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<document id="127CF2A3218E3BC3950EFF9F7D89D08B" ID-DOI="10.11606/1807-0205/2021.61.44" ID-ISSN="1807-0205" ID-Zenodo-Dep="5008615" ID-ZooBank="FE64EA84-C28D-4CE1-AE7A-5C973BBF4D45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624299270197" checkinUser="carolina" docAuthor="Fontenelle, José Heitzmann, Simone, Luiz Ricardo L. &amp; Cavallari, Daniel Caracanhas" docDate="2021" docId="90278786FFBEFF88FC14FC01FE2AFA4A" docLanguage="en" docName="PapAvZool.61.e20216144.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 61" docSource="http://dx.doi.org/10.11606/1807-0205/2021.61.44" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Megalobulimus dryades Fontenelle &amp; Simone &amp; Cavallari 2021, sp. nov." docType="treatment" docUuid="C413C457-BEF0-478E-8886-8F0CE9473724" docUuidSource="ZooBank" docVersion="3" lastPageNumber="10" masterDocId="6C1EFFFEFFBFFF81FF8CFF9BFFAEFF99" masterDocTitle="Megalobulimus dryades, a new species from the Atlantic Forest in southeastern Brazil, and redescription of Megalobulimus gummatus (Gastropoda: Strophocheilidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="2" updateTime="1698970361171" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="741E1ACD5F13F94544B55B63A693D8E9">Megalobulimus dryades, a new species from the Atlantic Forest in southeastern Brazil, and redescription of Megalobulimus gummatus (Gastropoda: Strophocheilidae)</mods:title>
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<mods:namePart id="4016DDFD328781AAE040FF4F07E06B33">Fontenelle, José Heitzmann</mods:namePart>
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<mods:namePart id="3F8599AA4478D88DEE5C945058AC12BC">Simone, Luiz Ricardo L.</mods:namePart>
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<mods:namePart id="DE18ED7F9262C5EEDA9A6B31611EA9C2">Cavallari, Daniel Caracanhas</mods:namePart>
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<taxonomicName id="DF8E4D13FFBEFF80FC14FC01FB02FC2A" ID-CoL="879WG" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[920,1196,922,947]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBEFF80FC14FC01FB02FC2A" bold="true" box="[920,1196,922,947]" italics="true" pageId="1" pageNumber="2">Megalobulimus dryades</emphasis>
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<taxonomicNameLabel id="31C957F9FFBEFF80FB3DFC01FAA5FC2A" box="[1201,1291,922,947]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="80B52A15FFBEFF80FB93FC21FB2CFC4A" box="[1055,1154,954,979]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure7" captionStart-7="Figure 8" captionStartId-0="1.[793,852,2042,2067]" captionStartId-1="2.[170,229,2042,2067]" captionStartId-2="3.[96,155,2074,2099]" captionStartId-3="4.[171,230,1224,1249]" captionStartId-4="4.[170,229,2074,2099]" captionStartId-5="5.[94,153,2074,2099]" captionStartId-6="6.[170,229,2074,2099]" captionStartId-7="7.[94,153,820,845]" captionTargetBox-0="[794,1418,1615,2012]" captionTargetBox-1="[227,1436,204,2012]" captionTargetBox-2="[94,1417,1118,2044]" captionTargetBox-3="[173,1489,534,1193]" captionTargetBox-4="[170,1493,1307,2044]" captionTargetBox-5="[95,1413,857,2041]" captionTargetBox-6="[172,1489,459,2037]" captionTargetBox-7="[96,1412,204,790]" captionTargetId-0="figure-742@1.[794,1418,1615,2012]" captionTargetId-1="figure-1@2.[227,1436,204,2012]" captionTargetId-2="figure-443@3.[94,1417,1118,2044]" captionTargetId-3="figure-156@4.[170,1493,532,1193]" captionTargetId-4="figure-187@4.[170,1493,1307,2044]" captionTargetId-5="figure-303@5.[94,1417,854,2045]" captionTargetId-6="figure-127@6.[170,1493,457,2044]" captionTargetId-7="figure-693@7.[94,1417,204,790]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionTargetPageId-6="6" captionTargetPageId-7="7" captionText-0="Figure 1. Megalobulimus dryades sp. nov., live specimen in the Atlantic Forest, Fazenda Intervales, São Miguel Arcanjo Municipality, São Paulo state (photo by B.M.Tomotani)." captionText-1="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." captionText-2="Figure 3. Megalobulimus dryades sp. nov. anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM." captionText-3="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." captionText-4="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." captionText-5="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." captionText-6="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." captionText-7="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5008619" figureDoi-1="http://doi.org/10.5281/zenodo.5008621" figureDoi-2="http://doi.org/10.5281/zenodo.5008623" figureDoi-3="http://doi.org/10.5281/zenodo.5008625" figureDoi-4="http://doi.org/10.5281/zenodo.5008627" figureDoi-5="http://doi.org/10.5281/zenodo.5008630" figureDoi-6="http://doi.org/10.5281/zenodo.5008634" figureDoi-7="http://doi.org/10.5281/zenodo.5008636" httpUri-0="https://zenodo.org/record/5008619/files/figure.png" httpUri-1="https://zenodo.org/record/5008621/files/figure.png" httpUri-2="https://zenodo.org/record/5008623/files/figure.png" httpUri-3="https://zenodo.org/record/5008625/files/figure.png" httpUri-4="https://zenodo.org/record/5008627/files/figure.png" httpUri-5="https://zenodo.org/record/5008630/files/figure.png" httpUri-6="https://zenodo.org/record/5008634/files/figure.png" httpUri-7="https://zenodo.org/record/5008636/files/figure.png" pageId="1" pageNumber="2">Figs. 1-8</figureCitation>
) http://zoobank.org/ 
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<taxonomicName id="DF8E4D13FFBEFF80FC96FB81FA88FBAA" ID-CoL="8QGPS" authority=": Morretes, 1949: 143" authorityName=": Morretes" authorityPageNumber="143" authorityYear="1949" box="[794,1318,1050,1075]" class="Gastropoda" family="Strophocheilidae" genus="Strophocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFBEFF80FC96FB81FB92FBAA" box="[794,1084,1050,1075]" italics="true" pageId="1" pageNumber="2">Strophocheilus gummatus:</emphasis>
<bibRefCitation id="7C1F4B61FFBEFF80FBC9FB81FA88FBAA" author="Morretes, F. L." box="[1093,1318,1050,1075]" pageId="1" pageNumber="2" pagination="5 - 216" refId="ref10099" refString="Morretes, F. L. 1949. Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paranaense, 7: 5 - 216." type="journal article" year="1949">Morretes, 1949: 143</bibRefCitation>
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(in part, non Hidalgo).
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<taxonomicName id="DF8E4D13FFBEFF80FC96FBC1FBBCFB0A" ID-CoL="7S3HW" authority=": Morretes, 1952: 113" authorityName=": Morretes" authorityPageNumber="113" authorityYear="1952" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="gummatus" subGenus="Phaiopharus">
<treatmentCitation id="992F1081FFBEFF80FC96FBC1FC03FB0A" author="Morretes, F. L." page="113" pageId="1" pageNumber="2" year="1952">
<emphasis id="2AFAEA82FFBEFF80FC96FBC1FAAAFBEA" box="[794,1284,1114,1139]" italics="true" pageId="1" pageNumber="2">Megalobulimus (Phaiopharus) gummatus:</emphasis>
<bibRefCitation id="7C1F4B61FFBEFF80FAADFBC1FC03FB0A" author="Morretes, F. L." pageId="1" pageNumber="2" pagination="109 - 128" refId="ref10126" refString="Morretes, F. L. 1952. Novas especies brasileiras de Strophocheilidae. Arquivos de Zoologia de Sao Paulo, 8: 109 - 128." type="journal article" year="1952">Morretes, 1952: 113</bibRefCitation>
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, 
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<bibRefCitation id="7C1F4B61FFBEFF80FC3AFBE1FBBCFB0A" author="Morretes, F. L." box="[950,1042,1146,1171]" pageId="1" pageNumber="2" pagination="37 - 76" refId="ref10153" refString="Morretes, F. L. 1953. Adenda e corrigenda ao ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paranaense, 10: 37 - 76." type="journal article" year="1953">1953: 65</bibRefCitation>
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(in part, non Hidalgo).
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<treatmentCitation id="992F1081FFBEFF80FC96FB01FA8AFB2A" author="Leme, J. L. M." box="[794,1316,1178,1203]" page="318" pageId="1" pageNumber="2" year="1973">
<taxonomicName id="DF8E4D13FFBEFF80FC96FB01FA8AFB2A" ID-CoL="7S3HW" authority=": Leme, 1973: 318" authorityName=": Leme" authorityPageNumber="318" authorityYear="1973" box="[794,1316,1178,1203]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFBEFF80FC96FB01FBE7FB2A" box="[794,1097,1178,1203]" italics="true" pageId="1" pageNumber="2">Megalobulimus gummatus:</emphasis>
<bibRefCitation id="7C1F4B61FFBEFF80FBD5FB01FA8AFB2A" author="Leme, J. L. M." box="[1113,1316,1178,1203]" pageId="1" pageNumber="2" pagination="295 - 337" refId="ref9592" refString="Leme, J. L. M. 1973. Anatomy and systematics of the neotropical Strophocheiloidea (Gastropoda, Pulmonata) with the description of a new family. Arquivos de Zoologia de Sao Paulo, 23: 295 - 337." type="journal article" year="1973">Leme, 1973: 318</bibRefCitation>
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, fig. 38; 
<bibRefCitation id="7C1F4B61FFBEFF80FCB3FB21FBE6FB4A" author="Sobreira, H. B. &amp; Molina, F. B." box="[831,1096,1210,1235]" pageId="1" pageNumber="2" pagination="139 - 141" refId="ref10643" refString="Sobreira, H. B. &amp; Molina, F. B. 2002. Observacoes preliminares sobre a biologia reprodutiva de Megalobulimus gummatus (Mollusca, Megalobulimidae) em laboratorio. Arquivos do Instituto BiolOgico, 69 (Supl.): 139 - 141." type="journal article" year="2002">Sobreira &amp; Molina, 2002</bibRefCitation>
; 
<treatmentCitation id="992F1081FFBEFF80FBDCFB21FA27FB4A" author="Salgado, N. C. &amp; Coelho, A. C. S." box="[1104,1417,1210,1235]" page="158" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="7C1F4B61FFBEFF80FBDCFB21FA27FB4A" author="Salgado, N. C. &amp; Coelho, A. C. S." box="[1104,1417,1210,1235]" pageId="1" pageNumber="2" pagination="149 - 189" refId="ref10375" refString="Salgado, N. C. &amp; Coelho, A. C. S. 2003. MoluscosterrestresdoBrasil (Gastropodes operculados ou nao exclusive Veronicellide, Milacidae e Limacidae). Revista de Biologia Tropical, 51 (Supl. 3): 149 - 189." type="journal article" year="2003">Salgado &amp; Coelho, 2003: 158</bibRefCitation>
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(in part); 
<treatmentCitation id="992F1081FFBEFF80FC29FB41FBDDFB6A" author="Simone, L. R. L." box="[933,1139,1242,1267]" page="211" pageId="1" pageNumber="2" year="2006">
<bibRefCitation id="7C1F4B61FFBEFF80FC29FB41FBDDFB6A" author="Simone, L. R. L." box="[933,1139,1242,1267]" pageId="1" pageNumber="2" refId="ref10534" refString="Simone, L. R. L. 2006. Land and Freshwater Molluscs of Brazil. Sao Paulo, EGB / FAPESP." type="book" year="2006">Simone, 2006: 211</bibRefCitation>
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, fig. 804 (in part); 
<treatmentCitation id="992F1081FFBEFF80FAB7FB41FC79FA8A" author="Thome, J. W. &amp; Arruda, J. O. &amp; Silva, L. F." page="22" pageId="1" pageNumber="2" year="2007">
<bibRefCitation id="7C1F4B61FFBEFF80FAB7FB41FC79FA8A" author="Thome, J. W. &amp; Arruda, J. O. &amp; Silva, L. F." pageId="1" pageNumber="2" pagination="9 - 38" refId="ref10759" refString="Thome, J. W.; Arruda, J. O. &amp; Silva, L. F. 2007. Moluscos terrestres no cone meridional da America do Sul. Ciencia e Ambiente, 35 (2): 9 - 38." type="journal article" year="2007">
Thomé 
<emphasis id="2AFAEA82FFBEFF80FCCDFB61FCD6FA8A" box="[833,888,1274,1299]" italics="true" pageId="1" pageNumber="2">et al.,</emphasis>
2007:22
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(in part); 
<treatmentCitation id="992F1081FFBEFF80FBB0FB61FAE6FA8A" author="Agudo-Padron, A." box="[1084,1352,1274,1299]" page="35" pageId="1" pageNumber="2" year="2010">
<bibRefCitation id="7C1F4B61FFBEFF80FBB0FB61FAE6FA8A" author="Agudo-Padron, A." box="[1084,1352,1274,1299]" pageId="1" pageNumber="2" pagination="32 - 37" refId="ref9013" refString="Agudo-Padron, A. 2010. The mollusc fauna of Santa Catarina state, Southern Brasil: knowledge gained from 13 years of research. Tentacle, 18: 32 - 37." type="journal article" year="2010">Agudo-Padrón, 2010:35</bibRefCitation>
</treatmentCitation>
, fig. 2, 2011:61, fig. 2; 
<treatmentCitation id="992F1081FFBEFF80FC68FA81FB49FAAA" author="Birckolz, C. &amp; Salvador, R. B. &amp; Cavallari, D. C. &amp; Simone, L. R. L." box="[996,1255,1306,1331]" page="150" pageId="1" pageNumber="2" year="2016">
<bibRefCitation id="7C1F4B61FFBEFF80FC68FA81FB49FAAA" author="Birckolz, C. &amp; Salvador, R. B. &amp; Cavallari, D. C. &amp; Simone, L. R. L." box="[996,1255,1306,1331]" pageId="1" pageNumber="2" pagination="133 - 150" refId="ref9126" refString="Birckolz, C.; Salvador, R. B.; Cavallari, D. C. &amp; Simone, L. R. L. 2016. Illustrated checklist of newly described (2006 - 2016) land and freshwater Gastropoda from Brazil. Archiv fur Molluskenkunde, 145: 133 - 150." type="journal article" year="2016">
Birckolz 
<emphasis id="2AFAEA82FFBEFF80FBCCFA81FBD6FAAA" box="[1088,1144,1306,1331]" italics="true" pageId="1" pageNumber="2">et al.,</emphasis>
2016: 150
</bibRefCitation>
</treatmentCitation>
(non Hidalgo).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFBEFF82FC96FAC1FAD5FE0A" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="18313690FFBEFF80FC96FAC1FC78FA0A" blockId="1.[794,1417,1370,1427]" pageId="1" pageNumber="2">
<emphasis id="2AFAEA82FFBEFF80FC96FAC1FC4EFAEA" bold="true" box="[794,992,1370,1395]" pageId="1" pageNumber="2">Type specimens:</emphasis>
<materialsCitation id="A8E63CCDFFBEFF80FC64FAC1FC78FA0A" collectingDate="1961-07-01" collectionCode="MZSP" collectorName="J. L. M. Leme" pageId="1" pageNumber="2" specimenCode="MZSP 120000" specimenCount="1" typeStatus="holotype">
<typeStatus id="C7358832FFBEFF80FC64FAC1FBFCFAEA" box="[1000,1106,1370,1395]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="48289EEBFFBEFF80FBD1FAC1FB59FAEA" box="[1117,1271,1370,1395]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 120000</specimenCode>
(
<collectorName id="B57B5346FFBEFF80FA8BFAC1FA27FAEA" box="[1287,1417,1370,1395]" pageId="1" pageNumber="2">J.L.M. Leme</collectorName>
col., 
<date id="6C301050FFBEFF80FCC7FAE1FC65FA0A" box="[843,971,1402,1427]" pageId="1" pageNumber="2" value="1961-07-01">
<collectingDate id="7C74E9B8FFBEFF80FCC7FAE1FC65FA0A" box="[843,971,1402,1427]" pageId="1" pageNumber="2" value="1961-07-01">01/vii/1961</collectingDate>
</date>
).
</materialsCitation>
</paragraph>
<paragraph id="18313690FFBEFF82FC96FA21FF3DFECA" blockId="1.[794,1419,1466,1587]" lastBlockId="3.[93,719,218,339]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<materialsCitation id="A8E63CCDFFBEFF80FC96FA21FB1BFA6A" collectingDate="1961-07-01" collectionCode="MZSP" collectorName="Iporanga &amp; J. L. M. Leme" country="Brazil" location="Sao Paulo" pageId="1" pageNumber="2" specimenCode="MZSP 15594, 13" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis id="2AFAEA82FFBEFF80FC96FA21FC39FA4A" bold="true" box="[794,919,1466,1491]" pageId="1" pageNumber="2">
<typeStatus id="C7358832FFBEFF80FC96FA21FC3DFA4A" box="[794,915,1466,1491]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="60997600FFBEFF80FC2CFA21FC5CFA4A" box="[928,1010,1466,1491]" name="Brazil" pageId="1" pageNumber="2">BRAZIL</collectingCountry>
. 
<emphasis id="2AFAEA82FFBEFF80FC71FA21FBD2FA4A" bold="true" box="[1021,1148,1466,1491]" pageId="1" pageNumber="2">
<collectingRegion id="DA4AF872FFBEFF80FC71FA21FBD9FA4A" box="[1021,1143,1466,1491]" country="Brazil" name="Sao Paulo" pageId="1" pageNumber="2">São Paulo</collectingRegion>
;
</emphasis>
<collectorName id="B57B5346FFBEFF80FB08FA21FB45FA4A" box="[1156,1259,1466,1491]" pageId="1" pageNumber="2">Iporanga</collectorName>
, 
<specimenCode id="48289EEBFFBEFF80FB7BFA21FA28FA4A" box="[1271,1414,1466,1491]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 15594</specimenCode>
, 
<specimenCode id="48289EEBFFBEFF80FC96FA41FC9BFA6A" box="[794,821,1498,1523]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">13</specimenCode>
spm (
<collectorName id="B57B5346FFBEFF80FCFBFA41FC59FA6A" box="[887,1015,1498,1523]" pageId="1" pageNumber="2">J.L.M. Leme</collectorName>
col., 
<date id="6C301050FFBEFF80FBA2FA41FB00FA6A" box="[1070,1198,1498,1523]" pageId="1" pageNumber="2" value="1961-07-01">
<collectingDate id="7C74E9B8FFBEFF80FBA2FA41FB00FA6A" box="[1070,1198,1498,1523]" pageId="1" pageNumber="2" value="1961-07-01">01/vii/1961</collectingDate>
</date>
)
</materialsCitation>
; 
<materialsCitation id="A8E63CCDFFBEFF80FB32FA41FA28F98A" collectingDate="1992-03-22" collectingDateMax="1992-03-26" collectingDateMin="1992-03-22" collectionCode="MZSP" collectorName="Guapiara &amp; Fazenda Intervales &amp; Exp." country="Brazil" pageId="1" pageNumber="2" specimenCode="MZSP 28925, 2" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectorName id="B57B5346FFBEFF80FB32FA41FA8BFA6A" box="[1214,1317,1498,1523]" pageId="1" pageNumber="2">Guapiará</collectorName>
, 
<collectorName id="B57B5346FFBEFF80FAA1FA41FC27F98A" pageId="1" pageNumber="2">Fazenda Intervales</collectorName>
, 
<specimenCode id="48289EEBFFBEFF80FC1EFA61FBB0F98A" box="[914,1054,1530,1555]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 28925</specimenCode>
, 
<specimenCode id="48289EEBFFBEFF80FBABFA61FB9BF98A" box="[1063,1077,1530,1555]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">2</specimenCode>
sh (
<collectionCode id="7E9FAE55FFBEFF80FBECFA61FB0CF98A" box="[1120,1186,1530,1555]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP</collectionCode>
<collectorName id="B57B5346FFBEFF80FB24FA61FB77F98A" box="[1192,1241,1530,1555]" pageId="1" pageNumber="2">Exp.</collectorName>
, 
<date id="6C301050FFBEFF80FB6DFA61FA2EF98A" box="[1249,1408,1530,1555]" pageId="1" pageNumber="2" value="1992-03-22" valueMax="1992-03-26" valueMin="1992-03-22">
<collectingDate id="7C74E9B8FFBEFF80FB6DFA61FA2EF98A" box="[1249,1408,1530,1555]" pageId="1" pageNumber="2" value="1992-03-22" valueMax="1992-03-26" valueMin="1992-03-22">22-26/iii/1992</collectingDate>
</date>
)
</materialsCitation>
; 
<materialsCitation id="A8E63CCDFFBEFF82FC96F981FD65FE8A" collectingDate="1997-10-03" collectionCode="MZSP" collectorName="Iguape &amp; Despraiado &amp; Rocha &amp; Bertani &amp; Mestre &amp; N. Moraccioli &amp; R. Krone" country="Brazil" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" specimenCode="MZSP 28773, 2, MZSP 28893, MZSP 29317" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectorName id="B57B5346FFBEFF80FC96F981FCC4F9AA" box="[794,874,1562,1587]" pageId="1" pageNumber="2">Iguape</collectorName>
,NÚcleo 
<collectorName id="B57B5346FFBEFF80FC48F981FBEBF9AA" box="[964,1093,1562,1587]" pageId="1" pageNumber="2">Despraiado</collectorName>
, 
<specimenCode id="48289EEBFFBEFF80FBC1F981FB76F9AA" box="[1101,1240,1562,1587]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 28773</specimenCode>
, 
<specimenCode id="48289EEBFFBEFF80FB6CF981FB40F9AA" box="[1248,1262,1562,1587]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">2</specimenCode>
sh (R.P. 
<collectorName id="B57B5346FFBEFF80FACCF981FA29F9AA" box="[1344,1415,1562,1587]" pageId="1" pageNumber="2">Rocha</collectorName>
, 
<collectorName id="B57B5346FFBCFF82FFD2FF41FF00FF6A" box="[94,174,218,243]" pageId="3" pageNumber="4">Bertani</collectorName>
&amp; 
<collectorName id="B57B5346FFBCFF82FF5DFF41FEB3FF6A" box="[209,285,218,243]" pageId="3" pageNumber="4">Mestre</collectorName>
cols., 
<date id="6C301050FFBCFF82FEE9FF41FE74FF6A" box="[357,474,218,243]" pageId="3" pageNumber="4" value="1997-10-03">
<collectingDate id="7C74E9B8FFBCFF82FEE9FF41FE74FF6A" box="[357,474,218,243]" pageId="3" pageNumber="4" value="1997-10-03">03/x/1997</collectingDate>
</date>
), 
<specimenCode id="48289EEBFFBCFF82FE60FF41FDD5FF6A" box="[492,635,218,243]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP 28893</specimenCode>
, 
<specimenCode id="48289EEBFFBCFF82FD0BFF41FD3BFF6A" box="[647,661,218,243]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">2</specimenCode>
spm (
<collectorName id="B57B5346FFBCFF82FFEAFF61FF5BFE8A" box="[102,245,250,275]" pageId="3" pageNumber="4">N. Moraccioli</collectorName>
col.), 
<specimenCode id="48289EEBFFBCFF82FEBDFF61FE12FE8A" box="[305,444,250,275]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP 29317</specimenCode>
, 
<specimenCode id="48289EEBFFBCFF82FE4FFF61FE7FFE8A" box="[451,465,250,275]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">2</specimenCode>
sh (
<collectorName id="B57B5346FFBCFF82FE76FF61FDF8FE8A" box="[506,598,250,275]" pageId="3" pageNumber="4">R. Krone</collectorName>
col., 1901)
</materialsCitation>
. 
<materialsCitation id="A8E63CCDFFBCFF82FFD1FE81FF21FECA" collectionCode="MZSP" collectorName="Lange" country="Brazil" location="Morretes" pageId="3" pageNumber="4" specimenCode="MZSP 16610, 2" specimenCount="1" stateProvince="Parana" typeStatus="paratype">
<emphasis id="2AFAEA82FFBCFF82FFD1FE81FF19FEAA" bold="true" box="[93,183,282,307]" pageId="3" pageNumber="4">
<collectingRegion id="DA4AF872FFBCFF82FFD1FE81FF1DFEAA" box="[93,179,282,307]" country="Brazil" name="Parana" pageId="3" pageNumber="4">Paraná</collectingRegion>
;
</emphasis>
<location id="1D51604BFFBCFF82FF32FE81FE8AFEAA" LSID="urn:lsid:plazi:treatment:90278786FFBEFF88FC14FC01FE2AFA4A:1D51604BFFBCFF82FF32FE81FE8AFEAA" box="[190,292,282,307]" country="Brazil" name="Morretes" pageId="3" pageNumber="4" stateProvince="Parana">Morretes</location>
, 
<specimenCode id="48289EEBFFBCFF82FEA1FE81FE14FEAA" box="[301,442,282,307]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP 16610</specimenCode>
, 
<specimenCode id="48289EEBFFBCFF82FE4FFE81FE7FFEAA" box="[451,465,282,307]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">2</specimenCode>
sh (
<collectorName id="B57B5346FFBCFF82FE70FE81FDECFEAA" box="[508,578,282,307]" pageId="3" pageNumber="4">Lange</collectorName>
de Morretes col.)
</materialsCitation>
.
</paragraph>
<caption id="4CF16618FFBEFF80FC95F861FC5EF7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008619" ID-Zenodo-Dep="5008619" httpUri="https://zenodo.org/record/5008619/files/figure.png" pageId="1" pageNumber="2" startId="1.[793,852,2042,2067]" targetBox="[794,1418,1615,2012]" targetPageId="1">
<paragraph id="18313690FFBEFF80FC95F861FC5EF7CA" blockId="1.[793,1417,2042,2131]" pageId="1" pageNumber="2">
Figure 1. 
<taxonomicName id="DF8E4D13FFBEFF80FCFAF861FB99F78A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[886,1079,2042,2067]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBEFF80FCFAF861FB99F78A" bold="true" box="[886,1079,2042,2067]" italics="true" pageId="1" pageNumber="2">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBEFF80FBB3F861FB25F78A" box="[1087,1163,2042,2067]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
, live specimen in the Atlantic Forest, Fazenda Intervales, São Miguel Arcanjo Municipality, São Paulo state (photo by B.M.Tomotani).
</paragraph>
</caption>
<caption id="4CF16618FFBDFF83FF26F861FADAF7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008621" ID-Zenodo-Dep="5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="2" pageNumber="3" startId="2.[170,229,2042,2067]" targetBox="[227,1436,204,2012]" targetPageId="2">
<paragraph id="18313690FFBDFF83FF26F861FADAF7CA" blockId="2.[169,1493,2042,2131]" pageId="2" pageNumber="3">
Figure 2. Shell of 
<taxonomicName id="DF8E4D13FFBDFF83FEB1F861FE55F78A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[317,507,2042,2067]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBDFF83FEB1F861FE55F78A" bold="true" box="[317,507,2042,2067]" italics="true" pageId="2" pageNumber="3">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBDFF83FE73F861FDE9F78A" box="[511,583,2042,2067]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM.
</paragraph>
</caption>
<paragraph id="18313690FFBCFF82FFD2FEE1FE9BFE4A" blockId="3.[94,719,378,467]" pageId="3" pageNumber="4">
<emphasis id="2AFAEA82FFBCFF82FFD2FEE1FEA3FE0A" bold="true" box="[94,269,378,403]" pageId="3" pageNumber="4">Type locality:</emphasis>
<collectingCountry id="60997600FFBCFF82FEAAFEE1FECAFE0A" box="[294,356,378,403]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
, 
<collectingRegion id="DA4AF872FFBCFF82FEF3FEE1FDE0FE0A" box="[383,590,378,403]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="4">São Paulo state</collectingRegion>
, Iporanga Municipality (center coordinates 
<geoCoordinate id="7DBA5057FFBCFF82FE7BFE01FDE2FE2A" box="[503,588,408,435]" degrees="24" direction="south" minutes="35" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-24.583334">
24°35 
<emphasis id="2AFAEA82FFBCFF82FDBAFE03FD92FE2B" bold="true" box="[566,572,408,434]" italics="true" pageId="3" pageNumber="4">′</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7DBA5057FFBCFF82FDE8FE01FD6AFE2A" box="[612,708,408,435]" degrees="48" direction="west" minutes="35" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="-48.583332">
48°35 
<emphasis id="2AFAEA82FFBCFF82FD2FFE03FD07FE2B" bold="true" box="[675,681,408,434]" italics="true" pageId="3" pageNumber="4">′</emphasis>
W
</geoCoordinate>
), ~ 
<quantity id="DF769B75FFBCFF82FFE7FE21FF0BFE4A" box="[107,165,442,467]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="m" value="80.0">80 m</quantity>
of elevation.
</paragraph>
<paragraph id="18313690FFBCFF82FFD2FE61FAD5FE0A" blockId="3.[93,720,506,1075]" lastBlockId="3.[793,1418,218,403]" pageId="3" pageNumber="4">
<emphasis id="2AFAEA82FFBCFF82FFD2FE61FE72FD8A" bold="true" box="[94,476,506,531]" pageId="3" pageNumber="4">Material examined (non-types):</emphasis>
<collectingCountry id="60997600FFBCFF82FE67FE61FD95FD8A" box="[491,571,506,531]" name="Brazil" pageId="3" pageNumber="4">BRAZIL</collectingCountry>
. 
<emphasis id="2AFAEA82FFBCFF82FDC0FE61FD60FD8A" bold="true" box="[588,718,506,531]" pageId="3" pageNumber="4">
<collectingRegion id="DA4AF872FFBCFF82FDC0FE61FD64FD8A" box="[588,714,506,531]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="4">São Paulo</collectingRegion>
;
</emphasis>
Sorocaba, 
<collectionCode id="7E9FAE55FFBCFF82FF59FD81FEBAFDAA" box="[213,276,538,563]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
62129 (1 sh); Mogi das Cruzes, 
<collectionCode id="7E9FAE55FFBCFF82FD03FD81FD60FDAA" box="[655,718,538,563]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
28962 (1 sh); Itapecerica da Serra, 
<collectionCode id="7E9FAE55FFBCFF82FE5DFDA1FDBEFDCA" box="[465,528,570,595]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
15724 (1 sh); São Sebastião, 
<collectionCode id="7E9FAE55FFBCFF82FF5EFDC1FEBFFDEA" box="[210,273,602,627]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
96475 (2 sh); Cubatão, 
<collectionCode id="7E9FAE55FFBCFF82FD89FDC1FDEAFDEA" box="[517,580,602,627]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
64563 (1 sh); Juquetiba, 
<collectionCode id="7E9FAE55FFBCFF82FF5AFDE1FEBBFD0A" box="[214,277,634,659]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
92832 (2 sh); Tapiraí, 
<collectionCode id="7E9FAE55FFBCFF82FE72FDE1FD93FD0A" box="[510,573,634,659]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
16039 (5 sh); São Vicente, 
<collectionCode id="7E9FAE55FFBCFF82FF78FD01FE9DFD2A" box="[244,307,666,691]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
16615 (1 sh); Itapeva, 
<collectionCode id="7E9FAE55FFBCFF82FDCDFD01FD2EFD2A" box="[577,640,666,691]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29320 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FF25FD21FF46FD4A" box="[169,232,698,723]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
96474 (2 sh); Itanhaém, 
<collectionCode id="7E9FAE55FFBCFF82FE76FD21FD97FD4A" box="[506,569,698,723]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
42350 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FFD2FD41FF33FD6A" box="[94,157,730,755]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
92833 (1 sh); Guapiará, 
<collectionCode id="7E9FAE55FFBCFF82FE2CFD41FE71FD6A" box="[416,479,730,755]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29316 (1 sh); Fazenda Intervales, 
<collectionCode id="7E9FAE55FFBCFF82FF57FD61FEB4FC8A" box="[219,282,762,787]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
30460 (1 sh); Miracatu, 
<collectionCode id="7E9FAE55FFBCFF82FDB3FD61FDD0FC8A" box="[575,638,762,787]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49935 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FF35FC81FF56FCAA" box="[185,248,794,819]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49936 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FE12FC81FE4FFCAA" box="[414,481,794,819]" pageId="3" pageNumber="4">FPZSP</collectionCode>
, 
<collectionCode id="7E9FAE55FFBCFF82FE64FC81FD89FCAA" box="[488,551,794,819]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49937,(1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FFD2FCA1FF33FCCA" box="[94,157,826,851]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49940 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FEDDFCA1FE3EFCCA" box="[337,400,826,851]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49941 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FDCFFCA1FD2CFCCA" box="[579,642,826,851]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49942 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FF37FCC1FF54FCEA" box="[187,250,858,883]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49944 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FE28FCC1FE4DFCEA" box="[420,483,858,883]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
49945 (1 spm); Pedro de Toledo, 
<collectionCode id="7E9FAE55FFBCFF82FF58FCE1FEBDFC0A" box="[212,275,890,915]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
92837 (1 sh); Itariri, 
<collectionCode id="7E9FAE55FFBCFF82FE67FCE1FD84FC0A" box="[491,554,890,915]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
48256 (1 spm), 
<collectionCode id="7E9FAE55FFBCFF82FFD3FC01FF30FC2A" box="[95,158,922,947]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
96477 (2 sh); Peruíbe, 
<collectionCode id="7E9FAE55FFBCFF82FE2AFC01FE4BFC2A" box="[422,485,922,947]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29309 (1 sh); Biguá, 
<collectionCode id="7E9FAE55FFBCFF82FFD2FC21FF33FC4A" box="[94,157,954,979]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
92836 (1 sh); Juquiá, 
<collectionCode id="7E9FAE55FFBCFF82FE05FC21FE66FC4A" box="[393,456,954,979]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
269 (3 sh), 
<collectionCode id="7E9FAE55FFBCFF82FDCAFC21FD2BFC4A" box="[582,645,954,979]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
90913 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FF29FC41FF4AFC6A" box="[165,228,986,1011]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
90914 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FEFAFC41FE1BFC6A" box="[374,437,986,1011]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
96478 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FDC4FC41FD29FC6A" box="[584,647,986,1011]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
96479 (1 sh); Registro, 
<collectionCode id="7E9FAE55FFBCFF82FE97FC61FEF4FB8A" box="[283,346,1018,1043]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
48523 (1 sh); Eldorado Paulista, 
<collectionCode id="7E9FAE55FFBCFF82FFD2FB81FF33FBAA" box="[94,157,1050,1075]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29315 (2 sh); Iporanga, 
<collectionCode id="7E9FAE55FFBCFF82FE3BFB81FE58FBAA" box="[439,502,1050,1075]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
7996 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FD03FB81FD60FBAA" box="[655,718,1050,1075]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
15603 (2 sh), 
<collectionCode id="7E9FAE55FFBCFF82FC2BFF41FC48FF6A" box="[935,998,218,243]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
15617 (4 sh), 
<collectionCode id="7E9FAE55FFBCFF82FBF5FF41FB16FF6A" box="[1145,1208,218,243]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
33031 (8 sh), 
<collectionCode id="7E9FAE55FFBCFF82FAC7FF41FA24FF6A" box="[1355,1418,218,243]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
43529 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FC2BFF61FC48FE8A" box="[935,998,250,275]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
43533 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FBF5FF61FB16FE8A" box="[1145,1208,250,275]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
43689 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FAC6FF61FA27FE8A" box="[1354,1417,250,275]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
43697 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FC29FE81FC4AFEAA" box="[933,996,282,307]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
92835 (1 sh); Jacupiranga, 
<collectionCode id="7E9FAE55FFBCFF82FA8EFE81FAEFFEAA" box="[1282,1345,282,307]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
32541 (1 sh); Iguape, 
<collectionCode id="7E9FAE55FFBCFF82FC49FEA1FBAAFECA" box="[965,1028,314,339]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29311 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FB20FEA1FB45FECA" box="[1196,1259,314,339]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29312 (4 sh), 
<collectionCode id="7E9FAE55FFBCFF82FC96FEC1FCF7FEEA" box="[794,857,346,371]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
71070 (1 sh); Pariquera-aÇÚ, 
<collectionCode id="7E9FAE55FFBCFF82FB27FEC1FB44FEEA" box="[1195,1258,346,371]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29313 (1 sh), 
<collectionCode id="7E9FAE55FFBCFF82FC96FEE1FCF7FE0A" box="[794,857,378,403]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
29318 (1 sh). 
<emphasis id="2AFAEA82FFBCFF82FC67FEE1FBEDFE0A" bold="true" box="[1003,1091,378,403]" pageId="3" pageNumber="4">
<collectingRegion id="DA4AF872FFBCFF82FC67FEE1FB91FE0A" box="[1003,1087,378,403]" country="Brazil" name="Parana" pageId="3" pageNumber="4">Paraná</collectingRegion>
;
</emphasis>
Sengés, 
<collectionCode id="7E9FAE55FFBCFF82FB2DFEE1FB4EFE0A" box="[1185,1248,378,403]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4" type="Museum">MZSP</collectionCode>
16603 (10 sh).
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFBCFF82FC96FE21FC7AFC8A" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="18313690FFBCFF82FC96FE21FC7AFC8A" blockId="3.[793,1418,442,787]" pageId="3" pageNumber="4">
<emphasis id="2AFAEA82FFBCFF82FC96FE21FC38FE4A" bold="true" box="[794,918,442,467]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
Shell large, with strong rugosities and malleated surface on teleoconch whorls; protoconch darker-colored with a clear lighter subsutural band, with microsculpture consisting of granules and macrosculpture of well-marked axial riblets that branch apically near suture. Peristome homogeneously white, columella oblique and slightly convex. Head-foot very light-colored, dirty white to grayish-bluish; free oviduct appendix present; penis long and uniform with convolutions, lacking developed transverse fold, bearing two short flagella of similar length.
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFBCFF86FC96FCA1FB30F80A" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4" type="description">
<paragraph id="18313690FFBCFF82FC96FCA1FC0AFCCA" blockId="3.[794,932,826,851]" box="[794,932,826,851]" pageId="3" pageNumber="4">
<emphasis id="2AFAEA82FFBCFF82FC96FCA1FC0AFCCA" bold="true" box="[794,932,826,851]" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="18313690FFBCFF84FC96FCE1FDEEFE0A" blockId="3.[794,1418,890,1075]" lastBlockId="5.[93,719,218,403]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<emphasis id="2AFAEA82FFBCFF82FC96FCE1FCFBFC0A" bold="true" box="[794,853,890,915]" pageId="3" pageNumber="4">Shell</emphasis>
(
<figureCitation id="80B52A15FFBCFF82FCEFFCE1FC6DFC0A" box="[867,963,890,915]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="1.[793,852,2042,2067]" captionStartId-1="2.[170,229,2042,2067]" captionTargetBox-0="[794,1418,1615,2012]" captionTargetBox-1="[227,1436,204,2012]" captionTargetId-0="figure-742@1.[794,1418,1615,2012]" captionTargetId-1="figure-1@2.[227,1436,204,2012]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figure 1. Megalobulimus dryades sp. nov., live specimen in the Atlantic Forest, Fazenda Intervales, São Miguel Arcanjo Municipality, São Paulo state (photo by B.M.Tomotani)." captionText-1="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi-0="http://doi.org/10.5281/zenodo.5008619" figureDoi-1="http://doi.org/10.5281/zenodo.5008621" httpUri-0="https://zenodo.org/record/5008619/files/figure.png" httpUri-1="https://zenodo.org/record/5008621/files/figure.png" pageId="3" pageNumber="4">Figs. 1-2</figureCitation>
): Large (L~ 
<quantity id="DF769B75FFBCFF82FBCCFCE1FB0AFC0A" box="[1088,1188,890,915]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="3" pageNumber="4" unit="mm" value="119.0">119 mm</quantity>
), oval, thick, of ~5.4 whorls, imperforate to semiperforate; profile strongly convex, moderately dorso-ventrally compressed (H:W = 0.8) (
<figureCitation id="80B52A15FFBCFF82FC2FFC41FC5DFC6A" box="[931,1011,986,1011]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="3" pageNumber="4">Fig. 2D</figureCitation>
); spire short and obtuse; usually not covered by periostracum in adults; color reddish ocher with yellowish-white subsutural band, ranging until last whorl. Protoconch (
<figureCitation id="80B52A15FFBBFF85FE05FF41FE45FF6A" box="[393,491,218,243]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="4" pageNumber="5">Fig. 2D, I</figureCitation>
) with ~3.2 whorls, weakly convex, usually darker than teleoconch, reddish-darkbrown with a clearly marked white-cream subsutural band; first 1¼ whorl smooth, followed by regular, complete prosocline riblets that branch apically near the suture, interspaces 2-3× wider than riblet width, presenting microscopic radial granulation that gradually increases in size toward final nepionic whorls (
<figureCitation id="80B52A15FFBBFF85FD4BFE21FCA0FE4A" box="[711,782,442,467]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="4" pageNumber="5">Fig. 2I</figureCitation>
). First ¼ teleoconch whorl with regular, thin riblets that are gradually replaced by delicate anastomosing rugosities and malleations that become larger, stronger, and coarser toward the last adult whorls (4½-5). Body whorl inflated, sculpture mostly consisting of strong rugosities and malleations (
<figureCitation id="80B52A15FFBBFF85FBAEFEC1FBF5FEEA" box="[1058,1115,346,371]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
G-H), but also presenting irregular, widely spaced axial riblets and short radial striae interconnected with the other sculpture elements. Aperture oval (
<figureCitation id="80B52A15FFBBFF85FC2BFE21FB96FE4A" box="[935,1080,442,467]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, E, G</figureCitation>
), length ~50% of shell length; peristome thickened and slightly reflected, outer lip arched, color white; columella thick, oblique,and slightly convex. Periostracum yellowish-brown, with dark radial bands in the interspaces between axial sculpture elements (
<figureCitation id="80B52A15FFBAFF84FFE9FEA1FF0DFECA" box="[101,163,314,339]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
G-H), and often darker colored in the malleation cavities;usually preserved on last two teleoconch whorls, with a glossy aspect when present (
<figureCitation id="80B52A15FFBAFF84FE6AFEE1FD9AFE0A" box="[486,564,378,403]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
).
</paragraph>
<caption id="4CF16618FFBCFF82FFECF781FDD5F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008623" ID-Zenodo-Dep="5008623" httpUri="https://zenodo.org/record/5008623/files/figure.png" pageId="3" pageNumber="4" startId="3.[96,155,2074,2099]" targetBox="[94,1417,1118,2044]" targetPageId="3">
<paragraph id="18313690FFBCFF82FFECF781FDD5F7CA" blockId="3.[95,1418,2074,2131]" pageId="3" pageNumber="4">
Figure 3. 
<taxonomicName id="DF8E4D13FFBCFF82FF39F781FEDAF7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[181,372,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBCFF82FF39F781FEDAF7AA" bold="true" box="[181,372,2074,2099]" italics="true" pageId="3" pageNumber="4">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBCFF82FEF4F781FE11F7AA" box="[376,447,2074,2099]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM.
</paragraph>
</caption>
<caption id="4CF16618FFBBFF85FF27FB53FDB5FA98" ID-DOI="http://doi.org/10.5281/zenodo.5008625" ID-Zenodo-Dep="5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="4" pageNumber="5" startId="4.[171,230,1224,1249]" targetBox="[173,1489,534,1193]" targetPageId="4">
<paragraph id="18313690FFBBFF85FF27FB53FDB5FA98" blockId="4.[171,1493,1224,1281]" pageId="4" pageNumber="5">
Figure 4. 
<taxonomicName id="DF8E4D13FFBBFF85FE8DFB53FE6EFB78" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[257,448,1224,1249]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBBFF85FE8DFB53FE6EFB78" bold="true" box="[257,448,1224,1249]" italics="true" pageId="4" pageNumber="5">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBBFF85FE48FB53FDA5FB78" box="[452,523,1224,1249]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm.
</paragraph>
</caption>
<caption id="4CF16618FFBBFF85FF26F781FBF9F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008627" ID-Zenodo-Dep="5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,229,2074,2099]" targetBox="[170,1493,1307,2044]" targetPageId="4">
<paragraph id="18313690FFBBFF85FF26F781FBF9F7CA" blockId="4.[170,1493,2074,2131]" pageId="4" pageNumber="5">
Figure 5. 
<taxonomicName id="DF8E4D13FFBBFF85FE8DF781FE6EF7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[257,448,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBBFF85FE8DF781FE6EF7AA" bold="true" box="[257,448,2074,2099]" italics="true" pageId="4" pageNumber="5">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBBFF85FE49F781FDA3F7AA" box="[453,525,2074,2099]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="18313690FFBAFF84FFD2FE21FE7EFD8A" blockId="5.[94,718,442,531]" pageId="5" pageNumber="6">
<emphasis id="2AFAEA82FFBAFF84FFD2FE21FF70FE4A" bold="true" box="[94,222,442,467]" pageId="5" pageNumber="6">Head-foot:</emphasis>
Tegument in living specimen very light-colored, from dirty white to grayish-bluish (
<figureCitation id="80B52A15FFBAFF84FD9AFE41FDE0FE6A" box="[534,590,474,499]" captionStart="Figure 1" captionStartId="1.[793,852,2042,2067]" captionTargetBox="[794,1418,1615,2012]" captionTargetId="figure-742@1.[794,1418,1615,2012]" captionTargetPageId="1" captionText="Figure 1. Megalobulimus dryades sp. nov., live specimen in the Atlantic Forest, Fazenda Intervales, São Miguel Arcanjo Municipality, São Paulo state (photo by B.M.Tomotani)." figureDoi="http://doi.org/10.5281/zenodo.5008619" httpUri="https://zenodo.org/record/5008619/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
); each buccal flange bearing seven papillae.
</paragraph>
<paragraph id="18313690FFBAFF84FFD2FDA1FBC4FE0A" blockId="5.[93,719,570,819]" lastBlockId="5.[794,1418,218,403]" pageId="5" pageNumber="6">
<emphasis id="2AFAEA82FFBAFF84FFD2FDA1FE3BFDCA" bold="true" box="[94,405,570,595]" pageId="5" pageNumber="6">Pulmonary (pallial) cavity:</emphasis>
Plexus of vessels containing divisions and anastomoses of collar vessel tributaries (
<figureCitation id="80B52A15FFBAFF84FFE8FDE1FF13FD0A" box="[100,189,634,659]" captionStart="Figure 3" captionStartId="3.[96,155,2074,2099]" captionTargetBox="[94,1417,1118,2044]" captionTargetId="figure-443@3.[94,1417,1118,2044]" captionTargetPageId="3" captionText="Figure 3. Megalobulimus dryades sp. nov. anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008623" httpUri="https://zenodo.org/record/5008623/files/figure.png" pageId="5" pageNumber="6">Figs. 3A</figureCitation>
, 
<figureCitation id="80B52A15FFBAFF84FF48FDE1FF4DFD0A" box="[196,227,634,659]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">4B</figureCitation>
:cv) triangular, of radicular aspect, occupying ~ 
<emphasis id="2AFAEA82FFBAFF84FFE2FD00FF2FFD2D" bold="true" box="[110,129,667,692]" italics="true" pageId="5" pageNumber="6">⅖</emphasis>
of cavity volume, located between mantle edge and septum. Cardiac area occupying ~ 
<emphasis id="2AFAEA82FFBAFF84FE6FFD20FE5BFD4D" bold="true" box="[483,501,699,724]" italics="true" pageId="5" pageNumber="6">⅓</emphasis>
of reno-pericardial area (
<figureCitation id="80B52A15FFBAFF84FF10FD41FF45FD6A" box="[156,235,730,755]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: ht), with equidistant insertions of tributaries of pericardial vessel (
<figureCitation id="80B52A15FFBAFF84FEEBFD61FE1BFC8A" box="[359,437,762,787]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: vp). Insertion of palio-diaphragmatic muscle (
<figureCitation id="80B52A15FFBAFF84FEDEFC81FE0CFCAA" box="[338,418,794,819]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: pv) in angular pulmonary cavity deep in kidney (
<figureCitation id="80B52A15FFBAFF84FBACFF41FBDFFF6A" box="[1056,1137,218,243]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: pd). Pulmonary septum (
<figureCitation id="80B52A15FFBAFF84FCACFF61FCD2FE8A" box="[800,892,250,275]" captionStart="Figure 3" captionStartId="3.[96,155,2074,2099]" captionTargetBox="[94,1417,1118,2044]" captionTargetId="figure-443@3.[94,1417,1118,2044]" captionTargetPageId="3" captionText="Figure 3. Megalobulimus dryades sp. nov. anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008623" httpUri="https://zenodo.org/record/5008623/files/figure.png" pageId="5" pageNumber="6">Figs. 3A</figureCitation>
, 
<figureCitation id="80B52A15FFBAFF84FC04FF61FC6BFE8A" box="[904,965,250,275]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">4A, B</figureCitation>
: se) separating excretory/respiratory areas, covered by plexus of vessels, extending through middle surface of reno-pericardial area (
<figureCitation id="80B52A15FFBAFF84FB71FEA1FAFDFECA" box="[1277,1363,314,339]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: ef). Pneumostome with strong anal folds anastomoses extending toward its inner edge.
</paragraph>
<paragraph id="18313690FFBAFF84FC96FE21FB73FD4A" blockId="5.[794,1419,442,723]" pageId="5" pageNumber="6">
<emphasis id="2AFAEA82FFBAFF84FC96FE21FC5FFE4A" bold="true" box="[794,1009,442,467]" pageId="5" pageNumber="6">Excretory system:</emphasis>
Kidney (
<figureCitation id="80B52A15FFBAFF84FBD0FE21FB1FFE4A" box="[1116,1201,442,467]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: ki) sub-triangular, occupying ~½ of pulmonary septum volume, with protuberance at edge close to nephrostome (
<figureCitation id="80B52A15FFBAFF84FA88FE61FAF8FD8A" box="[1284,1366,506,531]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: np) near transition between adrenal vessels (
<figureCitation id="80B52A15FFBAFF84FB52FD81FA82FDAA" box="[1246,1324,538,563]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
: nv) and ad-rectal vessels (
<figureCitation id="80B52A15FFBAFF84FC53FDA1FBFBFDCA" box="[991,1109,570,595]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Figs. 4A, B</figureCitation>
: rv). Ureter absent. Folds of urinary gutter (
<figureCitation id="80B52A15FFBAFF84FC49FDC1FB92FDEA" box="[965,1084,602,627]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Figs. 4A, B</figureCitation>
: ug) beginning near nephrostome level, flanking left side of rectum towards pneumostome (
<figureCitation id="80B52A15FFBAFF84FC18FD01FC4BFD2A" box="[916,997,666,691]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4B</figureCitation>
: pn), partially covering rectum at its final half (
<figureCitation id="80B52A15FFBAFF84FC09FD21FC7FFD4A" box="[901,977,698,723]" captionStart="Figure 4" captionStartId="4.[171,230,1224,1249]" captionTargetBox="[173,1489,534,1193]" captionTargetId="figure-156@4.[170,1493,532,1193]" captionTargetPageId="4" captionText="Figure 4. Megalobulimus dryades sp. nov. anatomy: (A) pallial chamber, renopericardial region, ventral view; (B) pneumostome region, ventral view, inner lip or pneumostome deflected.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008625" httpUri="https://zenodo.org/record/5008625/files/figure.png" pageId="5" pageNumber="6">Fig. 4B</figureCitation>
: rt), ending before anus.
</paragraph>
<paragraph id="18313690FFBAFF84FC96FD61FA24FCAA" blockId="5.[794,1418,762,819]" pageId="5" pageNumber="6">
<emphasis id="2AFAEA82FFBAFF84FC96FD61FC5CFC8A" bold="true" box="[794,1010,762,787]" pageId="5" pageNumber="6">Digestive system:</emphasis>
Buccal mass robust (
<figureCitation id="80B52A15FFBAFF84FB70FD61FAFDFC8A" box="[1276,1363,762,787]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
: ot), retractor muscle inserted laterally in radular pouch.
</paragraph>
<caption id="4CF16618FFBAFF84FFD2F781FAB8F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008630" ID-Zenodo-Dep="5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,153,2074,2099]" targetBox="[95,1413,857,2041]" targetPageId="5">
<paragraph id="18313690FFBAFF84FFD2F781FAB8F7CA" blockId="5.[94,1417,2074,2131]" pageId="5" pageNumber="6">
Figure 6. 
<taxonomicName id="DF8E4D13FFBAFF84FF3EF781FEDEF7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[178,368,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFBAFF84FF3EF781FEDEF7AA" bold="true" box="[178,368,2074,2099]" italics="true" pageId="5" pageNumber="6">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFBAFF84FEF8F781FE14F7AA" box="[372,442,2074,2099]" pageId="5" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
anatomy:(A)stomach,right view,opened longitudinally along dorsal surface;(B)same,left view;(C) intestine,transversally opened strip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="18313690FFB9FF86FF26FF41FD6EFBCA" blockId="6.[169,795,218,435]" lastBlockId="7.[94,719,922,1107]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Odontophore rounded (
<figureCitation id="80B52A15FFB9FF87FE33FF41FDBCFF6A" box="[447,530,218,243]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
: od), occupying ~¼ of buccal mass volume. Aperture of salivary gland ducts (
<figureCitation id="80B52A15FFB9FF87FF3DFE81FEAAFEAA" box="[177,260,282,307]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
: sa) located distally and laterally on ceiling of buccal mass (
<figureCitation id="80B52A15FFB9FF87FECBFEA1FE34FECA" box="[327,410,314,339]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
: bm). Jaw arc-shaped, rigid, elasmognate,pigmented (
<figureCitation id="80B52A15FFB9FF87FE13FEC1FE5BFEEA" box="[415,501,346,371]" captionStart="Figure 3" captionStartId="3.[96,155,2074,2099]" captionTargetBox="[94,1417,1118,2044]" captionTargetId="figure-443@3.[94,1417,1118,2044]" captionTargetPageId="3" captionText="Figure 3. Megalobulimus dryades sp. nov. anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008623" httpUri="https://zenodo.org/record/5008623/files/figure.png" pageId="6" pageNumber="7">Figs. 3B</figureCitation>
, 
<figureCitation id="80B52A15FFB9FF87FE71FEC1FDB0FEEA" box="[509,542,346,371]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">5A</figureCitation>
: jw), with ~35 irregular, narrow and well-marked axial columns and transverse growth lines. Pair of muscles m7 surrounding radular sac in region preceding exposed radula, anteriorly protected by hardened tissue (
<figureCitation id="80B52A15FFB9FF87FBCDFF61FB21FE8A" box="[1089,1167,250,275]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
: bt); dorsal surface of odontophore cartilages rigid and darker-colored (
<figureCitation id="80B52A15FFB9FF87FAB2FE81FAD7FEAA" box="[1342,1401,282,307]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
B-C: oc), with groove obstructed by elliptical scar. Horizontal muscle (
<figureCitation id="80B52A15FFB9FF87FC4FFEC1FBA2FEEA" box="[963,1036,346,371]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="6" pageNumber="7">Figs. 5</figureCitation>
B-C: m6) connecting midline of odontophore cartilages; length ~½ of cartilage length. Fusion of main and secondary dorsal tensor muscles of radula (
<figureCitation id="80B52A15FFB8FF86FFE9FC01FF1DFC2A" box="[101,179,922,947]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
: m5-m4) originating at anterior margin of cartilage, ventral tensor muscle of radula (
<figureCitation id="80B52A15FFB8FF86FE77FC21FDE9FC4A" box="[507,583,954,979]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5C</figureCitation>
: m11) along insertion of retractor muscle of buccal mass (
<figureCitation id="80B52A15FFB8FF86FDC5FC41FD3AFC6A" box="[585,660,986,1011]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
: m2), both inserting respectively in subradular membrane (
<figureCitation id="80B52A15FFB8FF86FFE9FB81FF31FBAA" box="[101,159,1050,1075]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
B-C: br). Jaw muscles (
<figureCitation id="80B52A15FFB8FF86FE1DFB81FE72FBAA" box="[401,476,1050,1075]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
: mj) originating at lateral margin of cartilage, surrounding oral tube dorsally.
</paragraph>
<caption id="4CF16618FFB9FF87FF26F781FA78F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008634" ID-Zenodo-Dep="5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="6" pageNumber="7" startId="6.[170,229,2074,2099]" targetBox="[172,1489,459,2037]" targetPageId="6">
<paragraph id="18313690FFB9FF87FF26F781FA78F7CA" blockId="6.[170,1494,2074,2131]" pageId="6" pageNumber="7">
Figure7. 
<taxonomicName id="DF8E4D13FFB9FF87FF70F781FE16F7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[252,440,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB9FF87FF70F781FE16F7AA" bold="true" box="[252,440,2074,2099]" italics="true" pageId="6" pageNumber="7">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFB9FF87FE37F781FE51F7AA" box="[443,511,2074,2099]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
anatomy:(A)genital system,complete dorsal view;(B)Carrefour region,ventral view;(C)albumen chamber opened,exposing its folds;(D)spermoviduct,cross-section at its middle region;(E)anterior region of genital ducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm.
</paragraph>
</caption>
<caption id="4CF16618FFB8FF86FFD2FCAFFCF2FCF4" ID-DOI="http://doi.org/10.5281/zenodo.5008636" ID-Zenodo-Dep="5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8" startId="7.[94,153,820,845]" targetBox="[96,1412,204,790]" targetPageId="7">
<paragraph id="18313690FFB8FF86FFD2FCAFFCF2FCF4" blockId="7.[94,1419,820,877]" pageId="7" pageNumber="8">
Figure 8. 
<taxonomicName id="DF8E4D13FFB8FF86FF3FFCAFFEDFFCD4" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[179,369,820,845]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB8FF86FF3FFCAFFEDFFCD4" bold="true" box="[179,369,820,845]" italics="true" pageId="7" pageNumber="8">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFB8FF86FEF8FCAFFE14FCD4" box="[372,442,820,845]" pageId="7" pageNumber="8" rank="species">sp.nov.</taxonomicNameLabel>
male structures:(A) penis and epiphallus,anterior view;(B) same,posterior view;(C) detail of epiphallus and its flagella; (D) penis and epiphallus opened longitudinally to expose their inner folds.Scale bars= 5 mm.
</paragraph>
</caption>
<paragraph id="18313690FFB8FF86FFD2FBE1FD61FAAA" blockId="7.[94,720,1146,2131]" pageId="7" pageNumber="8">
<emphasis id="2AFAEA82FFB8FF86FFD2FBE1FF1EFB0A" bold="true" box="[94,176,1146,1171]" pageId="7" pageNumber="8">Radula</emphasis>
(
<figureCitation id="80B52A15FFB8FF86FF30FBE1FEA4FB0A" box="[188,266,1146,1171]" captionStart="Figure 3" captionStartId="3.[96,155,2074,2099]" captionTargetBox="[94,1417,1118,2044]" captionTargetId="figure-443@3.[94,1417,1118,2044]" captionTargetPageId="3" captionText="Figure 3. Megalobulimus dryades sp. nov. anatomy:(A) pulmonary (pallial) cavity,ventral-inner view showing collar vessel (cv), septum (se),heart (ht); (B) jaw, anterior view,left side damaged;(C) radula,central region in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008623" httpUri="https://zenodo.org/record/5008623/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
): With ~130 rows of isolated unicuspidate teeth (64-1-64); central tooth 25% smaller than laterals, drop-shaped with posterior apex, basal plate with long apical extensions forming acute angles; lateral teeth elliptical, 2 times as long as wide, with single, short, obtuse cusp;basal plate coniform,with an acute lateral extension.
</paragraph>
<paragraph id="18313690FFB8FF86FF08FAA1FE78F8EA" blockId="7.[94,720,1146,2131]" pageId="7" pageNumber="8">
Anterior chamber of esophagus (
<figureCitation id="80B52A15FFB8FF86FD8FFAA1FDF8FACA" box="[515,598,1338,1363]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
: ae) occupying ~½ of esophagus volume, with thin, translucent wall; inner surface smooth, lacking folds. Salivary glands clustering around anterior esophagus. Middle esophagus (
<figureCitation id="80B52A15FFB8FF86FF1EFA21FF4EFA4A" box="[146,224,1466,1491]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig.5D</figureCitation>
: me) with strong longitudinal muscular folds, anastomosed distally. Posterior esophagus chamber-like (
<figureCitation id="80B52A15FFB8FF86FFE9FA61FF19F98A" box="[101,183,1530,1555]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
: po), with inner serpiginous folds increasing in number at final third, located at opening of duct to anterior lobe of digestive gland (
<figureCitation id="80B52A15FFB8FF86FE40F9A1FD8FF9CA" box="[460,545,1594,1619]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
: da) alongside emersion of esophageal typhlosole (
<figureCitation id="80B52A15FFB8FF86FE61F9C1FD95F9EA" box="[493,571,1626,1651]" captionStart="Figure 5" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[170,1493,1307,2044]" captionTargetId="figure-187@4.[170,1493,1307,2044]" captionTargetPageId="4" captionText="Figure 5. Megalobulimus dryades sp. nov. anatomy: (A) buccal mass opened longitudinally; (B) odontophore, dorsal view, radula and right muscles partially re- moved;(C) same,ventral view;(D) esophagus longitudinally opened,exposing its inner surface.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008627" httpUri="https://zenodo.org/record/5008627/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
: et). Stomach rounded, walls thick, muscular, internally (
<figureCitation id="80B52A15FFB8FF86FDDDF9E1FD33F90A" box="[593,669,1658,1683]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
A-B) covered by muscular folds and a bulging pyloric fold (
<figureCitation id="80B52A15FFB8FF86FFE9F921FF0EF94A" box="[101,160,1722,1747]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
A-B: pf) parallel to minor gastric curvature; pyloric fold surrounding both esophageal gutter (
<figureCitation id="80B52A15FFB8FF86FDDFF941FD0DF96A" box="[595,675,1754,1779]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
: sg) originating at end of esophageal typhlosole (
<figureCitation id="80B52A15FFB8FF86FDD5F961FD06F88A" box="[601,680,1786,1811]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
: et) and intestinal gutters (
<figureCitation id="80B52A15FFB8FF86FED5F881FE09F8AA" box="[345,423,1818,1843]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
: ig) originating at aperture of duct of posterior lobe of digestive gland (
<figureCitation id="80B52A15FFB8FF86FDC7F8A1FD34F8CA" box="[587,666,1850,1875]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
: dp), running into intestine (
<figureCitation id="80B52A15FFB8FF86FED1F8C1FE05F8EA" box="[349,427,1882,1907]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
: in).
</paragraph>
<paragraph id="18313690FFB8FF86FF08F8E1FBF0FB8A" blockId="7.[94,720,1146,2131]" lastBlockId="7.[793,1418,922,1043]" pageId="7" pageNumber="8">
Inner surface of proximal intestine with large intestinal typhlosole (
<figureCitation id="80B52A15FFB8FF86FE81F801FEFBF82A" box="[269,341,1946,1971]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
C-D: it) and parallel folds (p1-p5); distance between folds p1 and p2 slightly wider; fold p4 (
<figureCitation id="80B52A15FFB8FF86FF04F841FF60F86A" box="[136,206,2010,2035]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
C-D: p4) reaching pre-rectal valve (
<figureCitation id="80B52A15FFB8FF86FDC2F841FD30F86A" box="[590,670,2010,2035]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6D</figureCitation>
: va), continuing as post-valve central longitudinal fold, low and wide (
<figureCitation id="80B52A15FFB8FF86FF58F781FE8BF7AA" box="[212,293,2074,2099]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6D</figureCitation>
: p6), surrounded by oblique, post-valvar compact folds (
<figureCitation id="80B52A15FFB8FF86FEA2F7A1FEDDF7CA" box="[302,371,2106,2131]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
D-E: p7). At final third of rectum (
<figureCitation id="80B52A15FFB8FF86FCACFC01FCC2FC2A" box="[800,876,922,947]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6E</figureCitation>
), fold p6 extinguishing, p7 folds becoming anastomosed into four bulging rectal folds (
<figureCitation id="80B52A15FFB8FF86FB55FC21FA88FC4A" box="[1241,1318,954,979]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6E</figureCitation>
: rf), progressing in part beyond aperture of anus (
<figureCitation id="80B52A15FFB8FF86FB6FFC41FA83FC6A" box="[1251,1325,986,1011]" captionStart="Figure 6" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[95,1413,857,2041]" captionTargetId="figure-303@5.[94,1417,854,2045]" captionTargetPageId="5" captionText="Figure 6.Megalobulimus dryades sp.nov.anatomy:(A)stomach,rightview,openedlongitudinallyalong dorsal surface;(B)same,left view;(C) intestine,transversally openedstrip of its proximal region,inner surface exposed;(D) same,region of the pre-rectal valve;(E) same,rectum region.Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008630" httpUri="https://zenodo.org/record/5008630/files/figure.png" pageId="7" pageNumber="8">Fig. 6E</figureCitation>
: an) into inner edge of pneumostome.
</paragraph>
<paragraph id="18313690FFB8FF86FC96FBA1FA22F9EA" blockId="7.[793,1420,1082,1939]" pageId="7" pageNumber="8">
<emphasis id="2AFAEA82FFB8FF86FC96FBA1FC7BFBCA" bold="true" box="[794,981,1082,1107]" pageId="7" pageNumber="8">Genital system:</emphasis>
hermaphroditic duct (
<figureCitation id="80B52A15FFB8FF86FB56FBA1FAF8FBCA" box="[1242,1366,1082,1107]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, B</figureCitation>
: hd) inserting apically into talon (
<figureCitation id="80B52A15FFB8FF86FBDBFBC1FB0AFBEA" box="[1111,1188,1114,1139]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7B</figureCitation>
: ta) at base of accessory glandular sac (
<figureCitation id="80B52A15FFB8FF86FC7AFBE1FBEAFB0A" box="[1014,1092,1146,1171]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7B</figureCitation>
: al). Carrefour (
<figureCitation id="80B52A15FFB8FF86FB63FBE1FA93FB0A" box="[1263,1341,1146,1171]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7B</figureCitation>
) locat- ed medially in albumen gland (
<figureCitation id="80B52A15FFB8FF86FBF9FB01FB47FB2A" box="[1141,1257,1178,1203]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, B</figureCitation>
: ag). Albumen chamber (
<figureCitation id="80B52A15FFB8FF86FC00FB21FB8CFB4A" box="[908,1058,1210,1235]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, B, C</figureCitation>
: ac) with two inner voluminous folds (
<figureCitation id="80B52A15FFB8FF86FCD0FB41FC06FB6A" box="[860,936,1242,1267]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7C</figureCitation>
: fa), forming groove running toward uterus. Spermoviduct (
<figureCitation id="80B52A15FFB8FF86FC4EFB61FB98FA8A" box="[962,1078,1274,1299]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, C</figureCitation>
: eo, D) bearing accessory glandular groove (
<figureCitation id="80B52A15FFB8FF86FC3CFA81FC53FAAA" box="[944,1021,1306,1331]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig.7D</figureCitation>
:gg); uterus (
<figureCitation id="80B52A15FFB8FF86FB09FA81FB5BFAAA" box="[1157,1269,1306,1331]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs.7B,D</figureCitation>
: ut), spermatic gutter (
<figureCitation id="80B52A15FFB8FF86FC0EFAA1FC7EFACA" box="[898,976,1338,1363]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7D</figureCitation>
: eg) and narrow prostate (
<figureCitation id="80B52A15FFB8FF86FB62FAA1FACFFACA" box="[1262,1377,1338,1363]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7B, D</figureCitation>
: pt) occupying ~¼ of perimeter. Free oviduct and vagina (
<figureCitation id="80B52A15FFB8FF86FCACFAE1FC3DFA0A" box="[800,915,1402,1427]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7E, F</figureCitation>
: ol, vg) uniformly tubular, limited distally by robust free oviduct appendix (
<figureCitation id="80B52A15FFB8FF86FBF9FA01FB40FA2A" box="[1141,1262,1434,1459]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, E</figureCitation>
: ve), size well over half of free oviduct size. Inner muscle folds of free oviduct and vagina perpendicular and slightly directed to opening of duct of bursa copulatrix (
<figureCitation id="80B52A15FFB8FF86FB45FA61FABDF98A" box="[1225,1299,1530,1555]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7F</figureCitation>
: ob). Bursa copulatrix duct wide and flattened (
<figureCitation id="80B52A15FFB8FF86FB2EF981FABDF9AA" box="[1186,1299,1562,1587]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A, E</figureCitation>
: db); bursa copulatrix balloon-like (
<figureCitation id="80B52A15FFB8FF86FBAEF9A1FBDCF9CA" box="[1058,1138,1594,1619]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Fig. 7A</figureCitation>
: bc), with thick muscular walls, bearing internal folds transversal to duct insertion.
</paragraph>
<paragraph id="18313690FFB8FF86FCB3F9E1FB30F80A" blockId="7.[793,1420,1082,1939]" pageId="7" pageNumber="8">
Penis long and uniformly cylindrical (
<figureCitation id="80B52A15FFB8FF86FB5AF9E1FA81F90A" box="[1238,1327,1658,1683]" captionStart="Figure7" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[172,1489,459,2037]" captionTargetId="figure-127@6.[170,1493,457,2044]" captionTargetPageId="6" captionText="Figure7.Megalobulimusdryadessp.nov.anatomy:(A)genitalsystem,completedorsalview;(B)Carrefourregion,ventralview;(C)albumenchamberopened,exposing itsfolds;(D)spermoviduct,cross-section atits middle region;(E)anteriorregion of genitalducts,lateral view;(F)same,opened longitudinally.Scale bars=5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008634" httpUri="https://zenodo.org/record/5008634/files/figure.png" pageId="7" pageNumber="8">Figs. 7A</figureCitation>
, 
<figureCitation id="80B52A15FFB8FF86FABBF9E1FAF6F90A" box="[1335,1368,1658,1683]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">8A</figureCitation>
: pe), same diameter as free oviduct, drawn and coiled by traction of vas deferens (
<figureCitation id="80B52A15FFB8FF86FBB8F921FB2BF94A" box="[1076,1157,1722,1747]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Fig. 8A</figureCitation>
: dd). Inner penial folds (
<figureCitation id="80B52A15FFB8FF86FCACF941FCC0F96A" box="[800,878,1754,1779]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Fig. 8C</figureCitation>
: ip) longitudinal, with anastomoses, bearing low transverse fold near its tip (
<figureCitation id="80B52A15FFB8FF86FBDCF961FB31F88A" box="[1104,1183,1786,1811]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Fig. 8C</figureCitation>
: tf), not bulky or pedunculated. Epiphallus broad and short (
<figureCitation id="80B52A15FFB8FF86FB6DF881FA86F8AA" box="[1249,1320,1818,1843]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Figs. 8</figureCitation>
A-D: ep), ~ 
<emphasis id="2AFAEA82FFB8FF86FCA6F8A0FC92F8CD" bold="true" box="[810,828,1851,1876]" italics="true" pageId="7" pageNumber="8">⅕</emphasis>
of penis length, with two uniform and short flagella (
<figureCitation id="80B52A15FFB8FF86FCADF8C1FCD3F8EA" box="[801,893,1882,1907]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Figs. 8A</figureCitation>
, C-D: fl), and two inner folds (
<figureCitation id="80B52A15FFB8FF86FB40F8C1FAB2F8EA" box="[1228,1308,1882,1907]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Fig. 8D</figureCitation>
: el). Penis muscle sub-apical (
<figureCitation id="80B52A15FFB8FF86FC60F8E1FBCCF80A" box="[1004,1122,1914,1939]" captionStart="Figure 8" captionStartId="7.[94,153,820,845]" captionTargetBox="[96,1412,204,790]" captionTargetId="figure-693@7.[94,1417,204,790]" captionTargetPageId="7" captionText="Figure 8. Megalobulimus dryadessp.nov.male structures:(A) penisand epiphallus,anteriorview;(B) same,posterior view;(C) detailof epiphallus and its flagella; (D) penisand epiphallus opened longitudinallyto expose their inner folds.Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5008636" httpUri="https://zenodo.org/record/5008636/files/figure.png" pageId="7" pageNumber="8">Figs. 8A, D</figureCitation>
: mp).
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFB8FF89FC96F821FE4DFEEA" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="distribution">
<paragraph id="18313690FFB8FF86FC96F821FA25F7CA" blockId="7.[794,1419,1978,2131]" pageId="7" pageNumber="8">
<emphasis id="2AFAEA82FFB8FF86FC96F821FBCCF84A" bold="true" box="[794,1122,1978,2003]" pageId="7" pageNumber="8">Distribution and habitat:</emphasis>
<taxonomicName id="DF8E4D13FFB8FF86FBF6F821FA27F84A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[1146,1417,1978,2003]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB8FF86FBF6F821FA27F84A" box="[1146,1417,1978,2003]" italics="true" pageId="7" pageNumber="8">Megalobulimus dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB8FF86FC96F841FCDBF86A" bold="true" box="[794,885,2010,2035]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="31C957F9FFB8FF86FC96F841FCDBF86A" box="[794,885,2010,2035]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was initially recorded in the Vale do Ribeira region, in 
<collectingRegion id="DA4AF872FFB8FF86FCF8F861FB8EF78A" box="[884,1056,2042,2067]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo state</collectingRegion>
. It is found mainly in the middle and lower Rio Ribeira do Iguape Basin in southern 
<collectingRegion id="DA4AF872FFB8FF86FAEEF781FCF6F7CA" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo</collectingRegion>
and northeastern 
<collectingRegion id="DA4AF872FFB8FF86FBADF7A1FBC2F7CA" box="[1057,1132,2106,2131]" country="Brazil" name="Parana" pageId="7" pageNumber="8">Paraná</collectingRegion>
states (
<figureCitation id="80B52A15FFB8FF86FB31F7A1FB54F7CA" box="[1213,1274,2106,2131]" captionStart="Figure 9" captionStartId="8.[172,231,2074,2099]" captionTargetBox="[170,1492,817,2044]" captionTargetId="figure-299@8.[170,1493,816,2044]" captionTargetPageId="8" captionText="Figure 9.Megalobulimus dryades sp.nov. and M.gummatus in S-SE Brazil:Geographical distributions based on MZSP specimens.Megalobulimus dryades sp.nov. (circles;star indicating type locality,Iporanga) in south São Paulo state;M. gummatus (triangle) on the central coast of Rio de Janeiro state (type locality)." figureDoi="http://doi.org/10.5281/zenodo.5008638" httpUri="https://zenodo.org/record/5008638/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
). Apparently,
</paragraph>
<paragraph id="18313690FFB7FF89FF26FF41FE4DFEEA" blockId="8.[170,796,218,371]" pageId="8" pageNumber="9">
this species has a high environmental tolerance, and its range may have been expanded by humans who used its shells in handicrafts. 
<bibRefCitation id="7C1F4B61FFB7FF89FE37FE81FD06FEAA" author="Agudo-Padron, A." box="[443,680,282,307]" pageId="8" pageNumber="9" pagination="32 - 37" refId="ref9013" refString="Agudo-Padron, A. 2010. The mollusc fauna of Santa Catarina state, Southern Brasil: knowledge gained from 13 years of research. Tentacle, 18: 32 - 37." type="journal article" year="2010">Agudo-Padrón (2010</bibRefCitation>
, 
<bibRefCitation id="7C1F4B61FFB7FF89FD38FE81FD42FEAA" author="Agudo-Padron, A." box="[692,748,282,307]" pageId="8" pageNumber="9" pagination="59 - 66" refId="ref9046" refString="Agudo-Padron, A. 2011. Threatened freshwater and terrestrial molluscs (Mollusca, Gastropoda et Bivalvia) of Santa Catarina State, Southern Brazil: check list and evaluation of regional threats. Biodiversity Journal, 2 (2): 59 - 66." type="journal article" year="2011">2011</bibRefCitation>
) recorded its presence in 
<collectingRegion id="DA4AF872FFB7FF89FE2BFEA1FD29FECA" box="[423,647,314,339]" country="Brazil" name="Santa Catarina" pageId="8" pageNumber="9">Santa Catarina state</collectingRegion>
, identified as 
<taxonomicName id="DF8E4D13FFB7FF89FF27FEC1FE91FEEA" authorityName=": Leme" authorityYear="1973" box="[171,319,346,371]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB7FF89FF27FEC1FE91FEEA" box="[171,319,346,371]" italics="true" pageId="8" pageNumber="9">M. gummatus</emphasis>
</taxonomicName>
(non Hidalgo).
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFB7FF89FF26FE01FE48FD2A" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="18313690FFB7FF89FF26FE01FE48FD2A" blockId="8.[170,796,410,691]" pageId="8" pageNumber="9">
<emphasis id="2AFAEA82FFB7FF89FF26FE01FEA1FE2A" bold="true" box="[170,271,410,435]" pageId="8" pageNumber="9">Ecology:</emphasis>
<bibRefCitation id="7C1F4B61FFB7FF89FE98FE01FD8DFE2A" author="Sobreira, H. B. &amp; Molina, F. B." box="[276,547,410,435]" pageId="8" pageNumber="9" pagination="139 - 141" refId="ref10643" refString="Sobreira, H. B. &amp; Molina, F. B. 2002. Observacoes preliminares sobre a biologia reprodutiva de Megalobulimus gummatus (Mollusca, Megalobulimidae) em laboratorio. Arquivos do Instituto BiolOgico, 69 (Supl.): 139 - 141." type="journal article" year="2002">Sobreira &amp; Molina (2002)</bibRefCitation>
studied the reproduction of 
<taxonomicName id="DF8E4D13FFB7FF89FF73FE21FEDAFE4A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[255,372,442,467]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB7FF89FF73FE21FEDAFE4A" box="[255,372,442,467]" italics="true" pageId="8" pageNumber="9">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB7FF89FEF6FE21FE7BFE4A" bold="true" box="[378,469,442,467]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="31C957F9FFB7FF89FEF6FE21FE7BFE4A" box="[378,469,442,467]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(referred to as 
<taxonomicName id="DF8E4D13FFB7FF89FD09FE21FCB2FE4A" authorityName=": Leme" authorityYear="1973" box="[645,796,442,467]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB7FF89FD09FE21FCB2FE4A" box="[645,796,442,467]" italics="true" pageId="8" pageNumber="9">M. gummatus</emphasis>
</taxonomicName>
in their study) based on 
<specimenCount id="0E88FD19FFB7FF89FE49FE41FDD1FE6A" box="[453,639,474,499]" pageId="8" pageNumber="9" type="generic">three specimens</specimenCount>
from Vale do Ribeira. Egg-laying occurred in two periods, a longer one between April and May and a shorter one between September and November. Specimens laid 2 to 
<specimenCount id="0E88FD19FFB7FF89FD43FDA1FCB5FDCA" box="[719,795,570,595]" pageId="8" pageNumber="9" type="egg">5 eggs</specimenCount>
per clutch, with average hatching of 52.4% after a development period of 60 days (n = 21). The mean egg size was 28.6 × 
<quantity id="DF769B75FFB7FF89FEA8FD01FE2BFD2A" box="[292,389,666,691]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="8" pageNumber="9" unit="mm" value="21.3">21.3 mm</quantity>
(n = 26).
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFB7FF89FF26FD41FA7DFE6A" pageId="8" pageNumber="9" type="etymology">
<paragraph id="18313690FFB7FF89FF26FD41FCB4FC8A" blockId="8.[170,794,730,787]" pageId="8" pageNumber="9">
<emphasis id="2AFAEA82FFB7FF89FF26FD41FE81FD6A" bold="true" box="[170,303,730,755]" pageId="8" pageNumber="9">Etymology:</emphasis>
The comprehensive classification of the Brazilian phytogeographic regions proposed by Carl
</paragraph>
<paragraph id="18313690FFB7FF89FCE9FF41FA7DFE6A" blockId="8.[868,1496,218,499]" pageId="8" pageNumber="9">
Friedrich Philipp von 
<bibRefCitation id="7C1F4B61FFB7FF89FBEEFF41FAAEFF6A" author="Martius, C. F. P." box="[1122,1280,218,243]" pageId="8" pageNumber="9" refId="ref9929" refString="Martius, C. F. P. 1840 - 1869. Tabulae physiognomicae. In: Martius, C. F. P.; Eichler, A. G. &amp; Urban, I. (Eds.). Flora brasiliensis. Frid. Leipzig, Fleischer. vol. 1, pt. 1, 150 p." type="book" year="1840">Martius (1840</bibRefCitation>
-1869) divided the country into phytogeographic provinces named after nymphs from Greek mythology. The name Dryades was used to designate the region roughly corresponding to the current Brazilian Atlantic Forest. This biome has been reduced to about 15% of its original territorial coverage. The Vale do Ribeira region is the largest continuous reserve of this threatened biome, thus our inspiration for the specific epithet, which is used herein as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFB7FF89FCE9FD81FBCAFD0A" pageId="8" pageNumber="9" type="description">
<paragraph id="18313690FFB7FF89FCE9FD81FBCAFD0A" blockId="8.[869,1493,538,659]" pageId="8" pageNumber="9">
<emphasis id="2AFAEA82FFB7FF89FCE9FD81FBB2FDAA" bold="true" box="[869,1052,538,563]" pageId="8" pageNumber="9">Measurements:</emphasis>
<typeStatus id="C7358832FFB7FF89FBA1FD81FB38FDAA" box="[1069,1174,538,563]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
, L: 
<quantity id="DF769B75FFB7FF89FB42FD81FA99FDAA" box="[1230,1335,538,563]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="8" pageNumber="9" unit="mm" value="119.0">119 mm</quantity>
; W: 
<quantity id="DF769B75FFB7FF89FAFBFD81FA7CFDAA" box="[1399,1490,538,563]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="9" unit="mm" value="70.0">70 mm</quantity>
; H: 
<quantity id="DF769B75FFB7FF89FC0AFDA1FC74FDCA" box="[902,986,570,595]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="9" unit="mm" value="57.0">57 mm</quantity>
; S: 5.5; P: 3.5; 
<typeStatus id="C7358832FFB7FF89FB0CFDA1FB42FDCA" box="[1152,1260,570,595]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
(
<emphasis id="2AFAEA82FFB7FF89FB71FDA1FAA5FDCA" box="[1277,1291,570,595]" italics="true" pageId="8" pageNumber="9">n</emphasis>
= 23; ± standard deviation) L: 118 ± 
<quantity id="DF769B75FFB7FF89FBB2FDC1FB2CFDEA" box="[1086,1154,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="mm" value="8.0">8 mm</quantity>
; W: 72 ± 
<quantity id="DF769B75FFB7FF89FB64FDC1FA82FDEA" box="[1256,1324,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="4.0">4 mm</quantity>
; H: 54 ± 
<quantity id="DF769B75FFB7FF89FA02FDC1FA7CFDEA" box="[1422,1490,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="3.0">3 mm</quantity>
; S: 5.4 ± 0.2; P: 3.2 ± 0.2.
</paragraph>
</subSubSection>
<subSubSection id="5094651BFFB7FF88FCE9FD21FE2AFA4A" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="18313690FFB7FF88FCE9FD21FDDCFDAA" blockId="8.[869,1493,698,787]" lastBlockId="9.[94,721,217,1491]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="2AFAEA82FFB7FF89FCE9FD21FC7DFD4A" bold="true" box="[869,979,698,723]" pageId="8" pageNumber="9">Remarks:</emphasis>
The first shells of 
<taxonomicName id="DF8E4D13FFB7FF89FB3DFD21FA87FD4A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[1201,1321,698,723]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB7FF89FB3DFD21FA87FD4A" box="[1201,1321,698,723]" italics="true" pageId="8" pageNumber="9">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB7FF89FAB9FD21FA3DFD4A" bold="true" box="[1333,1427,698,723]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="31C957F9FFB7FF89FAB9FD21FA3DFD4A" box="[1333,1427,698,723]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected by naturalist Adolph Hempel and archaeologist Ricardo Krone (
<typeStatus id="C7358832FFB7FF89FBB2FD61FB0CFC8A" box="[1086,1186,762,787]" pageId="8" pageNumber="9" type="paratype">Paratype</typeStatus>
on 
<figureCitation id="80B52A15FFB7FF89FB42FD61FAA6FC8A" box="[1230,1288,762,787]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
E-F) in the Vale do Ribeira region in the late 19 
<superScript id="EFFB9BD8FFB6FF88FE14FF42FE08FF71" attach="left" box="[408,422,217,232]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
– early 20 
<superScript id="EFFB9BD8FFB6FF88FD97FF42FD87FF71" attach="left" box="[539,553,217,232]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
centuries, and were deposited in the Museu Paulista (currently MZSP). When 
<bibRefCitation id="7C1F4B61FFB6FF88FF22FE81FEF0FEAA" author="Morretes, F. L." box="[174,350,282,307]" pageId="9" pageNumber="10" pagination="5 - 216" refId="ref10099" refString="Morretes, F. L. 1949. Ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paranaense, 7: 5 - 216." type="journal article" year="1949">Morretes (1949</bibRefCitation>
, 
<bibRefCitation id="7C1F4B61FFB6FF88FEE0FE81FE0AFEAA" author="Morretes, F. L." box="[364,420,282,307]" pageId="9" pageNumber="10" pagination="37 - 76" refId="ref10153" refString="Morretes, F. L. 1953. Adenda e corrigenda ao ensaio de catalogo dos moluscos do Brasil. Arquivos do Museu Paranaense, 10: 37 - 76." type="journal article" year="1953">1953</bibRefCitation>
) published his catalog of mollusks from 
<collectingCountry id="60997600FFB6FF88FF72FEA1FE92FECA" box="[254,316,314,339]" name="Brazil" pageId="9" pageNumber="10">Brazil</collectingCountry>
,he considered those samples as representatives of southern populations of 
<taxonomicName id="DF8E4D13FFB6FF88FD9BFEC1FD05FEEA" authorityName=": Leme" authorityYear="1973" box="[535,683,346,371]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB6FF88FD9BFEC1FD05FEEA" box="[535,683,346,371]" italics="true" pageId="9" pageNumber="10">M. gummatus</emphasis>
</taxonomicName>
inhabiting the states of 
<collectingRegion id="DA4AF872FFB6FF88FEC1FEE1FE16FE0A" box="[333,440,378,403]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="10">São Paulo</collectingRegion>
and 
<collectingRegion id="DA4AF872FFB6FF88FE61FEE1FD95FE0A" box="[493,571,378,403]" country="Brazil" name="Parana" pageId="9" pageNumber="10">Paraná</collectingRegion>
.This concept was maintained in subsequent compilations (
<bibRefCitation id="7C1F4B61FFB6FF88FDD6FE01FF58FE4A" author="Salgado, N. C. &amp; Coelho, A. C. S." pageId="9" pageNumber="10" pagination="149 - 189" refId="ref10375" refString="Salgado, N. C. &amp; Coelho, A. C. S. 2003. MoluscosterrestresdoBrasil (Gastropodes operculados ou nao exclusive Veronicellide, Milacidae e Limacidae). Revista de Biologia Tropical, 51 (Supl. 3): 149 - 189." type="journal article" year="2003">Salgado &amp; Coelho, 2003</bibRefCitation>
; 
<bibRefCitation id="7C1F4B61FFB6FF88FE8EFE21FE31FE4A" author="Simone, L. R. L." box="[258,415,442,467]" pageId="9" pageNumber="10" refId="ref10534" refString="Simone, L. R. L. 2006. Land and Freshwater Molluscs of Brazil. Sao Paulo, EGB / FAPESP." type="book" year="2006">Simone, 2006</bibRefCitation>
). Meanwhile, the concomitant erroneous use of another name, 
<taxonomicName id="DF8E4D13FFB6FF88FE77FE41FEDAFD8A" authority="(Morch, 1852)" baseAuthorityName="Morch" baseAuthorityYear="1852" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="chionostoma">
<emphasis id="2AFAEA82FFB6FF88FE77FE41FF62FD8A" italics="true" pageId="9" pageNumber="10">Megalobulimus chionostoma</emphasis>
(
<bibRefCitation id="7C1F4B61FFB6FF88FF50FE61FEC1FD8A" author="Morch, O. A. L." box="[220,367,506,531]" pageId="9" pageNumber="10" refId="ref10066" refString="Morch, O. A. L. 1852. Catalogus conchyliorum quae reliquit D. Alphonso d'Aguirra &amp; Gadea Comes de Yoldi. Fasciculus primus. Cephalophora. Hafniae: Klein." type="book" year="1852">Mörch, 1852</bibRefCitation>
)
</taxonomicName>
, to refer to true 
<taxonomicName id="DF8E4D13FFB6FF88FDBAFE61FD60FD8A" authorityName=": Leme" authorityYear="1973" box="[566,718,506,531]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB6FF88FDBAFE61FD60FD8A" box="[566,718,506,531]" italics="true" pageId="9" pageNumber="10">M. gummatus</emphasis>
</taxonomicName>
specimens caused further taxonomic confusion.
</paragraph>
<caption id="4CF16618FFB7FF89FF20F781FA27F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5008638" ID-Zenodo-Dep="5008638" httpUri="https://zenodo.org/record/5008638/files/figure.png" pageId="8" pageNumber="9" startId="8.[172,231,2074,2099]" targetBox="[170,1492,817,2044]" targetPageId="8">
<paragraph id="18313690FFB7FF89FF20F781FA27F7CA" blockId="8.[170,1494,2074,2131]" pageId="8" pageNumber="9">
Figure 9. 
<taxonomicName id="DF8E4D13FFB7FF89FF73F781FE13F7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[255,445,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB7FF89FF73F781FE13F7AA" bold="true" box="[255,445,2074,2099]" italics="true" pageId="8" pageNumber="9">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFB7FF89FE4DF781FDA9F7AA" box="[449,519,2074,2099]" pageId="8" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
and 
<taxonomicName id="DF8E4D13FFB7FF89FDA2F781FD33F7AA" authorityName=": Leme" authorityYear="1973" box="[558,669,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB7FF89FDA2F781FD33F7AA" bold="true" box="[558,669,2074,2099]" italics="true" pageId="8" pageNumber="9">M.gummatus</emphasis>
</taxonomicName>
in S-SE Brazil:Geographical distributions based on MZSP specimens. 
<taxonomicName id="DF8E4D13FFB7FF89FB41F781FA25F7AA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[1229,1419,2074,2099]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB7FF89FB41F781FA25F7AA" bold="true" box="[1229,1419,2074,2099]" italics="true" pageId="8" pageNumber="9">Megalobulimus dryades</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31C957F9FFB7FF89FA1CF781FA78F7AA" box="[1424,1494,2074,2099]" pageId="8" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
(circles;star indicating type locality,Iporanga) in south São Paulo state; 
<taxonomicName id="DF8E4D13FFB7FF89FD7DF7A1FCCFF7CA" authorityName=": Leme" authorityYear="1973" box="[753,865,2106,2131]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB7FF89FD7DF7A1FCCFF7CA" bold="true" box="[753,865,2106,2131]" italics="true" pageId="8" pageNumber="9">M. gummatus</emphasis>
</taxonomicName>
(triangle) on the central coast of Rio de Janeiro state (type locality).
</paragraph>
</caption>
<paragraph id="18313690FFB6FF88FF08FDA1FED2FB8A" blockId="9.[94,721,217,1491]" pageId="9" pageNumber="10">
Apart from the white peristome and a glossy periostracum, the shells of 
<taxonomicName id="DF8E4D13FFB6FF88FED6FDC1FE7EFDEA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[346,464,602,627]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB6FF88FED6FDC1FE7EFDEA" box="[346,464,602,627]" italics="true" pageId="9" pageNumber="10">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB6FF88FE55FDC1FD98FDEA" bold="true" box="[473,566,602,627]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="31C957F9FFB6FF88FE55FDC1FD98FDEA" box="[473,566,602,627]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="DF8E4D13FFB6FF88FDFEFDC1FF0CFD0A" authorityName=": Leme" authorityYear="1973" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">
<emphasis id="2AFAEA82FFB6FF88FDFEFDC1FF0CFD0A" italics="true" pageId="9" pageNumber="10">M. gummatus</emphasis>
</taxonomicName>
present some coincidental features such as large size, a dorsoventrally compressed outline, similar protoconch macrosculpture, and pigmented shell matrix with a white subsutural band (
<figureCitation id="80B52A15FFB6FF88FEF2FD41FE77FD6A" box="[382,473,730,755]" captionStart="Figure 2" captionStartId="2.[170,229,2042,2067]" captionTargetBox="[227,1436,204,2012]" captionTargetId="figure-1@2.[227,1436,204,2012]" captionTargetPageId="2" captionText="Figure 2. Shell ofMegalobulimus dryades sp.nov.:(A-D) holotype (MZSP 120000,Iporanga,SP),(A) apertural view,(B)lateral view,(C)abapertural view,(D) apical view;C and D showing the shell measurements: L = length,W = width, H = height; (E-F) paratype (MZSP 29317, Iguape,SP), (E) apertural view, (F) abapertural view;(G-H) paratype (MZSP 28773,Iguape,SP),(G) apertural view,(H) abapertural view;(I) paratype (MZSP 15594),apical view of protoconch in SEM." figureDoi="http://doi.org/10.5281/zenodo.5008621" httpUri="https://zenodo.org/record/5008621/files/figure.png" pageId="9" pageNumber="10">Figs. 2D</figureCitation>
, 
<figureCitation id="80B52A15FFB6FF88FE68FD41FDACFD6A" box="[484,514,730,755]" captionStart="Figure 10" captionStartId="10.[170,229,2042,2067]" captionTargetBox="[217,1445,204,2012]" captionTargetId="figure-1@10.[217,1445,204,2012]" captionTargetPageId="10" captionText="Figure 10. Shells of Megalobulimus gummatus: (A-C) holotype MNCN 1505/3199 (Rio de Janeiro), (A) apertural view, (B) lateral view, (C) abapertural view; (D-G) ANSP 428461 (Rio de Janeiro),(D) abapertural view,(E) lateral view,(F) apertural view,(G) protoconch;(H) MZSP 66800 (Arraial do Cabo,RJ),apertural view; (I-J) FMNH 216409 (Macaé,RJ,det.Bequaert,1953),(I) apertural view,(J)protoconch.Arrows indicate location of spiral sulci." figureDoi="http://doi.org/10.5281/zenodo.5008642" httpUri="https://zenodo.org/record/5008642/files/figure.png" pageId="9" pageNumber="10">10</figureCitation>
). However, 
<taxonomicName id="DF8E4D13FFB6FF88FD08FD41FF3EFC8A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB6FF88FD08FD41FF3EFC8A" italics="true" pageId="9" pageNumber="10">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB6FF88FF19FD61FF40FC8A" bold="true" box="[149,238,762,787]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="31C957F9FFB6FF88FF19FD61FF40FC8A" box="[149,238,762,787]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs in having a darker protoconch with a clearly marked lighter subsutural band, with a sculpture lacking any distinct spiral elements, consisting of more pronounced axial riblets that branch apically near the suture, and microscopic granules. It also has a lower spire, a body whorl with a strongly corrugated surface with malleations, a less reflected but thicker peristome with a more rounded outer lip, and a thicker, oblique, and slightly convex columella.
</paragraph>
<paragraph id="18313690FFB6FF88FF08FB81FE2AFA4A" blockId="9.[94,721,217,1491]" pageId="9" pageNumber="10">
The soft parts of 
<taxonomicName id="DF8E4D13FFB6FF88FECBFB81FE13FBAA" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[327,445,1050,1075]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB6FF88FECBFB81FE13FBAA" box="[327,445,1050,1075]" italics="true" pageId="9" pageNumber="10">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB6FF88FE49FB81FD8FFBAA" bold="true" box="[453,545,1050,1075]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="31C957F9FFB6FF88FE49FB81FD8FFBAA" box="[453,545,1050,1075]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are lighter-colored externally compared to 
<emphasis id="2AFAEA82FFB6FF88FE3AFBA1FDF8FBCA" box="[438,598,1082,1107]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="DF8E4D13FFB6FF88FE3AFBA1FDFCFBCA" authorityName=": Leme" authorityYear="1973" box="[438,594,1082,1107]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">M. gummatus</taxonomicName>
.
</emphasis>
Internally, differences in the digestive system reside in the more robust mandible with narrow, well-marked columns, and the configuration of the radular teeth. The genital systems are also distinct, as 
<taxonomicName id="DF8E4D13FFB6FF88FEF0FB21FE5FFB4A" authority="Fontenelle &amp; Simone &amp; Cavallari, 2021" authorityName="Fontenelle &amp; Simone &amp; Cavallari" authorityYear="2021" box="[380,497,1210,1235]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="dryades" status="sp. nov.">
<emphasis id="2AFAEA82FFB6FF88FEF0FB21FE5FFB4A" box="[380,497,1210,1235]" italics="true" pageId="9" pageNumber="10">M. dryades</emphasis>
</taxonomicName>
<emphasis id="2AFAEA82FFB6FF88FE7AFB21FDFCFB4A" bold="true" box="[502,594,1210,1235]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="31C957F9FFB6FF88FE7AFB21FDFCFB4A" box="[502,594,1210,1235]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a long and uniform penis and an epiphallus bearing two flagella. The presence of a talon and a free oviduct appendix further distinguishes it from 
<emphasis id="2AFAEA82FFB6FF88FE17FA81FD9BFAAA" box="[411,565,1306,1331]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="DF8E4D13FFB6FF88FE17FA81FD9FFAAA" authorityName=": Leme" authorityYear="1973" box="[411,561,1306,1331]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">M. gummatus</taxonomicName>
,
</emphasis>
which in turn seems more closely related to species of the so-called “ 
<taxonomicName id="DF8E4D13FFB6FF88FFE4FAC1FF60FAEA" baseAuthorityName="O.F.Muller" baseAuthorityYear="1774" box="[104,206,1370,1395]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="ovatus">
<emphasis id="2AFAEA82FFB6FF88FFE4FAC1FF60FAEA" box="[104,206,1370,1395]" italics="true" pageId="9" pageNumber="10">M.ovatus</emphasis>
</taxonomicName>
species complex” 
<emphasis id="2AFAEA82FFB6FF88FE18FAC1FE7EFAEA" box="[404,464,1370,1395]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
<bibRefCitation id="7C1F4B61FFB6FF88FE5AFAC1FDF7FAEA" author="Leme, J. L. M." box="[470,601,1370,1395]" pageId="9" pageNumber="10" pagination="313 - 318" refId="ref9635" refString="Leme, J. L. M. 1989. Megalobulimus lopesi sp. nov., uma nova especie de Pulmonata terrestre da Mata Atlantica brasileira (Mollusca, Gastropoda, Megalobulimidae). MemOriasdoInstitutoOswaldoCruz, 84 (Supl. 4): 313 - 318." type="journal article" year="1989">Leme (1989</bibRefCitation>
, 
<bibRefCitation id="7C1F4B61FFB6FF88FDECFAC1FD36FAEA" author="Leme, J. L. M." box="[608,664,1370,1395]" pageId="9" pageNumber="10" pagination="95 - 105" refId="ref9683" refString="Leme, J. L. M. 1993. Estudo anatomico sobre Megalobulimus auritus (Sowerby, 1838) (Gastropoda, Megalobulimidae). Papeis Avulsos de Zoologia, 38: 95 - 105." type="journal article" year="1993">1993</bibRefCitation>
) due to the lack of these two last characters. Finally, both species present isolated distributions and distinct habitats (see 
<emphasis id="2AFAEA82FFB6FF88FF1DFA21FE85FA4A" box="[145,299,1466,1491]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="DF8E4D13FFB6FF88FF1DFA21FE89FA4A" authorityName=": Leme" authorityYear="1973" box="[145,295,1466,1491]" class="Gastropoda" family="Strophocheilidae" genus="Megalobulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="gummatus">M. gummatus</taxonomicName>
,
</emphasis>
below).
</paragraph>
</subSubSection>
</treatment>
</document>