449 lines
55 KiB
XML
449 lines
55 KiB
XML
<document id="E31DA5BEA79D4262C453A05695F62300" ID-CLB-Dataset="46246" ID-DOI="10.11646/zootaxa.3774.6.3" ID-GBIF-Dataset="5485c5f1-36b6-4d8c-b03b-c1d9fc230389" ID-ISSN="1175-5326" ID-Zenodo-Dep="253147" ID-ZooBank="A49D09BA-6134-4488-943C-D78418602AA3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460394987294" checkinUser="plazi" docAuthor="Gorochov, Andrej V. & Tan, Ming Kai" docDate="2014" docId="03CBD073FFD4FFC6498D9980FB02FE56" docLanguage="en" docName="zt03774p551.pdf" docOrigin="Zootaxa 3774 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Varitrella (Cantotrella) robusta Gorochov, sp. nov." docType="treatment" docVersion="9" lastPageNumber="549" masterDocId="FFF2A80BFFDCFFC8491A985CFFBDFFA3" masterDocTitle="Species of the subgenus Cantotrella (Orthoptera: Gryllidae: Podoscirtinae: Varitrella) from Singapore, Malaysia and Indonesia" masterLastPageNumber="551" masterPageNumber="535" pageNumber="543" updateTime="1698298647871" updateUser="ExternalLinkService">
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<mods:mods id="F4428BA20B2FE047F6A2793B98725ADA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="35C0A74052BB8408DA41207DCE10A071">
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<mods:title id="717940A4A1EDC5704232D16333C92E83">Species of the subgenus Cantotrella (Orthoptera: Gryllidae: Podoscirtinae: Varitrella) from Singapore, Malaysia and Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="CE180F97533A15BBC5DCC9A271E8AB05" type="personal">
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<mods:role id="F160A1177B6D63DAD893BF1885725644">
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<mods:roleTerm id="86A6739E727C78DF95260147D8B52C78">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E0618127E5363C9F47CC587BDB0BC9FB">Gorochov, Andrej V.</mods:namePart>
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</mods:name>
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<mods:name id="ACD335EF364B3501D64840DB5F16E8A9" type="personal">
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<mods:role id="DF3BE34B60B7BC85A32BF3CEE5FDDB32">
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<mods:roleTerm id="001A962820B19EF76E77CE6B857E8156">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1071D903E37C056B031607D845D3CE50">Tan, Ming Kai</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0D07516CE1D3DCDC35EC56E7ABC220FC">text</mods:typeOfResource>
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<mods:titleInfo id="8104AD2EE0AC27F2C5F6D2DF1EE9DDDB">
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<mods:title id="BAE5065EC70D1282207D44E8AA2547A9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5404053BBFCD8D6E26195ECB6E282620">
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<mods:date id="89AF19CA45D32A41C69A1B6B8B82888B">2014</mods:date>
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<mods:detail id="E4F4360BC307AF1D4542EFD23C15C62C" type="volume">
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<mods:number id="C61DF3421C67E3B2C02DF971BB2DA177">3774</mods:number>
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</mods:detail>
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<mods:detail id="660D7C593B0E44F96C160E1B010B5A0B" type="issue">
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<mods:number id="0DCDFFB2FB058745B077032D1F6A2D37">6</mods:number>
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</mods:detail>
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<mods:start id="18B4F3B56BD8AF4620CD60C9F38EC3F6">535</mods:start>
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<mods:end id="02AF97EB2436B37DD91AFE6D8D3B3A48">551</mods:end>
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</mods:extent>
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<mods:classification id="57E691B5635BBC069A7D7DE4F94BAEEF">journal article</mods:classification>
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<mods:identifier id="D36BC1D5372F596858CC8DAD5C81B9E8" type="CLB-Dataset">46246</mods:identifier>
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<mods:identifier id="019F74C8F032E46A2F385606CCA81AB6" type="DOI">10.11646/zootaxa.3774.6.3</mods:identifier>
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<mods:identifier id="561DAE8DD395E75D574ACFCE2022F4BB" type="GBIF-Dataset">5485c5f1-36b6-4d8c-b03b-c1d9fc230389</mods:identifier>
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<mods:identifier id="87B29960CF74C3D83D1717276FB3B92B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0DF0F40F4758C0B56BD08D875977868B" type="Zenodo-Dep">253147</mods:identifier>
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<mods:identifier id="B5419CAAEA6D868E7B36A32E62E50413" type="ZooBank">A49D09BA-6134-4488-943C-D78418602AA3</mods:identifier>
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</mods:mods>
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<treatment id="03CBD073FFD4FFC6498D9980FB02FE56" ID-DOI="http://doi.org/10.5281/zenodo.6139053" ID-GBIF-Taxon="119417934" ID-Zenodo-Dep="6139053" LSID="urn:lsid:plazi:treatment:03CBD073FFD4FFC6498D9980FB02FE56" httpUri="http://treatment.plazi.org/id/03CBD073FFD4FFC6498D9980FB02FE56" lastPageId="14" lastPageNumber="549" pageId="8" pageNumber="543">
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<subSubSection id="C37832EEFFD4FFC0498D9980FE7CFDB4" pageId="8" pageNumber="543" type="nomenclature">
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<paragraph id="8BDD6165FFD4FFC0498D9980FD46FE55" blockId="8.[151,763,476,535]" box="[151,763,476,502]" pageId="8" pageNumber="543">
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<heading id="D095D609FFD4FFC0498D9980FD46FE55" bold="true" box="[151,763,476,502]" fontSize="11" level="1" pageId="8" pageNumber="543" reason="1">
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<emphasis id="B916BD77FFD4FFC0498D9980FD46FE55" bold="true" box="[151,763,476,502]" pageId="8" pageNumber="543">
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<taxonomicName id="4C621AE6FFD4FFC0498D9980FD2CFE55" authority="Gorochov" authorityName="Gorochov" box="[151,657,476,502]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov." subGenus="Cantotrella">
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<emphasis id="B916BD77FFD4FFC0498D9980FEB4FE55" bold="true" box="[151,265,476,502]" italics="true" pageId="8" pageNumber="543">Varitrella</emphasis>
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||
(
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<emphasis id="B916BD77FFD4FFC048009980FE1EFE55" bold="true" box="[282,419,476,502]" italics="true" pageId="8" pageNumber="543">Cantotrella</emphasis>
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)
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<emphasis id="B916BD77FFD4FFC048AE9980FDB2FE55" bold="true" box="[436,527,476,502]" italics="true" pageId="8" pageNumber="543">robusta</emphasis>
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Gorochov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A225000CFFD4FFC04B849980FD46FE55" box="[670,763,476,502]" pageId="8" pageNumber="543" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDD6165FFD4FFC0498D99A2FE7CFDB4" blockId="8.[151,763,476,535]" box="[151,449,510,535]" pageId="8" pageNumber="543">
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(
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<figureCitation id="13597DE0FFD4FFC0498599A2FF5AFDB4" box="[159,231,510,535]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1900,1922]" captionTargetBox="[237,1352,199,1875]" captionTargetId="figure@3.[232,1356,193,1879]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Head and pronotum of male from above: 2, Varitrella orion sp. nov.; 3, V. trusmadi sp. nov.; 4, V. striata sp. nov.; 5, V. ro bu st a sp. nov.; 6, V. suk au sp. nov.; 7. V. tawau sp. nov.; 8, V. amoena sp. nov.; 9, V. manukan sp. nov." httpUri="https://zenodo.org/record/253149/files/figure.png" pageId="8" pageNumber="543">Figs 5</figureCitation>
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,
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<figureCitation id="13597DE0FFD4FFC049E999A2FE86FDB4" box="[243,315,510,535]" captionStart="FIGURES 16 – 19" captionStartId="9.[151,264,1791,1813]" captionTargetBox="[303,1277,199,1764]" captionTargetId="figure@9.[302,1285,193,1770]" captionTargetPageId="9" captionText="FIGURES 16 – 19. Left and right male tegmina: 16, 17, Varitrella robusta sp. nov.; 18, V. s uk a u sp. nov.; 19, V. m a nu k a n sp. nov." httpUri="https://zenodo.org/record/253152/files/figure.png" pageId="8" pageNumber="543">16, 17</figureCitation>
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,
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<figureCitation id="13597DE0FFD4FFC0485D99A2FED8FDB4" box="[327,357,510,535]" captionStart="FIGURES 20 – 31" captionStartId="10.[151,264,1764,1786]" captionTargetBox="[157,1433,196,1737]" captionTargetId="figure@10.[151,1436,193,1742]" captionTargetPageId="10" captionText="FIGURES 20 – 31. Varitrella, male: 20 – 22, V. orion sp. nov., stridulatory vein of right tegmen from below (20), abdominal apex from above (21), and genital plate from below (22); 23 – 28, anal plate partly from above and partly from behind in V. trusmadi sp. nov. (23), V. striata sp. nov. (24), V. robusta sp. nov. (25), V. tawau sp. nov. (26), V. suk au sp. nov. (27) and V. amoena sp. nov. (28); 29 – 31, genitalia of V. palawanensis (29), V. mindoroensis (30) and V. quadrata (31) from side and from below, with names of morphological structures. Abbreviations: ds, dorsal epiphallic spine; ec, ectoparamere; ea, endoparameral apodeme; en, endoparamere; ep, epiphallus; f, formula (= mold of spermatophore attachment plate); lo, lateral lobe of rachis; mr, median sclerotized ribbon fused with formula; r, rachis (= guiding rod); ra, ramus." httpUri="https://zenodo.org/record/253153/files/figure.png" pageId="8" pageNumber="543">25</figureCitation>
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,
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<figureCitation id="13597DE0FFD4FFC0486B99A2FE04FDB4" box="[369,441,510,535]" captionStart="FIGURES 44 – 58" captionStartId="12.[151,264,1928,1950]" captionTargetBox="[202,1387,199,1879]" captionTargetId="figure@12.[179,1391,193,1897]" captionTargetPageId="12" captionText="FIGURES 44 – 58. Varitrella: 44 – 52, V. robusta sp. nov. (44 – 46, holotype); 53 – 58, V. manukan sp. nov. Male genitalia from above (44, 47, 53), from below (45, 48, 54) and from side (46, 49, 55); left half of distal part of ovipositor from above (50, 57), from below (51, 58) and from side (52, 56)." httpUri="https://zenodo.org/record/253154/files/figure.png" pageId="8" pageNumber="543">44–52</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37832EEFFD4FFC0498D9A1AFB43FCB0" pageId="8" pageNumber="543" type="materials_examined">
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<paragraph id="8BDD6165FFD4FFC0498D9A1AFEEAFD04" blockId="8.[151,1437,582,1975]" pageId="8" pageNumber="543">
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<emphasis id="B916BD77FFD4FFC0498D9A1AFE3DFDFC" bold="true" box="[151,384,582,607]" pageId="8" pageNumber="543">Material examined.</emphasis>
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<typeStatus id="54D9DFC7FFD4FFC0489C9A1AFE53FDFC" box="[390,494,582,607]" pageId="8" pageNumber="543" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="F37521F5FFD4FFC04B569A1AFD7CFDFC" box="[588,705,582,607]" name="Indonesia" pageId="8" pageNumber="543">
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<emphasis id="B916BD77FFD4FFC04B569A1AFD7CFDFC" bold="true" box="[588,705,582,607]" pageId="8" pageNumber="543">Indonesia</emphasis>
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</collectingCountry>
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, Sulawesi Utara Prov., Bunaken Island not far from Manado City (Sulawesi), Bunaken Marine National Park, secondary forest, on leaf of bush at night,
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<date id="FFDC47A5FFD4FFC04D669A36FACFFD20" box="[1148,1394,618,643]" pageId="8" pageNumber="543" value="2011-02-18" valueMax="2011-02-25" valueMin="2011-02-18">18–25 February 2011</date>
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, A. Gorochov (
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<collectionCode id="ED73F9A0FFD4FFC0480C9AD3FEF7FD04" box="[278,330,655,679]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="543">ZIN</collectionCode>
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).
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</paragraph>
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<paragraph id="8BDD6165FFD4FFC049DD9AEFFB43FCB0" blockId="8.[151,1437,582,1975]" pageId="8" pageNumber="543">
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<typeStatus id="54D9DFC7FFD4FFC049DD9AEFFE85FD68" box="[199,312,691,715]" pageId="8" pageNumber="543" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D64AAECFFD4FFC048599AEFFE24FD68" box="[323,409,690,715]" pageId="8" pageNumber="543" type="male">2 males</specimenCount>
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and
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<specimenCount id="9D64AAECFFD4FFC048CA9AEFFDFDFD68" box="[464,576,690,715]" pageId="8" pageNumber="543" type="female">2 females</specimenCount>
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, same data as for
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<typeStatus id="54D9DFC7FFD4FFC04A129AEEFCD7FD68" box="[776,874,690,715]" pageId="8" pageNumber="543" type="holotype">holotype</typeStatus>
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||
(
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<collectionCode id="ED73F9A0FFD4FFC04A609AEFFC13FD68" box="[890,942,691,715]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="543">ZIN</collectionCode>
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);
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<specimenCount id="9D64AAECFFD4FFC04AD89AEFFB9BFD68" box="[962,1062,690,715]" pageId="8" pageNumber="543" type="female">1 female</specimenCount>
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, same province, about
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<quantity id="4C9ACC80FFD4FFC04C329AEFFAD2FD68" box="[1320,1391,690,715]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="543" unit="km" value="40.0">40 km</quantity>
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NE of Manado City, Tangkoko National Park on eastern coast of Minahassa Peninsula (Sulawesi), environs of Tangkoko Lodge, secondary forest, on leaf of bush at night,
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<date id="FFDC47A5FFD4FFC04A209AA6FBACFCB0" box="[826,1041,762,787]" pageId="8" pageNumber="543" value="2011-02-03" valueMax="2011-02-06" valueMin="2011-02-03">3–6 February 2011</date>
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, A. Gorochov (
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<collectionCode id="ED73F9A0FFD4FFC04DA79AA7FB4CFCB0" box="[1213,1265,763,787]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="543">ZIN</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37832EEFFD4FFC649DD9B42FB02FE56" lastPageId="14" lastPageNumber="549" pageId="8" pageNumber="543" type="description">
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<paragraph id="8BDD6165FFD4FFC049DD9B42FDEAFA40" blockId="8.[151,1437,582,1975]" pageId="8" pageNumber="543">
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<emphasis id="B916BD77FFD4FFC049DD9B42FEE4FC94" bold="true" box="[199,345,798,823]" pageId="8" pageNumber="543">Description.</emphasis>
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Male (
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<typeStatus id="54D9DFC7FFD4FFC048AB9B42FDA4FC94" box="[433,537,798,823]" pageId="8" pageNumber="543" type="holotype">holotype</typeStatus>
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). General appearance similar to that of
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<taxonomicName id="4C621AE6FFD4FFC04AEE9B43FBC5FC95" box="[1012,1144,798,822]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
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<emphasis id="B916BD77FFD4FFC04AEE9B43FBC5FC95" box="[1012,1144,798,822]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C621AE6FFD4FFC04DA29B43FA9CFC95" box="[1208,1313,799,822]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="striata">
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<emphasis id="B916BD77FFD4FFC04DA29B43FA9CFC95" box="[1208,1313,799,822]" italics="true" pageId="8" pageNumber="543">V. striata</emphasis>
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</taxonomicName>
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, but body somewhat larger, slightly more robust, and with shorter hind wings. Coloration light brown with following marks: head with darker (greyish brown) dorsum (
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<figureCitation id="13597DE0FFD4FFC04B679B3AFD7CFCDC" box="[637,705,870,895]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1900,1922]" captionTargetBox="[237,1352,199,1875]" captionTargetId="figure@3.[232,1356,193,1879]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Head and pronotum of male from above: 2, Varitrella orion sp. nov.; 3, V. trusmadi sp. nov.; 4, V. striata sp. nov.; 5, V. ro bu st a sp. nov.; 6, V. suk au sp. nov.; 7. V. tawau sp. nov.; 8, V. amoena sp. nov.; 9, V. manukan sp. nov." httpUri="https://zenodo.org/record/253149/files/figure.png" pageId="8" pageNumber="543">Fig. 5</figureCitation>
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) and small spot at region of each dorsolateral corner of clypeus; pronotal disc brown, darker than lateral pronotal lobes (
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<figureCitation id="13597DE0FFD4FFC04A199BD6FCF5FC00" box="[771,840,906,931]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1900,1922]" captionTargetBox="[237,1352,199,1875]" captionTargetId="figure@3.[232,1356,193,1879]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Head and pronotum of male from above: 2, Varitrella orion sp. nov.; 3, V. trusmadi sp. nov.; 4, V. striata sp. nov.; 5, V. ro bu st a sp. nov.; 6, V. suk au sp. nov.; 7. V. tawau sp. nov.; 8, V. amoena sp. nov.; 9, V. manukan sp. nov." httpUri="https://zenodo.org/record/253149/files/figure.png" pageId="8" pageNumber="543">Fig. 5</figureCitation>
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); tegmina with brown dorsal field (but majority of its membranes slightly lighter than venation), yellowish white both humeral stripe and spot in corner between stock of
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<emphasis id="B916BD77FFD4FFC0498D9B8FFEAFFC48" box="[151,274,979,1003]" italics="true" pageId="8" pageNumber="543">MP+CuA1</emphasis>
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and distal part of mirror, dark brown area between
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<emphasis id="B916BD77FFD4FFC04A549B8FFCD4FC49" box="[846,873,979,1002]" italics="true" pageId="8" pageNumber="543">Sc</emphasis>
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||
and
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<emphasis id="B916BD77FFD4FFC04A849B8FFC12FC49" box="[926,943,979,1002]" italics="true" pageId="8" pageNumber="543">R</emphasis>
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(excepting distal part of this area), and light and semitransparent other parts of lateral field (but with light brown branches of Sc, few small dark spots between their proximal parts, and several whitish crossveins in proximal half of this field;
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<figureCitation id="13597DE0FFD4FFC04D739C46FB4EFB90" box="[1129,1267,1050,1075]" captionStart="FIGURES 16 – 19" captionStartId="9.[151,264,1791,1813]" captionTargetBox="[303,1277,199,1764]" captionTargetId="figure@9.[302,1285,193,1770]" captionTargetPageId="9" captionText="FIGURES 16 – 19. Left and right male tegmina: 16, 17, Varitrella robusta sp. nov.; 18, V. s uk a u sp. nov.; 19, V. m a nu k a n sp. nov." httpUri="https://zenodo.org/record/253152/files/figure.png" pageId="8" pageNumber="543">Figs 16, 17</figureCitation>
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); legs without darkened spots and dots but with slightly darker dorsal surface of fore tibiae and brownish grey ventral surface of hind tibiae; thoracic venter and abdomen almost yellowish with brown dorsum of tergites and characteristic area on anal plate (
|
||
<figureCitation id="13597DE0FFD4FFC0480B9CDAFED8FB3C" box="[273,357,1158,1183]" captionStart="FIGURES 20 – 31" captionStartId="10.[151,264,1764,1786]" captionTargetBox="[157,1433,196,1737]" captionTargetId="figure@10.[151,1436,193,1742]" captionTargetPageId="10" captionText="FIGURES 20 – 31. Varitrella, male: 20 – 22, V. orion sp. nov., stridulatory vein of right tegmen from below (20), abdominal apex from above (21), and genital plate from below (22); 23 – 28, anal plate partly from above and partly from behind in V. trusmadi sp. nov. (23), V. striata sp. nov. (24), V. robusta sp. nov. (25), V. tawau sp. nov. (26), V. suk au sp. nov. (27) and V. amoena sp. nov. (28); 29 – 31, genitalia of V. palawanensis (29), V. mindoroensis (30) and V. quadrata (31) from side and from below, with names of morphological structures. Abbreviations: ds, dorsal epiphallic spine; ec, ectoparamere; ea, endoparameral apodeme; en, endoparamere; ep, epiphallus; f, formula (= mold of spermatophore attachment plate); lo, lateral lobe of rachis; mr, median sclerotized ribbon fused with formula; r, rachis (= guiding rod); ra, ramus." httpUri="https://zenodo.org/record/253153/files/figure.png" pageId="8" pageNumber="543">Fig. 25</figureCitation>
|
||
). External structure of body also similar to that of
|
||
<taxonomicName id="4C621AE6FFD4FFC04A8E9CDBFBAFFB3D" box="[916,1042,1158,1182]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD4FFC04A8E9CDBFBAFFB3D" box="[916,1042,1158,1182]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD4FFC04D5D9CDBFB16FB3D" box="[1095,1195,1159,1182]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD4FFC04D5D9CDBFB16FB3D" box="[1095,1195,1159,1182]" italics="true" pageId="8" pageNumber="543">V. striata</emphasis>
|
||
</taxonomicName>
|
||
, but scape almost 1.7 times as wide as rostral apex, pronotum with more parallel lateral sides (
|
||
<figureCitation id="13597DE0FFD4FFC04ADF9CF6FBB5FB60" box="[965,1032,1194,1219]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1900,1922]" captionTargetBox="[237,1352,199,1875]" captionTargetId="figure@3.[232,1356,193,1879]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Head and pronotum of male from above: 2, Varitrella orion sp. nov.; 3, V. trusmadi sp. nov.; 4, V. striata sp. nov.; 5, V. ro bu st a sp. nov.; 6, V. suk au sp. nov.; 7. V. tawau sp. nov.; 8, V. amoena sp. nov.; 9, V. manukan sp. nov." httpUri="https://zenodo.org/record/253149/files/figure.png" pageId="8" pageNumber="543">Fig. 5</figureCitation>
|
||
), tegmina with apical area of dorsal field almost as in
|
||
<taxonomicName id="4C621AE6FFD4FFC048729C93FE54FB45" box="[360,489,1230,1254]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD4FFC048729C93FE54FB45" box="[360,489,1230,1254]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13597DE0FFD4FFC048E09C92FD3DFB44" box="[506,640,1230,1255]" captionStart="FIGURES 16 – 19" captionStartId="9.[151,264,1791,1813]" captionTargetBox="[303,1277,199,1764]" captionTargetId="figure@9.[302,1285,193,1770]" captionTargetPageId="9" captionText="FIGURES 16 – 19. Left and right male tegmina: 16, 17, Varitrella robusta sp. nov.; 18, V. s uk a u sp. nov.; 19, V. m a nu k a n sp. nov." httpUri="https://zenodo.org/record/253152/files/figure.png" pageId="8" pageNumber="543">Figs 16, 17</figureCitation>
|
||
), hind wings insignificantly longer than tegmina; and anal plate with central membranous area almost round (
|
||
<figureCitation id="13597DE0FFD4FFC04B4D9CAEFD16FAA8" box="[599,683,1266,1291]" captionStart="FIGURES 20 – 31" captionStartId="10.[151,264,1764,1786]" captionTargetBox="[157,1433,196,1737]" captionTargetId="figure@10.[151,1436,193,1742]" captionTargetPageId="10" captionText="FIGURES 20 – 31. Varitrella, male: 20 – 22, V. orion sp. nov., stridulatory vein of right tegmen from below (20), abdominal apex from above (21), and genital plate from below (22); 23 – 28, anal plate partly from above and partly from behind in V. trusmadi sp. nov. (23), V. striata sp. nov. (24), V. robusta sp. nov. (25), V. tawau sp. nov. (26), V. suk au sp. nov. (27) and V. amoena sp. nov. (28); 29 – 31, genitalia of V. palawanensis (29), V. mindoroensis (30) and V. quadrata (31) from side and from below, with names of morphological structures. Abbreviations: ds, dorsal epiphallic spine; ec, ectoparamere; ea, endoparameral apodeme; en, endoparamere; ep, epiphallus; f, formula (= mold of spermatophore attachment plate); lo, lateral lobe of rachis; mr, median sclerotized ribbon fused with formula; r, rachis (= guiding rod); ra, ramus." httpUri="https://zenodo.org/record/253153/files/figure.png" pageId="8" pageNumber="543">Fig. 25</figureCitation>
|
||
). Genitalia similar to those of
|
||
<taxonomicName id="4C621AE6FFD4FFC04AE09CAFFBE3FAA9" box="[1018,1118,1267,1290]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD4FFC04AE09CAFFBE3FAA9" box="[1018,1118,1267,1290]" italics="true" pageId="8" pageNumber="543">V. striata</emphasis>
|
||
</taxonomicName>
|
||
, but dorsal epiphallic spines somewhat shorter and with slightly wider (longer) bases, proximal part of epiphallus (before these spines) longer than in
|
||
<taxonomicName id="4C621AE6FFD4FFC049F19D67FEF2FAF1" box="[235,335,1339,1362]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD4FFC049F19D67FEF2FAF1" box="[235,335,1339,1362]" italics="true" pageId="8" pageNumber="543">V. striata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD4FFC0489C9D67FDB8FAF0" box="[390,517,1339,1363]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD4FFC0489C9D67FDB8FAF0" box="[390,517,1339,1363]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
and with distinctly convex dorsal edge in profile, distal part of epiphallus (behind spines) somewhat shorter than in both latter species, ectoparameres slightly S-shaped in profile and with longer distal (curved) part (but this part shorter than in
|
||
<taxonomicName id="4C621AE6FFD4FFC04BA29DDFFCADFA39" box="[696,784,1411,1434]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="orion">
|
||
<emphasis id="B916BD77FFD4FFC04BA29DDFFCADFA39" box="[696,784,1411,1434]" italics="true" pageId="8" pageNumber="543">V. orion</emphasis>
|
||
</taxonomicName>
|
||
), rachis and formula approximately as long as in
|
||
<emphasis id="B916BD77FFD4FFC04C599DDFFA24FA39" box="[1347,1433,1411,1434]" italics="true" pageId="8" pageNumber="543">V. o r i o n</emphasis>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD4FFC049DC9DFBFEF8FA1D" box="[198,325,1446,1470]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD4FFC049DC9DFBFEF8FA1D" box="[198,325,1446,1470]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
but with distal (widened) part of rachis somewhat shorter than in
|
||
<taxonomicName id="4C621AE6FFD4FFC04D3B9DFBFBC4FA1D" box="[1057,1145,1447,1470]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="orion">
|
||
<emphasis id="B916BD77FFD4FFC04D3B9DFBFBC4FA1D" box="[1057,1145,1447,1470]" italics="true" pageId="8" pageNumber="543">V. orion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD4FFC04DB49DFBFAAFFA1D" box="[1198,1298,1447,1470]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD4FFC04DB49DFBFAAFFA1D" box="[1198,1298,1447,1470]" italics="true" pageId="8" pageNumber="543">V. striata</emphasis>
|
||
</taxonomicName>
|
||
and slightly longer than in
|
||
<taxonomicName id="4C621AE6FFD4FFC048239D97FE04FA40" box="[313,441,1483,1507]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD4FFC048239D97FE04FA40" box="[313,441,1483,1507]" italics="true" pageId="8" pageNumber="543">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13597DE0FFD4FFC048DD9D96FDF6FA40" box="[455,587,1482,1507]" captionStart="FIGURES 44 – 58" captionStartId="12.[151,264,1928,1950]" captionTargetBox="[202,1387,199,1879]" captionTargetId="figure@12.[179,1391,193,1897]" captionTargetPageId="12" captionText="FIGURES 44 – 58. Varitrella: 44 – 52, V. robusta sp. nov. (44 – 46, holotype); 53 – 58, V. manukan sp. nov. Male genitalia from above (44, 47, 53), from below (45, 48, 54) and from side (46, 49, 55); left half of distal part of ovipositor from above (50, 57), from below (51, 58) and from side (52, 56)." httpUri="https://zenodo.org/record/253154/files/figure.png" pageId="8" pageNumber="543">Figs 44–46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD6165FFD4FFC049DD9DB2FA24F9D0" blockId="8.[151,1437,582,1975]" pageId="8" pageNumber="543">
|
||
Variation. Other males with slightly lighter coloration of pronotal disc and of most part of dorsal tegminal field; one male with light spot near anterior edge of median ocellus and with uniformly light fore tibiae; another male with a pair of small light spots near eyes (not far from hind parts of lateral ocelli), with rose area on mesonotum, with slightly darkened dorsal surface of hind tibiae (variability in genitalia insignificant;
|
||
<figureCitation id="13597DE0FFD4FFC04C119E06FA30F9D0" box="[1291,1421,1626,1651]" captionStart="FIGURES 44 – 58" captionStartId="12.[151,264,1928,1950]" captionTargetBox="[202,1387,199,1879]" captionTargetId="figure@12.[179,1391,193,1897]" captionTargetPageId="12" captionText="FIGURES 44 – 58. Varitrella: 44 – 52, V. robusta sp. nov. (44 – 46, holotype); 53 – 58, V. manukan sp. nov. Male genitalia from above (44, 47, 53), from below (45, 48, 54) and from side (46, 49, 55); left half of distal part of ovipositor from above (50, 57), from below (51, 58) and from side (52, 56)." httpUri="https://zenodo.org/record/253154/files/figure.png" pageId="8" pageNumber="543">Figs 47–49</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD6165FFD4FFC049DD9E22FB52F814" blockId="8.[151,1437,582,1975]" pageId="8" pageNumber="543">
|
||
Female. General appearance similar to that of males, but with following differences: areas between Sc and M in tegmina more or less uniformly light brown (almost semitransparent); rest of dorsal tegminal field greyish brown or light brown and with slightly darker venation, narrow dark stripe along lateral edge of this field, and several distinct or almost indistinct very small whitish marks along this edge; lateral tegminal field sometimes with few additional dark dots between middle parts of
|
||
<emphasis id="B916BD77FFD4FFC04BAF9F53FD72F885" box="[693,719,1807,1830]" italics="true" pageId="8" pageNumber="543">Sc</emphasis>
|
||
branches or without darkened spots and dots between these branches; dorsal tegminal field with 15–17 longitudinal (somewhat oblique) veins and very numerous crossveins; lateral tegminal field with12–14 branches on
|
||
<emphasis id="B916BD77FFD4FFC04B8E9F0BFD12F8CD" box="[660,687,1879,1902]" italics="true" pageId="8" pageNumber="543">Sc</emphasis>
|
||
and numerous (but often weakly distinct) crossveins. Genital plate weakly elongate, slightly narrowing to apex, with rather deep posteromedian notch and rounded lobes around it; ovipositor approximately 1.3 times as long as hind femur and with drilling apical part (
|
||
<figureCitation id="13597DE0FFD4FFC04D7E9FC2FB59F814" box="[1124,1252,1950,1975]" captionStart="FIGURES 44 – 58" captionStartId="12.[151,264,1928,1950]" captionTargetBox="[202,1387,199,1879]" captionTargetId="figure@12.[179,1391,193,1897]" captionTargetPageId="12" captionText="FIGURES 44 – 58. Varitrella: 44 – 52, V. robusta sp. nov. (44 – 46, holotype); 53 – 58, V. manukan sp. nov. Male genitalia from above (44, 47, 53), from below (45, 48, 54) and from side (46, 49, 55); left half of distal part of ovipositor from above (50, 57), from below (51, 58) and from side (52, 56)." httpUri="https://zenodo.org/record/253154/files/figure.png" pageId="8" pageNumber="543">Fig. 50–52</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1D31EDFFD5FFC1498D9EA3FF7EF890" httpUri="https://zenodo.org/record/253152/files/figure.png" pageId="9" pageNumber="544" targetBox="[303,1277,199,1764]" targetPageId="9">
|
||
<paragraph id="8BDD6165FFD5FFC1498D9EA3FF7EF890" blockId="9.[151,1436,1791,1843]" pageId="9" pageNumber="544">
|
||
<emphasis id="B916BD77FFD5FFC1498D9EA3FEE8F8B6" bold="true" box="[151,341,1791,1813]" pageId="9" pageNumber="544">FIGURES 16–19.</emphasis>
|
||
Left and right male tegmina: 16, 17,
|
||
<taxonomicName id="4C621AE6FFD5FFC14BC29EA3FC31F8B6" box="[728,908,1791,1813]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="544" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD5FFC14BC29EA3FC31F8B6" box="[728,908,1791,1813]" italics="true" pageId="9" pageNumber="544">Varitrella robusta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD5FFC14A899EA3FC59F8B6" bold="true" box="[915,996,1791,1813]" pageId="9" pageNumber="544">
|
||
<taxonomicNameLabel id="A225000CFFD5FFC14A899EA3FC59F8B6" box="[915,996,1791,1813]" pageId="9" pageNumber="544" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 18,
|
||
<emphasis id="B916BD77FFD5FFC14D019F5CFBD0F8B6" box="[1051,1133,1791,1813]" italics="true" pageId="9" pageNumber="544">V. s uk a u</emphasis>
|
||
<emphasis id="B916BD77FFD5FFC14D629EA3FB74F8B6" bold="true" box="[1144,1225,1791,1813]" pageId="9" pageNumber="544">sp. nov.</emphasis>
|
||
; 19,
|
||
<emphasis id="B916BD77FFD5FFC14DE59F5CFAC9F8B6" box="[1279,1396,1791,1813]" italics="true" pageId="9" pageNumber="544">V. m a nu k a n</emphasis>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDD6165FFD5FFC149DD9F3CFEE0F862" blockId="9.[151,1437,1888,1985]" pageId="9" pageNumber="544">Length in mm. Body: male 19–21, female 19–24; body with wings: male 25–27, female 30–32; pronotum: male 3.4–3.7, female 4.2–4.6; tegmina: male 17–19, female 21–22.5; hind femora: male 14–15.5, female 16–18; ovipositor 21–23.</paragraph>
|
||
<caption id="DF1D31EDFFD6FFC2498D9EB8FC39F811" httpUri="https://zenodo.org/record/253153/files/figure.png" pageId="10" pageNumber="545" targetBox="[157,1433,196,1737]" targetPageId="10">
|
||
<paragraph id="8BDD6165FFD6FFC2498D9EB8FC39F811" blockId="10.[151,1436,1764,1970]" pageId="10" pageNumber="545">
|
||
<emphasis id="B916BD77FFD6FFC2498D9EB8FEEEF959" bold="true" box="[151,339,1764,1786]" pageId="10" pageNumber="545">FIGURES 20–31.</emphasis>
|
||
<taxonomicName id="4C621AE6FFD6FFC248429EB8FE05F959" box="[344,440,1764,1786]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BD77FFD6FFC248429EB8FE05F959" box="[344,440,1764,1786]" italics="true" pageId="10" pageNumber="545">Varitrella</emphasis>
|
||
</taxonomicName>
|
||
, male: 20–22,
|
||
<taxonomicName id="4C621AE6FFD6FFC24B579EB9FD21F959" box="[589,668,1765,1786]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="orion" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC24B579EB9FD21F959" box="[589,668,1765,1786]" italics="true" pageId="10" pageNumber="545">V. orion</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225000CFFD6FFC24BB89EB8FD4FF959" box="[674,754,1764,1786]" pageId="10" pageNumber="545" rank="species">
|
||
<emphasis id="B916BD77FFD6FFC24BB89EB8FD4FF959" bold="true" box="[674,754,1764,1786]" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, stridulatory vein of right tegmen from below (20), abdominal apex from above (21), and genital plate from below (22); 23–28, anal plate partly from above and partly from behind in
|
||
<taxonomicName id="4C621AE6FFD6FFC24C339F58FA21F8BA" box="[1321,1436,1795,1817]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="trusmadi" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC24C339F58FA21F8BA" box="[1321,1436,1795,1817]" italics="true" pageId="10" pageNumber="545">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225000CFFD6FFC2498D9F7DFF55F894" box="[151,232,1825,1847]" pageId="10" pageNumber="545" rank="species">
|
||
<emphasis id="B916BD77FFD6FFC2498D9F7DFF55F894" bold="true" box="[151,232,1825,1847]" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(23),
|
||
<taxonomicName id="4C621AE6FFD6FFC2483C9F7EFE3CF894" box="[294,385,1826,1847]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="striata" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC2483C9F7EFE3CF894" box="[294,385,1826,1847]" italics="true" pageId="10" pageNumber="545">V. striata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225000CFFD6FFC248929F7DFE65F894" box="[392,472,1825,1847]" pageId="10" pageNumber="545" rank="species">
|
||
<emphasis id="B916BD77FFD6FFC248929F7DFE65F894" bold="true" box="[392,472,1825,1847]" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(24),
|
||
<taxonomicName id="4C621AE6FFD6FFC24B0D9F7EFDC1F894" box="[535,636,1825,1847]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC24B0D9F7EFDC1F894" box="[535,636,1825,1847]" italics="true" pageId="10" pageNumber="545">V. robusta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225000CFFD6FFC24B999F7DFD6EF894" box="[643,723,1825,1847]" pageId="10" pageNumber="545" rank="species">
|
||
<emphasis id="B916BD77FFD6FFC24B999F7DFD6EF894" bold="true" box="[643,723,1825,1847]" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(25),
|
||
<taxonomicName id="4C621AE6FFD6FFC24A089F7EFCD4F894" box="[786,873,1826,1847]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="tawau" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC24A089F7EFCD4F894" box="[786,873,1826,1847]" italics="true" pageId="10" pageNumber="545">V. tawau</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD6FFC24A6A9F7DFC7DF894" bold="true" box="[880,960,1825,1847]" pageId="10" pageNumber="545">
|
||
<taxonomicNameLabel id="A225000CFFD6FFC24A6A9F7DFC7DF894" box="[880,960,1825,1847]" pageId="10" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(26),
|
||
<emphasis id="B916BD77FFD6FFC24AE59F7EFBEDF894" box="[1023,1104,1825,1847]" italics="true" pageId="10" pageNumber="545">
|
||
<taxonomicName id="4C621AE6FFD6FFC24AE59F7EFB8AF894" box="[1023,1079,1825,1847]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="suk">V. suk</taxonomicName>
|
||
au
|
||
</emphasis>
|
||
<emphasis id="B916BD77FFD6FFC24D409F7DFB17F894" bold="true" box="[1114,1194,1825,1847]" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
(27) and
|
||
<taxonomicName id="4C621AE6FFD6FFC24C179F7EFACBF894" box="[1293,1398,1826,1847]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="amoena" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD6FFC24C179F7EFACBF894" box="[1293,1398,1826,1847]" italics="true" pageId="10" pageNumber="545">V. amoena</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225000CFFD6FFC24C679F7DFF7EF8F5" pageId="10" pageNumber="545" rank="species">
|
||
<emphasis id="B916BD77FFD6FFC24C679F7DFF7EF8F5" bold="true" pageId="10" pageNumber="545">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(28); 29–31, genitalia of
|
||
<taxonomicName id="4C621AE6FFD6FFC248D29F1DFDD6F8F5" box="[456,619,1856,1878]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="palawanensis">
|
||
<emphasis id="B916BD77FFD6FFC248D29F1DFDD6F8F5" box="[456,619,1856,1878]" italics="true" pageId="10" pageNumber="545">V. palawanensis</emphasis>
|
||
</taxonomicName>
|
||
(29),
|
||
<taxonomicName id="4C621AE6FFD6FFC24BB39F1DFCF4F8F5" box="[681,841,1856,1878]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="mindoroensis">
|
||
<emphasis id="B916BD77FFD6FFC24BB39F1DFCF4F8F5" box="[681,841,1856,1878]" italics="true" pageId="10" pageNumber="545">V. mindoroensis</emphasis>
|
||
</taxonomicName>
|
||
(30) and
|
||
<taxonomicName id="4C621AE6FFD6FFC24AB79F1DFB9EF8F5" box="[941,1059,1856,1878]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="quadrata">
|
||
<emphasis id="B916BD77FFD6FFC24AB79F1DFB9EF8F5" box="[941,1059,1856,1878]" italics="true" pageId="10" pageNumber="545">V. quadrata</emphasis>
|
||
</taxonomicName>
|
||
(31) from side and from below, with names of morphological structures. Abbreviations:
|
||
<emphasis id="B916BD77FFD6FFC24BBE9F02FD06F8D7" box="[676,699,1886,1908]" italics="true" pageId="10" pageNumber="545">ds</emphasis>
|
||
, dorsal epiphallic spine;
|
||
<emphasis id="B916BD77FFD6FFC24AA69F3CFC6EF8D6" box="[956,979,1888,1909]" italics="true" pageId="10" pageNumber="545">ec</emphasis>
|
||
, ectoparamere;
|
||
<emphasis id="B916BD77FFD6FFC24D6C9F3CFB33F8D6" box="[1142,1166,1888,1909]" italics="true" pageId="10" pageNumber="545">ea</emphasis>
|
||
, endoparameral apodeme;
|
||
<emphasis id="B916BD77FFD6FFC2498D9F22FF12F830" box="[151,175,1918,1939]" italics="true" pageId="10" pageNumber="545">en</emphasis>
|
||
, endoparamere;
|
||
<emphasis id="B916BD77FFD6FFC248439F22FECCF830" box="[345,369,1918,1939]" italics="true" pageId="10" pageNumber="545">ep</emphasis>
|
||
, epiphallus;
|
||
<emphasis id="B916BD77FFD6FFC248EE9F21FE46F830" box="[500,507,1917,1939]" italics="true" pageId="10" pageNumber="545">f</emphasis>
|
||
, formula (=mold of spermatophore attachment plate);
|
||
<emphasis id="B916BD77FFD6FFC24D339F21FB80F830" box="[1065,1085,1917,1939]" italics="true" pageId="10" pageNumber="545">lo</emphasis>
|
||
, lateral lobe of rachis;
|
||
<emphasis id="B916BD77FFD6FFC24C339F22FAF9F830" box="[1321,1348,1918,1939]" italics="true" pageId="10" pageNumber="545">mr</emphasis>
|
||
, median sclerotized ribbon fused with formula;
|
||
<emphasis id="B916BD77FFD6FFC24B3B9FC1FD97F811" box="[545,554,1949,1970]" italics="true" pageId="10" pageNumber="545">r</emphasis>
|
||
, rachis (=guiding rod);
|
||
<emphasis id="B916BD77FFD6FFC24A019FC1FC8FF811" box="[795,818,1949,1970]" italics="true" pageId="10" pageNumber="545">ra</emphasis>
|
||
, ramus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDD6165FFD7FFC3498D9FD3FEE9F841" pageId="11" pageNumber="546">
|
||
<table id="F96293C5FFD70037498D9FD3FA21F841" box="[151,1436,1935,2018]" gridcols="3" gridrows="3" pageId="11" pageNumber="546">
|
||
<tr id="35526327FFD70037498D9FD3FA21F806" box="[151,1436,1935,1957]" gridrow="0" pageId="11" pageNumber="546">
|
||
<th id="76830A5BFFD70037498D9FD3FD5FF806" box="[151,738,1935,1957]" gridcol="0" gridrow="0" pageId="11" pageNumber="546">
|
||
<emphasis id="B916BD77FFD7FFC3498D9FD3FEEAF806" bold="true" box="[151,343,1935,1957]" pageId="11" pageNumber="546">FIGURES 32–43.</emphasis>
|
||
<taxonomicName id="4C621AE6FFD7FFC3487B9FD3FE7CF806" box="[353,449,1935,1957]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BD77FFD7FFC3487B9FD3FE7CF806" box="[353,449,1935,1957]" italics="true" pageId="11" pageNumber="546">Varitrella</emphasis>
|
||
</taxonomicName>
|
||
, male: 32–34,
|
||
<taxonomicName id="4C621AE6FFD7FFC34B7E9FCCFD0AF806" box="[612,695,1936,1957]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="species" species="orion" status="sp.">
|
||
<emphasis id="B916BD77FFD7FFC34B7E9FCCFD0AF806" box="[612,695,1936,1957]" italics="true" pageId="11" pageNumber="546">V. orion</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD7FFC34BDB9FD3FD5DF806" bold="true" box="[705,736,1935,1957]" pageId="11" pageNumber="546">
|
||
<taxonomicNameLabel id="A225000CFFD7FFC34BDB9FD3FD5DF806" box="[705,736,1935,1957]" pageId="11" pageNumber="546">sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76830A5BFFD700374BF39FD3FB19F806" box="[745,1188,1935,1957]" gridcol="1" gridrow="0" pageId="11" pageNumber="546">
|
||
<emphasis id="B916BD77FFD7FFC34BF39FD3FCA8F806" bold="true" box="[745,789,1935,1957]" pageId="11" pageNumber="546">nov.</emphasis>
|
||
; 35–38,
|
||
<taxonomicName id="4C621AE6FFD7FFC34A6C9FCCFC50F806" box="[886,1005,1935,1957]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="species" species="trusmadi" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD7FFC34A6C9FCCFC50F806" box="[886,1005,1935,1957]" italics="true" pageId="11" pageNumber="546">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD7FFC34AED9FD3FBF6F806" bold="true" box="[1015,1099,1935,1957]" pageId="11" pageNumber="546">
|
||
<taxonomicNameLabel id="A225000CFFD7FFC34AED9FD3FBF6F806" box="[1015,1099,1935,1957]" pageId="11" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 39–43,
|
||
</th>
|
||
<th id="76830A5BFFD700374DB69FD3FA21F806" box="[1196,1436,1935,1957]" gridcol="2" gridrow="0" pageId="11" pageNumber="546">
|
||
<taxonomicName id="4C621AE6FFD7FFC34DB69FCCFAB6F806" box="[1196,1291,1936,1957]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="species" species="striata" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD7FFC34DB69FCCFAB6F806" box="[1196,1291,1936,1957]" italics="true" pageId="11" pageNumber="546">V. striata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD7FFC34C0F9FD3FAD4F806" bold="true" box="[1301,1385,1935,1957]" pageId="11" pageNumber="546">
|
||
<taxonomicNameLabel id="A225000CFFD7FFC34C0F9FD3FAD4F806" box="[1301,1385,1935,1957]" pageId="11" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(43,
|
||
</th>
|
||
</tr>
|
||
<tr id="35526327FFD70037498D9FF1FA21F860" box="[151,1436,1965,1987]" gridrow="1" pageId="11" pageNumber="546">
|
||
<th id="76830A5BFFD70037498D9FF1FD5FF860" box="[151,738,1965,1987]" gridcol="0" gridrow="1" pageId="11" pageNumber="546">holotype). Genitalia from above (32, 35, 39), from below</th>
|
||
<td id="76830A5BFFD700374BF39FF1FB19F860" box="[745,1188,1965,1987]" gridcol="1" gridrow="1" pageId="11" pageNumber="546">(33, 36, 40) and from side (34, 37, 41, 43);</td>
|
||
<td id="76830A5BFFD700374DB69FF1FA21F860" box="[1196,1436,1965,1987]" gridcol="2" gridrow="1" pageId="11" pageNumber="546">spermatophore ampulla</td>
|
||
</tr>
|
||
<tr id="35526327FFD70037498D9F90FA21F841" box="[151,1436,1996,2018]" gridrow="2" pageId="11" pageNumber="546" rowspan-1="1" rowspan-2="1">
|
||
<th id="76830A5BFFD70037498D9F90FD5FF841" box="[151,738,1996,2018]" gridcol="0" gridrow="2" pageId="11" pageNumber="546">from side (38, 42).</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF1D31EDFFD0FFC4498D9FD4FDECF87F" httpUri="https://zenodo.org/record/253154/files/figure.png" pageId="12" pageNumber="547" targetBox="[202,1387,199,1879]" targetPageId="12">
|
||
<paragraph id="8BDD6165FFD0FFC4498D9FD4FDECF87F" blockId="12.[151,1436,1928,2012]" pageId="12" pageNumber="547">
|
||
<emphasis id="B916BD77FFD0FFC4498D9FD4FEE8F83D" bold="true" box="[151,341,1928,1950]" pageId="12" pageNumber="547">FIGURES 44–58.</emphasis>
|
||
<taxonomicName id="4C621AE6FFD0FFC448479FD4FE00F83D" box="[349,445,1928,1950]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="547" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BD77FFD0FFC448479FD4FE00F83D" box="[349,445,1928,1950]" italics="true" pageId="12" pageNumber="547">Varitrella</emphasis>
|
||
</taxonomicName>
|
||
: 44–52,
|
||
<taxonomicName id="4C621AE6FFD0FFC44B039FD6FD3DF83D" box="[537,640,1928,1951]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="547" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD0FFC44B039FD6FD3DF83D" box="[537,640,1928,1951]" italics="true" pageId="12" pageNumber="547">V. robusta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD0FFC44B929FD4FD67F83D" bold="true" box="[648,730,1928,1950]" pageId="12" pageNumber="547">
|
||
<taxonomicNameLabel id="A225000CFFD0FFC44B929FD4FD67F83D" box="[648,730,1928,1950]" pageId="12" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(44–46, holotype); 53–58,
|
||
<taxonomicName id="4C621AE6FFD0FFC44AEC9FD6FBD3F83D" box="[1014,1134,1928,1951]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="547" phylum="Arthropoda" rank="species" species="manukan" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD0FFC44AEC9FD6FBD3F83D" box="[1014,1134,1928,1951]" italics="true" pageId="12" pageNumber="547">V. manukan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD0FFC44D6F9FD4FB7AF83D" bold="true" box="[1141,1223,1928,1950]" pageId="12" pageNumber="547">
|
||
<taxonomicNameLabel id="A225000CFFD0FFC44D6F9FD4FB7AF83D" box="[1141,1223,1928,1950]" pageId="12" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male genitalia from above (44, 47, 53), from below (45, 48, 54) and from side (46, 49, 55); left half of distal part of ovipositor from above (50, 57), from below (51, 58) and from side (52, 56).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1D31EDFFD1FFC5498D9FFFFF7AF855" httpUri="https://zenodo.org/record/253155/files/figure.png" pageId="13" pageNumber="548" targetBox="[185,1390,202,1895]" targetPageId="13">
|
||
<paragraph id="8BDD6165FFD1FFC5498D9FFFFF7AF855" blockId="13.[151,1436,1955,2038]" pageId="13" pageNumber="548">
|
||
<emphasis id="B916BD77FFD1FFC5498D9FFFFEE8F81A" bold="true" box="[151,341,1955,1977]" pageId="13" pageNumber="548">FIGURES 59–70.</emphasis>
|
||
<taxonomicName id="4C621AE6FFD1FFC548469FFFFE01F81A" box="[348,444,1955,1977]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="548" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916BD77FFD1FFC548469FFFFE01F81A" box="[348,444,1955,1977]" italics="true" pageId="13" pageNumber="548">Varitrella</emphasis>
|
||
</taxonomicName>
|
||
, male: 59–62,
|
||
<taxonomicName id="4C621AE6FFD1FFC54B429FF8FD0DF81A" box="[600,688,1956,1977]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="548" phylum="Arthropoda" rank="species" species="tawau" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD1FFC54B429FF8FD0DF81A" box="[600,688,1956,1977]" italics="true" pageId="13" pageNumber="548">V. tawau</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD1FFC54BA29FFFFCB7F81A" bold="true" box="[696,778,1955,1977]" pageId="13" pageNumber="548">
|
||
<taxonomicNameLabel id="A225000CFFD1FFC54BA29FFFFCB7F81A" box="[696,778,1955,1977]" pageId="13" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 63–66,
|
||
<emphasis id="B916BD77FFD1FFC54A7D9FF8FC07F81A" box="[871,954,1955,1977]" italics="true" pageId="13" pageNumber="548">V. su k a u</emphasis>
|
||
<emphasis id="B916BD77FFD1FFC54ADE9FFFFBA8F81A" bold="true" box="[964,1045,1955,1977]" pageId="13" pageNumber="548">sp. nov.</emphasis>
|
||
; 67–70,
|
||
<taxonomicName id="4C621AE6FFD1FFC54D699FF8FB60F81A" box="[1139,1245,1956,1977]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="548" phylum="Arthropoda" rank="species" species="amoena" status="sp. nov.">
|
||
<emphasis id="B916BD77FFD1FFC54D699FF8FB60F81A" box="[1139,1245,1956,1977]" italics="true" pageId="13" pageNumber="548">V. amoena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B916BD77FFD1FFC54DFF9FFFFA8AF81A" bold="true" box="[1253,1335,1955,1977]" pageId="13" pageNumber="548">
|
||
<taxonomicNameLabel id="A225000CFFD1FFC54DFF9FFFFA8AF81A" box="[1253,1335,1955,1977]" pageId="13" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Genitalia from above (59, 63, 67), from below (60, 64, 68) and from side (61, 65, 69); spermatophore (66, 70) and its ampulla (62) from side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDD6165FFD2FFC649DD98CBFB02FE56" blockId="14.[151,1437,151,501]" pageId="14" pageNumber="549">
|
||
<emphasis id="B916BD77FFD2FFC649DD98CBFEDCFF13" bold="true" box="[199,353,151,176]" pageId="14" pageNumber="549">Comparison.</emphasis>
|
||
The new species is most similar to
|
||
<taxonomicName id="4C621AE6FFD2FFC64A1A98C5FC34FF13" box="[768,905,152,176]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="variabilis">
|
||
<emphasis id="B916BD77FFD2FFC64A1A98C5FC34FF13" box="[768,905,152,176]" italics="true" pageId="14" pageNumber="549">V. variabilis</emphasis>
|
||
</taxonomicName>
|
||
but distinguished by the following characters: epiphallus is with somewhat higher distolateral parts in profile and wider (longer) dorsal spines as well as with a deeper posteromedian notch (this notch reaches the proximal edge of these spines;
|
||
<emphasis id="B916BD77FFD2FFC64D5A98BDFBEAFF5B" box="[1088,1111,225,248]" italics="true" pageId="14" pageNumber="549">vs</emphasis>
|
||
, it reaches the distal edge of these spines); epiphallic proximal half is with a more convex dorsal edge in profile; ectoparameres and rachis are clearly longer; and formula is distinctly wider. From
|
||
<taxonomicName id="4C621AE6FFD2FFC64BF49975FCD3FEE3" box="[750,878,296,320]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="trusmadi">
|
||
<emphasis id="B916BD77FFD2FFC64BF49975FCD3FEE3" box="[750,878,296,320]" italics="true" pageId="14" pageNumber="549">V. trusmadi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD2FFC64ABF9975FC42FE9C" box="[933,1023,296,320]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="orion">
|
||
<emphasis id="B916BD77FFD2FFC64ABF9975FC42FE9C" box="[933,1023,296,320]" italics="true" pageId="14" pageNumber="549">V. orion</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs in distinctly shorter dorsal epiphallic spines, a convex dorsal edge of the proximal epiphallic part (before these spines) in profile, and clearly shorter ectoparameres; from
|
||
<taxonomicName id="4C621AE6FFD2FFC64BD9992DFC97FE24" box="[707,810,368,392]" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD2FFC64BD9992DFC97FE24" box="[707,810,368,392]" italics="true" pageId="14" pageNumber="549">V. striata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C621AE6FFD2FFC64A7F992DFC72FE2B" box="[869,975,368,392]" class="Insecta" family="Haglotettigoniidae" genus="Noctitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="glabra">
|
||
<emphasis id="B916BD77FFD2FFC64A7F992DFC72FE2B" box="[869,975,368,392]" italics="true" pageId="14" pageNumber="549">V. glabra</emphasis>
|
||
</taxonomicName>
|
||
, in the same characters of epiphallus as well as more strong and slightly S-shaped ectoparameres (
|
||
<emphasis id="B916BD77FFD2FFC64A3C99C9FC80FE0F" box="[806,829,405,428]" italics="true" pageId="14" pageNumber="549">vs</emphasis>
|
||
. not S-shaped), and a distinctly longer rachis (from
|
||
<taxonomicName id="4C621AE6FFD2FFC64C9D99C9FF5DFE6C" class="Insecta" family="Haglotettigoniidae" genus="Varitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B916BD77FFD2FFC64C9D99C9FF5DFE6C" italics="true" pageId="14" pageNumber="549">V. striata</emphasis>
|
||
</taxonomicName>
|
||
) or a comparatively deeper posteromedian epiphallic notch reaching the epiphallic spine proximal edge (from
|
||
<taxonomicName id="4C621AE6FFD2FFC649C79981FEF9FE57" box="[221,324,476,500]" class="Insecta" family="Haglotettigoniidae" genus="Noctitrella" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="549" phylum="Arthropoda" rank="species" species="glabra">
|
||
<emphasis id="B916BD77FFD2FFC649C79981FEF9FE57" box="[221,324,476,500]" italics="true" pageId="14" pageNumber="549">V. glabra</emphasis>
|
||
</taxonomicName>
|
||
; the latter species has this notch reaching the epiphallic spine distal edge only).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |