441,1666,1903]" box="[826,927,1666,1687]" pageId="10" pageNumber="380">
<emphasis id="02A3EAA1FFDDFFA10D39F94A5DE57FEC" box="[826,927,1666,1687]" italics="true" pageId="10" pageNumber="380">Remarks</emphasis>
</paragraph>
<paragraph id="306836B3FFDDFFA10D39F9695BDB7E14" blockId="10.[826,1441,1666,1903]" pageId="10" pageNumber="380">
The taxonomy of 
<taxonomicName id="F7D74D30FFDDFFA10A06F96A5A2F7FCD" authorityName="Boheman" authorityYear="1836" box="[1029,1109,1697,1719]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="380" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDDFFA10A06F96A5A6D7FCC" box="[1029,1047,1698,1719]" italics="true" pageId="10" pageNumber="380">E</emphasis>
. 
<emphasis id="02A3EAA1FFDDFFA10A25F9695A2F7FCD" box="[1062,1109,1697,1718]" italics="true" pageId="10" pageNumber="380">afra</emphasis>
</taxonomicName>
is somewhat complicated as the existing 
<typeStatus id="EF6C8811FFDDFFA10DC9F9085D817FAE" box="[970,1019,1728,1749]" pageId="10" pageNumber="380">type</typeStatus>
is a male. The original female – not that selected by Thomson, and later examined by Ferrière – certainly belongs to the same species as the male. The original description mentions ‘antennis breviter monoliformibus’ and ‘prothorace utrinque flavomaculato’, which all together are distinct features of
</paragraph>
<caption id="64A8663BFFDCFFA00E93F9235C677E08" ID-DOI="http://doi.org/10.5281/zenodo.6987032" ID-Zenodo-Dep="6987032" httpUri="https://zenodo.org/record/6987032/files/figure.png" pageId="11" pageNumber="381" startId="11.[144,223,1771,1790]" subCaptionStartIDs="11.[1319,1421,1829,1848]" subCaptionStarts="Abbr" targetBox="[197,1371,195,1740]" targetPageId="11">
<paragraph id="306836B3FFDCFFA00E93F9235C677E08" blockId="11.[144,1421,1771,1907]" pageId="11" pageNumber="381">
<emphasis id="02A3EAA1FFDCFFA00E93F9235E827F85" bold="true" box="[144,248,1771,1790]" pageId="11" pageNumber="381">Figure 4.</emphasis>
A–O, 
<taxonomicName id="F7D74D30FFDCFFA00F43F9235F897F85" authorityName="Yano" authorityYear="1918" box="[320,499,1771,1790]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="381" phylum="Arthropoda" rank="species" species="laricis">
<emphasis id="02A3EAA1FFDCFFA00F43F9235F897F85" box="[320,499,1771,1790]" italics="true" pageId="11" pageNumber="381">Eurytoma laricis</emphasis>
</taxonomicName>
(D, F, H, I, J); 
<taxonomicName id="F7D74D30FFDCFFA00C99F9235D3C7F86" authorityName="Boheman" authorityYear="1836" box="[666,838,1771,1789]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="381" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="02A3EAA1FFDCFFA00C99F9235D3C7F86" box="[666,838,1771,1789]" italics="true" pageId="11" pageNumber="381">Eurytoma morio</emphasis>
</taxonomicName>
(G, M, N); 
<taxonomicName id="F7D74D30FFDCFFA00DC7F9235AF67F85" authorityName="Ratzeburg" authorityYear="1848" box="[964,1164,1771,1790]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="381" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="02A3EAA1FFDCFFA00DC7F9235AF67F85" box="[964,1164,1771,1790]" italics="true" pageId="11" pageNumber="381">Eurytoma striolata</emphasis>
</taxonomicName>
(A, B, C, L, O). A, back of head; B, postgena; C, postoccipital region; D–E, clypeus; F–G, lower face; H, male scape; I, scape plate with pores (arrows); J, male antenna; K, mesoscutum; L, female propodeum; M, procoxa; N, metacoxa; O, female petiole. Abbreviations: ATP, anterior tentorial pits; GC, genal carina; LFP, lateral formaminal plate; PGG, postgenal groove, PGL, postgenal lamina; PTP, posterior tentorial pits.
</paragraph>
</caption>
<caption id="64A8663BFFDBFFA70EA7F8C85B287E10" ID-DOI="http://doi.org/10.5281/zenodo.5306185" ID-Zenodo-Dep="5306185" httpUri="https://zenodo.org/record/5306185/files/figure.png" pageId="12" pageNumber="382" startId="12.[164,243,1792,1811]" targetBox="[209,1398,206,1701]" targetPageId="12">
<paragraph id="306836B3FFDBFFA70EA7F8C85B287E10" blockId="12.[164,1442,1792,1899]" pageId="12" pageNumber="382">
<emphasis id="02A3EAA1FFDBFFA70EA7F8C85F767E68" bold="true" box="[164,268,1792,1811]" pageId="12" pageNumber="382">Figure 5.</emphasis>
A–K, 
<taxonomicName id="F7D74D30FFDBFFA70F55F8C95FC47E68" authorityName="Illiger" authorityYear="1807" box="[342,446,1793,1811]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="02A3EAA1FFDBFFA70F55F8C95FC47E68" box="[342,446,1793,1811]" italics="true" pageId="12" pageNumber="382">Eurytoma</emphasis>
</taxonomicName>
(continued). 
<taxonomicName id="F7D74D30FFDBFFA70C4DF8C95D5E7E68" authorityName="Hedqvist" authorityYear="1976" box="[590,804,1792,1811]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="asphodeli">
<emphasis id="02A3EAA1FFDBFFA70C4DF8C95D5E7E68" box="[590,804,1792,1811]" italics="true" pageId="12" pageNumber="382">Eurytoma asphodeli</emphasis>
</taxonomicName>
(A); 
<taxonomicName id="F7D74D30FFDBFFA70D5DF8C95A527E68" authorityName="Ratzeburg" authorityYear="1848" box="[862,1064,1792,1811]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="02A3EAA1FFDBFFA70D5DF8C95A527E68" box="[862,1064,1792,1811]" italics="true" pageId="12" pageNumber="382">Eurytoma striolata</emphasis>
</taxonomicName>
(B); 
<taxonomicName id="F7D74D30FFDBFFA70A61F8C95B607E68" authorityName="Boheman" authorityYear="1836" box="[1122,1306,1793,1811]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="02A3EAA1FFDBFFA70A61F8C95B607E68" box="[1122,1306,1793,1811]" italics="true" pageId="12" pageNumber="382">Eurytoma maura</emphasis>
</taxonomicName>
(C, D, E, F, G); 
<taxonomicName id="F7D74D30FFDBFFA70ECAF8D65F147E4B" authorityName="Boheman" authorityYear="1836" box="[201,366,1822,1840]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="02A3EAA1FFDBFFA70ECAF8D65F147E4B" box="[201,366,1822,1840]" italics="true" pageId="12" pageNumber="382">Eurytoma morio</emphasis>
</taxonomicName>
(H); 
<taxonomicName id="F7D74D30FFDBFFA70FA1F8D65C2C7E4B" authorityName="Hedqvist" authorityYear="1974" box="[418,598,1821,1840]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="annilai">
<emphasis id="02A3EAA1FFDBFFA70FA1F8D65C2C7E4B" box="[418,598,1821,1840]" italics="true" pageId="12" pageNumber="382">Eurytoma annilai</emphasis>
</taxonomicName>
(I); 
<taxonomicName id="F7D74D30FFDBFFA70C82F8D65D727E4A" box="[641,776,1822,1841]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="02A3EAA1FFDBFFA70C82F8D65C9C7E4B" box="[641,742,1822,1840]" italics="true" pageId="12" pageNumber="382">Eurytoma</emphasis>
sp.
</taxonomicName>
(= 
<taxonomicName id="F7D74D30FFDBFFA70D2BF8D65D987E4B" authorityName="Hedqvist" authorityYear="1966" box="[808,994,1821,1840]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="kangasi">
<emphasis id="02A3EAA1FFDBFFA70D2BF8D65D987E4B" box="[808,994,1821,1840]" italics="true" pageId="12" pageNumber="382">Eurytoma kangasi</emphasis>
</taxonomicName>
allotype) (J); 
<emphasis id="02A3EAA1FFDBFFA70A6DF8D65BD87E4B" bold="true" box="[1134,1442,1821,1841]" pageId="12" pageNumber="382">
<taxonomicName id="F7D74D30FFDBFFA70A6DF8D65B307E4B" authority="Delvare &amp; Gebiola &amp; Zeiri &amp; Garonna, 2014" authorityName="Delvare &amp; Gebiola &amp; Zeiri &amp; Garonna" authorityYear="2014" box="[1134,1354,1821,1841]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="382" phylum="Arthropoda" rank="species" species="saliphila" status="sp. nov.">
<emphasis id="02A3EAA1FFDBFFA70A6DF8D65B307E4B" bold="true" box="[1134,1354,1821,1841]" italics="true" pageId="12" pageNumber="382">Eurytoma saliphila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="199057DAFFDBFFA70B4CF8D55BD87E4B" box="[1359,1442,1821,1840]" pageId="12" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(K). A, B, head in lateral view; C, female pedicel; D, male scape; E, male antenna; F, head in frontal view; G, pronotum; H, propodeum; I–J, mesopleuron; K, base of forewing. Abbreviations: GC, genal carina; PGL, postgenal lamina.
</paragraph>
</caption>
<caption id="64A8663BFFDAFFA60E93FB9C5C7D7DD9" ID-DOI="http://doi.org/10.5281/zenodo.5306189" ID-Zenodo-Dep="5306189" httpUri="https://zenodo.org/record/5306189/files/figure.png" pageId="13" pageNumber="383" startId="13.[144,220,1108,1127]" targetBox="[303,1263,195,1079]" targetPageId="13">
<paragraph id="306836B3FFDAFFA60E93FB9C5C7D7DD9" blockId="13.[144,1421,1108,1186]" pageId="13" pageNumber="383">
<emphasis id="02A3EAA1FFDAFFA60E93FB9C5E8E7D1C" bold="true" box="[144,244,1108,1127]" pageId="13" pageNumber="383">Figure 6.</emphasis>
A–F, 
<taxonomicName id="F7D74D30FFDAFFA60F30FB9D5FED7D1C" authorityName="Illiger" authorityYear="1807" box="[307,407,1109,1127]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="02A3EAA1FFDAFFA60F30FB9D5FED7D1C" box="[307,407,1109,1127]" italics="true" pageId="13" pageNumber="383">Eurytoma</emphasis>
</taxonomicName>
(continued). 
<taxonomicName id="F7D74D30FFDAFFA60C18FB9D5D707D1C" authorityName="Walker" authorityYear="1879" box="[539,778,1108,1127]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="melanoneura">
<emphasis id="02A3EAA1FFDAFFA60C18FB9D5D707D1C" box="[539,778,1108,1127]" italics="true" pageId="13" pageNumber="383">Eurytoma melanoneura</emphasis>
</taxonomicName>
(A); 
<taxonomicName id="F7D74D30FFDAFFA60D38FB9D5D807D1C" authorityName="Ratzeburg" authorityYear="1848" box="[827,1018,1108,1127]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="02A3EAA1FFDAFFA60D38FB9D5D807D1C" box="[827,1018,1108,1127]" italics="true" pageId="13" pageNumber="383">Eurytoma striolata</emphasis>
</taxonomicName>
(B, D); 
<taxonomicName id="F7D74D30FFDAFFA60A4AFB9D5A827D1C" authorityName="Boheman" authorityYear="1836" box="[1097,1272,1109,1127]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="02A3EAA1FFDAFFA60A4AFB9D5A827D1C" box="[1097,1272,1109,1127]" italics="true" pageId="13" pageNumber="383">Eurytoma maura</emphasis>
</taxonomicName>
(C); 
<taxonomicName id="F7D74D30FFDAFFA60B2AFB9D5EB67DFF" authorityName="Boheman" authorityYear="1836" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="02A3EAA1FFDAFFA60B2AFB9D5EB67DFF" italics="true" pageId="13" pageNumber="383">Eurytoma morio</emphasis>
</taxonomicName>
(E); 
<emphasis id="02A3EAA1FFDAFFA60F07FBBA5C597DFE" bold="true" box="[260,547,1137,1157]" pageId="13" pageNumber="383">
<taxonomicName id="F7D74D30FFDAFFA60F07FBBA5FBC7DFF" authority="Delvare &amp; Gebiola &amp; Zeiri &amp; Garonna, 2014" authorityName="Delvare &amp; Gebiola &amp; Zeiri &amp; Garonna" authorityYear="2014" box="[260,454,1137,1157]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="ithma" status="sp. nov.">
<emphasis id="02A3EAA1FFDAFFA60F07FBBA5FBC7DFF" bold="true" box="[260,454,1137,1157]" italics="true" pageId="13" pageNumber="383">Eurytoma ithma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="199057DAFFDAFFA60FCEFBBA5C597DFE" box="[461,547,1138,1157]" pageId="13" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(F). A, B, costal cell of forewing; C, D, apical venation of forewing; E, pronotum and mesoscutum, F, male antenna.
</paragraph>
</caption>
<paragraph id="306836B3FFDAFFA60E93FB1D5C8D7F40" blockId="13.[144,759,1236,1717]" pageId="13" pageNumber="383">
<taxonomicName id="F7D74D30FFDAFFA60E93FB1D5E9C7D91" authorityName="Boheman" authorityYear="1836" box="[144,230,1237,1258]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDAFFA60E93FB1D5ED87D91" box="[144,162,1237,1258]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60EB0FB1D5E9C7D91" box="[179,230,1237,1258]" italics="true" pageId="13" pageNumber="383">afra</emphasis>
</taxonomicName>
<emphasis id="02A3EAA1FFDAFFA60EE5FB1D5E907D91" box="[230,234,1237,1258]" italics="true" pageId="13" pageNumber="383">.</emphasis>
<bibRefCitation id="54464B42FFDAFFA60EF7FB1C5FDE7D91" author="Ferriere C" box="[244,420,1236,1258]" pageId="13" pageNumber="383" pagination="377 - 410" refId="ref22575" refString="Ferriere C. 1950. Notes sur les Eurytoma (Hym., Chalcidoidea). I. les types de Thomson et de Mayr. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 23: 377 - 410." type="journal article" year="1950">Ferrière (1950)</bibRefCitation>
erroneously synonymized 
<taxonomicName id="F7D74D30FFDAFFA60CDDFB1D5F637C73" authorityName="Mayr" authorityYear="1878" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="saliciperdae">
<emphasis id="02A3EAA1FFDAFFA60CDDFB1D5C8A7D91" box="[734,752,1237,1258]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60E93FB3B5F637C73" box="[144,281,1267,1288]" italics="true" pageId="13" pageNumber="383">saliciperdae</emphasis>
</taxonomicName>
with it. The confusion was later recognized by 
<bibRefCitation id="54464B42FFDAFFA60EF9FADA5FD77C53" author="Graham MWR de V." box="[250,429,1298,1320]" pageId="13" pageNumber="383" pagination="125 - 152" refId="ref22855" refString="Graham MWR de V. 1970. Taxonomic notes on some western Palaearctic Eurytomidae (Hymenoptera: Chalcidoidea). Proceedings of the Royal Entomological Society of London (B) 39: 125 - 152." type="journal article" year="1970">Graham (1970)</bibRefCitation>
. Part of the 
<typeStatus id="EF6C8811FFDAFFA60C41FADA5C097C5C" box="[578,627,1298,1319]" pageId="13" pageNumber="383">type</typeStatus>
series of 
<taxonomicName id="F7D74D30FFDAFFA60CE5FADA5F687C3D" authorityName="Boucek &amp; Novicky" authorityYear="1954" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFDAFFA60CE5FADA5C8A7C5C" box="[742,752,1298,1319]" italics="true" pageId="13" pageNumber="383">I</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60E93FAF95F687C3D" box="[144,274,1329,1350]" italics="true" pageId="13" pageNumber="383">spessivtsevi</emphasis>
</taxonomicName>
had previously been correctly identified by Erdös as belonging to 
<taxonomicName id="F7D74D30FFDAFFA60F80FA985FB57C1F" authorityName="Boheman" authorityYear="1836" box="[387,463,1359,1381]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDAFFA60F80FA985FEF7C1E" box="[387,405,1360,1381]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60FA3FA875FB57C1F" box="[416,463,1359,1380]" italics="true" pageId="13" pageNumber="383">afra</emphasis>
</taxonomicName>
(Boucˇek &amp; 
<bibRefCitation id="54464B42FFDAFFA60C4DFA875C967C1E" author="Novicky S" box="[590,748,1359,1381]" pageId="13" pageNumber="383" pagination="266 - 271" refId="ref22101" refString="Boucˇek Z, Novicky S. 1954. Ipideurytoma spessivtsevi n. g., n. sp., ein neuer Borlenkaferparasit. Entomologisk Tidskrift 75: 266 - 271." type="journal article" year="1954">Novicky, 1954</bibRefCitation>
). 
<bibRefCitation id="54464B42FFDAFFA60E93FAA65F517CFF" author="Zerova MD" box="[144,299,1390,1412]" pageId="13" pageNumber="383" pagination="32" refId="ref24564" refString="Zerova MD. 1998. [The identity of Eurytoma spessivtsevi (Boucek et Novicky, 1956) to Decatomoidea polygraphi (Ashmead, 1894) (Hymenoptera, Eurytomidae)]. Vestnik Zoologii, Kiev 32: 32. [in Russian]." type="journal article" year="1998">Zerova (1998)</bibRefCitation>
rightly synonymized 
<taxonomicName id="F7D74D30FFDAFFA60C21FAA65CC67CF8" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[546,700,1390,1411]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFDAFFA60C21FAA65C567CF8" box="[546,556,1390,1411]" italics="true" pageId="13" pageNumber="383">I</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60C39FAA65CC67CF8" box="[570,700,1390,1411]" italics="true" pageId="13" pageNumber="383">spessivtsevi</emphasis>
</taxonomicName>
with 
<taxonomicName id="F7D74D30FFDAFFA60E93FA445FDC7CDA" authorityName="Ashmead" authorityYear="1894" box="[144,422,1420,1441]" class="Insecta" family="Eurytomidae" genus="Decatomidea" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="polygraphi">
<emphasis id="02A3EAA1FFDAFFA60E93FA445FDC7CDA" box="[144,422,1420,1441]" italics="true" pageId="13" pageNumber="383">Decatomidea polygraphi</emphasis>
</taxonomicName>
, but followed Ferrière in the interpretation of 
<taxonomicName id="F7D74D30FFDAFFA60F48FA635FEC7CBB" authorityName="Boheman" authorityYear="1836" box="[331,406,1451,1472]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDAFFA60F48FA635F267CBB" box="[331,348,1451,1472]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60F6BFA635FEC7CBB" box="[360,406,1451,1472]" italics="true" pageId="13" pageNumber="383">afra</emphasis>
</taxonomicName>
. As an implication of the species name synonymy, Zerova also synonymized 
<taxonomicName id="F7D74D30FFDAFFA60C62FA015C8C7CA5" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[609,758,1481,1502]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="02A3EAA1FFDAFFA60C62FA015C8C7CA5" box="[609,758,1481,1502]" italics="true" pageId="13" pageNumber="383">Ipideurytoma</emphasis>
</taxonomicName>
with 
<taxonomicName id="F7D74D30FFDAFFA60ECDFA215F457C85" authorityName="Illiger" authorityYear="1807" box="[206,319,1513,1534]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="02A3EAA1FFDAFFA60ECDFA215F457C85" box="[206,319,1513,1534]" italics="true" pageId="13" pageNumber="383">Eurytoma</emphasis>
</taxonomicName>
. Her suggestion is validated here by our molecular results. 
<taxonomicName id="F7D74D30FFDAFFA60F97F9CF5C477F60" authorityName="Boheman" authorityYear="1836" box="[404,573,1542,1564]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDAFFA60F97F9CF5C477F60" box="[404,573,1542,1564]" italics="true" pageId="13" pageNumber="383">Eurytoma afra</emphasis>
</taxonomicName>
always appears as the sister group of 
<taxonomicName id="F7D74D30FFDAFFA60F97F9EE5F857F40" authorityName="Boheman" authorityYear="1836" box="[404,511,1574,1595]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="02A3EAA1FFDAFFA60F97F9EE5FDC7F40" box="[404,422,1574,1595]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60FB0F9EE5F857F40" box="[435,511,1574,1595]" italics="true" pageId="13" pageNumber="383">maura</emphasis>
</taxonomicName>
with strong support.
</paragraph>
<paragraph id="306836B3FFDAFFA60EABF98C5F857FCE" blockId="13.[144,759,1236,1717]" pageId="13" pageNumber="383">
The species was well described and illustrated by Boucˇek &amp; 
<bibRefCitation id="54464B42FFDAFFA60F05F9AA5FC97F03" author="Novicky S" box="[262,435,1634,1656]" pageId="13" pageNumber="383" pagination="266 - 271" refId="ref22101" refString="Boucˇek Z, Novicky S. 1954. Ipideurytoma spessivtsevi n. g., n. sp., ein neuer Borlenkaferparasit. Entomologisk Tidskrift 75: 266 - 271." type="journal article" year="1954">Novicky (1954)</bibRefCitation>
. Because of the shape of the head, especially in the female (
<figureCitation id="A8EC2A36FFDAFFA60FE7F9495C1B7FEC" box="[484,609,1665,1687]" captionStart="Figure 10" captionStartId="16.[164,243,799,818]" captionTargetBox="[323,1283,195,770]" captionTargetId="figure-430@16.[323,1283,195,770]" captionTargetPageId="16" captionText="Figure 10. A–D, Eurytoma afra: A–C, female; D, male. A, head in frontal view; B, head and pronotum in lateral view; C, head and mesosoma in dorsal view; D, antenna." figureDoi="http://doi.org/10.5281/zenodo.5306201" httpUri="https://zenodo.org/record/5306201/files/figure.png" pageId="13" pageNumber="383">Fig. 10A–C</figureCitation>
), it was originally proposed as a new genus.
</paragraph>
</subSubSection>
<subSubSection id="78CD6538FFDAFFA60E93F9175D9F7F28" pageId="13" pageNumber="383" type="biology_ecology">
<paragraph id="306836B3FFDAFFA60E93F9175EB57F8F" blockId="13.[144,759,1759,1903]" box="[144,207,1759,1780]" pageId="13" pageNumber="383">
<emphasis id="02A3EAA1FFDAFFA60E93F9175EB57F8F" box="[144,207,1759,1780]" italics="true" pageId="13" pageNumber="383">Hosts</emphasis>
</paragraph>
<paragraph id="306836B3FFDAFFA60E93F9355A257C1E" blockId="13.[144,759,1759,1903]" lastBlockId="13.[806,1422,1236,1381]" pageId="13" pageNumber="383">
The 
<typeStatus id="EF6C8811FFDAFFA60EC7F9355F5F7E68" box="[196,293,1789,1811]" pageId="13" pageNumber="383" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="F7D74D30FFDAFFA60F49F9355C147E69" authorityName="Boucek &amp; Novicky" authorityYear="1954" box="[330,622,1789,1810]" class="Insecta" family="Eurytomidae" genus="Ipideurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="spessivtsevi">
<emphasis id="02A3EAA1FFDAFFA60F49F9355C147E69" box="[330,622,1789,1810]" italics="true" pageId="13" pageNumber="383">Ipideurytoma spessivtsevi</emphasis>
</taxonomicName>
was reared from 
<taxonomicName id="F7D74D30FFDAFFA60EDAF8D35C8C7E49" authority="(Fabricius, 1787)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[217,758,1819,1842]" class="Insecta" family="Curculionidae" genus="Trypodendron" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="signatum">
<emphasis id="02A3EAA1FFDAFFA60EDAF8D35C717E4A" box="[217,523,1819,1841]" italics="true" pageId="13" pageNumber="383">Trypodendron signatum</emphasis>
(Fabricius, 1787)
</taxonomicName>
(
<taxonomicName id="F7D74D30FFDAFFA60E94F8F25F407E2B" authorityName="Latreille" authorityYear="1802" box="[151,314,1850,1872]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
: 
<taxonomicName id="F7D74D30FFDAFFA60F45F8F25FC67E2B" box="[326,444,1850,1872]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
) (Boucˇek &amp; 
<bibRefCitation id="54464B42FFDAFFA60C49F8F25C917E2B" author="Novicky S" box="[586,747,1850,1872]" pageId="13" pageNumber="383" pagination="266 - 271" refId="ref22101" refString="Boucˇek Z, Novicky S. 1954. Ipideurytoma spessivtsevi n. g., n. sp., ein neuer Borlenkaferparasit. Entomologisk Tidskrift 75: 266 - 271." type="journal article" year="1954">Novicky, 1954</bibRefCitation>
). A list was provided by 
<bibRefCitation id="54464B42FFDAFFA60F91F8915C157E14" author="Hedqvist KJ" box="[402,623,1881,1903]" pageId="13" pageNumber="383" pagination="1 - 176" refId="ref23071" refString="Hedqvist KJ. 1963. Die Feinde der Borkenkafer in Schweden. I. Erzwespen (Chalcidoidea). Studia Forestalia Suecica 11: 1 - 176." type="journal article" year="1963">Hedqvist (1963: 47)</bibRefCitation>
from initial data published by 
<bibRefCitation id="54464B42FFDAFFA60A16FB1C5A9C7D91" author="Jamnicky J" box="[1045,1254,1236,1258]" pageId="13" pageNumber="383" pagination="597 - 604" refId="ref23177" refString="Jamnicky J. 1957. Prispekov k poznaniu parazitov korovcov (Scolytidae) z radu blanokridlovcov (Hymenoptera). Biologia 12: 597 - 604." type="journal article" year="1957">Jamnický (1957)</bibRefCitation>
. This list includes 
<taxonomicName id="F7D74D30FFDAFFA60D7EFB3B5B2E7C72" authority="(Olivier, 1795)" baseAuthorityName="Olivier" baseAuthorityYear="1795" box="[893,1364,1267,1289]" class="Insecta" family="Curculionidae" genus="Trypodendron" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="lineatum">
<emphasis id="02A3EAA1FFDAFFA60D7EFB3B5AE37C73" box="[893,1177,1267,1288]" italics="true" pageId="13" pageNumber="383">Trypodendron lineatum</emphasis>
(Olivier, 1795)
</taxonomicName>
and 
<taxonomicName id="F7D74D30FFDAFFA60D25FADA5B607C5C" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[806,1306,1297,1320]" class="Insecta" family="Curculionidae" genus="Trypodendron" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="domesticum">
<emphasis id="02A3EAA1FFDAFFA60D25FADA5A2B7C5C" box="[806,1105,1298,1319]" italics="true" pageId="13" pageNumber="383">Trypodendron domesticum</emphasis>
(Linnaeus, 1758)
</taxonomicName>
. 
<bibRefCitation id="54464B42FFDAFFA60B25FADA5DAC7C3D" author="Eichhorn O &amp; Graf P" pageId="13" pageNumber="383" pagination="129 - 135" refId="ref22507" refString="Eichhorn O, Graf P. 1974. Uber einige Nutzholzborkenkafer und ihre Feinde. Anzeiger fur Schadlingskunde, Pflanzen- und Umweltschutz 47: 129 - 135." type="journal article" year="1974">Eichhorn &amp; Graf (1974)</bibRefCitation>
mentioned 
<taxonomicName id="F7D74D30FFDAFFA60A6CFAF95AB97C3E" authorityName="Boheman" authorityYear="1836" box="[1135,1219,1328,1350]" class="Insecta" family="Eurytomidae" genus="Eurytoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="383" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="02A3EAA1FFDAFFA60A6CFAF95AFB7C3D" box="[1135,1153,1329,1350]" italics="true" pageId="13" pageNumber="383">E</emphasis>
. 
<emphasis id="02A3EAA1FFDAFFA60A92FAF85AB97C3E" box="[1169,1219,1328,1349]" italics="true" pageId="13" pageNumber="383">afra</emphasis>
</taxonomicName>
as the primary parasitoid of these beetles.
</paragraph>
<paragraph id="306836B3FFDAFFA60D25FA4D5DCB7CE1" blockId="13.[806,1421,1413,1619]" box="[806,945,1413,1434]" pageId="13" pageNumber="383">
<emphasis id="02A3EAA1FFDAFFA60D25FA4D5DCB7CE1" box="[806,945,1413,1434]" italics="true" pageId="13" pageNumber="383">Distribution</emphasis>
</paragraph>
<paragraph id="306836B3FFDAFFA60D25FA6C5D9F7F28" blockId="13.[806,1421,1413,1619]" pageId="13" pageNumber="383">
The species is widely distributed, being present in the Nearctic and Palaearctic regions. It is from Northern and Central Europe (
<bibRefCitation id="54464B42FFDAFFA60A23FA295ACB7C8C" author="Noyes JS" box="[1056,1201,1505,1527]" pageId="13" pageNumber="383" refId="ref23581" refString="Noyes JS. 2013. Universal Chalcidoidea Database. World Wide Web electronic publication. Available at: http: // www. nhm. ac. uk / research-curation / projects / chalcidoids /" type="url" year="2013">Noyes, 2013</bibRefCitation>
). The westernmost specimens were collected in the Juliana Alps, close to the Slovenian border (see Appendix S4 and 
<figureCitation id="A8EC2A36FFDAFFA60B2CF9D65BF77F4F" box="[1327,1421,1566,1588]" captionStart="Figure 10" captionStartId="16.[164,243,799,818]" captionTargetBox="[323,1283,195,770]" captionTargetId="figure-430@16.[323,1283,195,770]" captionTargetPageId="16" captionText="Figure 10. A–D, Eurytoma afra: A–C, female; D, male. A, head in frontal view; B, head and pronotum in lateral view; C, head and mesosoma in dorsal view; D, antenna." figureDoi="http://doi.org/10.5281/zenodo.5306201" httpUri="https://zenodo.org/record/5306201/files/figure.png" pageId="13" pageNumber="383">
Fig. S 
<quantity id="F72F9B56FFDAFFA60B72F9D75BF77F4F" box="[1393,1421,1567,1588]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="383" unit="in" value="10.0">10</quantity>
</figureCitation>
in Appendix S8).
</paragraph>
</subSubSection>
</treatment>
</document>