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<document id="EB5C4FA2E1769E3CF3E0FA80D18B297B" ID-DOI="10.1649/582" ID-ISSN="1938-4394" ID-Zenodo-Dep="10103782" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630163904379" checkinUser="tatiana" docAuthor="Stebnicka, Zdzisława Teresa" docDate="2003" docId="03EA6A15B10F97523C5AFB1061454225" docLanguage="en" docName="582.pdf.imf" docOrigin="The Coleopterists Bulletin 57 (4)" docSource="http://dx.doi.org/10.1649/582" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Saprositellus ariquemes Stebnicka 2003, new species" docType="treatment" docVersion="3" lastPageNumber="455" masterDocId="FFD3126DB10D97563D3FFFD462174367" masterDocTitle="The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini)" masterLastPageNumber="457" masterPageNumber="451" pageNumber="453" updateTime="1699627149666" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="18B8FC0BB8F37679CC97F14D0A9F98C5">The Genus Saprositellus Balthasar, with Descriptions of Three New Neotropical Species (Coleoptera: Scarabaeidae: Odontolochini)</mods:title>
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<mods:namePart id="ACBE70B2C1F7D8CCCC93FC81D34EBA03">Stebnicka, Zdzisława Teresa</mods:namePart>
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<mods:title id="82342136F359D1BB3F3D411530A9F152">The Coleopterists Bulletin</mods:title>
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<mods:part id="CF74CD48310F892A4662CB8A37C0926E">
<mods:date id="54372F55327B190D9DA2EE0C18E1FEB6">2003</mods:date>
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<mods:number id="B94F464F0683C51D8A6B5E90E9B75CD0">2003-12-31</mods:number>
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<mods:number id="7C86764EEE0380715E344BF01FF04750">57</mods:number>
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<mods:number id="EF12DA6C756F414324C4C0A8517B23AD">4</mods:number>
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<mods:classification id="B3BABE9BAA1309F487EE014C20E2BAA8">journal article</mods:classification>
<mods:identifier id="EB37A2EDA2931666A5BA9ADB2002D5D0" type="DOI">10.1649/582</mods:identifier>
<mods:identifier id="85B2BE2A51302116E572E04307C1C8E8" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03EA6A15B10F97523C5AFB1061454225" LSID="urn:lsid:plazi:treatment:03EA6A15B10F97523C5AFB1061454225" httpUri="http://treatment.plazi.org/id/03EA6A15B10F97523C5AFB1061454225" lastPageId="4" lastPageNumber="455" pageId="2" pageNumber="453">
<subSubSection id="C3598888B10F97543C5AFB10612147BF" box="[357,822,1219,1241]" pageId="2" pageNumber="453" type="nomenclature">
<paragraph id="8BFCDB03B10F97543C5AFB10612147BF" blockId="2.[357,822,1219,1267]" box="[357,822,1219,1241]" pageId="2" pageNumber="453">
<heading id="D0B46C6FB10F97543C5AFB10612147BF" box="[357,822,1219,1241]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="453" reason="2">
<taxonomicName id="4C43A080B10F97543C5AFB1060A347BF" authority="Stebnicka" authorityName="Stebnicka" authorityYear="2003" box="[357,692,1219,1241]" class="Insecta" family="Aphodiidae" genus="Saprositellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="ariquemes" status="sp. nov.">
<emphasis id="B9370711B10F97543C5AFB10605B47BE" box="[357,588,1220,1241]" italics="true" pageId="2" pageNumber="453">Saprositellus ariquemes</emphasis>
Stebnicka
</taxonomicName>
,
<emphasis id="B9370711B10F97543F80FB17612147BF" bold="true" box="[703,822,1219,1240]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A204BA6AB10F97543F80FB17612147BF" box="[703,822,1219,1240]" pageId="2" pageNumber="453" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3598888B10F97553F33FB0A63544545" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="description">
<paragraph id="8BFCDB03B10F97543F33FB0A60874794" blockId="2.[357,822,1219,1267]" box="[524,656,1246,1267]" pageId="2" pageNumber="453">
(
<figureCitation id="1378C786B10F97543F2CFB0A604F4794" box="[531,600,1246,1267]" captionStart="Figs" captionStartId="2.[197,236,922,941]" captionTargetBox="[179,1012,197,901]" captionTargetId="figure-236@2.[175,1013,194,901]" captionTargetPageId="2" captionText="Figs. 14. 1) Saprositellus denticulatus Balthasar, habitus; 24) left half of body; 2) S. ariquemes n. sp.; 3) S. santaritae n. sp. 4) S. peruanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.10103784" httpUri="https://zenodo.org/record/10103784/files/figure.png" pageId="2" pageNumber="453">Figs. 2</figureCitation>
,
<figureCitation id="1378C786B10F97543F5BFB0A609E4794" box="[612,649,1246,1267]" captionStart="Figs" captionStartId="3.[169,208,1052,1071]" captionTargetBox="[170,955,195,1030]" captionTargetId="figure-208@3.[170,955,194,1030]" captionTargetPageId="3" captionText="Figs. 58. Saprositellus denticulatus Balthasar, right posterior leg; 67) S. ariquemes n. sp.;" figureDoi="http://doi.org/10.5281/zenodo.10103786" httpUri="https://zenodo.org/record/10103786/files/figure.png" pageId="2" pageNumber="453">67</figureCitation>
)
</paragraph>
<paragraph id="8BFCDB03B10F97543DFAFADC61F94545" blockId="2.[175,1006,1288,1570]" pageId="2" pageNumber="453">
<emphasis id="B9370711B10F97543DFAFADC6324467C" bold="true" box="[197,307,1288,1307]" pageId="2" pageNumber="453">Description.</emphasis>
Length
<quantity id="4CBB76E6B10F97543C40FADC63FE467C" box="[383,489,1288,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="2" pageNumber="453" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.62.8 mm</quantity>
, greatest width 1.0 mm. Form (
<figureCitation id="1378C786B10F97543FC7FADC613C467C" box="[760,811,1288,1307]" captionStart="Figs" captionStartId="2.[197,236,922,941]" captionTargetBox="[179,1012,197,901]" captionTargetId="figure-236@2.[175,1013,194,901]" captionTargetPageId="2" captionText="Figs. 14. 1) Saprositellus denticulatus Balthasar, habitus; 24) left half of body; 2) S. ariquemes n. sp.; 3) S. santaritae n. sp. 4) S. peruanus n. sp." figureDoi="http://doi.org/10.5281/zenodo.10103784" httpUri="https://zenodo.org/record/10103784/files/figure.png" pageId="2" pageNumber="453">Fig. 2</figureCitation>
) elongate oval, widest at apical third of elytra; color dark brown, legs reddish brown, surface microreticulate. Clypeus truncate anteriorly, edge on each side with 67 denticles, surface just above shining and here minutely transversely granulate; remaining surface of head with fine, very close, nearly contiguous punctures; frontal suture marked by slightly thickened line; gena small rounded. Pronotum strongly convex, side slightly arcuate, finely margined, lateral tooth acutely prominent, posterior angle excised, base sinuate, finely crenate with fine marginal line; pronotal disc with shallow longitudinal furrow and small fovea laterally, surface punctures everywhere very close, almost contiguous, rugose. Elytra distinctly arcuate, apex with minute setae visible under high magnification; base as wide as pronotal base, humeral denticle strong, directed upward; striae deep, slightly wider than intervals with deep, coarse punctures crenating inner margins of intervals, punctures separate each other by about 1/3 of their diameter; intervals carinate, microreticulate.
</paragraph>
<caption id="DF3C8B8BB10E97553D96FBC861C64748" ID-DOI="http://doi.org/10.5281/zenodo.10103786" ID-Zenodo-Dep="10103786" box="[169,977,1052,1071]" httpUri="https://zenodo.org/record/10103786/files/figure.png" pageId="3" pageNumber="454" startId="3.[169,208,1052,1071]" targetBox="[170,955,195,1030]" targetPageId="3">
<paragraph id="8BFCDB03B10E97553D96FBC861C64748" blockId="3.[148,977,1052,1095]" box="[169,977,1052,1071]" pageId="3" pageNumber="454">
<emphasis id="B9370711B10E97553D96FBC863174748" bold="true" box="[169,256,1052,1071]" pageId="3" pageNumber="454">Figs. 58.</emphasis>
<taxonomicName id="4C43A080B10E97553C2AFBC8605F4748" authority="Balthasar" authorityName="Balthasar" authorityYear="1967" box="[277,584,1052,1071]" class="Insecta" family="Aphodiidae" genus="Saprositellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B9370711B10E97553C2AFBC863E64748" box="[277,497,1052,1071]" italics="true" pageId="3" pageNumber="454">Saprositellus denticulatus</emphasis>
Balthasar
</taxonomicName>
, right posterior leg;
<emphasis id="B9370711B10E97553FC6FBC861374748" bold="true" box="[761,800,1052,1071]" pageId="3" pageNumber="454">67)</emphasis>
<taxonomicName id="4C43A080B10E97553E18FBC861824748" authorityName="Stebnicka" authorityYear="2003" box="[807,917,1052,1071]" class="Insecta" family="Aphodiidae" genus="Saprositellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="ariquemes" status="sp. nov.">
<emphasis id="B9370711B10E97553E18FBC861824748" box="[807,917,1052,1071]" italics="true" pageId="3" pageNumber="454">S. ariquemes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204BA6AB10E97553EA3FBC861D94748" box="[924,974,1052,1071]" pageId="3" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
;
</paragraph>
</caption>
<paragraph id="8BFCDB03B10E97553DABFBE7618E4720" blockId="3.[148,977,1052,1095]" box="[148,921,1075,1095]" pageId="3" pageNumber="454">
<emphasis id="B9370711B10E97553DABFBE762B14721" bold="true" box="[148,166,1075,1094]" pageId="3" pageNumber="454">6)</emphasis>
epipharynx;
<emphasis id="B9370711B10E97553C27FBE7633E4721" bold="true" box="[280,297,1075,1094]" pageId="3" pageNumber="454">7)</emphasis>
male genitalia in lateral view;
<emphasis id="B9370711B10E97553F07FBE7605D4721" bold="true" box="[568,586,1075,1094]" pageId="3" pageNumber="454">8)</emphasis>
<taxonomicName id="4C43A080B10E97553F6FFBE060A14720" authorityName="Stebnicka" authorityYear="2003" box="[592,694,1076,1095]" class="Insecta" family="Aphodiidae" genus="Saprositellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="peruanus" status="sp. nov.">
<emphasis id="B9370711B10E97553F6FFBE060A14720" box="[592,694,1076,1095]" italics="true" pageId="3" pageNumber="454">S. peruanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204BA6AB10E97553F82FBE060F94720" box="[701,750,1076,1095]" pageId="3" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
, right posterior leg.
</paragraph>
<paragraph id="8BFCDB03B10E97553DABFBAC61C6461C" blockId="3.[148,978,1144,1450]" pageId="3" pageNumber="454">
Ventral sclerites alutaceous, surface punctate, punctures gradually decreasing in size from mesosternum to penultimate sternite, generally separated by about one diameter; prosternal process widely triangular, punctate; mesosternum lower than metasternum, mestasternal midline shallow, lateral metasternal triangle marked by very shallow concavity; abdominal sternites finely fluted along sutures, punctate from side to side; disc of pygidium with fine punctures same size as those on penultimate sternite. Profemur wide, perimarginal groove deep, surface punctures same size as those of penultimate sternite; meso- and metafemora narrow, slender, posterior femoral lines strong, complete, posterior edge slightly sinuate; meso- and metatibiae as long as femora, slightly sinuate, minutely and scarcely setigerous; metatibia with fine external spine and seta-like apical spurs, tarsus short, segments subquadrate; basal tarsomere of metatarsus two times shorter than upper tibial spur and equal in length to second tarsomere. Epipharyngeal structures as in
<figureCitation id="1378C786B10E97553EBBFABC61D9461C" box="[900,974,1384,1403]" captionStart="Figs" captionStartId="3.[169,208,1052,1071]" captionTargetBox="[170,955,195,1030]" captionTargetId="figure-208@3.[170,955,194,1030]" captionTargetPageId="3" captionText="Figs. 58. Saprositellus denticulatus Balthasar, right posterior leg; 67) S. ariquemes n. sp.;" figureDoi="http://doi.org/10.5281/zenodo.10103786" httpUri="https://zenodo.org/record/10103786/files/figure.png" pageId="3" pageNumber="454">Figure 6</figureCitation>
.
</paragraph>
<paragraph id="8BFCDB03B10E97553D96FAAB63EC46CD" blockId="3.[148,978,1144,1450]" pageId="3" pageNumber="454">
In the male, the penultimate abdominal sternite is shorter than in the female, and the disc of pygidium longer; genitalia as in
<figureCitation id="1378C786B10E97553C92FA4363E046CD" box="[429,503,1431,1450]" captionStart="Figs" captionStartId="3.[169,208,1052,1071]" captionTargetBox="[170,955,195,1030]" captionTargetId="figure-208@3.[170,955,194,1030]" captionTargetPageId="3" captionText="Figs. 58. Saprositellus denticulatus Balthasar, right posterior leg; 67) S. ariquemes n. sp.;" figureDoi="http://doi.org/10.5281/zenodo.10103786" httpUri="https://zenodo.org/record/10103786/files/figure.png" pageId="3" pageNumber="454">Figure 7</figureCitation>
.
</paragraph>
<paragraph id="8BFCDB03B10E97553D96FA6963544545" blockId="3.[148,977,1469,1570]" pageId="3" pageNumber="454">
<emphasis id="B9370711B10E97553D96FA6962E846B5" bold="true" box="[169,255,1469,1490]" pageId="3" pageNumber="454">Affinity.</emphasis>
The new species is most similar to
<taxonomicName id="4C43A080B10E97553F58FA6A614846B4" authorityName="Balthasar" authorityYear="1967" box="[615,863,1470,1491]" class="Insecta" family="Aphodiidae" genus="Saprositellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B9370711B10E97553F58FA6A614846B4" box="[615,863,1470,1491]" italics="true" pageId="3" pageNumber="454">Saprositellus denticulatus</emphasis>
</taxonomicName>
, but differs from that species by having the pronotal punctures denser and pronotal base margined, the elytral humeral denticles significantly larger and the elytral striae wider, with coarser punctures.
</paragraph>
</subSubSection>
<subSubSection id="C3598888B10997523DFAFF16618A4240" pageId="4" pageNumber="455" type="materials_examined">
<paragraph id="8BFCDB03B10997523DFAFF16618A4240" blockId="4.[175,1005,194,322]" pageId="4" pageNumber="455">
<emphasis id="B9370711B10997523DFAFF16634843B0" bold="true" box="[197,351,194,215]" pageId="4" pageNumber="455">Type Material.</emphasis>
<materialsCitation id="3B2BD15EB10997523C54FF16619A4395" collectingDate="1994-11-08" collectingDateMax="1994-11-20" collectingDateMin="1994-11-08" collectionCode="FSCA" collectorName="J. Eger &amp; C. O'Brien" country="Brazil" location="Fzda Rancho Grande" pageId="4" pageNumber="455" specimenCount="1" stateProvince="Rondonia" typeStatus="holotype">
<typeStatus id="54F865A1B10997523C54FF1663D443B0" box="[363,451,194,215]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5243C6B10997523F39FF16605143B0" box="[518,582,194,215]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="455" type="Museum">FSCA</collectionCode>
),
<collectingCountry id="F3549B93B10997523F5EFF1660B643B0" box="[609,673,194,215]" name="Brazil" pageId="4" pageNumber="455">Brazil</collectingCountry>
,
<collectingRegion id="498715E1B10997523F94FF16611D43B0" box="[683,778,194,215]" country="Brazil" name="Rondonia" pageId="4" pageNumber="455">Rondonia</collectingRegion>
,
<locationDeviation id="9A4EC221B10997523E2AFF1661FB43B0" box="[789,1004,194,215]" location="Ariquemes" pageId="4" pageNumber="455">
<quantity id="4CBB76E6B10997523E2AFF16614543B0" box="[789,850,194,215]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="455" unit="km" value="62.0">62 km</quantity>
SW Ariquemes
</locationDeviation>
nr
<location id="8E9C8DD8B10997523DF3FF09638A4395" LSID="urn:lsid:plazi:treatment:03EA6A15B10F97523C5AFB1061454225:8E9C8DD8B10997523DF3FF09638A4395" box="[204,413,221,242]" country="Brazil" name="Fzda Rancho Grande" pageId="4" pageNumber="455" stateProvince="Rondonia">Fzda Rancho Grande</location>
,
<date id="FFFDFDC3B10997523C95FF0960264395" box="[426,561,221,242]" pageId="4" pageNumber="455" value="1994-11-08" valueMax="1994-11-20" valueMin="1994-11-08">
<collectingDate id="EFB9042BB10997523C95FF0960264395" box="[426,561,221,242]" pageId="4" pageNumber="455" value="1994-11-08" valueMax="1994-11-20" valueMin="1994-11-08">820.XI.1994</collectingDate>
</date>
, black light,
<collectorName id="26B6BED5B10997523F8BFF0960ED4395" box="[692,762,221,242]" pageId="4" pageNumber="455">J. Eger</collectorName>
&amp;
<collectorName id="26B6BED5B10997523E21FF09619A4395" box="[798,909,221,242]" pageId="4" pageNumber="455">C. OBrien</collectorName>
</materialsCitation>
.
<materialsCitation id="3B2BD15EB10997523EA5FF09602E426A" collectingDate="1994-11-08" collectingDateMax="1994-11-20" collectingDateMin="1994-11-08" collectionCode="FSCA" collectorName="J. Eger &amp; C. O'Brien" country="Brazil" location="Fzda Rancho Grande" pageId="4" pageNumber="455" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<typeStatus id="54F865A1B10997523EA5FF0961FA4395" box="[922,1005,221,242]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
female, same data as holotype (
<collectionCode id="ED5243C6B10997523CCEFF2C6025426A" box="[497,562,248,269]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="455" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2BD15EB10997523F7BFF2C618E4240" collectingDate="1998-02-20" collectionCode="ISEA" collectorName="L. Stange'" country="Bolivia" elevation="380" latitude="-7.27" location="Buena Vista" longLatPrecision="782" longitude="-63.39" pageId="4" pageNumber="455" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54F865A1B10997523F7BFF2C6080426A" box="[580,663,248,269]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
male
<collectingCountry id="F3549B93B10997523FE1FF2C613D426A" box="[734,810,248,269]" name="Bolivia" pageId="4" pageNumber="455">Bolivia</collectingCountry>
,
<collectingRegion id="498715E1B10997523E09FF2C61B4426A" box="[822,931,248,269]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="455">Santa Cruz</collectingRegion>
,
<location id="8E9C8DD8B10997523E8FFF2C62F64240" LSID="urn:lsid:plazi:treatment:03EA6A15B10F97523C5AFB1061454225:8E9C8DD8B10997523E8FFF2C62F64240" country="Bolivia" latitude="-7.27" longLatPrecision="782" longitude="-63.39" name="Buena Vista" pageId="4" pageNumber="455" stateProvince="Santa Cruz">Buena Vista</location>
<quantity id="4CBB76E6B10997523DD6FEC663304240" box="[233,295,274,295]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="455" unit="m" value="380.0">
<elevation id="006E3C30B10997523DD6FEC663304240" box="[233,295,274,295]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="455" unit="m" value="380.0">380 m</elevation>
</quantity>
elev.,
<geoCoordinate id="EE77BDC4B10997523C55FEC663B44240" box="[362,419,274,295]" degrees="7.27" direction="south" orientation="latitude" pageId="4" pageNumber="455" precision="555" value="-7.27">7.27S</geoCoordinate>
,
<geoCoordinate id="EE77BDC4B10997523C90FEC663E94240" box="[431,510,274,295]" degrees="63.39" direction="west" orientation="longitude" pageId="4" pageNumber="455" precision="555" value="-63.39">63.39W</geoCoordinate>
,
<date id="FFFDFDC3B10997523F36FEC660794240" box="[521,622,274,295]" pageId="4" pageNumber="455" value="1998-02-20">
<collectingDate id="EFB9042BB10997523F36FEC660794240" box="[521,622,274,295]" pageId="4" pageNumber="455" value="1998-02-20">20.II.1998</collectingDate>
</date>
,
<collectingMethod id="5202A314B10997523F45FEC660C34240" box="[634,724,274,295]" pageId="4" pageNumber="455">light trap</collectingMethod>
,
<collectorName id="26B6BED5B10997523FDFFEC661514240" box="[736,838,274,295]" pageId="4" pageNumber="455">L. Stange</collectorName>
(
<collectionCode id="ED5243C6B10997523E66FEC661854240" box="[857,914,274,295]" pageId="4" pageNumber="455">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3598888B10997523DFAFEF861454225" box="[197,850,300,322]" pageId="4" pageNumber="455" type="etymology">
<paragraph id="8BFCDB03B10997523DFAFEF861454225" blockId="4.[175,1005,194,322]" box="[197,850,300,322]" pageId="4" pageNumber="455">
<emphasis id="B9370711B10997523DFAFEF8632E4226" bold="true" box="[197,313,300,321]" pageId="4" pageNumber="455">Etymology.</emphasis>
Derived from the name of
<typeStatus id="54F865A1B10997523F75FEF960644225" box="[586,627,301,322]" pageId="4" pageNumber="455">type</typeStatus>
locality Ariquemes.
</paragraph>
</subSubSection>
</treatment>
</document>