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<document id="3F90E419012B25A9C9238ECD205C0BF4" ID-ISSN="1175­5334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K. A." docDate="2008" docId="BC22C32217E5A9C9989DF99E7932FED6" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Cheumatopsyche tiokalamkita Olah &amp; Johanson 2008, new species" docType="treatment" docVersion="2" lastPageNumber="216" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="213" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="9CFEDB342CCF5C9806CEA72722CE168A">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="AAB959C1906F3F8DF69D5707C7ADA5F5">Oláh, J.</mods:namePart>
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<mods:namePart id="3EC988A33F1369F9673E3ACCB05D8A62">Johanson, K. A.</mods:namePart>
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<mods:date id="1BE29A02B51C5C6ABF54CE63A5788A57">2008</mods:date>
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<mods:number id="39C9E1E33AF589EF0A3F28D4B04D1801">2008-06-16</mods:number>
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<mods:number id="91113270E1CEE04541F5E0CBD15F3288">1802</mods:number>
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<mods:classification id="0B3B172BD73B2C9DC611833C82F676E1">journal article</mods:classification>
<mods:identifier id="77C578C073E46726A185AC9045B53D56" type="ISSN">1175­5334</mods:identifier>
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<treatment id="BC22C32217E5A9C9989DF99E7932FED6" LSID="urn:lsid:plazi:treatment:BC22C32217E5A9C9989DF99E7932FED6" httpUri="http://treatment.plazi.org/id/BC22C32217E5A9C9989DF99E7932FED6" lastPageId="215" lastPageNumber="216" pageId="212" pageNumber="213">
<subSubSection id="7C9121BF17E5A9CA989DF99E7BC6F992" box="[151,901,1620,1647]" pageId="212" pageNumber="213" type="nomenclature">
<paragraph id="3434723417E5A9CA989DF99E7BC6F992" blockId="212.[151,901,1620,1687]" box="[151,901,1620,1647]" pageId="212" pageNumber="213">
<heading id="6F7CC55817E5A9CA989DF99E7BC6F992" bold="true" box="[151,901,1620,1647]" fontSize="11" level="1" pageId="212" pageNumber="213" reason="1">
<emphasis id="06FFAE2617E5A9CA989DF99E7BC6F992" bold="true" box="[151,901,1620,1647]" pageId="212" pageNumber="213">
<taxonomicName id="F38B09B717E5A9CA989DF99E7AAAF992" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[151,745,1620,1647]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="tiokalamkita" status="new species">
<emphasis id="06FFAE2617E5A9CA989DF99E7A47F993" bold="true" box="[151,516,1621,1647]" italics="true" pageId="212" pageNumber="213">Cheumatopsyche tiokalamkita</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="1DCC135D17E5A9CA9AFCF99F7BC6F992" box="[758,901,1620,1646]" pageId="212" pageNumber="213" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7C9121BF17E5A9C8989DF9B67BC9F8E6" lastPageId="214" lastPageNumber="215" pageId="212" pageNumber="213" type="description">
<paragraph id="3434723417E5A9CA989DF9B67970F96B" blockId="212.[151,901,1620,1687]" box="[151,307,1661,1687]" pageId="212" pageNumber="213">
<figureCitation id="ACB06EB117E5A9CA989DF9B67970F96B" box="[151,307,1661,1687]" captionStart-0="FIGURES 358–361" captionStart-1="FIGURE 362" captionStartId-0="215.[151,269,1136,1160]" captionStartId-1="215.[151,255,1988,2012]" captionTargetBox-0="[177,1410,352,1093]" captionTargetBox-1="[432,1207,1250,1956]" captionTargetId-0="figure-108@215.[151,1436,323,1110]" captionTargetId-1="figure-113@215.[349,1257,1201,2001]" captionTargetPageId-0="215" captionTargetPageId-1="215" captionText-0="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." captionText-1="FIGURE 362. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, paratype, female. 362 —genitalia, lateral." pageId="212" pageNumber="213">Fig. 358–362</figureCitation>
</paragraph>
<paragraph id="3434723417E5A9CA989DF9067B9AF87B" blockId="212.[151,1436,1741,2007]" pageId="212" pageNumber="213">
This dark animal bimaculated in white, is very similar to
<taxonomicName id="F38B09B717E5A9CA9B3CF9067CF9F91B" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[822,1210,1741,1767]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E5A9CA9B3CF9067BA4F91A" box="[822,999,1741,1766]" italics="true" pageId="212" pageNumber="213">C. ranomafana</emphasis>
Oláh &amp; Johanson
</taxonomicName>
in the male genitalia, and both species collected in the same Malaise trap.
<taxonomicName id="F38B09B717E5A9CA9B2CF93E7CC7F8F2" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[806,1156,1781,1806]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E5A9CA9B2CF93E7CC7F8F2" box="[806,1156,1781,1806]" italics="true" pageId="212" pageNumber="213">Cheumatopsyche ranomafana</emphasis>
</taxonomicName>
has uniformly coloured forewing membranes without the remarkable white patch being characteristic for the new species. Although there are some individuals of
<taxonomicName id="F38B09B717E5A9CA99F0F88E7AF3F8A2" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[506,688,1861,1886]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="tiokalamkita">
<emphasis id="06FFAE2617E5A9CA99F0F88E7AF3F8A2" box="[506,688,1861,1886]" italics="true" pageId="212" pageNumber="213">C. tiokalamkita</emphasis>
</taxonomicName>
with less pronounced white patch, both males and females,
<taxonomicName id="F38B09B717E5A9CA9D8BF88E796AF87A" authorityName="Olah &amp; Johanson" authorityYear="2008" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="tiokalamkita">
<emphasis id="06FFAE2617E5A9CA9D8BF88E796AF87A" italics="true" pageId="212" pageNumber="213">C. tiokalamkita</emphasis>
</taxonomicName>
are easily distinguished by the presence of this pale patch.
</paragraph>
<paragraph id="3434723417E5A9CB98CFF85E7C7AFED6" blockId="212.[151,1436,1741,2007]" lastBlockId="213.[151,1436,152,2018]" lastPageId="213" lastPageNumber="214" pageId="212" pageNumber="213">
<taxonomicName id="F38B09B717E5A9CA98CFF85E7A6AF852" authority="Oláh &amp; Johanson, 2008" authorityName="Oláh &amp; Johanson" authorityYear="2008" box="[197,553,1941,1966]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="tiokalamkita" status="new species">
<emphasis id="06FFAE2617E5A9CA98CFF85E7A6AF852" box="[197,553,1941,1966]" italics="true" pageId="212" pageNumber="213">Cheumatopsyche tiokalamkita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="1DCC135D17E5A9CA9A30F85E7A8AF853" box="[570,713,1941,1967]" pageId="212" pageNumber="213" rank="species">new species</taxonomicNameLabel>
is separated from
<taxonomicName id="F38B09B717E5A9CA9BA1F85E7C1CF852" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[939,1119,1941,1966]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E5A9CA9BA1F85E7C1CF852" box="[939,1119,1941,1966]" italics="true" pageId="212" pageNumber="213">C. ranomafana</emphasis>
</taxonomicName>
in the female genitalia by the absence of an elongated, ventrocaudal angle of tergite VIII. It is separated from
<taxonomicName id="F38B09B717E5A9CA9C69F8767D57F82A" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[1123,1300,1981,2006]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E5A9CA9C69F8767D57F82A" box="[1123,1300,1981,2006]" italics="true" pageId="212" pageNumber="213">C. ranomafana</emphasis>
</taxonomicName>
in the male genitalia by the rounded, more robust apicoventral setose lobes, being more slender in
<taxonomicName id="F38B09B717E4A9CB9C8DFF537D7BFF4D" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[1159,1336,152,177]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E4A9CB9C8DFF537D7BFF4D" box="[1159,1336,152,177]" italics="true" pageId="213" pageNumber="214">C. ranomafana</emphasis>
</taxonomicName>
; and the apical lobes of segment IX is rounded, while triangular in
<taxonomicName id="F38B09B717E4A9CB9B40FF0B7BBFFF25" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[842,1020,192,217]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E4A9CB9B40FF0B7BBFFF25" box="[842,1020,192,217]" italics="true" pageId="213" pageNumber="214">C. ranomafana</emphasis>
</taxonomicName>
. In the over-cleared
<typeStatus id="EB30CC9617E4A9CB9CF8FF0B7D1BFF26" box="[1266,1368,192,218]" pageId="213" pageNumber="214">holotype</typeStatus>
of
<taxonomicName id="F38B09B717E4A9CB9D8BFF0B7965FEFD" authorityName="Olah &amp; Johanson" authorityYear="2008" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="06FFAE2617E4A9CB9D8BFF0B7965FEFD" italics="true" pageId="213" pageNumber="214">C. ranomafana</emphasis>
</taxonomicName>
the left apical lobe do not looks triangular, as it was also drawn, but it is triangular on the right side of the
<typeStatus id="EB30CC9617E4A9CB9912FEDB793DFED6" box="[280,382,272,298]" pageId="213" pageNumber="214">holotype</typeStatus>
and on both sides of all subsequently examined specimens.
</paragraph>
<paragraph id="3434723417E4A9CB98CFFEF37D25FDE6" blockId="213.[151,1436,152,2018]" pageId="213" pageNumber="214">
Male (
<figureCitation id="ACB06EB117E4A9CB9918FEF379F5FEAE" box="[274,438,312,338]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358–361</figureCitation>
). Body and wings dark brown with darker pubescence; legs and antennae with paler pubescence. Maxillary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum dark chestnut brown, with 7 dark chestnut brown setal warts. Mesolateral warts dividing into mesad orienting row of 3–4 warts gradually decreasing in size. Swollen setal wart absent on proepisternum. Setal wart present on precoxale. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Mid- and hind-leg claws asymmetrical. Spur fomula 244. Protibiae with posteroapical spur slightly smaller than anteroapical spur.
</paragraph>
<paragraph id="3434723417E4A9CB98CFFDE37BEDFD1E" blockId="213.[151,1436,152,2018]" pageId="213" pageNumber="214">Wings. Forewing evenly brown with slightly darker veins and setae. Large, white, compact, rounded, patch located between first radial forks and M veins at point where discoidal and median cells start, more at basis of median cells. Forewing crossveins m-cu and cu tangential. Sc and R separate before costa. Cu2 and A1 confluent, meeting before wing margin. Hind wing Sc and R meeting after crossvein r. Crossvein r located posteriorly of crossvein s. Fork 1 present. Forewing length 7.1 mm.</paragraph>
<paragraph id="3434723417E4A9CB98CFFD3B79EBF936" blockId="213.[151,1436,152,2018]" pageId="213" pageNumber="214">
Male genitalia. Abdominal segment IX fused annularly, tergum much shorter than sternum (
<figureCitation id="ACB06EB117E4A9CB9D2EFD3B7DCCFCF6" box="[1316,1423,752,778]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
), reduced to narrow strip constricting at middle in dorsal view (
<figureCitation id="ACB06EB117E4A9CB9B7AFCD37B94FCCE" box="[880,983,792,818]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 359</figureCitation>
). Anterior margins of segment IX regularly bow-shaped; dorsally sinuate in lateral view (
<figureCitation id="ACB06EB117E4A9CB9B1AFC8B7B38FCA6" box="[784,891,832,858]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
). Apical lobe on posterolateral margins right angled, rounded; situated above less sclerotized articulation cavity of gonocoxites. Spine row on posterior margins of segment IX almost continuous, heterogeneous. Spines on dorsolateral spiny lobes longer than spines on apical lobes; dorsolateral spiny lobes reduced in dorsal view (
<figureCitation id="ACB06EB117E4A9CB9BDDFC737C7EFC2E" box="[983,1085,952,978]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 359</figureCitation>
), present on posterior slope of tergum IX and ventrad into intersegmental depression (
<figureCitation id="ACB06EB117E4A9CB9B26FC2B7BD6FC06" box="[812,917,992,1018]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
). Intersegmental step between segments IX and X sloping. Segment X elongate, with rounded, with slightly dorsad orienting apex (
<figureCitation id="ACB06EB117E4A9CB9CC8FBC37D6FFBDE" box="[1218,1324,1032,1058]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
); dorsum strongly concave in lateral view; nearly quadrangular in dorsal view (
<figureCitation id="ACB06EB117E4A9CB9BFDFBFB7C20FBB6" box="[1015,1123,1072,1098]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 359</figureCitation>
); slightly broadening and rounded at middle; apicoventral setal lobes wide; forming bilobed segment X in dorsal view (
<figureCitation id="ACB06EB117E4A9CB9CE0FB937D11FB8E" box="[1258,1362,1112,1138]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 359</figureCitation>
); with narrow and deep dorsal interlobular gap. Sutures broad; transverse sutures crossing segment X obliquely; longitudinal sutures starting from apicoventral setose lobes; fusing with transverse sutures into Y. Smooth mesocaudal plate with reduced central part filling basal half of interlobular gap in dorsal view (
<figureCitation id="ACB06EB117E4A9CB9D2BFB1B7DD3FB16" box="[1313,1424,1232,1258]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 359</figureCitation>
). Apicoventral setose lobes modified into large, laterally broad, posterodorsad orienting, rounded lobes being setose on apex and basis (
<figureCitation id="ACB06EB117E4A9CB99DAFAEB7A79FAC6" box="[464,570,1312,1338]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
). Lateral setose areas (superior or preanal appendages) forming vertical, elongated, elevated setal warts centrally on segment X; reaching slightly above dorsum in lateral view (
<figureCitation id="ACB06EB117E4A9CB9D67FA837884FA76" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
); surrounded basally by longitudinal sutures, and anteriorly by transverse sutures. Coxopodites slightly exceeding apex of segment X, forming straight rod dilating gradually apicad (
<figureCitation id="ACB06EB117E4A9CB9C56FA537C8BFA4E" box="[1116,1224,1432,1458]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
); weakly curving mesad in ventral view (
<figureCitation id="ACB06EB117E4A9CB99ADFA0B7A4EFA26" box="[423,525,1472,1498]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 360</figureCitation>
). Harpagones broad, nearly parallel-sided in lateral view (
<figureCitation id="ACB06EB117E4A9CB9CBBFA0B7D5BFA26" box="[1201,1304,1472,1498]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 358</figureCitation>
); in ventral view (
<figureCitation id="ACB06EB117E4A9CB98D5FA237900F9FE" box="[223,323,1512,1538]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 360</figureCitation>
) apically tapering from after mid-length, slightly concave mesally. Phallotheca (
<figureCitation id="ACB06EB117E4A9CB9CE7FA237D13F9FE" box="[1261,1360,1512,1538]" captionStart="FIGURES 358–361" captionStartId="215.[151,269,1136,1160]" captionTargetBox="[177,1410,352,1093]" captionTargetId="figure-108@215.[151,1436,323,1110]" captionTargetPageId="215" captionText="FIGURES 358–361. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, holotype, male. 358 — genitalia, lateral; 359 — genitalia, dorsal; 360 — gonocoxite, ventral; 361 — phallus, lateral." pageId="213" pageNumber="214">Fig. 361</figureCitation>
) welldeveloped, broad, ventrad curving; dorsally concave.
<taxonomicName id="F38B09B717E4A9CB9B31F9DB7B8EF9D6" authorityName="M. Zang &amp; R.H. Petersen" authorityYear="1989" box="[827,973,1552,1578]" class="Agaricomycetes" family="Phallaceae" genus="Endophallus" kingdom="Fungi" order="Phallales" pageId="213" pageNumber="9" phylum="Basidiomycota" rank="genus">Endophallus</taxonomicName>
forming long, narrow tube, running through entire phallotheca, ending in sclerotized ring at gonopore; ventral enlarged part difficult to discern. Chitinized endothecal process circular, located apicodorsally on apex. Phallotremal sclerites large, broad, rounded in lateral view. Vestigial, membranous, ventral endothecal lobes situated together with more sclerotized ventromesal lobe.
</paragraph>
<paragraph id="3434723417E4A9CB98CFF9137D48F81E" blockId="213.[151,1436,152,2018]" pageId="213" pageNumber="214">Female. Body and wings dark brown with darker pubescence; legs and antennae with paler pubescence. Maxillary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum dark chestnut brown, with 7 dark chestnut brown setal warts. Mesolateral warts forming mesad orienting row of 3–4 warts gradually decreasing in size. Swollen setal wart absent on proepisternum. Setal wart present on precoxale. Pretarsal claws symmetrical, not laterally flanked by setal bundle. Claws on mid- and hind legs symmetrical. Spur fomula 244. Protibiae with posteroapical spur slightly smaller than anteroapical spur. Spotreticulated circular plaque present on both sides of membranous pleura of abdominal segment IV.</paragraph>
<paragraph id="3434723417E7A9C898CFFF537C49FEAE" blockId="214.[151,1436,152,2018]" pageId="214" pageNumber="215">Wings. Forewing membrane ground color evenly brown, with slightly darker veins and setae; large, white compact, rounded patch located between first forks of R and M where Dc and median cells starting, slightly more on basis of median cells. Forewing crossveins m-cu and cu on line. Sc and R running separately before costa. Cu2 and A1 meeting before wing margin. Hind wing Sc and R meeting after crossvein r. Crossvein r located posteriorly of crossvein s. Fork 1 present. Forewing length 7.6 mm.</paragraph>
<paragraph id="3434723417E7A9C898CFFEAB7BC9F8E6" blockId="214.[151,1436,152,2018]" pageId="214" pageNumber="215">
Female genitalia (
<figureCitation id="ACB06EB117E7A9C8999CFEAB79BDFE86" box="[406,510,352,378]" captionStart="FIGURE 362" captionStartId="215.[151,255,1988,2012]" captionTargetBox="[432,1207,1250,1956]" captionTargetId="figure-113@215.[349,1257,1201,2001]" captionTargetPageId="215" captionText="FIGURE 362. Cheumatopsyche tiokalamkita Oláh &amp; Johanson, new species, paratype, female. 362 —genitalia, lateral." pageId="214" pageNumber="215">Fig. 362</figureCitation>
). Anteroventral corners remaining covered by sternum VIII. Abdomen tapering slightly more in lateral than in dorsal and ventral views, sagittally narrowing posterad. Tergum VIII quadrangular, narrowing slightly apicad; with small ventrocaudal lobes or triangular corners in lateral view. Lobes with wide, shallowly sinuate, mesal dorsocaudal incision visible in dorsal view. Tergal surface smooth, produced, provided by tiny, closely set microtrichiae; setae absent, except on ventrocaudal corners being covered by mesad curving setae. Tergite VIII producing into long, slender anterolateral apodemes. Pleurosternum (
<taxonomicName id="F38B09B717E7A9C898ABFD9B79D1FD96" authority="Denning 1943" authorityName="Denning" authorityYear="1943" box="[161,402,592,618]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C898ABFD9B78A1FD95" box="[161,226,592,617]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C898E2FD9B79D1FD96" author="Denning, D. G." box="[232,402,592,618]" pageId="214" pageNumber="215" pagination="101 - 171" refId="ref107007" refString="Denning, D. G. (1943) The Hydropsychidae of Minnesota. Entomologica Americana (New Series), 23 (2), 101 - 171." type="journal article" year="1943">Denning 1943</bibRefCitation>
</taxonomicName>
), or posterolateral apodeme (
<taxonomicName id="F38B09B717E7A9C89AEFFD9B7C76FD96" authority="Schefter &amp; Ward 2002" authorityName="Schefter &amp; Ward" authorityYear="2002" box="[741,1077,592,618]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89AEFFD9B7B65FD95" box="[741,806,592,617]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89B27FD9B7C76FD96" author="Schefter, P. W. &amp; Ward, J. B." box="[813,1077,592,618]" pageId="214" pageNumber="215" pagination="18 - 31" refId="ref112062" refString="Schefter, P. W. &amp; Ward, J. B. (2002) Revision of the New Caledonian endemic genus Caledopsyche (Trichoptera: Hydropsychidae: Hydropsychinae), with the description of five new species. Records of the Canterbury Museum, 16, 18 - 31." type="journal article" year="2002">Schefter &amp; Ward 2002</bibRefCitation>
</taxonomicName>
) narrow, more strongly sclerotized anterior and posteriorly than at centre; parallel-sided; fusing to dorsal margin of sternum VIII; separated by well-developed sutures; setose posteriorly. Sternum VIII rounded, nearly quadrangular; freely flanking ventrocaudal lobe-like corners, ventral plate (
<taxonomicName id="F38B09B717E7A9C89AC2FD037BFBFD1E" authority="Nielsen 1980" authorityName="Nielsen" authorityYear="1980" box="[712,952,712,738]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89AC2FD037B4AFD1D" box="[712,777,712,737]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89B1CFD037BFBFD1E" author="Nielsen, A." box="[790,952,712,738]" pageId="214" pageNumber="215" pagination="1 - 200" refId="ref111172" refString="Nielsen, A. (1980) A comparative study of the genital segments and the genital chamber in female Trichoptera. Biologiske Skifter, 23 (1), 1 - 200." type="journal article" year="1980">Nielsen 1980</bibRefCitation>
</taxonomicName>
) or lateral plate (
<taxonomicName id="F38B09B717E7A9C89C92FD037DD7FD1E" authority="Denning 1943" authorityName="Denning" authorityYear="1943" box="[1176,1428,712,738]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89C92FD037C9AFD1D" box="[1176,1241,712,737]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89CECFD037DD7FD1E" author="Denning, D. G." box="[1254,1428,712,738]" pageId="214" pageNumber="215" pagination="101 - 171" refId="ref107007" refString="Denning, D. G. (1943) The Hydropsychidae of Minnesota. Entomologica Americana (New Series), 23 (2), 101 - 171." type="journal article" year="1943">Denning 1943</bibRefCitation>
</taxonomicName>
) formed by deep, wide, mesal V-shaped incision. Lobe-like corners, together with distal margin densely covered by long setae flanking fusing membranous sternal part of segment IX. Internal surfaces densely packed with minute hooks; heavily setose ventrocaudal lobes forming second support of copulation chelum of segment VIII. Long setae densely packed on ventrocaudal lobes; posterior half of sternum VIII covered by short setae; basal part devoid of setae. Tergum IX vertically elongate in lateral view; orienting obliquely to main axis of abdomen; narrowing ventrad together with lateral lobe on ventrodistal corners of segment. Narrow antecosta present on anterior margins, with acrotergite on dorsum. Dorsal margin rounded. Posterodorsal margin of segment IX with small rim-like lobe on both sides; with 3 long, stiff setae. Sclerotized ridge running along central part of tergum IX from apicodorsal corners and from small clasper receptacles, ventrad to onethird of segment; delineating shallow, flat, depressed, distal cavity of clasper grooves. Clasper grooves forming small clasper areas being depressions on apicodorsal corners of tergum IX; delineated anteriorly by ridge with 3 setae on anterior margins; delineated ventrally by elevated lateral lobes; delineated dorsally by dorsal rim housing clasper receptacles. Elevating ventral part producing into small triangular, lateral lobe (
<taxonomicName id="F38B09B717E7A9C89D51FB1B7903FAEE" authority="Denning 1943" authorityName="Denning" authorityYear="1943" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89D51FB1B7DDFFB15" box="[1371,1436,1232,1257]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C8989DFB337903FAEE" author="Denning, D. G." box="[151,320,1272,1298]" pageId="214" pageNumber="215" pagination="101 - 171" refId="ref107007" refString="Denning, D. G. (1943) The Hydropsychidae of Minnesota. Entomologica Americana (New Series), 23 (2), 101 - 171." type="journal article" year="1943">Denning 1943</bibRefCitation>
</taxonomicName>
) or tongue-like lamellae (
<taxonomicName id="F38B09B717E7A9C89A7EFB337B1EFAEE" authority="Nielsen 1980" authorityName="Nielsen" authorityYear="1980" box="[628,861,1272,1298]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89A7EFB337AF6FAED" box="[628,693,1272,1297]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89AB7FB337B1EFAEE" author="Nielsen, A." box="[701,861,1272,1298]" pageId="214" pageNumber="215" pagination="1 - 200" refId="ref111172" refString="Nielsen, A. (1980) A comparative study of the genital segments and the genital chamber in female Trichoptera. Biologiske Skifter, 23 (1), 1 - 200." type="journal article" year="1980">Nielsen 1980</bibRefCitation>
</taxonomicName>
); with 2–3 long apical setae. Clasper receptacles reduced into small, sinuous, cavities. Distinctly marked suture delineating less pigmented, short tergum X along distal margin of tergite IX. Tergum X representing body of upper lip of atrium (
<taxonomicName id="F38B09B717E7A9C89CA3FA837DD7FA9E" authority="Nielsen 1980" authorityName="Nielsen" authorityYear="1980" box="[1193,1428,1352,1378]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89CA3FA837CA9FA9D" box="[1193,1258,1352,1377]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89CFEFA837DD7FA9E" author="Nielsen, A." box="[1268,1428,1352,1378]" pageId="214" pageNumber="215" pagination="1 - 200" refId="ref111172" refString="Nielsen, A. (1980) A comparative study of the genital segments and the genital chamber in female Trichoptera. Biologiske Skifter, 23 (1), 1 - 200." type="journal article" year="1980">Nielsen 1980</bibRefCitation>
</taxonomicName>
) composed of pair of foliate, principal, flat lobes covered by short setae nested in large wart-like alveoli; and tridactylate short lobes; dorsal lobe being largest; middle process (cercus
<taxonomicName id="F38B09B717E7A9C89C08FA537CA9FA4E" authority="Nielsen 1980" authorityName="Nielsen" authorityYear="1980" box="[1026,1258,1432,1458]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89C08FA537C00FA4D" box="[1026,1091,1432,1457]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89C46FA537CA9FA4E" author="Nielsen, A." box="[1100,1258,1432,1458]" pageId="214" pageNumber="215" pagination="1 - 200" refId="ref111172" refString="Nielsen, A. (1980) A comparative study of the genital segments and the genital chamber in female Trichoptera. Biologiske Skifter, 23 (1), 1 - 200." type="journal article" year="1980">Nielsen 1980</bibRefCitation>
</taxonomicName>
) digitate, slen- der, armed with 2 apical setae. Dorsal and ventral processes blunt, with tiny, peg-like setae on dome-like alveolar membranes in deep alveoli, visible at high magnification. Sclerotized, setose, foliate, small, flat, lobe concealed under foliate, principal setose lobes of tergite X; possibly representing rudimentary sternal lobes, or the lateral tergal plate (
<taxonomicName id="F38B09B717E7A9C899A5F9F37AE4F9AE" authority="Denning 1943" authorityName="Denning" authorityYear="1943" box="[431,679,1592,1618]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C899A5F9F379B3F9AD" box="[431,496,1592,1617]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C899F3F9F37AE4F9AE" author="Denning, D. G." box="[505,679,1592,1618]" pageId="214" pageNumber="215" pagination="101 - 171" refId="ref107007" refString="Denning, D. G. (1943) The Hydropsychidae of Minnesota. Entomologica Americana (New Series), 23 (2), 101 - 171." type="journal article" year="1943">Denning 1943</bibRefCitation>
</taxonomicName>
). Lobes connected by smooth, membranous, triangular, median body surrounding anovaginal dorsal parts of opening; connecting basally to lower lip. Lower lip of atrium (
<taxonomicName id="F38B09B717E7A9C898ABF94379CAF95E" authority="Nielsen 1980" authorityName="Nielsen" authorityYear="1980" box="[161,393,1672,1698]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C898ABF94378A1F95D" box="[161,226,1672,1697]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C898E3F94379CAF95E" author="Nielsen, A." box="[233,393,1672,1698]" pageId="214" pageNumber="215" pagination="1 - 200" refId="ref111172" refString="Nielsen, A. (1980) A comparative study of the genital segments and the genital chamber in female Trichoptera. Biologiske Skifter, 23 (1), 1 - 200." type="journal article" year="1980">Nielsen 1980</bibRefCitation>
</taxonomicName>
), or vulvar scale (
<taxonomicName id="F38B09B717E7A9C89A57F9437B06F95E" authority="Schmid 1998" authorityName="Schmid" authorityYear="1998" box="[605,837,1672,1698]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="06FFAE2617E7A9C89A57F9437ADDF95D" box="[605,670,1672,1697]" italics="true" pageId="214" pageNumber="215">sensu</emphasis>
<bibRefCitation id="501A0FC517E7A9C89AAFF9437B06F95E" author="Schmid, F." box="[677,837,1672,1698]" pageId="214" pageNumber="215" refId="ref112510" refString="Schmid, F. (1998) The insects and arachnids of Canada Part 7. Genera of the Trichoptera of Canada and adjoining or adjacent United States. NRC Research Press, Ottawa, 289 pp." type="book" year="1998">Schmid 1998</bibRefCitation>
</taxonomicName>
), representing sternal portions of segment IX, visible mostly as distended bag-like structure; mostly covered by sternum VIII. Vaginal apparatus in mostly membranous, slightly sclerotized, sinuate, thin, trip connecting lower lip with sclerotized tergum of segment IX, as well as pair of slightly sclerotized plates on dorsum of lower lip.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17E7A9C898CFF8EC78A8F846" pageId="214" pageNumber="215" type="materials_examined">
<paragraph id="3434723417E7A9C898CFF8EC7C07F896" blockId="214.[151,1436,152,2018]" pageId="214" pageNumber="215">
<materialsCitation id="84E3786917E7A9C898CFF8EC7C07F896" collectingDate="1990-03-26" collectingDateMax="1990-03-29" collectingDateMin="1990-03-26" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="214" pageNumber="215" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="06FFAE2617E7A9C898CFF8EC7A2CF8BD" bold="true" box="[197,623,1831,1857]" pageId="214" pageNumber="215">
<typeStatus id="EB30CC9617E7A9C898CFF8EC7974F8BD" box="[197,311,1831,1857]" pageId="214" pageNumber="215">Holotype</typeStatus>
male:
<collectingCountry id="4C9C32A417E7A9C89984F8EC7A29F8BD" box="[398,618,1831,1857]" name="Madagascar" pageId="214" pageNumber="215">MADAGASCAR</collectingCountry>
:
</emphasis>
<collectingRegion id="F64FBCD617E7A9C89A72F8E37BC1F8BE" box="[632,898,1832,1858]" country="Madagascar" name="Fianarantsoa" pageId="214" pageNumber="215">Fianarantsoa Province</collectingRegion>
,
<quantity id="F373DFD117E7A9C89B85F8E37B88F8BE" box="[911,971,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="214" pageNumber="215" unit="km" value="7.0">7 km</quantity>
W
<location id="315424EF17E7A9C89BF0F8E37CD2F8BE" LSID="urn:lsid:plazi:treatment:BC22C32217E5A9C9989DF99E7932FED6:315424EF17E7A9C89BF0F8E37CD2F8BE" box="[1018,1169,1832,1858]" country="Madagascar" name="Ranomafana" pageId="214" pageNumber="215" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="F373DFD117E7A9C89C94F8E37CB8F8BE" box="[1182,1275,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="214" pageNumber="215" unit="m" value="1000.0">
<elevation id="BFA6950717E7A9C89C94F8E37CB8F8BE" box="[1182,1275,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="214" pageNumber="215" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<collectingMethod id="EDCA0A2317E7A9C89D03F8E37DDFF8BE" box="[1289,1436,1832,1858]" pageId="214" pageNumber="215">Malaise trap</collectingMethod>
in small clearing, mountain rainforest,
<date id="403554F417E7A9C89A54F89B7B4AF896" box="[606,777,1872,1898]" pageId="214" pageNumber="215" value="1990-03-26" valueMax="1990-03-29" valueMin="1990-03-26">
<collectingDate id="5071AD1C17E7A9C89A54F89B7B4AF896" box="[606,777,1872,1898]" pageId="214" pageNumber="215" value="1990-03-26" valueMax="1990-03-29" valueMin="1990-03-26">26–29.iii.1990</collectingDate>
</date>
[
<collectorName id="997E17E217E7A9C89B1DF89B7BECF896" box="[791,943,1872,1898]" pageId="214" pageNumber="215">W.E. Steiner</collectorName>
] - (
<collectionCode id="529AEAF117E7A9C89BDDF89B7C7BF896" box="[983,1080,1872,1898]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="214" pageNumber="215" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="3434723417E7A9C898CFF8B378A8F846" blockId="214.[151,1436,152,2018]" pageId="214" pageNumber="215">
<materialsCitation id="84E3786917E7A9C898CFF8B378A5F846" collectingDate="1990-03-26" collectingDateMax="1990-03-29" collectingDateMin="1990-03-26" collectionCode="NMNH, OPC" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="214" pageNumber="215" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="EB30CC9617E7A9C898CFF8B3797FF86E" box="[197,316,1912,1938]" pageId="214" pageNumber="215">Paratypes</typeStatus>
:
<emphasis id="06FFAE2617E7A9C89940F8BC7A68F86D" bold="true" box="[330,555,1911,1937]" pageId="214" pageNumber="215">
<collectingCountry id="4C9C32A417E7A9C89940F8BC7A65F86D" box="[330,550,1911,1937]" name="Madagascar" pageId="214" pageNumber="215">MADAGASCAR</collectingCountry>
:
</emphasis>
same data as holotype -
<specimenCount id="228DB9BD17E7A9C89B6FF8B37BF8F86E" box="[869,955,1912,1938]" pageId="214" pageNumber="215" type="male">1 male</specimenCount>
,
<specimenCount id="228DB9BD17E7A9C89BC1F8B37C03F86E" box="[971,1088,1912,1938]" pageId="214" pageNumber="215" type="female">5 females</specimenCount>
(
<collectionCode id="529AEAF117E7A9C89C5EF8B37CF6F86E" box="[1108,1205,1912,1938]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="214" pageNumber="215" type="Museum">NMNH</collectionCode>
);
<specimenCount id="228DB9BD17E7A9C89CC6F8B37D61F86E" box="[1228,1314,1912,1938]" pageId="214" pageNumber="215" type="male">1 male</specimenCount>
,
<specimenCount id="228DB9BD17E7A9C89D38F8B37DDFF86E" box="[1330,1436,1912,1938]" pageId="214" pageNumber="215" type="female">1 female</specimenCount>
(
<collectionCode id="529AEAF117E7A9C89895F86B789DF846" box="[159,222,1952,1978]" pageId="214" pageNumber="215">OPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17E7A9C898CFF80C7A43F81E" box="[197,512,1991,2018]" pageId="214" pageNumber="215" type="distribution">
<paragraph id="3434723417E7A9C898CFF80C7A43F81E" blockId="214.[151,1436,152,2018]" box="[197,512,1991,2018]" pageId="214" pageNumber="215">
<emphasis id="06FFAE2617E7A9C898CFF80C7926F81D" bold="true" box="[197,357,1991,2017]" pageId="214" pageNumber="215">Distribution.</emphasis>
<collectingCountry id="4C9C32A417E7A9C89966F80379BFF81E" box="[364,508,1992,2018]" name="Madagascar" pageId="214" pageNumber="215">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17E6A9C998CFFF5C7DD5FF26" pageId="215" pageNumber="216" type="etymology">
<paragraph id="3434723417E6A9C998CFFF5C7DD5FF26" blockId="215.[151,1436,151,298]" pageId="215" pageNumber="216">
<emphasis id="06FFAE2617E6A9C998CFFF5C7912FF4D" bold="true" box="[197,337,151,177]" pageId="215" pageNumber="216">Etymology.</emphasis>
<taxonomicName id="F38B09B717E6A9C99955FF5379B2FF4D" authorityName="Oláh &amp; Johanson" authorityYear="2008" box="[351,497,152,177]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" species="tiokalamkita">
<emphasis id="06FFAE2617E6A9C99955FF5379B2FF4D" box="[351,497,152,177]" italics="true" pageId="215" pageNumber="216">tiokalamkita</emphasis>
</taxonomicName>
, from “tio”, meaning bi; and “kalamkita”, meaning maculated in Sanskrit, named after the remarkable, single white spot in the forewing centre, giving it an elegant, bimaculate feature.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17E6A9C998CFFF2C7932FED6" pageId="215" pageNumber="216" type="discussion">
<paragraph id="3434723417E6A9C998CFFF2C7932FED6" blockId="215.[151,1436,151,298]" pageId="215" pageNumber="216">
<emphasis id="06FFAE2617E6A9C998CFFF2C797DFEFD" bold="true" box="[197,318,231,257]" pageId="215" pageNumber="216">Remarks.</emphasis>
Segments IX and X slightly extruded with pincers from partial coverage of segment VIII for proper illustration.
</paragraph>
</subSubSection>
</treatment>
</document>
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