158 lines
22 KiB
XML
158 lines
22 KiB
XML
<document id="2FADFAABFC570073B4B8451DECB9F9F7" ID-ISSN="11755334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. & Johanson, K. A." docDate="2008" docId="BC22C3221788A9A4989DFA917A72F872" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Cheumatopsyche anonima Olah & Johanson 2008, new species" docType="treatment" docVersion="3" lastPageNumber="187" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="186" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="58C7B6B35EB1B8F6F71850C63F7CC847">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="A4E5B34327088C8FE0B4E284C330DB24">Oláh, J.</mods:namePart>
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<mods:namePart id="2B79B8D33FFF3296C3B735437A7B652C">Johanson, K. A.</mods:namePart>
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<mods:title id="79AE9089F91C3881305D928D684CAD61">Zootaxa</mods:title>
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<mods:part id="6C4CD207771E4AD3E9A25DFF657D9AB6">
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<mods:date id="6939768D5EFAEF6F3F43EA9C49DCCAFE">2008</mods:date>
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<mods:number id="2E9EB245C0347C4ADDEA04719032A763">2008-06-16</mods:number>
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<mods:number id="CB5ACF576D6D8EF13EBD09EE19CA9396">1802</mods:number>
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<mods:classification id="71BAC1D9EA3DDF554EBDC6A0C88D6FFC">journal article</mods:classification>
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<mods:identifier id="617B696C12363FC151B0D0FB73FF97B6" type="ISSN">11755334</mods:identifier>
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<treatment id="BC22C3221788A9A4989DFA917A72F872" ID-DOI="http://doi.org/10.5281/zenodo.5126156" ID-Zenodo-Dep="5126156" LSID="urn:lsid:plazi:treatment:BC22C3221788A9A4989DFA917A72F872" httpUri="http://treatment.plazi.org/id/BC22C3221788A9A4989DFA917A72F872" lastPageId="186" lastPageNumber="187" pageId="185" pageNumber="186">
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<subSubSection id="7C9121BF1788A9A7989DFA917B19FA88" box="[151,858,1370,1396]" pageId="185" pageNumber="186" type="nomenclature">
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<paragraph id="343472341788A9A7989DFA917B19FA88" blockId="185.[151,858,1370,1436]" box="[151,858,1370,1396]" pageId="185" pageNumber="186">
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<heading id="6F7CC5581788A9A7989DFA917B19FA88" bold="true" box="[151,858,1370,1396]" fontSize="11" level="1" pageId="185" pageNumber="186" reason="1">
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<emphasis id="06FFAE261788A9A7989DFA917B19FA88" bold="true" box="[151,858,1370,1396]" pageId="185" pageNumber="186">
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<taxonomicName id="F38B09B71788A9A7989DFA917AFDFA88" authority="Olah & Johanson" authorityName="Olah & Johanson" authorityYear="2008" box="[151,702,1370,1396]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="anonima" status="new species">
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<emphasis id="06FFAE261788A9A7989DFA91799BFA88" bold="true" box="[151,472,1370,1396]" italics="true" pageId="185" pageNumber="186">Cheumatopsyche anonima</emphasis>
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Oláh & Johanson
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</taxonomicName>
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,
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<taxonomicNameLabel id="1DCC135D1788A9A79AC0FA917B19FA88" box="[714,858,1370,1396]" pageId="185" pageNumber="186" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="7C9121BF1788A9A4989DFA497B3BF912" lastPageId="186" lastPageNumber="187" pageId="185" pageNumber="186" type="description">
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<paragraph id="343472341788A9A7989DFA497970FA60" blockId="185.[151,858,1370,1436]" box="[151,307,1410,1436]" pageId="185" pageNumber="186">
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<figureCitation id="ACB06EB11788A9A7989DFA497970FA60" box="[151,307,1410,1436]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="185" pageNumber="186">Fig. 307–310</figureCitation>
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</paragraph>
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<paragraph id="343472341788A9A7989DFA197886F948" blockId="185.[151,1437,1490,1996]" pageId="185" pageNumber="186">
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This species is recognized by having a combination of trilobed segment X; shortened apicoventral setose lobes; and a short, convex smooth mesocaudal lobe. The species is closely related to
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<taxonomicName id="F38B09B71788A9A79C90FA307DDFF9E8" authority="Mey" authorityName="Mey" box="[1178,1436,1530,1556]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="nathanbanksi">
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<emphasis id="06FFAE261788A9A79C90FA307D1EF9E8" box="[1178,1373,1531,1556]" italics="true" pageId="185" pageNumber="186">C. nathanbanksi</emphasis>
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Mey
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</taxonomicName>
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and
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<taxonomicName id="F38B09B71788A9A798C1F9E8799CF9C0" authority="Mey" authorityName="Mey" box="[203,479,1570,1596]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="longinosnavasi">
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<emphasis id="06FFAE261788A9A798C1F9E879E3F9C0" box="[203,416,1571,1596]" italics="true" pageId="185" pageNumber="186">C. longinosnavasi</emphasis>
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Mey
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</taxonomicName>
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from the
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<collectingCountry id="4C9C32A41788A9A79A52F9E97AA3F9C0" box="[600,736,1570,1596]" name="Philippines" pageId="185" pageNumber="186">Philippines</collectingCountry>
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.
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<taxonomicName id="F38B09B71788A9A79AE7F9E87C66F9C0" authorityName="Olah & Johanson" authorityYear="2008" box="[749,1061,1571,1596]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="anonima">
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<emphasis id="06FFAE261788A9A79AE7F9E87C66F9C0" box="[749,1061,1571,1596]" italics="true" pageId="185" pageNumber="186">Cheumatopsyche anonima</emphasis>
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</taxonomicName>
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is easily distinguished from
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<taxonomicName id="F38B09B71788A9A79D8BF9E8790BF998" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="longinosnavasi">
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<emphasis id="06FFAE261788A9A79D8BF9E8790BF998" italics="true" pageId="185" pageNumber="186">C. longinosnavasi</emphasis>
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</taxonomicName>
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by the straight harpagones; and from
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<taxonomicName id="F38B09B71788A9A79B03F9807B8AF998" box="[777,969,1611,1636]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="nathanbanksi">
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<emphasis id="06FFAE261788A9A79B03F9807B8AF998" box="[777,969,1611,1636]" italics="true" pageId="185" pageNumber="186">C. nathanbanksi</emphasis>
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</taxonomicName>
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by the more ventral position of preanal appendages, by the smaller and rounded apicoventral setose lobes, and by the convex dorsum of the phallotheca.
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</paragraph>
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<paragraph id="343472341788A9A798CFF90979C4F8A8" blockId="185.[151,1437,1490,1996]" pageId="185" pageNumber="186">Male. Body brown. Maxillary palp formula I-(III-IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum dark brown, with 9 setal warts brown. Swollen setal wart absent on proepisternum. Setal surface present on each precoxale. Pretarsal claw asymmetrical, laterally flanked by setal bundle. Mid- and hind-leg claws asymmetrical.</paragraph>
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<paragraph id="343472341788A9A798CFF8A97C7DF830" blockId="185.[151,1437,1490,1996]" pageId="185" pageNumber="186">Wings. Forewing membranes evenly brown, with little darker veins. Crossveins m-cu and cu meeting in oblique line. Cu2 and A1 running separately before costa. Hind wing Sc and R meeting at crossvein s. Crossvein r located anteriorly of crossvein s. Fork 1 absent. Forewing length 6.5 mm.</paragraph>
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<caption id="60F422BC178BA9A4989DFCA47BFCFC55" pageId="186" pageNumber="187" startId="186.[151,269,879,903]" targetBox="[170,1413,206,835]" targetPageId="186">
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<paragraph id="34347234178BA9A4989DFCA47BFCFC55" blockId="186.[151,1436,879,937]" pageId="186" pageNumber="187">
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<emphasis id="06FFAE26178BA9A4989DFCA4793BFC7B" bold="true" box="[151,376,879,903]" pageId="186" pageNumber="187">FIGURES 307–310.</emphasis>
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<taxonomicName id="F38B09B7178BA9A49974FCA47B1AFC7A" authority="Olah & Johanson" authorityName="Olah & Johanson" authorityYear="2008" box="[382,857,879,902]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="anonima" status="new species">
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<emphasis id="06FFAE26178BA9A49974FCA47AD4FC7A" box="[382,663,879,902]" italics="true" pageId="186" pageNumber="187">Cheumatopsyche anonima</emphasis>
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Oláh & Johanson
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</taxonomicName>
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,
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<taxonomicNameLabel id="1DCC135D178BA9A49B69FCA47BA6FC7A" box="[867,997,879,902]" pageId="186" pageNumber="187" rank="species">new species</taxonomicNameLabel>
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, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral.
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</paragraph>
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</caption>
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<paragraph id="34347234178BA9A498CFFC177B3BF912" blockId="186.[151,1436,988,1934]" pageId="186" pageNumber="187">
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Male genitalia. Abdominal segment IX fused annularly, tergum as long as sternum (
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<figureCitation id="ACB06EB1178BA9A49CB3FC177D61FC0A" box="[1209,1314,988,1014]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307</figureCitation>
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). Anterior margins of segment IX regularly bow-shaped, drawn slightly ventrally in lateral aspect (
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<figureCitation id="ACB06EB1178BA9A49C90FBCF7D43FBE2" box="[1178,1280,1028,1054]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307</figureCitation>
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). Apical lobe on posterolateral margins blunt right-angled; located immediately above less sclerotized articulation cavity of gonocoxites. Spine row on posterior margins of segment IX intermittent, heterogeneous. Spines on dorsolateral spiny lobes 2 times longer than spines on apical lobes; in dorsal view, dorsolateral spiny lobes fusing into continuous, rim-like plate, with tiny, mesoapical triangular process (
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<figureCitation id="ACB06EB1178BA9A49BBDFB6F7C5DFB42" box="[951,1054,1188,1214]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 308</figureCitation>
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). Step between segments IX and X stepwise, shallow, sloping. Segment X short (
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<figureCitation id="ACB06EB1178BA9A49AC4FB077B76FB1A" box="[718,821,1228,1254]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307</figureCitation>
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). Small step on dorsum present at mid-length, distal margin of mesocaudal smooth plate slightly curving dorsad. Segment X trilobed in dorsal aspect (
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<figureCitation id="ACB06EB1178BA9A49D2CFB3F7DCDFAF2" box="[1318,1422,1268,1294]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 308</figureCitation>
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); dorsal, interlobular gap almost absent, filled by mesocaudal plate. Lateral sutures well developed; transverse; crossing segment X obliquely; orienting toward step corners. Both longitudinal sutures broad, forming continuation of apicoventral setose lobes; meeting transverse suture into Y. Smooth mesocaudal plate reduced, ending in triangular margin; slightly curving dorsad, forming rim in lateral view (
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<figureCitation id="ACB06EB1178BA9A49C3FFA5F7CDEFA52" box="[1077,1181,1428,1454]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307</figureCitation>
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). Apicoventral setose lobes short, orienting posterad, rounded apically (
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<figureCitation id="ACB06EB1178BA9A49AF9FA777BDFFA2A" box="[755,924,1468,1494]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307, 308</figureCitation>
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). Lateral setose areas (superior or preanal appendages) forming small elevated warts located near ventral margin of segment X. Coxopodites exceeding apex of segment X, forming straight rod, dilating gradually towards apex (
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<figureCitation id="ACB06EB1178BA9A49BF4F9C77C27F9DA" box="[1022,1124,1548,1574]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307</figureCitation>
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); curving mesad in ventral view (
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<figureCitation id="ACB06EB1178BA9A498E9F9FF790FF9B2" box="[227,332,1588,1614]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 309</figureCitation>
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). Harpagones slender, filiform, nearly straight (
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<figureCitation id="ACB06EB1178BA9A49B82F9FF7C6CF9B2" box="[904,1071,1588,1614]" captionStart="FIGURES 307–310" captionStartId="186.[151,269,879,903]" captionTargetBox="[170,1413,206,835]" captionTargetId="figure-371@186.[151,1436,197,852]" captionTargetPageId="186" captionText="FIGURES 307–310. Cheumatopsyche anonima Oláh & Johanson, new species, holotype, male. 307 — genitalia, lateral; 308 — genitalia, dorsal; 309 — gonocoxite, ventral; 310 — phallus, lateral." pageId="186" pageNumber="187">Fig. 307, 309</figureCitation>
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). Phallotheca with short, shallowly broadening, ventrad curving basal part; horizontal part with convex dorsum; ventrum concave; producing slightly ventrad before apex.
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<taxonomicName id="F38B09B7178BA9A49A13F94F7AE8F962" authorityName="M. Zang & R.H. Petersen" authorityYear="1989" box="[537,683,1668,1694]" class="Agaricomycetes" family="Phallaceae" genus="Endophallus" kingdom="Fungi" order="Phallales" pageId="186" pageNumber="9" phylum="Basidiomycota" rank="genus">Endophallus</taxonomicName>
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long, running through entire phallotheca; ending in narrow tube at gonopore. Chitinized endothecal process cut distally; phallotremal sclerites vertically rounded in lateral view. Vestigial, membranous,ventral endothecal lobes present.
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</paragraph>
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</subSubSection>
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<subSubSection id="7C9121BF178BA9A498CFF9377BCEF8EA" box="[197,909,1788,1814]" pageId="186" pageNumber="187" type="materials_examined">
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<paragraph id="34347234178BA9A498CFF9377BCEF8EA" blockId="186.[151,1436,988,1934]" box="[197,909,1788,1814]" pageId="186" pageNumber="187">
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<materialsCitation id="84E37869178BA9A498CFF9377BCEF8EA" box="[197,909,1788,1814]" collectorName="Phil. I." country="Philippines" location="PHILIPPINES" pageId="186" pageNumber="187" specimenCount="1" typeStatus="holotype">
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<emphasis id="06FFAE26178BA9A498CFF9377A05F8EA" bold="true" box="[197,582,1788,1814]" pageId="186" pageNumber="187">
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<typeStatus id="EB30CC96178BA9A498CFF9377974F8EA" box="[197,311,1788,1814]" pageId="186" pageNumber="187">Holotype</typeStatus>
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male:
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<collectingCountry id="4C9C32A4178BA9A49981F9377A05F8EA" box="[395,582,1788,1814]" name="Philippines" pageId="186" pageNumber="187">PHILIPPINES</collectingCountry>
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</emphasis>
|
||
: Thru Miss Ludlow,
|
||
<collectorName id="997E17E2178BA9A49B34F9377BCEF8EA" box="[830,909,1788,1814]" pageId="186" pageNumber="187">Phil. I.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C9121BF178BA9A498CFF8EF79BBF8C2" box="[197,504,1828,1854]" pageId="186" pageNumber="187" type="distribution">
|
||
<paragraph id="34347234178BA9A498CFF8EF79BBF8C2" blockId="186.[151,1436,988,1934]" box="[197,504,1828,1854]" pageId="186" pageNumber="187">
|
||
<emphasis id="06FFAE26178BA9A498CFF8EF7926F8C2" bold="true" box="[197,357,1828,1854]" pageId="186" pageNumber="187">Distribution.</emphasis>
|
||
<collectingCountry id="4C9C32A4178BA9A49966F8EF79B7F8C2" box="[364,500,1828,1854]" name="Philippines" pageId="186" pageNumber="187">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C9121BF178BA9A498CFF8877A72F872" pageId="186" pageNumber="187" type="etymology">
|
||
<paragraph id="34347234178BA9A498CFF8877A72F872" blockId="186.[151,1436,988,1934]" pageId="186" pageNumber="187">
|
||
<emphasis id="06FFAE26178BA9A498CFF8877912F89A" bold="true" box="[197,337,1868,1894]" pageId="186" pageNumber="187">Etymology.</emphasis>
|
||
<taxonomicName id="F38B09B7178BA9A49953F88679FCF89A" authorityName="Olah & Johanson" authorityYear="2008" box="[345,447,1869,1894]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="anonima">
|
||
<emphasis id="06FFAE26178BA9A49953F88679FCF89A" box="[345,447,1869,1894]" italics="true" pageId="186" pageNumber="187">anonima</emphasis>
|
||
</taxonomicName>
|
||
, from anonymous, named after the limited label data available, i.e. no sampling date or locality name in the
|
||
<collectingCountry id="4C9C32A4178BA9A499AFF8BF7A6EF872" box="[421,557,1908,1934]" name="Philippines" pageId="186" pageNumber="187">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |