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<document id="D60B128932493D35C284E4B098DF177F" ID-CLB-Dataset="6148" ID-DOI="10.11646/zootaxa.4975.3.2" ID-GBIF-Dataset="330b4589-7947-450b-857f-812cb72beb6d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4808307" ID-ZooBank="222B81FA-E0F5-43A9-960D-5C9ACC2ADD0A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622026211124" checkinUser="plazi" docAuthor="Martynov, Alexander V., Selvakumar, C., Subramanian, K. A., Sivaramakrishnan, K. G., Vasanth, M., Sinha, Bikramjit &amp; Jacobus, Luke M." docDate="2021" docId="1D7287CDFFC2D660FF09086A0852F8AE" docLanguage="en" docName="zootaxa.4975.3.2.pdf" docOrigin="Zootaxa 4975 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torleya dibruensis Selvakumar, Martynov &amp; Jacobus 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="E14BFFB5FFD2D673FF9E0C080A62FFC5" masterDocTitle="Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions" masterLastPageNumber="482" masterPageNumber="451" pageNumber="467" updateTime="1698991974093" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F46E6945243D4DFAC7A03522BD22677E">Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions</mods:title>
</mods:titleInfo>
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<mods:namePart id="611D70B75214ACE46AE9211F2FAC4002">Martynov, Alexander V.</mods:namePart>
<mods:nameIdentifier id="EDF555CBB187ECE497F7339190FFC05F" type="ORCID">0000-0002-6506-5134</mods:nameIdentifier>
<mods:affiliation id="71ED32A03EFFCE3317F21F3F4F5008B6">National Museum of Natural History, National Academy of Sciences of Ukraine, Bohdan Khmelnytsky Str., 15, 01030, Kyiv, Ukraine. martynov _ av @ ukr. net; centroptilum @ gmail. com; https: // orcid. org / 0000 - 0002 - 6506 - 5134</mods:affiliation>
<mods:nameIdentifier id="AB0764E848AF495500A342752860B29D" type="email">martynov_av@ukr.net</mods:nameIdentifier>
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<mods:namePart id="9307E09C3DBECCC29C1DB3E12B009932">Selvakumar, C.</mods:namePart>
<mods:affiliation id="1758E4FD453107F88383381F67F979FD">Department of Zoology, The Madura College (Autonomous), Madurai- 625011, Tamil Nadu, India.</mods:affiliation>
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<mods:namePart id="8CC5FB289607CC41D2A215A41F7D6DC0">Subramanian, K. A.</mods:namePart>
<mods:nameIdentifier id="AAB19AB55A016F0057AA5229DD1A0C95" type="ORCID">0000-0003-0872-9771</mods:nameIdentifier>
<mods:affiliation id="D7F02B7283EDCB95EE17BFA6B7A5E9C9">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India. &amp; subbuka. zsi @ gmail. com; https: // orcid. org / 0000 - 0003 - 0872 - 9771</mods:affiliation>
<mods:nameIdentifier id="4A20F3EAAD5FC27CB938483703461127" type="email">subbuka.zsi@gmail.com</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="859423D73627055F5DDE697E48C028AD">Sivaramakrishnan, K. G.</mods:namePart>
<mods:affiliation id="E9122835D048BC0EA42310ADA0A995C2">Flat 3, Gokulam Apartments, No. 7, Gokulam Colony, West Mambalam, Chennai- 600 033, India.</mods:affiliation>
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<mods:namePart id="EE8DD6640A37E92D4ADE38B869C69C50">Vasanth, M.</mods:namePart>
<mods:nameIdentifier id="199BCEECCD8AB94CC29C2EA7D65370B7" type="ORCID">0000-0003-4446-4889</mods:nameIdentifier>
<mods:affiliation id="E346E69762F01D79BE68E3EAF56E2438">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India. &amp; vasan 071994 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4446 - 4889</mods:affiliation>
<mods:nameIdentifier id="D8DF11678840ACB00FC7CA1D694167C2" type="email">vasan071994@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D3C43C57FFDB0E3CB524922B8E014C59">Sinha, Bikramjit</mods:namePart>
<mods:nameIdentifier id="65D15E0E0A7F45FE1D5EB01744ACA680" type="ORCID">0000-0002-9904-4964</mods:nameIdentifier>
<mods:affiliation id="24753A86F91C3DBE363617B2D08B256F">Zoological Survey of India, Arunachal Pradesh Regional Centre, Itanagar- 791113, Arunachal Pradesh, India. sinhabj @ gmail. com; https: // orcid. org / 0000 - 0002 - 9904 - 4964</mods:affiliation>
<mods:nameIdentifier id="C540C322EA03172E5026C5BC6640ACD7" type="email">sinhabj@gmail.com</mods:nameIdentifier>
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<mods:namePart id="63EFA13CB38B447D9371725CA2A00730">Jacobus, Luke M.</mods:namePart>
<mods:nameIdentifier id="813A78F5C845AC02C6C1E418FCEE6C27" type="ORCID">0000-0002-7201-3922</mods:nameIdentifier>
<mods:affiliation id="0D82CDC7F5BFF7B3440C4AFF8B414C82">Division of Science, Indiana University Purdue University Columbus, Columbus, Indiana, 47203, USA. lukemjacobus @ alumni. purdue. edu; https: // orcid. org / 0000 - 0002 - 7201 - 3922</mods:affiliation>
<mods:nameIdentifier id="6DE2907721B0F3EBBFAB787314451872" type="email">lukemjacobus@alumni.purdue.edu</mods:nameIdentifier>
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<mods:typeOfResource id="8781F3746EA9595F9AF5BEF7DE93AC8C">text</mods:typeOfResource>
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<mods:titleInfo id="829013FD855B8FD615146A041E2BE748">
<mods:title id="0DBC360824304F2566D211236B623326">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D3307B5BB6B4ED668B2BC2621D13145E">
<mods:date id="2283B9C6A54960F44EA03F842E5F4354">2021</mods:date>
<mods:detail id="AD571DCCD053BD73F05D572483379B7D" type="pubDate">
<mods:number id="C750938271F0F5462532A553A7731C74">2021-05-26</mods:number>
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<mods:detail id="2DB2C3761260D1D505AA926950ED2D33" type="volume">
<mods:number id="B86EC2A48F0AEE5F5BB05D4D880E9874">4975</mods:number>
</mods:detail>
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<mods:number id="87F236433A75B1B9BE178B91416E238E">3</mods:number>
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<mods:classification id="AB1BE7C7B722796D2DA9E3BE3377F494">journal article</mods:classification>
<mods:identifier id="5B1555FA5430914BB0E5B9ECD4B5B25A" type="CLB-Dataset">6148</mods:identifier>
<mods:identifier id="F7941A417715335F09F169C64FBD4528" type="DOI">10.11646/zootaxa.4975.3.2</mods:identifier>
<mods:identifier id="1C7A18C8EEB5751DA98DB913AE63B638" type="GBIF-Dataset">330b4589-7947-450b-857f-812cb72beb6d</mods:identifier>
<mods:identifier id="AAF2D462F7D3E46A981C27F3B514D941" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="1D7287CDFFC2D660FF09086A0852F8AE" ID-DOI="http://doi.org/10.5281/zenodo.4808325" ID-GBIF-Taxon="182651341" ID-Zenodo-Dep="4808325" LSID="urn:lsid:plazi:treatment:1D7287CDFFC2D660FF09086A0852F8AE" httpUri="http://treatment.plazi.org/id/1D7287CDFFC2D660FF09086A0852F8AE" lastPageId="19" lastPageNumber="470" pageId="16" pageNumber="467">
<subSubSection id="DDC16550FFC2D663FF09086A09F5FBB8" box="[151,919,1122,1149]" pageId="16" pageNumber="467" type="nomenclature">
<paragraph id="956436DBFFC2D663FF09086A09F5FBB8" blockId="16.[151,919,1122,1185]" box="[151,919,1122,1149]" pageId="16" pageNumber="467">
<heading id="CE2C81B7FFC2D663FF09086A09F5FBB8" bold="true" box="[151,919,1122,1149]" fontSize="11" level="1" pageId="16" pageNumber="467" reason="1">
<emphasis id="A7AFEAC9FFC2D663FF09086A09F5FBB8" bold="true" box="[151,919,1122,1149]" pageId="16" pageNumber="467">
<taxonomicName id="52DB4D58FFC2D663FF09086A094FFBB8" authority="Selvakumar, Martynov &amp; Jacobus" authorityName="Selvakumar, Martynov &amp; Jacobus" authorityYear="2021" box="[151,813,1122,1149]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC2D663FF09086A0B15FBB8" bold="true" box="[151,375,1122,1149]" italics="true" pageId="16" pageNumber="467">Torleya dibruensis</emphasis>
Selvakumar, Martynov &amp; Jacobus
</taxonomicName>
,
<taxonomicNameLabel id="BC9C57B2FFC2D663FCA7086B09F5FBB8" box="[825,919,1123,1149]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC2D663FF09088F0B44FB64" box="[151,294,1159,1185]" pageId="16" pageNumber="467" type="description">
<paragraph id="956436DBFFC2D663FF09088F0B44FB64" blockId="16.[151,919,1122,1185]" box="[151,294,1159,1185]" pageId="16" pageNumber="467">
(
<figureCitation id="0DE02A5EFFC2D663FF01088F0B7FFB64" box="[159,285,1159,1185]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="17.[151,250,1981,2005]" captionStartId-1="18.[151,250,805,829]" captionStartId-2="19.[151,250,1587,1611]" captionTargetBox-0="[165,1421,181,1938]" captionTargetBox-1="[151,1436,181,780]" captionTargetBox-2="[151,1436,181,1560]" captionTargetId-0="figure-20@17.[165,1421,181,1957]" captionTargetId-1="figure-19@18.[151,1436,181,781]" captionTargetId-2="figure-20@19.[151,1436,181,1563]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." captionText-1="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." captionText-2="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi-0="http://doi.org/10.5281/zenodo.4808355" figureDoi-1="http://doi.org/10.5281/zenodo.4808359" figureDoi-2="http://doi.org/10.5281/zenodo.4808363" httpUri-0="https://zenodo.org/record/4808355/files/figure.png" httpUri-1="https://zenodo.org/record/4808359/files/figure.png" httpUri-2="https://zenodo.org/record/4808363/files/figure.png" pageId="16" pageNumber="467">Figs 11–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC2D663FF0908C708F1FABC" pageId="16" pageNumber="467" type="materials_examined">
<paragraph id="956436DBFFC2D663FF0908C708F1FABC" blockId="16.[151,1437,1230,1941]" pageId="16" pageNumber="467">
<emphasis id="A7AFEAC9FFC2D663FF0908C70B2BFB2D" bold="true" box="[151,329,1230,1257]" pageId="16" pageNumber="467">Type material.</emphasis>
<materialsCitation id="25B33C86FFC2D663FECA08C70FFAFAC8" ID-GBIF-Occurrence="3128883324" collectingDate="2015-04-22" collectionCode="ZSI" collectorName="K. A. Subramanian &amp; Bikramjit Sinha " country="India" elevation="128" latitude="27.147655" location="Dibru River" longLatPrecision="1" longitude="93.74908" municipality="Papum Pare District" pageId="16" pageNumber="467" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="A7AFEAC9FFC2D663FECA08C70BA3FB2C" bold="true" box="[340,449,1231,1257]" pageId="16" pageNumber="467">
<typeStatus id="4A608879FFC2D663FECA08C70BA3FB2C" box="[340,449,1231,1257]" pageId="16" pageNumber="467" type="holotype">Holotype</typeStatus>
</emphasis>
: larva,
<collectingCountry id="EDCC764BFFC2D663FD8008C70812FB2C" box="[542,624,1231,1257]" name="India" pageId="16" pageNumber="467">INDIA</collectingCountry>
,
<collectingRegion id="571FF839FFC2D663FDE308C7093BFB2C" box="[637,857,1231,1257]" country="India" name="Arunachal Pradesh" pageId="16" pageNumber="467">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="7500ACA1FFC2D663FCF608C70E36FB2C" box="[872,1108,1231,1257]" pageId="16" pageNumber="467">Papum Pare District</collectingMunicipality>
,
<location id="90046000FFC2D663FBFD08C70E92FB2C" LSID="urn:lsid:plazi:treatment:1D7287CDFFC2D660FF09086A0852F8AE:90046000FFC2D663FBFD08C70E92FB2C" box="[1123,1264,1231,1257]" country="India" latitude="27.147655" longLatPrecision="1" longitude="93.74908" municipality="Papum Pare District" name="Dibru River" pageId="16" pageNumber="467" stateProvince="Arunachal Pradesh">Dibru River</location>
,
<geoCoordinate id="F0EF501CFFC2D663FB6108C60FFAFB2C" box="[1279,1432,1230,1257]" degrees="27.147655" direction="north" orientation="latitude" pageId="16" pageNumber="467" precision="1" value="27.147655">27.147655°N</geoCoordinate>
,
<geoCoordinate id="F0EF501CFFC2D663FF0908FB0B42FAC8" box="[151,288,1267,1293]" degrees="93.74908" direction="east" orientation="longitude" pageId="16" pageNumber="467" precision="1" value="93.74908">93.74908°E</geoCoordinate>
, h ~
<elevation id="1EF6D1E8FFC2D663FEC808FB0BBBFAC8" box="[342,473,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="16" pageNumber="467" unit="m" value="128.0">
<quantity id="52239B3EFFC2D663FEC808FB0BFFFAC8" box="[342,413,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="16" pageNumber="467" unit="m" value="128.0">128 m</quantity>
a.s.l.
</elevation>
,
<date id="E165101BFFC2D663FE7808FA083DFAC8" box="[486,607,1266,1293]" pageId="16" pageNumber="467" value="2015-04-22">
<collectingDate id="F121E9F3FFC2D663FE7808FA083DFAC8" box="[486,607,1266,1293]" pageId="16" pageNumber="467" value="2015-04-22">22.iv.2015</collectingDate>
</date>
,
<collectorName id="382E530DFFC2D663FDF408FB09E6FAC8" box="[618,900,1267,1293]" pageId="16" pageNumber="467">Colls. K.A. Subramanian</collectorName>
&amp;
<collectorName id="382E530DFFC2D663FC3608FB0E39FAC8" box="[936,1115,1267,1293]" pageId="16" pageNumber="467">Bikramjit Sinha</collectorName>
—
<collectorName id="382E530DFFC2D663FBE708FB0EB9FAC8" box="[1145,1243,1267,1293]" pageId="16" pageNumber="467">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFC2D663FB7C08FA0F30FAC9" box="[1250,1362,1266,1292]" italics="true" pageId="16" pageNumber="467">5606/H13</emphasis>
[
<collectionCode id="F3CAAE1EFFC2D663FAFE08FA0FF3FAC8" box="[1376,1425,1266,1293]" pageId="16" pageNumber="467">ZSI</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="25B33C86FFC2D663FF09091F09C4FAF4" ID-GBIF-Occurrence="3128883316" box="[151,934,1302,1329]" collectingDate="2015-04-22" collectionCode="ZSI" collectorName="Reg. No." country="India" elevation="128" latitude="27.147655" location="Dibru River" longLatPrecision="1" longitude="93.74908" municipality="Papum Pare District" pageId="16" pageNumber="467" specimenCount="1" specimenCount-larva="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="A7AFEAC9FFC2D663FF09091F0B6DFAF4" bold="true" box="[151,271,1303,1329]" pageId="16" pageNumber="467">
<typeStatus id="4A608879FFC2D663FF09091F0B6DFAF4" box="[151,271,1303,1329]" pageId="16" pageNumber="467" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="83DDFD52FFC2D663FE83091F0B0EFAF4" box="[285,364,1303,1329]" pageId="16" pageNumber="467" type="larva">1 larva</specimenCount>
, same data as holotype—
<collectorName id="382E530DFFC2D663FD14091F0889FAF4" box="[650,747,1303,1329]" pageId="16" pageNumber="467">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFC2D663FD6F091E0903FAF5" box="[753,865,1302,1328]" italics="true" pageId="16" pageNumber="467">5607/H13</emphasis>
[
<collectionCode id="F3CAAE1EFFC2D663FCF0091E09FDFAF4" box="[878,927,1302,1329]" pageId="16" pageNumber="467">ZSI</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="25B33C86FFC2D663FC2F091F08EDFABC" ID-GBIF-Occurrence="3128883330" collectingDate="2017-06-14" collectionCode="ZSI" collectorName="K. A. Subramanian &amp; M. Vasanth " country="India" elevation="993" latitude="27.81791" location="Paniya" longLatPrecision="1" longitude="94.09502" municipality="Lower Subansiri District" pageId="16" pageNumber="467" specimenCount="2" specimenCount-larva="2" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<specimenCount id="83DDFD52FFC2D663FC2F091F0E6FFAF4" box="[945,1037,1303,1329]" pageId="16" pageNumber="467" type="larva">2 larvae</specimenCount>
,
<collectingCountry id="EDCC764BFFC2D663FB89091F0E0BFAF4" box="[1047,1129,1303,1329]" name="India" pageId="16" pageNumber="467">INDIA</collectingCountry>
,
<collectingRegion id="571FF839FFC2D663FBEC091F0F2BFAF4" box="[1138,1353,1303,1329]" country="India" name="Arunachal Pradesh" pageId="16" pageNumber="467">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="7500ACA1FFC2D663FACD091F0B3DFA90" pageId="16" pageNumber="467">Lower Subansiri District</collectingMunicipality>
,
<location id="90046000FFC2D663FEF409330BD4FA90" LSID="urn:lsid:plazi:treatment:1D7287CDFFC2D660FF09086A0852F8AE:90046000FFC2D663FEF409330BD4FA90" box="[362,438,1339,1365]" country="India" latitude="27.81791" longLatPrecision="1" longitude="94.09502" municipality="Lower Subansiri District" name="Paniya" pageId="16" pageNumber="467" stateProvince="Arunachal Pradesh">Paniya</location>
stream,
<geoCoordinate id="F0EF501CFFC2D663FD88093308C3FA90" box="[534,673,1339,1365]" degrees="27.81791" direction="north" orientation="latitude" pageId="16" pageNumber="467" precision="1" value="27.81791">27.81791°N</geoCoordinate>
,
<geoCoordinate id="F0EF501CFFC2D663FD3309320957FA90" box="[685,821,1338,1365]" degrees="94.09502" direction="east" orientation="longitude" pageId="16" pageNumber="467" precision="1" value="94.09502">94.09502°E</geoCoordinate>
, h ~
<elevation id="1EF6D1E8FFC2D663FCF20933098DFA90" box="[876,1007,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="9.93" pageId="16" pageNumber="467" unit="m" value="993.0">
<quantity id="52239B3EFFC2D663FCF2093309D1FA90" box="[876,947,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="9.93" pageId="16" pageNumber="467" unit="m" value="993.0">993 m</quantity>
a.s.l.
</elevation>
,
<date id="E165101BFFC2D663FC6209330E14FA90" box="[1020,1142,1339,1365]" pageId="16" pageNumber="467" value="2017-06-14">
<collectingDate id="F121E9F3FFC2D663FC6209330E14FA90" box="[1020,1142,1339,1365]" pageId="16" pageNumber="467" value="2017-06-14">14.vi.2017</collectingDate>
</date>
,
<collectorName id="382E530DFFC2D663FB1C09330FFFFA90" box="[1154,1437,1339,1365]" pageId="16" pageNumber="467">Colls. K.A. Subramanian</collectorName>
&amp;
<collectorName id="382E530DFFC2D663FF2A09570B54FABC" box="[180,310,1375,1401]" pageId="16" pageNumber="467">M. Vasanth</collectorName>
—
<collectorName id="382E530DFFC2D663FECD09570BD6FABC" box="[339,436,1375,1401]" pageId="16" pageNumber="467">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFC2D663FE240957082BFABC" box="[442,585,1375,1401]" italics="true" pageId="16" pageNumber="467">SRC-I/E 510</emphasis>
[
<collectionCode id="F3CAAE1EFFC2D663FDC9095608EAFABC" box="[599,648,1374,1401]" pageId="16" pageNumber="467">ZSI</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC2D661FF59098A0BF9F993" lastPageId="18" lastPageNumber="469" pageId="16" pageNumber="467" type="description">
<paragraph id="956436DBFFC2D663FF59098A0F3FFA20" blockId="16.[151,1437,1230,1941]" pageId="16" pageNumber="467">
<emphasis id="A7AFEAC9FFC2D663FF59098A0B3BFA59" bold="true" box="[199,345,1410,1436]" pageId="16" pageNumber="467">Description.</emphasis>
Late larval instar: Length of body
<quantity id="52239B3EFFC2D663FD45098A0907FA58" box="[731,869,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="16" pageNumber="467" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.5–3.8 mm</quantity>
; cercus and median caudal filament subequal, each
<quantity id="52239B3EFFC2D663FF0909AE0B40FA04" box="[151,290,1446,1473]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="16" pageNumber="467" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
. General color light brown, with variable brown shading and markings; front of head, outer and inner margins of legs and lateral margins of abdominal terga covered with long, hair-like setae (
<figureCitation id="0DE02A5EFFC2D663FB1809C30E92FA20" box="[1158,1264,1483,1509]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Figs 11A</figureCitation>
,
<figureCitation id="0DE02A5EFFC2D663FB6209C30F2DFA20" box="[1276,1359,1483,1509]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="16" pageNumber="467">12A, B</figureCitation>
).
</paragraph>
<paragraph id="956436DBFFC2D663FF5909E70BD2F850" blockId="16.[151,1437,1230,1941]" pageId="16" pageNumber="467">
<emphasis id="A7AFEAC9FFC2D663FF5909E70B66F9CC" box="[199,260,1519,1545]" italics="true" pageId="16" pageNumber="467">Head</emphasis>
: Head with one pair of distinct, blunt occipital tubercles bear small short stout setae. The same scattered stout setae cover head. Color light brown with variable brown markings; frons pale; two dark brown, rounded spots situated between occipital protuberances and compound eyes. Areas around paired ocelli yellowish. Antennal scape and pedicel light brown; flagellar segments yellowish-brown, with fine, hair-like setae at apex of each segment; hair-like setae nearly one-third length of respective segment. Clypeus and genae densely covered with long hair-like setae (
<figureCitation id="0DE02A5EFFC2D663FF430AAB0B26F978" box="[221,324,1699,1725]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="16" pageNumber="467">Fig. 12A</figureCitation>
). Labrum (
<figureCitation id="0DE02A5EFFC2D663FE230AAB087DF978" box="[445,543,1699,1725]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Fig. 11B</figureCitation>
) brown; anterior notch shallow and wide; anterolateral angles weakly expressed, rounded; dorsal surface with dense, transverse row of cilia-like setae. Superlinguae of hypopharynx with row of setae on anterior margin; lingua with very sparse and tiny setae on dorsal and ventral surfaces, apex convex (
<figureCitation id="0DE02A5EFFC2D663FAB30AE30FEDF8C0" box="[1325,1423,1771,1797]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Fig. 11C</figureCitation>
). Mandible external margin with scattered hair-like setae basally and pair of long cilia-like setae medially (
<figureCitation id="0DE02A5EFFC2D663FA930B070FF2F8EC" box="[1293,1424,1807,1833]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Fig. 11D, E</figureCitation>
). Maxilla (
<figureCitation id="0DE02A5EFFC2D663FE9E0B3B0B00F888" box="[256,354,1843,1869]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Fig. 11F</figureCitation>
) with few distal hair-like setae; palp 3-segmented, with visible articulation; spinous processes short. Labium with cilia-like setae on ventral surface; segment III of maxillary palp rounded, about one-third length of segment II (
<figureCitation id="0DE02A5EFFC2D663FEA20B730BC1F850" box="[316,419,1915,1941]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="16" pageNumber="467">Fig. 11G</figureCitation>
).
</paragraph>
<caption id="C1A46653FFC3D662FF090BB50E3BF83C" ID-DOI="http://doi.org/10.5281/zenodo.4808355" ID-Zenodo-Dep="4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="17" pageNumber="468" startId="17.[151,250,1981,2005]" targetBox="[165,1421,181,1938]" targetPageId="17">
<paragraph id="956436DBFFC3D662FF090BB50E3BF83C" blockId="17.[151,1436,1981,2042]" pageId="17" pageNumber="468">
<emphasis id="A7AFEAC9FFC3D662FF090BB50B7AF810" bold="true" box="[151,280,1981,2005]" pageId="17" pageNumber="468">FIGURE 11</emphasis>
. Larva of
<taxonomicName id="52DB4D58FFC3D662FE1E0BB509FFF810" authority="Selvakumar, Martynov &amp; Jacobus" authorityName="Selvakumar, Martynov &amp; Jacobus" authorityYear="2021" box="[384,925,1981,2006]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC3D662FE1E0BB5085FF810" box="[384,573,1981,2005]" italics="true" pageId="17" pageNumber="468">Torleya dibruensis</emphasis>
Selvakumar, Martynov &amp; Jacobus
</taxonomicName>
,
<emphasis id="A7AFEAC9FFC3D662FC390BB60995F813" bold="true" box="[935,1015,1982,2006]" pageId="17" pageNumber="468">
<taxonomicNameLabel id="BC9C57B2FFC3D662FC390BB60995F813" box="[935,1015,1982,2006]" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, type specimens.
<emphasis id="A7AFEAC9FFC3D662FB350BB50EA6F810" bold="true" box="[1195,1220,1981,2005]" pageId="17" pageNumber="468">A.</emphasis>
—total view of larva;
<emphasis id="A7AFEAC9FFC3D662FF090BEA0ACCF83F" bold="true" box="[151,174,2018,2042]" pageId="17" pageNumber="468">B.</emphasis>
—labrum;
<emphasis id="A7AFEAC9FFC3D662FE830BE90B57F83C" bold="true" box="[285,309,2017,2041]" pageId="17" pageNumber="468">C.</emphasis>
—hypopharynx;
<emphasis id="A7AFEAC9FFC3D662FE7C0BEA087AF83F" bold="true" box="[482,536,2018,2042]" pageId="17" pageNumber="468">D, E.</emphasis>
—mandibles;
<emphasis id="A7AFEAC9FFC3D662FD390BEA08D8F83F" bold="true" box="[679,698,2018,2042]" pageId="17" pageNumber="468">F.</emphasis>
—maxilla;
<emphasis id="A7AFEAC9FFC3D662FCB30BE90925F83C" bold="true" box="[813,839,2017,2041]" pageId="17" pageNumber="468">G.</emphasis>
—labium. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="C1A46653FFC0D661FF090F2D080AFCA7" ID-DOI="http://doi.org/10.5281/zenodo.4808359" ID-Zenodo-Dep="4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469" startId="18.[151,250,805,829]" targetBox="[151,1436,181,780]" targetPageId="18">
<paragraph id="956436DBFFC0D661FF090F2D080AFCA7" blockId="18.[151,1436,805,866]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF090F2D0B7BFCF8" bold="true" box="[151,281,805,829]" pageId="18" pageNumber="469">FIGURE 12</emphasis>
. Larva of
<taxonomicName id="52DB4D58FFC0D661FE1D0F2E09C0FCFB" authority="Selvakumar, Martynov &amp; Jacobus" authorityName="Selvakumar, Martynov &amp; Jacobus" authorityYear="2021" box="[387,930,806,830]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC0D661FE1D0F2E0822FCFB" box="[387,576,806,830]" italics="true" pageId="18" pageNumber="469">Torleya dibruensis</emphasis>
Selvakumar, Martynov &amp; Jacobus
</taxonomicName>
,
<emphasis id="A7AFEAC9FFC0D661FC350F2E099EFCFB" bold="true" box="[939,1020,806,830]" pageId="18" pageNumber="469">
<taxonomicNameLabel id="BC9C57B2FFC0D661FC350F2E099EFCFB" box="[939,1020,806,830]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, type specimens.
<emphasis id="A7AFEAC9FFC0D661FB2F0F2D0EA1FCF8" bold="true" box="[1201,1219,805,829]" pageId="18" pageNumber="469">A</emphasis>
.—head, dorsal view;
<emphasis id="A7AFEAC9FFC0D661FF090F420ACCFCA7" bold="true" box="[151,174,842,866]" pageId="18" pageNumber="469">B.</emphasis>
—abdomen and part of thorax, dorsal view.
</paragraph>
</caption>
<paragraph id="956436DBFFC0D661FF590F980FEAFB47" blockId="18.[151,1437,912,1982]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF590F980B7AFC6F" box="[199,280,912,938]" italics="true" pageId="18" pageNumber="469">Thorax</emphasis>
: Pronotum without distinct protuberances. Mesonotum with several small, protuberances. Mesonotum with transverse brown shading medially (
<figureCitation id="0DE02A5EFFC0D661FDC30FBC08A0FC0B" box="[605,706,948,974]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Fig. 11A</figureCitation>
); tip of fore wingpads pale to white. Legs pale, distinctly flattened, especially fore femur (
<figureCitation id="0DE02A5EFFC0D661FE0E0FD0087AFC37" box="[400,536,984,1010]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13B–D</figureCitation>
). Dorsal surface of forefemur with irregular transverse row of long, pointed, stout setae proximally; outer and inner margins with rows of long hair-like setae and long, pointed, stout setae (
<figureCitation id="0DE02A5EFFC0D661FAAF0FF40ACEFBFF" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13A, B</figureCitation>
); dorsal surface of middle and hind femora smoky brown, with a few short stout setae; outer and inner margins with long, pointed, stout setae and long, hair-like setae (
<figureCitation id="0DE02A5EFFC0D661FC8F084C09FEFB9B" box="[785,924,1092,1118]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13C, D</figureCitation>
). Tibiae of all legs distinctly wider than tarsi. Each claw with 2–3 medial denticles, palisade of four to five long subdistal denticles, and subapical seta (
<figureCitation id="0DE02A5EFFC0D661FA8708600F19FB47" box="[1305,1403,1128,1154]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13E</figureCitation>
).
</paragraph>
<paragraph id="956436DBFFC0D661FF5908840BFCF9F7" blockId="18.[151,1437,912,1982]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF5908840B53FB63" box="[199,305,1164,1190]" italics="true" pageId="18" pageNumber="469">Abdomen</emphasis>
: Terga with dark longitudinal medial line and lateral shading (
<figureCitation id="0DE02A5EFFC0D661FC7108840E38FB63" box="[1007,1114,1164,1190]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Figs 11A</figureCitation>
,
<figureCitation id="0DE02A5EFFC0D661FBF808840EFAFB63" box="[1126,1176,1164,1190]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">12B</figureCitation>
). Dorsal lamella of gill III rounded, extending to middle of tergum VIII (
<figureCitation id="0DE02A5EFFC0D661FD2508B80942FB0F" box="[699,800,1200,1226]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">Fig. 12B</figureCitation>
), apically rounded, with fine distal setae, brown, trilobed pattern indistinct (
<figureCitation id="0DE02A5EFFC0D661FEFE08DC0BA0FB2B" box="[352,450,1236,1262]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13F</figureCitation>
); gill III operculate; ventral lamellae of gills III–VI bifurcate and multifoliate (
<figureCitation id="0DE02A5EFFC0D661FAB708DC0FF2FB2B" box="[1321,1424,1236,1262]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13G</figureCitation>
). Terga I–V and VIII–X mostly pale; terga VI–VII usually brown (
<figureCitation id="0DE02A5EFFC0D661FCED08F009B8FAD7" box="[883,986,1272,1298]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">Fig. 12B</figureCitation>
). Terga V–IX with spatulate, stout setae on posterior margins (setae progressively shorter on segments VI–VIII); terga VII and VIII with hair-like and spatulate, stout setae; terga IV–VII and IX with paired posterior protuberances, bears small stout setae with rounded apices; terga VIII and X without paired posterior protuberances; posterior protuberances very small on tergum IV and IX; largest on terga V–VII, usually bearing 4–6 short spatulate stout setae; Lateral parts of posterior margin of tergum VIII with row of spatulate, stout setae. Segments IV–IX with posterolateral projections. Segment IX distinctly narrower than segment VIII (
<figureCitation id="0DE02A5EFFC0D661FDD509D808D3FA2F" box="[587,689,1488,1514]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">Fig. 12B</figureCitation>
). Sterna pale, with few short, hair-like setae. Caudal filaments pale to white, with broad, dark brown band medially; tips dark brown; apex of each segment with spatulate, stout setae and fine hair-like setae.
</paragraph>
<paragraph id="956436DBFFC0D661FF590A340BF9F993" blockId="18.[151,1437,912,1982]" box="[199,411,1596,1622]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF590A340B7CF993" bold="true" box="[199,286,1596,1622]" pageId="18" pageNumber="469">Adults.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC0D661FF590A680BF3F85F" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="956436DBFFC0D661FF590A680BF3F85F" blockId="18.[151,1437,912,1982]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF590A680B20F9BF" bold="true" box="[199,322,1632,1658]" pageId="18" pageNumber="469">Diagnosis.</emphasis>
Larvae of
<taxonomicName id="52DB4D58FFC0D661FE230A6808ECF9BF" authority="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus, 2021" authorityName="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus" authorityYear="2021" box="[445,654,1632,1658]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC0D661FE230A6808ECF9BF" box="[445,654,1632,1658]" italics="true" pageId="18" pageNumber="469">Torleya dibruensis</emphasis>
</taxonomicName>
<emphasis id="A7AFEAC9FFC0D661FD0B0A68088DF9BF" bold="true" box="[661,751,1632,1658]" pageId="18" pageNumber="469">
<taxonomicNameLabel id="BC9C57B2FFC0D661FD0B0A68088DF9BF" box="[661,751,1632,1658]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other representatives of the genus by the following combination of characters: (i) head with one pair of blunt occipital tubercles; (ii) head, legs and abdomen with areas densely covered with long, hair-like setae (
<figureCitation id="0DE02A5EFFC0D661FCE90AA00986F907" box="[887,996,1704,1730]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Figs 11A</figureCitation>
,
<figureCitation id="0DE02A5EFFC0D661FC6D0AA00E2BF907" box="[1011,1097,1704,1730]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">12A, B</figureCitation>
,
<figureCitation id="0DE02A5EFFC0D661FBC90AA00ECCF907" box="[1111,1198,1704,1730]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">13A–D</figureCitation>
); (iii) claw with 2–3 medial denticles, palisade of four to five long subdistal denticles on the inner margin, and subapical seta (
<figureCitation id="0DE02A5EFFC0D661FAB40AC40FECF923" box="[1322,1422,1740,1766]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13E</figureCitation>
); (iv) terga V–VII with paired blunt protuberances, usually bearing four to six short spatulate, stout setae (
<figureCitation id="0DE02A5EFFC0D661FAB60AF80FEDF8CF" box="[1320,1423,1776,1802]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Fig. 11A</figureCitation>
); (v) gills rounded, gill III entirely cover following gills (
<figureCitation id="0DE02A5EFFC0D661FD670B1C0900F8EB" box="[761,866,1812,1838]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Figs 11A</figureCitation>
,
<figureCitation id="0DE02A5EFFC0D661FCF30B1C09FCF8EB" box="[877,926,1812,1838]" captionStart="FIGURE 12" captionStartId="18.[151,250,805,829]" captionTargetBox="[151,1436,181,780]" captionTargetId="figure-19@18.[151,1436,181,781]" captionTargetPageId="18" captionText="FIGURE 12. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—head, dorsal view; B.—abdomen and part of thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4808359" httpUri="https://zenodo.org/record/4808359/files/figure.png" pageId="18" pageNumber="469">12B</figureCitation>
,
<figureCitation id="0DE02A5EFFC0D661FC360B1C0994F8EB" box="[936,1014,1812,1838]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">13F, G</figureCitation>
); (vi) foreleg with irregular transverse row of long pointed, stout setae at proximal part of dorsal surface; same setae presented on outer and inner margins (
<figureCitation id="0DE02A5EFFC0D661FF3E0B540B4AF8B3" box="[160,296,1884,1910]" captionStart="FIGURE 13" captionStartId="19.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure-20@19.[151,1436,181,1563]" captionTargetPageId="19" captionText="FIGURE 13. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—setation of basal half of fore femur; B.—fore leg; C.—middle leg; D.—hind leg; E.—tarsal claw; F.—gill III; G.—gill IV." figureDoi="http://doi.org/10.5281/zenodo.4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="18" pageNumber="469">Fig. 13A, B</figureCitation>
); (vii) maxillary palp present, short (
<figureCitation id="0DE02A5EFFC0D661FD590B540948F8B3" box="[711,810,1884,1910]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Fig. 11F</figureCitation>
); (viii) labial palp segment III about one-third length of segment II (
<figureCitation id="0DE02A5EFFC0D661FE830B880BE7F85F" box="[285,389,1920,1946]" captionStart="FIGURE 11" captionStartId="17.[151,250,1981,2005]" captionTargetBox="[165,1421,181,1938]" captionTargetId="figure-20@17.[165,1421,181,1957]" captionTargetPageId="17" captionText="FIGURE 11. Larva of Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov., type specimens. A.—total view of larva; B.—labrum; C.—hypopharynx; D, E.—mandibles; F.—maxilla; G.—labium. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808355" httpUri="https://zenodo.org/record/4808355/files/figure.png" pageId="18" pageNumber="469">Fig. 11G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC0D661FF590BAC0EDBF87B" box="[199,1209,1956,1982]" pageId="18" pageNumber="469" type="etymology">
<paragraph id="956436DBFFC0D661FF590BAC0EDBF87B" blockId="18.[151,1437,912,1982]" box="[199,1209,1956,1982]" pageId="18" pageNumber="469">
<emphasis id="A7AFEAC9FFC0D661FF590BAC0B2CF87B" bold="true" box="[199,334,1956,1982]" pageId="18" pageNumber="469">Etymology.</emphasis>
This species is named after the
<typeStatus id="4A608879FFC0D661FD2A0BAC0886F87B" box="[692,740,1956,1982]" pageId="18" pageNumber="469">type</typeStatus>
locality, Dibru River,
<collectingRegion id="571FF839FFC0D661FC430BAC0ED7F87B" box="[989,1205,1956,1982]" country="India" name="Arunachal Pradesh" pageId="18" pageNumber="469">Arunachal Pradesh</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="C1A46653FFC1D660FF090A3B0EA1F9AB" ID-DOI="http://doi.org/10.5281/zenodo.4808363" ID-Zenodo-Dep="4808363" httpUri="https://zenodo.org/record/4808363/files/figure.png" pageId="19" pageNumber="470" startId="19.[151,250,1587,1611]" targetBox="[151,1436,181,1560]" targetPageId="19">
<paragraph id="956436DBFFC1D660FF090A3B0EA1F9AB" blockId="19.[151,1436,1587,1647]" pageId="19" pageNumber="470">
<emphasis id="A7AFEAC9FFC1D660FF090A3B0B78F98E" bold="true" box="[151,282,1587,1611]" pageId="19" pageNumber="470">FIGURE 13</emphasis>
. Larva of
<taxonomicName id="52DB4D58FFC1D660FE160A3B09CFF98E" authority="Selvakumar, Martynov &amp; Jacobus" authorityName="Selvakumar, Martynov &amp; Jacobus" authorityYear="2021" box="[392,941,1587,1611]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC1D660FE160A3B0824F98E" box="[392,582,1587,1611]" italics="true" pageId="19" pageNumber="470">Torleya dibruensis</emphasis>
Selvakumar, Martynov &amp; Jacobus
</taxonomicName>
,
<emphasis id="A7AFEAC9FFC1D660FC260A3B0E68F98E" bold="true" box="[952,1034,1587,1611]" pageId="19" pageNumber="470">
<taxonomicNameLabel id="BC9C57B2FFC1D660FC260A3B0E68F98E" box="[952,1034,1587,1611]" pageId="19" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, type specimens.
<emphasis id="A7AFEAC9FFC1D660FB5D0A3B0EBEF98E" bold="true" box="[1219,1244,1587,1611]" pageId="19" pageNumber="470">A.</emphasis>
—setation of basal half of fore femur;
<emphasis id="A7AFEAC9FFC1D660FEC70A5F0B12F9AA" bold="true" box="[345,368,1623,1647]" pageId="19" pageNumber="470">B.</emphasis>
—fore leg;
<emphasis id="A7AFEAC9FFC1D660FE7B0A5F0B9CF9AA" bold="true" box="[485,510,1623,1647]" pageId="19" pageNumber="470">C.</emphasis>
—middle leg;
<emphasis id="A7AFEAC9FFC1D660FD0E0A5F08CBF9AA" bold="true" box="[656,681,1623,1647]" pageId="19" pageNumber="470">D.</emphasis>
—hind leg;
<emphasis id="A7AFEAC9FFC1D660FCBC0A5F095BF9AA" bold="true" box="[802,825,1623,1647]" pageId="19" pageNumber="470">E.</emphasis>
—tarsal claw;
<emphasis id="A7AFEAC9FFC1D660FC530A5F0982F9AA" bold="true" box="[973,992,1623,1647]" pageId="19" pageNumber="470">F.</emphasis>
—gill III;
<emphasis id="A7AFEAC9FFC1D660FBD70A5F0E01F9AA" bold="true" box="[1097,1123,1623,1647]" pageId="19" pageNumber="470">G.</emphasis>
—gill IV.
</paragraph>
</caption>
<subSubSection id="DDC16550FFC1D660FF590A950B90F93A" pageId="19" pageNumber="470" type="distribution">
<paragraph id="956436DBFFC1D660FF590A9508F1F972" blockId="19.[151,1437,1693,1900]" box="[199,659,1693,1719]" pageId="19" pageNumber="470">
<emphasis id="A7AFEAC9FFC1D660FF590A950B03F972" bold="true" box="[199,353,1693,1719]" pageId="19" pageNumber="470">Distribution.</emphasis>
India-China border region.
</paragraph>
<paragraph id="956436DBFFC1D660FF590AC90B90F93A" blockId="19.[151,1437,1693,1900]" pageId="19" pageNumber="470">
<emphasis id="A7AFEAC9FFC1D660FF590AC90B41F91E" bold="true" box="[199,291,1729,1755]" pageId="19" pageNumber="470">Habitat</emphasis>
. Larvae of the species were collected in rivers and large streams with cobble and sandy bottom and numerous boulders (
<figureCitation id="0DE02A5EFFC1D660FEE20AED0B84F93A" box="[380,486,1765,1791]" captionStart="FIGURE 20" captionStartId="28.[151,250,1070,1094]" captionTargetBox="[151,1436,181,1043]" captionTargetId="figure-19@28.[151,1436,181,1046]" captionTargetPageId="28" captionText="FIGURE 20. Habitats of Torleya simbalbarensis Selvakumar, Subramanian, Martynov &amp; Jacobus, sp. nov. (A), Serratella palatovi Martynov, Selvakumar &amp; Jacobus, sp. nov. (B, C) and Torleya dibruensis Selvakumar, Martynov &amp; Jacobus, sp. nov. (D). A.—stream in Simbalbara Wildlife Sanctuary, Sirmour District, Himachal Pradesh, India; B.—stream-main source of the Klang Phat River, Chom Thong District, Chiang Mai Province, Thailand; C.—stream in Tale Valley, Lower Subansiri District, Arunachal Pradesh, India; D.—Dibru River, Papum Pare District, Arunachal Pradesh, India." figureDoi="http://doi.org/10.5281/zenodo.4808393" httpUri="https://zenodo.org/record/4808393/files/figure.png" pageId="19" pageNumber="470">Fig. 20D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC1D660FF590B010852F8AE" pageId="19" pageNumber="470" type="discussion">
<paragraph id="956436DBFFC1D660FF590B010852F8AE" blockId="19.[151,1437,1693,1900]" pageId="19" pageNumber="470">
<emphasis id="A7AFEAC9FFC1D660FF590B010B59F8E6" bold="true" box="[199,315,1801,1827]" pageId="19" pageNumber="470">Remarks.</emphasis>
<taxonomicName id="52DB4D58FFC1D660FEDF0B01087DF8E6" baseAuthorityName="Gui, Zhou &amp; Su" baseAuthorityYear="1999" box="[321,543,1801,1827]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="longforceps">
<emphasis id="A7AFEAC9FFC1D660FEDF0B01087DF8E6" box="[321,543,1801,1827]" italics="true" pageId="19" pageNumber="470">Torleya longforceps</emphasis>
</taxonomicName>
from
<collectingRegion id="571FF839FFC1D660FDFF0B0108CEF8E6" box="[609,684,1801,1827]" country="China" name="Fujian" pageId="19" pageNumber="470">Fujian</collectingRegion>
, in far eastern
<collectingCountry id="EDCC764BFFC1D660FCD30B0109F2F8E6" box="[845,912,1801,1827]" name="China" pageId="19" pageNumber="470">China</collectingCountry>
(
<bibRefCitation id="F14A4B2AFFC1D660FC030B010E2AF8E6" author="Gui, H. &amp; Zhou, C. - F. &amp; Su, C. - R." box="[925,1096,1801,1827]" pageId="19" pageNumber="470" refId="ref14572" refString="Gui, H., Zhou, C. - F. &amp; Su, C. - R. (1999) Ephemeroptera. In: Huang, B. - K. (Ed.), Fujian Insect Fauna. Vol. 1. Fujian Science &amp; Technology Press, Fuzhou, pp. 324 - 346." type="book" year="1999">
Gui
<emphasis id="A7AFEAC9FFC1D660FC510B020E64F8E6" box="[975,1030,1801,1827]" italics="true" pageId="19" pageNumber="470">et al.</emphasis>
1999
</bibRefCitation>
), is the only species of
<taxonomicName id="52DB4D58FFC1D660FAD60B010FFFF8E6" authorityName="Lestage" authorityYear="1917" box="[1352,1437,1801,1827]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC1D660FAD60B010FFFF8E6" box="[1352,1437,1801,1827]" italics="true" pageId="19" pageNumber="470">Torleya</emphasis>
</taxonomicName>
unknown in the larval stage, and we consider it unlikely (based on biogeography) to be conspecific with this new species, which is unknown as alates.
</paragraph>
</subSubSection>
</treatment>
</document>
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