treatments-xml/data/78/5F/AD/785FAD8B3F23144876A32FE04397F386.xml
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<mods:title id="60F13AE7EC577A34C457C9C84AEFCBC5">A new tarantula species from northern Australia (Araneae, Theraphosidae)</mods:title>
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<taxonomicName id="A7F8706AAAD2F26FC795D99E88D39335" authority="Simon 1889" authorityName="Simon" authorityYear="1889" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">
Selenocosmiinae
<bibRefCitation id="50D4DA1364F593D68832FC46814F477B" author="Simon, E." box="[206,1039,930,979]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="169 - 220" part="(6) 9" refId="ref6001" refString="Simon, E. (1889) Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides. Annales de la Societe entomologique de France, (6) 9, 169 - 220." title="Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides" type="journal article" year="1889">Simon 1889</bibRefCitation>
</taxonomicName>
</paragraph>
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<taxonomicName id="F1BABF31227CA666F75B26D462B2786B" authority="Simon 1889: 204" authorityName="Simon" authorityPageNumber="204" authorityYear="1889" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">
Selenocosmiinae
<bibRefCitation id="A1392E5BE79EE49FCE59E93B523C921C" author="Simon, E." box="[206,1039,930,979]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="169 - 220" part="(6) 9" refId="ref6001" refString="Simon, E. (1889) Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides. Annales de la Societe entomologique de France, (6) 9, 169 - 220." title="Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides" type="journal article" year="1889">Simon 1889: 204</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="C3D55CAF936870CA5D734366C2C565C1" author="Simon, E." box="[206,878,986,1004]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="1 - 256" part="vol. 1, part 1" refId="ref6048" refString="Simon, E. (1892) Histoire naturelle des araignees. Paris, vol. 1, part 1, pp. 1 - 256." title="Histoire naturelle des araignees" type="book chapter" year="1892">Simon 1892: 147</bibRefCitation>
;
<bibRefCitation id="B5F2475876C84D513F01369F748E03DB" author="Raven, R. J." box="[206,1032,480,523]" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="1 - 180" part="182" refId="ref5663" refString="Raven, R. J. (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics" type="journal article" year="1985">Raven 1985: 118</bibRefCitation>
.
</paragraph>
<paragraph id="F473CE74672350AB2D62A61C1F1E13F5" box="[206,643,922,940]" pageId="1" pageNumber="16" type="mainText">
<taxonomicName id="63AAE8D949DF709614F02D8E00B54462" authority="Simon 1889: 204" authorityName="Simon" authorityPageNumber="204" authorityYear="1889" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Poecilotherieae">
Poecilotherieae
<bibRefCitation id="4B0CA6B7190933C0A6FA522D0E62A04A" author="Simon, E." box="[206,1039,930,979]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="169 - 220" part="(6) 9" refId="ref6001" refString="Simon, E. (1889) Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides. Annales de la Societe entomologique de France, (6) 9, 169 - 220." title="Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire, Arachnides" type="journal article" year="1889">Simon 1889: 204</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="4843D92448CEDD1A56A79A3C3C5F74B9" author="Simon, E." box="[206,878,986,1004]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="1 - 256" part="vol. 1, part 1" refId="ref6048" refString="Simon, E. (1892) Histoire naturelle des araignees. Paris, vol. 1, part 1, pp. 1 - 256." title="Histoire naturelle des araignees" type="book chapter" year="1892">Simon 1892: 144</bibRefCitation>
.
</paragraph>
<paragraph id="D3DD9758C5ED72B2E533B566F055C579" box="[206,780,947,965]" pageId="1" pageNumber="16" type="mainText">
<taxonomicName id="3767E67899BE50992EEC6DE67421C70F" authority="Simon 1892: 144" authorityName="Simon" authorityPageNumber="144" authorityYear="1892" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Phlogieae">
Phlogieae
<bibRefCitation id="8FBDFD464AC25F62F8BECE65696A0951" author="Simon, E." box="[206,878,986,1004]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="1 - 256" part="vol. 1, part 1" refId="ref6048" refString="Simon, E. (1892) Histoire naturelle des araignees. Paris, vol. 1, part 1, pp. 1 - 256." title="Histoire naturelle des araignees" type="book chapter" year="1892">Simon 1892: 144</bibRefCitation>
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. First synonymised by
<bibRefCitation id="47609C9F6807BE4E50E7851E512E1A83" author="Simon, E." box="[206,910,1011,1029]" journalOrPublisher="Annales de la Societe entomologique de France" pagination="669 - 1080" part="vol. 2, part 4" refId="ref6075" refString="Simon, E. (1903) Histoire naturelle des araignees. Paris, vol. 2, part 4, pp. 669 - 1080." title="Histoire naturelle des araignees" type="book chapter" year="1903">Simon 1903: 953</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B06217B7D0D33AE1DF631BC1720CB1AC" pageId="1" pageNumber="16" type="diagnosis">
<paragraph id="0BBDA04F94A0507B0727041D05ADE972" box="[206,1034,1003,1054]" pageId="1" pageNumber="16" type="mainText">Diagnosis: With ovoid lyra on anterior face of maxillae which may be reduced or absent, legs weakly spinose, if at all; males lacking coupling spurs on tibia I.</paragraph>
</subSubSection>
<subSubSection id="68335CA9028A1D6091052C2613B10895" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="discussion">
<paragraph id="78BFAE7ACA4CCD5EF664B66105C5DE75" box="[206,1033,1065,1242]" pageId="1" pageNumber="16" type="mainText">
Remarks: The placement of taxa which lack a maxillary lyra (e. g., Yamia; see
<bibRefCitation id="0DDA624C753E9FFE7411163B771807D0" author="Haupt, J. &amp; Schmidt, G. E. W." box="[206,1033,330,398]" journalOrPublisher="Spixiana" pagination="199 - 204" part="27 (3)" refId="ref5551" refString="Haupt, J. &amp; Schmidt, G. E. W. (2004) Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae). Spixiana, 27 (3), 199 - 204." title="Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae)" type="journal article" year="2004">Haupt and Schmidt 2004</bibRefCitation>
) within the
<taxonomicName id="67537F398DAE6B634278DACA896A31EB" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">Selenocosmiinae</taxonomicName>
only presents a phylogenetic problem if the alyrate taxa are considered the sister group of all of the other lyrate species. However, if the lyra has been secondarily lost in Yamia then that species may be better placed in
<taxonomicName id="4F9AA57327D9325BC488279713A3D839" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">Phlogiellus</taxonomicName>
. In either case, the monothetic definition of the subfamily remains an unresolved issue. No phylogenetic analysis has ever been presented which could resolve this question.
</paragraph>
<paragraph id="15F81DC6FA1B5480CD7E383A77293824" box="[206,1033,1253,1461][206,1033,222,429]" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="mainText">
Curiously,
<bibRefCitation id="1FD5183F6C479EC7C245B0E4E41383C6" author="Simon, E." box="[206,878,986,1004]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="1" pageNumber="16" pagination="1 - 256" part="vol. 1, part 1" refId="ref6048" refString="Simon, E. (1892) Histoire naturelle des araignees. Paris, vol. 1, part 1, pp. 1 - 256." title="Histoire naturelle des araignees" type="book chapter" year="1892">Simon (1892)</bibRefCitation>
diagnosed the tribes
<taxonomicName id="4F253B5DAE2F7413F29E48350D93C8E2" class="Arachnida" family="Theraphosidae" genus="Phlogieae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus" tribe="Phlogieae">Phlogieae</taxonomicName>
(and others) as differing from
<taxonomicName id="F984C08390231A3C5EDEE8258DA72EBB" class="Arachnida" family="Theraphosidae" genus="Selenocosmieae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus" tribe="Selenocosmieae">Selenocosmieae</taxonomicName>
in that at least the posterior tarsi had the scopula divided by setae but later (1903) rescinded the separation.
<bibRefCitation id="C6C46B769C13015C05EC639849EF51A6" author="Haupt, J. &amp; Schmidt, G. E. W." box="[206,1033,330,398]" journalOrPublisher="Spixiana" pageId="1" pageNumber="16" pagination="199 - 204" part="27 (3)" refId="ref5551" refString="Haupt, J. &amp; Schmidt, G. E. W. (2004) Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae). Spixiana, 27 (3), 199 - 204." title="Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae)" type="journal article" year="2004">Haupt and Schmidt (2004: 202)</bibRefCitation>
lept from that kind of distinction to a more generalised but unsupported conclusion about the value of the division of the tarsal scopula: &quot; Certain characters do not seem to be very useful for classification:
<bibRefCitation id="89BEC5EA4317BAF7EF934F2CD0BD395B" author="Simon, E." box="[206,878,986,1004]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="1" pageNumber="16" pagination="1 - 256" part="vol. 1, part 1" refId="ref6048" refString="Simon, E. (1892) Histoire naturelle des araignees. Paris, vol. 1, part 1, pp. 1 - 256." title="Histoire naturelle des araignees" type="book chapter" year="1892">Simon (1892)</bibRefCitation>
followed by
<bibRefCitation id="603AE33FA24A754820BDB7DE5F93FEC4" author="Raven, R. J." box="[206,1032,480,523]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="1" pageNumber="16" pagination="1 - 180" part="182" refId="ref5663" refString="Raven, R. J. (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics" type="journal article" year="1985">Raven (1985)</bibRefCitation>
used the division of metatarsal and tarsal scopulae to distinguish different genera. &quot; In fact,
<bibRefCitation id="95B4A374B7C00E3A1B5229E127721573" author="Raven, R. J." box="[206,1032,480,523]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="1" pageNumber="16" pagination="1 - 180" part="182" refId="ref5663" refString="Raven, R. J. (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." title="The spider infraorder Mygalomorphae (Araneae): cladistics and systematics" type="journal article" year="1985">Raven (1985)</bibRefCitation>
never used the division of the
<pageBreakToken id="1AF666360DD90470D314B0698BFA8F98" pageId="2" pageNumber="17" start="start">metatarsal</pageBreakToken>
scopula but its extent and the tarsal division was not used to distinguish between genera (no diagnoses of theraphosid genera were provided) but in an identification key in which the differing states of the tarsal scopula were accommodated by having the genus appear twice in the overall key, e. g.
<taxonomicName id="1DC2340C6B48AD355F8864457C903AF9" class="Arachnida" family="Theraphosidae" genus="Ischnocolus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="17" phylum="Arthropoda" rank="genus">Ischnocolus</taxonomicName>
appeared in Group I and Group II. Furthermore, in a paper presented to the International Congress of Arachnology, Ghent, 2004, Guadanucci (in press) showed unequivocally that the division of the tarsal scopula is informative, provided it is not used to identify juveniles.
</paragraph>
</subSubSection>
</treatment>
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