506 lines
75 KiB
XML
506 lines
75 KiB
XML
<document id="A2EFE3FA9490947C3EFB16AEC3697DEA" ID-CLB-Dataset="24872" ID-DOI="10.11646/zootaxa.4700.1.4" ID-GBIF-Dataset="5d620328-25b8-42e3-a7e9-c641e9031332" ID-ISSN="1175-5326" ID-Zenodo-Dep="3545358" ID-ZooBank="622016E8-D658-4AC6-971F-8C5D9BEFC666" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1574060906900" checkinUser="plazi" docAuthor="Tauber, Catherine A." docDate="2019" docId="6C62879C6624022955A0FD33300C5990" docLanguage="en" docName="zootaxa.4700.1.4.pdf" docOrigin="Zootaxa 4700 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ungla pseudomeleoma Tauber 2019" docType="treatment" docVersion="6" lastPageNumber="96" masterDocId="905BFFE4662502205537FFEA35065E7F" masterDocTitle="A new species of Ungla (Neuroptera: Chrysopidae) that exhibits remarkable homoplasy in male secondary sexual characters" masterLastPageNumber="101" masterPageNumber="87" pageNumber="88" updateTime="1698767467248" updateUser="ExternalLinkService">
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<mods:mods id="BA35620822E22B50F782390A31CE4F8F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="859D18376AE463433CBB402E2FD6557F">
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<mods:title id="281A997782B52F5C732F399886B96E93">A new species of Ungla (Neuroptera: Chrysopidae) that exhibits remarkable homoplasy in male secondary sexual characters</mods:title>
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<mods:name id="A6C6C2EC5D8F9B1046AB6F4378577BF7" type="personal">
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<mods:roleTerm id="028F239D9AA622043CDA15E49D6B3A1E">Author</mods:roleTerm>
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<mods:namePart id="0E0E0CB004BE34C9CFBAD60E906E89F7">Tauber, Catherine A.</mods:namePart>
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<mods:typeOfResource id="1E0D48D12B5C059A5742207EF9ED6D0F">text</mods:typeOfResource>
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<mods:title id="BDD969FC908B649192BEA3E727C97259">Zootaxa</mods:title>
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<mods:part id="D1E2A78961CDEE25D5E4BF705188B723">
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<mods:date id="560F92FE8BC5E64959C9F2F9AF864393">2019</mods:date>
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<mods:number id="FA722DB010E522869C9456DC8D96D5E6">2019-11-18</mods:number>
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<mods:number id="304712B1A07C329DE01816B0B0490989">4700</mods:number>
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<mods:detail id="2A1E352F5AFE82CC8F70786EB565CCCD" type="issue">
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<mods:classification id="E2A98E64521DE1CEBD6F0583FBEA7317">journal article</mods:classification>
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<mods:identifier id="4F8542F2839B49AD8F0A2D2FFFED1EA1" type="DOI">10.11646/zootaxa.4700.1.4</mods:identifier>
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</mods:mods>
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<treatment id="6C62879C6624022955A0FD33300C5990" ID-GBIF-Taxon="160449244" LSID="urn:lsid:plazi:treatment:6C62879C6624022955A0FD33300C5990" httpUri="http://treatment.plazi.org/id/6C62879C6624022955A0FD33300C5990" lastPageId="9" lastPageNumber="96" pageId="1" pageNumber="88">
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<subSubSection id="ACD165016624022155A0FD33370B5C8B" box="[151,525,729,756]" pageId="1" pageNumber="88" type="nomenclature">
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<paragraph id="E474368A6624022155A0FD33370B5C8B" blockId="1.[151,525,729,756]" box="[151,525,729,756]" pageId="1" pageNumber="88">
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<heading id="BF3C81E66624022155A0FD33370B5C8B" bold="true" box="[151,525,729,756]" fontSize="11" level="1" pageId="1" pageNumber="88" reason="1">
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<taxonomicName id="23CB4D096624022155A0FD33370B5C8B" authority="Tauber" authorityName="Tauber" authorityYear="2019" box="[151,525,729,756]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudomeleoma">
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<emphasis id="D6BFEA986624022155A0FD33370B5C8B" bold="true" box="[151,525,729,756]" pageId="1" pageNumber="88">
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<emphasis id="D6BFEA986624022155A0FD3334AD5C8B" bold="true" box="[151,427,729,756]" italics="true" pageId="1" pageNumber="88">Ungla pseudomeleoma</emphasis>
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Tauber
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ACD165016624022155A0FCC8309C5DD8" pageId="1" pageNumber="88" type="materials_examined">
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<paragraph id="E474368A6624022155A0FCC8309C5DD8" blockId="1.[151,1437,801,2016]" pageId="1" pageNumber="88">
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<materialsCitation id="54A33CD76624022155A0FCC835ED5D1F" ID-GBIF-Occurrence="2556249866" collectionCode="FSCA" country="Peru" pageId="1" pageNumber="88" typeStatus="holotype">
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<emphasis id="D6BFEA986624022155A0FCC8340D5D43" bold="true" box="[151,267,802,828]" pageId="1" pageNumber="88">
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<typeStatus id="3B7088286624022155A0FCC834005D43" box="[151,262,802,828]" pageId="1" pageNumber="88" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="0D154E06662402215425FCCB34385D44" box="[274,318,801,827]" pageId="1" pageNumber="88">The</collectingCounty>
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<typeStatus id="3B708828662402215472FCCB34AC5D44" box="[325,426,801,827]" pageId="1" pageNumber="88" type="holotype">holotype</typeStatus>
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, a male, is in the
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<collectingMunicipality id="0410ACF0662402215747FCCB31245D44" box="[624,1058,801,827]" pageId="1" pageNumber="88">Florida State Collection of Arthropods</collectingMunicipality>
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(
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<collectionCode id="82DAAE4F662402215106FCCB317B5D44" box="[1073,1149,801,827]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="88" type="Museum">FSCA</collectionCode>
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), Gainesville, FL (
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<figureCitation id="7CF02A0F662402215079FCCB30935D44" box="[1358,1429,801,827]" captionStart="FIGURE 1" captionStartId="2.[151,250,1837,1862]" captionTargetBox="[151,1436,342,1809]" captionTargetId="figure@2.[151,1436,342,1809]" captionTargetPageId="2" captionText="FIGURE 1. Ungla pseudomeleoma, sp. nov., Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel." figureDoi="http://doi.org/10.5281/zenodo.3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="1" pageNumber="88">Figs 1</figureCitation>
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,
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<figureCitation id="7CF02A0F6624022155A0FCAF35A05D20" box="[151,166,837,863]" captionStart="FIGURE 3" captionStartId="4.[151,250,1610,1635]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure@4.[151,1436,181,1585]" captionTargetPageId="4" captionText="FIGURE 3. Ungla pseudomeleoma, sp. nov., Exterior head and body (male, Peru: La Libertad). A. Holotype, head (ventral, FSCA); B. Paratype, head, prothorax (dorsal, TRC); C. Holotype, body (lateral, FSCA); D. Holotype, head and thorax (lateral, FSCA); E. Holotype, abdomen (lateral, FSCA); F. Holotype, terminus (lateral, FSCA). c.c., callus cerci; S4, fourth sternite; S8+9, fused eighth and ninth sternites, sc, scape; sp, spiracular openings." figureDoi="http://doi.org/10.5281/zenodo.3545364" httpUri="https://zenodo.org/record/3545364/files/figure.png" pageId="1" pageNumber="88">3</figureCitation>
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,
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<figureCitation id="7CF02A0F662402215583FCAF35C25D1F" box="[180,196,837,864]" captionStart="FIGURE 5" captionStartId="6.[151,250,1715,1740]" captionTargetBox="[151,1436,237,1690]" captionTargetId="figure@6.[151,1436,237,1690]" captionTargetPageId="6" captionText="FIGURE 5. Ungla pseudomeleoma, sp. nov., Holotype, cleared abdomen (male, Peru: La Libertad, FSCA). A. Abdomen (lateral); B. Abdominal terminus (lateral); C. Terminal abdominal segments and genitalia (lateral); D. Fused ninth tergite and ectoproct (dorsal); E. Fused eighth and ninth sternites (ventral, suture on dorsal integument between T8 and T9 visible). c.c., callus cerci; g.ap., gonarcal apodeme; g.br., gonarcal bridge; inv, invaginated dorsal cleft in T9+e, above anus; S3, third sternite; S8+9, fused eighth and ninth sternites, s.p., setose subanal plate; sp, spiracular opening; T4, T6, T8, T9, fourth, sixth, eighth, ninth tergites; T9+e, fused ninth tergite and ectoproct." figureDoi="http://doi.org/10.5281/zenodo.3545368" httpUri="https://zenodo.org/record/3545368/files/figure.png" pageId="1" pageNumber="88">5</figureCitation>
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,
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<figureCitation id="7CF02A0F6624022155E6FCAF35E75D1F" box="[209,225,837,864]" captionStart="FIGURE 6" captionStartId="7.[151,250,1754,1779]" captionTargetBox="[151,1436,273,1727]" captionTargetId="figure@7.[151,1436,273,1729]" captionTargetPageId="7" captionText="FIGURE 6. Ungla pseudomeleoma, sp. nov., Holotype, genitalia (male, Peru: La Libertad, FSCA). A. Gonarcal complex (anterolateral); B. Gonarcus (posterodorsal); C. Gonarcal complex (frontal); D. gonarcus (caudal); E. Gonarcal complex (frontoventral). c, comes; gsac, gonosaccus; g.ap., gonarcal apodeme; g.br., gonarcal bridge; h.i., hypandrium internum; mu, mediuncus; pr, unarticulated process on frontal margin of gonarcal apodeme; s.p., setose subanal plate; s.pl., paired setose plate of gonosaccus." figureDoi="http://doi.org/10.5281/zenodo.3545370" httpUri="https://zenodo.org/record/3545370/files/figure.png" pageId="1" pageNumber="88">6</figureCitation>
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)
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</materialsCitation>
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.
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<materialsCitation id="54A33CD76624022155C0FCAC37E55DFC" ID-GBIF-Occurrence="2556249867" collectingDate="1975-07-12" country="Peru" latitude="-7.9833336" longitude="-78.683334" pageId="1" pageNumber="88" stateProvince="La Libertad">
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<location id="E11460516624022155C0FCAC34155D1F" LSID="urn:lsid:plazi:treatment:6C62879C6624022955A0FD33300C5990:E11460516624022155C0FCAC34155D1F" box="[247,275,838,864]" country="Peru" county="The" latitude="-7.9833336" longLatPrecision="1302" longitude="-78.683334" municipality="Florida State Collection of Arthropods" name="Its" pageId="1" pageNumber="88" stateProvince="La Libertad">Its</location>
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labels read: [1] “
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<collectingCountry id="9CDC761A6624022154E1FCAF37225D20" box="[470,548,837,863]" name="Peru" pageId="1" pageNumber="88">PERU</collectingCountry>
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: Samme, ca. /
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<locationDeviation id="F5C62FA86624022157FEFCAF369C5D20" box="[713,922,837,864]" location="Trujillo" pageId="1" pageNumber="88">
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<locationDeviation id="F5C62FA86624022157FEFCAF36175D20" box="[713,785,837,863]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="88" unit="km" value="40.0">40 km</locationDeviation>
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</quantity>
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||
NE Trujillo
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</locationDeviation>
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/ Prov.
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<collectingRegion id="260FF8686624022156C5FCAC31715D20" box="[1010,1143,837,864]" country="Peru" name="La Libertad" pageId="1" pageNumber="88">La Libertad</collectingRegion>
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/
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<geoCoordinate id="81FF504D6624022151B9FCAF31D85D1F" box="[1166,1246,837,864]" degrees="7" direction="south" minutes="59" orientation="latitude" pageId="1" pageNumber="88" precision="925" value="-7.9833336">7°59’S</geoCoordinate>
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,
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<geoCoordinate id="81FF504D6624022151DCFCAF30575D20" box="[1259,1361,837,863]" degrees="78" direction="west" minutes="41" orientation="longitude" pageId="1" pageNumber="88" precision="925" value="-78.683334">78°41’W</geoCoordinate>
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/ elev. ca.
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||
<quantity id="23339B6F66240221558AFC8334125DFB" box="[189,276,873,900]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="88" unit="m" value="1500.0">
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<elevation id="6FE6D1B966240221558AFC8334125DFB" box="[189,276,873,900]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="88" unit="m" value="1500.0">1500 m</elevation>
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</quantity>
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. /
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<date id="9075104A66240221541BFC8334E35DFB" box="[300,485,873,900]" pageId="1" pageNumber="88" value="1975-07-12" valueMax="1975-07-17" valueMin="1975-07-12">
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<collectingDate id="8031E9A266240221541BFC8334E35DFB" box="[300,485,873,900]" pageId="1" pageNumber="88" value="1975-07-12" valueMax="1975-07-17" valueMin="1975-07-12">12–17 July 1975</collectingDate>
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</date>
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||
/
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<collectorName id="493E535C6624022154CEFC8337595DFB" box="[505,607,873,900]" pageId="1" pageNumber="88">C. Porter</collectorName>
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,
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<collectorName id="493E535C66240221575EFC8037DF5DFC" box="[617,729,873,900]" pageId="1" pageNumber="88">L. Stange</collectorName>
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”
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="54A33CD76624022157D9FC83309C5DD8" ID-GBIF-Occurrence="2556249865" country="Peru" pageId="1" pageNumber="88" stateProvince="La Libertad" typeStatus="holotype">
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[2] “
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<typeStatus id="3B708828662402215629FC8336B35DFC" box="[798,949,873,899]" pageId="1" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="23CB4D096624022156FEFC8331CF5DFC" authorityName="Tauber" authorityYear="2019" box="[969,1225,873,899]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudomeleoma">
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<emphasis id="D6BFEA986624022156FEFC8331CF5DFC" box="[969,1225,873,899]" italics="true" pageId="1" pageNumber="88">Ungla pseudomeleoma</emphasis>
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</taxonomicName>
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||
/ des.
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<collectorName id="493E535C662402215027FC83309A5DFC" box="[1296,1436,873,900]" pageId="1" pageNumber="88">C. A. Tauber</collectorName>
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2019”. The specimen is in good condition; the right labial palpus is partially covered with a white, frothy material.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="ACD165016624022155C0FC58343D5874" pageId="1" pageNumber="88" type="etymology">
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||
<paragraph id="E474368A6624022155C0FC58317D5D8F" blockId="1.[151,1437,801,2016]" pageId="1" pageNumber="88">
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||
<emphasis id="D6BFEA986624022155C0FC5834785DB3" bold="true" box="[247,382,946,972]" pageId="1" pageNumber="88">Etymology.</emphasis>
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The species name
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<taxonomicName id="23CB4D09662402215778FC5B36055DB4" authorityName="Tauber" authorityYear="2019" box="[591,771,945,971]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudomeleoma">
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<emphasis id="D6BFEA98662402215778FC5B36055DB4" box="[591,771,945,971]" italics="true" pageId="1" pageNumber="88">pseudomeleoma</emphasis>
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</taxonomicName>
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recognizes the elaborate
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<taxonomicName id="23CB4D0966240221512BFC5B31825DB4" authorityName="Fitch" authorityYear="1855" box="[1052,1156,945,971]" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="D6BFEA9866240221512BFC5B31825DB4" box="[1052,1156,945,971]" italics="true" pageId="1" pageNumber="88">Meleoma</emphasis>
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</taxonomicName>
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-like antennal and frontal modifications expressed by this species. It is a noun in apposition with the generic name.
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</paragraph>
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<paragraph id="E474368A6624022155F0FC13343D5874" blockId="1.[151,1437,801,2016]" pageId="1" pageNumber="88">
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<emphasis id="D6BFEA986624022155F0FC13373A5A6B" bold="true" box="[199,572,1017,1044]" pageId="1" pageNumber="88">Support for generic placement.</emphasis>
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In addition to the crucial genital characters described below, this species expresses several externally visible features that support its placement in the genus
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<taxonomicName id="23CB4D09662402215124FBF7315F5A48" authorityName="Navas" authorityYear="1914" box="[1043,1113,1053,1079]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="D6BFEA98662402215124FBF7315F5A48" box="[1043,1113,1053,1079]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
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</taxonomicName>
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(c.f.,
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<bibRefCitation id="805A4B7B6624022151AFFBF730625A48" author="Tauber, C. A. & Sosa, F. & Albuquerque, G. S. & Tauber, M. J." box="[1176,1380,1053,1079]" pageId="1" pageNumber="88" pagination="1 - 188" refId="ref7000" refString="Tauber, C. A., Sosa, F., Albuquerque, G. S. & Tauber, M. J. (2017) Revision of the Neotropical green lacewing genus Ungla (Neuroptera, Chrysopidae). ZooKeys, 674, 1 - 188. https: // doi. org / 10.3897 / zookeys. 674.11435" type="journal article" year="2017">
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Tauber
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<emphasis id="D6BFEA986624022151DCFBF430245A48" box="[1259,1314,1053,1079]" italics="true" pageId="1" pageNumber="88">et al.</emphasis>
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2017
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</bibRefCitation>
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). For both males and females, the wing venation—especially the insertion of the basal inner gradate on a radial vein and not on the Psm—is more typical of
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<taxonomicName id="23CB4D09662402215710FB8F376B5A00" authorityName="Navas" authorityYear="1914" box="[551,621,1125,1151]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662402215710FB8F376B5A00" box="[551,621,1125,1151]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
|
||
</taxonomicName>
|
||
species than those in most other chrysopine genera, including
|
||
<taxonomicName id="23CB4D0966240221501AFB8F30935A00" authorityName="Fitch" authorityYear="1855" box="[1325,1429,1125,1151]" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA9866240221501AFB8F30935A00" box="[1325,1429,1125,1151]" italics="true" pageId="1" pageNumber="88">Meleoma</emphasis>
|
||
</taxonomicName>
|
||
. In addition, males have enlarged abdominal spiracles; T9+ect is truncated so that it and the dorsal apodeme do not extend proximally below T8; T9+ect has a deep anal invagination; sternites 8 and 9 are fully fused, and S8+9 is elongate, plate-like, and terminates broadly with distal margin upturned. All of the above features characterize males of
|
||
<taxonomicName id="23CB4D096624022155A0FB1F35DB5B70" authorityName="Navas" authorityYear="1914" box="[151,221,1269,1295]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA986624022155A0FB1F35DB5B70" box="[151,221,1269,1295]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
|
||
</taxonomicName>
|
||
species. Internally, the structure and composition of the male genitalia is typical of all
|
||
<taxonomicName id="23CB4D09662402215192FB1F31ED5B70" authorityName="Navas" authorityYear="1914" box="[1189,1259,1269,1295]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662402215192FB1F31ED5B70" box="[1189,1259,1269,1295]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
|
||
</taxonomicName>
|
||
species: simple gonarcus without gonocornua, with an unarticulated frontal process on the lateral apodeme; mediuncus narrow, elongate, attached to gonarcal bridge via a flexible hinge; gonosaccus with two patches of robust setae on opposing robust plates; and the absence of a tignum, gonapsis, pseudopenis, gonocristae, and parameres. Female genital characters also are consistent with the genus
|
||
<taxonomicName id="23CB4D096624022157A1FA6F37DA5BE0" authorityName="Navas" authorityYear="1914" box="[662,732,1413,1439]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA986624022157A1FA6F37DA5BE0" box="[662,732,1413,1439]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
|
||
</taxonomicName>
|
||
: spermatheca round, pillbox-shaped, with tall vellum opening directly to the bursa copulatrix via a dorsal slit; spermathecal duct with two to three curves, extending posteriorly, with a distal filamentous section; subgenitale bilobed distally, with simple pocket or lobe below; and the absence of a praegenitale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ACD165016624022155F0F9FF35CE5978" pageId="1" pageNumber="88" type="diagnosis">
|
||
<paragraph id="E474368A6624022155F0F9FF35CE5978" blockId="1.[151,1437,801,2016]" pageId="1" pageNumber="88">
|
||
<emphasis id="D6BFEA986624022155F0F9FF343C584F" bold="true" box="[199,314,1557,1584]" pageId="1" pageNumber="88">Diagnosis</emphasis>
|
||
. Both males and females of
|
||
<taxonomicName id="23CB4D0966240221574BF9FC36555850" authorityName="Tauber" authorityYear="2019" box="[636,851,1557,1583]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudomeleoma">
|
||
<emphasis id="D6BFEA9866240221574BF9FC36555850" box="[636,851,1557,1583]" italics="true" pageId="1" pageNumber="88">U. pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
are distinguished from other
|
||
<taxonomicName id="23CB4D09662402215197F9FF31E05850" authorityName="Navas" authorityYear="1914" box="[1184,1254,1557,1583]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662402215197F9FF31E05850" box="[1184,1254,1557,1583]" italics="true" pageId="1" pageNumber="88">Ungla</emphasis>
|
||
</taxonomicName>
|
||
species by their elongate scapes with longitudinal red marks on the dorsum and by the distinctive red lines around their antennal fossae (
|
||
<figureCitation id="7CF02A0F6624022155DBF9B734C95808" box="[236,463,1629,1655]" captionStart="FIGURE 1" captionStartId="2.[151,250,1837,1862]" captionTargetBox="[151,1436,342,1809]" captionTargetId="figure@2.[151,1436,342,1809]" captionTargetPageId="2" captionText="FIGURE 1. Ungla pseudomeleoma, sp. nov., Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel." figureDoi="http://doi.org/10.5281/zenodo.3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="1" pageNumber="88">Figs 1B, 1D, 1E, 1F</figureCitation>
|
||
,
|
||
<figureCitation id="7CF02A0F6624022154EBF9B737595808" box="[476,607,1629,1655]" captionStart="FIGURE 2" captionStartId="3.[151,250,1946,1971]" captionTargetBox="[151,1436,561,1921]" captionTargetId="figure@3.[151,1436,561,1921]" captionTargetPageId="3" captionText="FIGURE 2. Ungla pseudomeleoma, sp. nov., Paratypes (female, Peru: La Libertad).A. Head, frontal (FSCA); B. Head, prothorax (dorsal, FSCA); C. Head, prothorax, mesothorax (dorsal, TRC); D.Antenna (FSCA). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; ped, pedicel; sc, scape; tor, torulus.." figureDoi="http://doi.org/10.5281/zenodo.3545362" httpUri="https://zenodo.org/record/3545362/files/figure.png" pageId="1" pageNumber="88">2A, 2B, 2C</figureCitation>
|
||
). Males also are easily recognized by the unique shapes and setal patterns of the scape and basal flagellar segments (
|
||
<figureCitation id="7CF02A0F662402215758F96B37B358E4" box="[623,693,1665,1691]" captionStart="FIGURE 1" captionStartId="2.[151,250,1837,1862]" captionTargetBox="[151,1436,342,1809]" captionTargetId="figure@2.[151,1436,342,1809]" captionTargetPageId="2" captionText="FIGURE 1. Ungla pseudomeleoma, sp. nov., Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel." figureDoi="http://doi.org/10.5281/zenodo.3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="1" pageNumber="88">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="7CF02A0F6624022157F4F96B37D458E4" box="[707,722,1665,1691]" captionStart="FIGURE 3" captionStartId="4.[151,250,1610,1635]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure@4.[151,1436,181,1585]" captionTargetPageId="4" captionText="FIGURE 3. Ungla pseudomeleoma, sp. nov., Exterior head and body (male, Peru: La Libertad). A. Holotype, head (ventral, FSCA); B. Paratype, head, prothorax (dorsal, TRC); C. Holotype, body (lateral, FSCA); D. Holotype, head and thorax (lateral, FSCA); E. Holotype, abdomen (lateral, FSCA); F. Holotype, terminus (lateral, FSCA). c.c., callus cerci; S4, fourth sternite; S8+9, fused eighth and ninth sternites, sc, scape; sp, spiracular openings." figureDoi="http://doi.org/10.5281/zenodo.3545364" httpUri="https://zenodo.org/record/3545364/files/figure.png" pageId="1" pageNumber="88">3</figureCitation>
|
||
), the deeply ridged interantennal surface (
|
||
<figureCitation id="7CF02A0F66240221519CF96B305858E4" box="[1195,1374,1665,1691]" captionStart="FIGURE 1" captionStartId="2.[151,250,1837,1862]" captionTargetBox="[151,1436,342,1809]" captionTargetId="figure@2.[151,1436,342,1809]" captionTargetPageId="2" captionText="FIGURE 1. Ungla pseudomeleoma, sp. nov., Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel." figureDoi="http://doi.org/10.5281/zenodo.3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="1" pageNumber="88">Figs 1C, 1E, 1F</figureCitation>
|
||
), and the highly ornamented frons (
|
||
<figureCitation id="7CF02A0F6624022154D3F94F372D58C0" box="[484,555,1701,1727]" captionStart="FIGURE 1" captionStartId="2.[151,250,1837,1862]" captionTargetBox="[151,1436,342,1809]" captionTargetId="figure@2.[151,1436,342,1809]" captionTargetPageId="2" captionText="FIGURE 1. Ungla pseudomeleoma, sp. nov., Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel." figureDoi="http://doi.org/10.5281/zenodo.3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="1" pageNumber="88">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="7CF02A0F66240221570EF94F374E58C0" box="[569,584,1701,1727]" captionStart="FIGURE 3" captionStartId="4.[151,250,1610,1635]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure@4.[151,1436,181,1585]" captionTargetPageId="4" captionText="FIGURE 3. Ungla pseudomeleoma, sp. nov., Exterior head and body (male, Peru: La Libertad). A. Holotype, head (ventral, FSCA); B. Paratype, head, prothorax (dorsal, TRC); C. Holotype, body (lateral, FSCA); D. Holotype, head and thorax (lateral, FSCA); E. Holotype, abdomen (lateral, FSCA); F. Holotype, terminus (lateral, FSCA). c.c., callus cerci; S4, fourth sternite; S8+9, fused eighth and ninth sternites, sc, scape; sp, spiracular openings." figureDoi="http://doi.org/10.5281/zenodo.3545364" httpUri="https://zenodo.org/record/3545364/files/figure.png" pageId="1" pageNumber="88">3</figureCitation>
|
||
,
|
||
<figureCitation id="7CF02A0F662402215762F94F376358BF" box="[597,613,1701,1728]" captionStart="FIGURE 5" captionStartId="6.[151,250,1715,1740]" captionTargetBox="[151,1436,237,1690]" captionTargetId="figure@6.[151,1436,237,1690]" captionTargetPageId="6" captionText="FIGURE 5. Ungla pseudomeleoma, sp. nov., Holotype, cleared abdomen (male, Peru: La Libertad, FSCA). A. Abdomen (lateral); B. Abdominal terminus (lateral); C. Terminal abdominal segments and genitalia (lateral); D. Fused ninth tergite and ectoproct (dorsal); E. Fused eighth and ninth sternites (ventral, suture on dorsal integument between T8 and T9 visible). c.c., callus cerci; g.ap., gonarcal apodeme; g.br., gonarcal bridge; inv, invaginated dorsal cleft in T9+e, above anus; S3, third sternite; S8+9, fused eighth and ninth sternites, s.p., setose subanal plate; sp, spiracular opening; T4, T6, T8, T9, fourth, sixth, eighth, ninth tergites; T9+e, fused ninth tergite and ectoproct." figureDoi="http://doi.org/10.5281/zenodo.3545368" httpUri="https://zenodo.org/record/3545368/files/figure.png" pageId="1" pageNumber="88">5</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="23CB4D096624022157C9F94C36D358C0" authorityName="Tauber" authorityYear="2019" box="[766,981,1701,1727]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="pseudomeleoma">
|
||
<emphasis id="D6BFEA986624022157C9F94C36D358C0" box="[766,981,1701,1727]" italics="true" pageId="1" pageNumber="88">U. pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
do not exhibit the extreme male modifications; however, their elongated scapes and the paired ridges between the scapes are distinctive features (
|
||
<figureCitation id="7CF02A0F66240221500DF92335BD5978" captionStart="FIGURE 2" captionStartId="3.[151,250,1946,1971]" captionTargetBox="[151,1436,561,1921]" captionTargetId="figure@3.[151,1436,561,1921]" captionTargetPageId="3" captionText="FIGURE 2. Ungla pseudomeleoma, sp. nov., Paratypes (female, Peru: La Libertad).A. Head, frontal (FSCA); B. Head, prothorax (dorsal, FSCA); C. Head, prothorax, mesothorax (dorsal, TRC); D.Antenna (FSCA). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; ped, pedicel; sc, scape; tor, torulus.." figureDoi="http://doi.org/10.5281/zenodo.3545362" httpUri="https://zenodo.org/record/3545362/files/figure.png" pageId="1" pageNumber="88">Figs 2A, 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ACD165016624022955F0F8FB300C5990" lastPageId="9" lastPageNumber="96" pageId="1" pageNumber="88" type="description">
|
||
<paragraph id="E474368A6624022255F0F8FB348B5F3C" blockId="1.[151,1437,801,2016]" lastBlockId="2.[151,1436,153,323]" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
|
||
<emphasis id="D6BFEA986624022155F0F8FB345F5953" bold="true" box="[199,345,1809,1836]" pageId="1" pageNumber="88">Description.</emphasis>
|
||
<emphasis id="D6BFEA98662402215456F8FB34A15954" box="[353,423,1809,1835]" italics="true" pageId="1" pageNumber="88">Head:</emphasis>
|
||
Width
|
||
<quantity id="23339B6F6624022154C9F8FB37505953" box="[510,598,1809,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="88" unit="mm" value="1.4">1.4 mm</quantity>
|
||
(female),
|
||
<quantity id="23339B6F6624022157FAF8FB36575953" box="[717,849,1809,1836]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="88" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
|
||
(male); ratio of head width: eye width, 2.4-2.6: 1 (female), 1.8–1.9: 1 (male). Male vertex, frons, antennal base (scape, pedicel, basal flagellar segments) with elaborate modification. Vertex with scape surrounded by large, delineated fossae, bases of scapes located close to each other, within much less than width of scape; surface of vertex between toruli, scapes longitudinally furrowed; upper surface of frons also furrowed anteriorly, bending forward, forming interantennal horn consisting of connected longitudinal ridges that merge into a medial, bifid ledge with setose lower surface. Frons invaginated below inter- antennal horn and scapes, forming distinct, rounded, shallow cavity; lateral edges of cavity extending towards eyes; lower part of frons with two elevated, rounded ridges, each bearing pair of triangular patches of dense, elongate, dark brown setae curved upward, appearing to be a lower horn. Female face only lightly sculptured, mostly smooth, without elaborate modifications, except vertex with pair of distinct, longitudinal ridges between scapes, extending to upper part of frons.
|
||
</paragraph>
|
||
<caption id="B0B466026627022255A0F8C7314759F1" ID-DOI="http://doi.org/10.5281/zenodo.3545360" ID-Zenodo-Dep="3545360" httpUri="https://zenodo.org/record/3545360/files/figure.png" pageId="2" pageNumber="89" startId="2.[151,250,1837,1862]" targetBox="[151,1436,342,1809]" targetPageId="2">
|
||
<paragraph id="E474368A6627022255A0F8C7314759F1" blockId="2.[151,1437,1837,1934]" pageId="2" pageNumber="89">
|
||
<emphasis id="D6BFEA986627022255A0F8C734125939" bold="true" box="[151,276,1837,1862]" pageId="2" pageNumber="89">FIGURE 1.</emphasis>
|
||
<taxonomicName id="23CB4D0966270222542BF8C437005939" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[284,518,1838,1862]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA9866270222542BF8C437005939" box="[284,518,1838,1862]" italics="true" pageId="2" pageNumber="89">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662702225723F8C437605939" bold="true" box="[532,614,1838,1862]" pageId="2" pageNumber="89">
|
||
<taxonomicNameLabel id="CD8C57E3662702225723F8C437605939" box="[532,614,1838,1862]" pageId="2" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype (male, Peru: La Libertad, FSCA). A. Head, prothorax (frontolateral); B. Head, prothorax (lateral); C. Head (frontal); D. Head (lateral); E. Face (frontal); F. Head, prothorax (dorsal). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; lo.h., lower horn; ped, pedicel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A6627022255F0F857309A59A8" blockId="2.[151,1436,1979,2043]" box="[199,1436,1979,2007]" pageId="2" pageNumber="89">
|
||
<emphasis id="D6BFEA986627022255F0F857342859A8" box="[199,302,1981,2007]" italics="true" pageId="2" pageNumber="89">Antenna:</emphasis>
|
||
Length 12.0–
|
||
<quantity id="23339B6F6627022254FDF856373659A8" box="[458,560,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="2" pageNumber="89" unit="mm" value="12.4">12.4 mm</quantity>
|
||
. Scape slender, elongate (ratio of length: width, 1.7–1.8: 1 (female), 2.3–2.5:
|
||
</paragraph>
|
||
<paragraph id="E474368A6627022355A0F80A368B5E84" blockId="2.[151,1436,1979,2043]" lastBlockId="3.[151,1437,153,539]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
|
||
1 (male); few short setae dorsally, more numerous, long setae ventrally; sides straight (male), slightly curved later- ally (female). Pedicel (male, female), basal flagellar segment (female) without distinct modification; basal flagellar segment (male,
|
||
<figureCitation id="7CF02A0F66260223547BFF5734A35EA8" box="[332,421,189,215]" captionStart="FIGURE 8" captionStartId="9.[151,250,997,1022]" captionTargetBox="[213,1375,181,971]" captionTargetId="figure@9.[213,1375,181,971]" captionTargetPageId="9" captionText="FIGURE 8. Similarity in the antennal features of Ungla pseudomeleoma with those of three species of Meleoma. A. Ungla pseudomeleoma, sp. nov., Holotype (Peru: La Libertad, FSCA); B. Meleoma emuncta Fitch (USA: California, TRC); C. Meleoma undescribed sp. #1 (USA:Arizona, TRC); D. Meleoma stangei Penny (Mexico: Baja California, San Diego County Museum of Natural History). f1, first flagellomere; h, horn extending from scape; ped, pedicel; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.3545374" httpUri="https://zenodo.org/record/3545374/files/figure.png" pageId="3" pageNumber="90">Fig. 8A</figureCitation>
|
||
) enlarged, bulbous, highly setose frontally, extending posteriorly from pedicel; second to fourth flagellar segments tapering, with setae of decreasing lengths.
|
||
</paragraph>
|
||
<paragraph id="E474368A6626022355F0FEEF31BF5C64" blockId="3.[151,1437,153,539]" pageId="3" pageNumber="90">
|
||
<emphasis id="D6BFEA986626022355F0FEEF34A15F60" box="[199,423,261,287]" italics="true" pageId="3" pageNumber="90">Coloration of head:</emphasis>
|
||
Head cream-colored (probably green in life) with dark to light red markings; vertex with pair of distinct red spots behind anterior ridge, red margin around base of torulus, dorsal fossa; female with thin red line along anterior fossa, frons with red mark along frontoclypeal margin; male without frontal or clypeal marks; gena with bright red to dark reddish black lateral stripe almost reaching eye posteriorly, extending distally to clypeal margin; clypeus with small or large black stripe laterally. Antenna with scape cream-colored, with irregular, longitudinal red stripe; pedicel, flagellum cream-colored; maxillary palpus with distal two segments dark brown, middle segment light brown, two basal segments pale, articulations pale; labial palpus with basal two segments pale, distal segment dark brown, articulation pale. Corner of cervix with small, dark brown to black mark.
|
||
</paragraph>
|
||
<caption id="B0B466026626022355A0F870367F5984" ID-DOI="http://doi.org/10.5281/zenodo.3545362" ID-Zenodo-Dep="3545362" httpUri="https://zenodo.org/record/3545362/files/figure.png" pageId="3" pageNumber="90" startId="3.[151,250,1946,1971]" targetBox="[151,1436,561,1921]" targetPageId="3">
|
||
<paragraph id="E474368A6626022355A0F870367F5984" blockId="3.[151,1437,1946,2043]" pageId="3" pageNumber="90">
|
||
<emphasis id="D6BFEA986626022355A0F870341459CC" bold="true" box="[151,274,1946,1971]" pageId="3" pageNumber="90">FIGURE 2.</emphasis>
|
||
<taxonomicName id="23CB4D0966260223542FF870370659CD" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[280,512,1946,1970]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA9866260223542FF870370659CD" box="[280,512,1946,1970]" italics="true" pageId="3" pageNumber="90">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA9866260223573BF871375A59CC" bold="true" box="[524,604,1947,1971]" pageId="3" pageNumber="90">
|
||
<taxonomicNameLabel id="CD8C57E366260223573BF871375A59CC" box="[524,604,1947,1971]" pageId="3" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Paratypes (female, Peru: La Libertad). A. Head, frontal (FSCA); B. Head, prothorax (dorsal, FSCA); C. Head, prothorax, mesothorax (dorsal, TRC); D. Antenna (FSCA). f1, first flagellomere; in.fu., interantennal furrow; in.h., interantennal horn; ped, pedicel; sc, scape; tor, torulus..
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B0B466026621022455A0F9A0367C58B0" ID-DOI="http://doi.org/10.5281/zenodo.3545364" ID-Zenodo-Dep="3545364" httpUri="https://zenodo.org/record/3545364/files/figure.png" pageId="4" pageNumber="91" startId="4.[151,250,1610,1635]" targetBox="[151,1436,181,1585]" targetPageId="4">
|
||
<paragraph id="E474368A6621022455A0F9A0367C58B0" blockId="4.[151,1436,1610,1743]" pageId="4" pageNumber="91">
|
||
<emphasis id="D6BFEA986621022455A0F9A03412581C" bold="true" box="[151,276,1610,1635]" pageId="4" pageNumber="91">FIGURE 3.</emphasis>
|
||
<taxonomicName id="23CB4D0966210224542BF9A13701581C" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[284,519,1611,1635]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA9866210224542BF9A13701581C" box="[284,519,1611,1635]" italics="true" pageId="4" pageNumber="91">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662102245722F9A13761581C" bold="true" box="[533,615,1611,1635]" pageId="4" pageNumber="91">
|
||
<taxonomicNameLabel id="CD8C57E3662102245722F9A13761581C" box="[533,615,1611,1635]" pageId="4" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Exterior head and body (male, Peru: La Libertad). A. Holotype, head (ventral, FSCA); B. Paratype, head, prothorax (dorsal, TRC); C. Holotype, body (lateral, FSCA); D. Holotype, head and thorax (lateral, FSCA); E. Holotype, abdomen (lateral, FSCA); F. Holotype, terminus (lateral, FSCA). c.c., callus cerci; S4, fourth sternite; S8+9, fused eighth and ninth sternites, sc, scape; sp, spiracular openings.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A6621022455F0F917346759FC" blockId="4.[151,1437,1789,2032]" pageId="4" pageNumber="91">
|
||
<emphasis id="D6BFEA986621022455F0F91734275968" box="[199,289,1789,1815]" italics="true" pageId="4" pageNumber="91">Thorax:</emphasis>
|
||
Prothorax length
|
||
<quantity id="23339B6F6621022454C6F91737715967" box="[497,631,1789,1816]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="91" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6-0.8 mm</quantity>
|
||
, width
|
||
<quantity id="23339B6F6621022457F9F91736535967" box="[718,853,1789,1816]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="4" pageNumber="91" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9-1.1 mm</quantity>
|
||
; pronotum with furrow down midline, longitudinal furrows across midsection and near posterior margin; diffused markings in irregular pattern along lateral surface, broad cream-colored band mesally; surface with numerous elongate, cream setae. Mesothorax, metathorax without distinct markings.
|
||
</paragraph>
|
||
<paragraph id="E474368A6621022555F0F86437B25C64" blockId="4.[151,1437,1789,2032]" lastBlockId="5.[151,1437,153,539]" lastPageId="5" lastPageNumber="92" pageId="4" pageNumber="91">
|
||
<emphasis id="D6BFEA986621022455F0F864341359D8" box="[199,277,1934,1959]" italics="true" pageId="4" pageNumber="91">Wings:</emphasis>
|
||
Forewing (
|
||
<figureCitation id="7CF02A0F6621022454AEF86734F759D8" box="[409,497,1933,1959]" captionStart="FIGURE 4" captionStartId="5.[151,250,1537,1562]" captionTargetBox="[151,1436,561,1512]" captionTargetId="figure@5.[151,1436,561,1512]" captionTargetPageId="5" captionText="FIGURE 4. Ungla pseudomeleoma, sp. nov., Paratype (female, Peru: La Libertad, TRC). Wings. A. Forewing B. Hindwing. b2, b3, second, third upper Banksian cells (cells below RP that do not include a gradate vein); b’2, b’4, second, fourth lower Banksian cells (cells between Psm and Psc; gc3, third gradate cell (cell with inner and outer gradates at the two ends); icu1, first intracubital cell; 1ig, first inner gradate vein, im1, first intramedian cell; Psc, Pseudocubitus; Psm, Pseudomedius; rarp1, rarp9, first, ninth closed cells between RA and RP; RA, anterior branch of Radius; RP, posterior branch of Radius." figureDoi="http://doi.org/10.5281/zenodo.3545366" httpUri="https://zenodo.org/record/3545366/files/figure.png" pageId="4" pageNumber="91">Fig. 4A</figureCitation>
|
||
) slender, apex broadly subacute to rounded, membrane clear, hyaline, with light to very light suffusion of brown surrounding gradate veins; stigma very slightly opaque; longitudinal veins green, marked brown at intersections with other veins; transverse veins brown or marked brown at ends; forewing with veins robust to slender, not crassate; RP straight; first intramedian cell ovate; basal inner gradate vein not meeting Psm; gradate veins dark brown, costal crossveins, ra-rp crossveins, intracubital crossveins brown or marked brown basally. Hindwing (
|
||
<figureCitation id="7CF02A0F662002255441FF0B34C95E84" box="[374,463,225,251]" captionStart="FIGURE 4" captionStartId="5.[151,250,1537,1562]" captionTargetBox="[151,1436,561,1512]" captionTargetId="figure@5.[151,1436,561,1512]" captionTargetPageId="5" captionText="FIGURE 4. Ungla pseudomeleoma, sp. nov., Paratype (female, Peru: La Libertad, TRC). Wings. A. Forewing B. Hindwing. b2, b3, second, third upper Banksian cells (cells below RP that do not include a gradate vein); b’2, b’4, second, fourth lower Banksian cells (cells between Psm and Psc; gc3, third gradate cell (cell with inner and outer gradates at the two ends); icu1, first intracubital cell; 1ig, first inner gradate vein, im1, first intramedian cell; Psc, Pseudocubitus; Psm, Pseudomedius; rarp1, rarp9, first, ninth closed cells between RA and RP; RA, anterior branch of Radius; RP, posterior branch of Radius." figureDoi="http://doi.org/10.5281/zenodo.3545366" httpUri="https://zenodo.org/record/3545366/files/figure.png" pageId="5" pageNumber="92">Fig. 4B</figureCitation>
|
||
) narrow, apex subacute, membrane clear, hyaline, without markings; venation green, except ra-rp crossveins, intersections with RP brown at least basally. Forewing length
|
||
<quantity id="23339B6F662002255167FEEF31E95F60" box="[1104,1263,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.34" metricValueMin="1.16" pageId="5" pageNumber="92" unit="mm" value="12.5" valueMax="13.4" valueMin="11.6">11.6-13.4 mm</quantity>
|
||
, height 3.7-4.4; ratio of length to height, 3.0-3.1: 1; height of tallest costal cell
|
||
<quantity id="23339B6F66200225566BFEC336D95F3C" box="[860,991,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="5" pageNumber="92" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8-0.9 mm</quantity>
|
||
(cell number 5); width of first intramedian cell
|
||
<quantity id="23339B6F6620022555CBFEA734865F18" box="[252,384,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="5" pageNumber="92" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7-0.8 mm</quantity>
|
||
; 9-10
|
||
<emphasis id="D6BFEA986620022554F3FEA434F05F18" box="[452,502,334,359]" italics="true" pageId="5" pageNumber="92">rarp</emphasis>
|
||
cells (closed cells between RA and RP); third gradate cell (third cell from wing base with inner and outer gradate veins at either end)
|
||
<quantity id="23339B6F662002255781FE9B363E5FF4" box="[694,824,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="5" pageNumber="92" unit="mm" value="1.05" valueMax="1.3" valueMin="0.8">0.8-1.3 mm</quantity>
|
||
long,
|
||
<quantity id="23339B6F662002255649FE9A31065FF4" box="[894,1024,368,395]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="92" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4-0.5 mm</quantity>
|
||
wide, ratio of length: width, 2.0-2.7: 1); fourth gradate cell 1.0 mm long,
|
||
<quantity id="23339B6F662002255776FE7F379E5FD0" box="[577,664,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="92" unit="mm" value="0.4">0.4 mm</quantity>
|
||
wide (ratio of length: width, 2.4: 1); 4 b cells (closed cells beneath RP that do not include a gradate vein), 4 b’ cells (closed cells between Psm and Psc); 3-4 inner gradates, 4-5 outer gradates. Hindwing length
|
||
<quantity id="23339B6F6620022554F0FE3737285F88" box="[455,558,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="92" unit="mm" value="12.2">12.2 mm</quantity>
|
||
, height
|
||
<quantity id="23339B6F6620022557B0FE37360C5F88" box="[647,778,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="5" pageNumber="92" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.2-3.9 mm</quantity>
|
||
(ratio of length: height, 3.1: 1), 9-11
|
||
<emphasis id="D6BFEA9866200225518AFE3431E95F88" box="[1213,1263,478,503]" italics="true" pageId="5" pageNumber="92">rarp</emphasis>
|
||
cells, 3 b cells, 4 b’ cells, 2-3 inner gradates, 4-5 outer gradates.
|
||
</paragraph>
|
||
<caption id="B0B466026620022555A0F9EB31DC58D5" ID-DOI="http://doi.org/10.5281/zenodo.3545366" ID-Zenodo-Dep="3545366" httpUri="https://zenodo.org/record/3545366/files/figure.png" pageId="5" pageNumber="92" startId="5.[151,250,1537,1562]" targetBox="[151,1436,561,1512]" targetPageId="5">
|
||
<paragraph id="E474368A6620022555A0F9EB31DC58D5" blockId="5.[151,1437,1537,1706]" pageId="5" pageNumber="92">
|
||
<emphasis id="D6BFEA986620022555A0F9EB34125865" bold="true" box="[151,276,1537,1562]" pageId="5" pageNumber="92">FIGURE 4.</emphasis>
|
||
<taxonomicName id="23CB4D0966200225542BF9EB37005866" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[284,518,1537,1561]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA9866200225542BF9EB37005866" box="[284,518,1537,1561]" italics="true" pageId="5" pageNumber="92">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662002255723F9E837605865" bold="true" box="[532,614,1538,1562]" pageId="5" pageNumber="92">
|
||
<taxonomicNameLabel id="CD8C57E3662002255723F9E837605865" box="[532,614,1538,1562]" pageId="5" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Paratype (female, Peru: La Libertad, TRC). Wings. A. Forewing B. Hindwing.
|
||
<emphasis id="D6BFEA986620022555A0F9CF35DE5842" box="[151,216,1573,1597]" italics="true" pageId="5" pageNumber="92">b2, b3</emphasis>
|
||
, second, third upper Banksian cells (cells below RP that do not include a gradate vein);
|
||
<emphasis id="D6BFEA9866200225515EF9CF31BC5842" box="[1129,1210,1573,1597]" italics="true" pageId="5" pageNumber="92">b’2, b’4</emphasis>
|
||
, second, fourth lower Banksian cells (cells between Psm and Psc;
|
||
<emphasis id="D6BFEA98662002255765F9A33770581E" box="[594,630,1609,1633]" italics="true" pageId="5" pageNumber="92">gc3</emphasis>
|
||
, third gradate cell (cell with inner and outer gradates at the two ends);
|
||
<emphasis id="D6BFEA9866200225500BF9A3306B581E" box="[1340,1389,1609,1633]" italics="true" pageId="5" pageNumber="92">icu1,</emphasis>
|
||
first intracubital cell; 1ig, first inner gradate vein,
|
||
<emphasis id="D6BFEA98662002255768F987378D58FA" box="[607,651,1645,1669]" italics="true" pageId="5" pageNumber="92">im1,</emphasis>
|
||
first intramedian cell; Psc, Pseudocubitus; Psm, Pseudomedius;
|
||
<emphasis id="D6BFEA98662002255021F987309058FA" box="[1302,1430,1645,1669]" italics="true" pageId="5" pageNumber="92">rarp1, rarp9</emphasis>
|
||
, first, ninth closed cells between RA and RP; RA, anterior branch of Radius; RP, posterior branch of Radius.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A6620022655F0F93234365EA8" blockId="5.[151,1436,1751,2030]" lastBlockId="6.[151,1437,152,215]" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
|
||
<emphasis id="D6BFEA986620022555F0F9323476588D" box="[199,368,1752,1778]" italics="true" pageId="5" pageNumber="92">Male abdomen</emphasis>
|
||
(
|
||
<figureCitation id="7CF02A0F662002255448F93234CF588D" box="[383,457,1751,1778]" captionStart="FIGURE 5" captionStartId="6.[151,250,1715,1740]" captionTargetBox="[151,1436,237,1690]" captionTargetId="figure@6.[151,1436,237,1690]" captionTargetPageId="6" captionText="FIGURE 5. Ungla pseudomeleoma, sp. nov., Holotype, cleared abdomen (male, Peru: La Libertad, FSCA). A. Abdomen (lateral); B. Abdominal terminus (lateral); C. Terminal abdominal segments and genitalia (lateral); D. Fused ninth tergite and ectoproct (dorsal); E. Fused eighth and ninth sternites (ventral, suture on dorsal integument between T8 and T9 visible). c.c., callus cerci; g.ap., gonarcal apodeme; g.br., gonarcal bridge; inv, invaginated dorsal cleft in T9+e, above anus; S3, third sternite; S8+9, fused eighth and ninth sternites, s.p., setose subanal plate; sp, spiracular opening; T4, T6, T8, T9, fourth, sixth, eighth, ninth tergites; T9+e, fused ninth tergite and ectoproct." figureDoi="http://doi.org/10.5281/zenodo.3545368" httpUri="https://zenodo.org/record/3545368/files/figure.png" pageId="5" pageNumber="92">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="7CF02A0F6620022554E1F93234E3588C" box="[470,485,1752,1779]" captionStart="FIGURE 6" captionStartId="7.[151,250,1754,1779]" captionTargetBox="[151,1436,273,1727]" captionTargetId="figure@7.[151,1436,273,1729]" captionTargetPageId="7" captionText="FIGURE 6. Ungla pseudomeleoma, sp. nov., Holotype, genitalia (male, Peru: La Libertad, FSCA). A. Gonarcal complex (anterolateral); B. Gonarcus (posterodorsal); C. Gonarcal complex (frontal); D. gonarcus (caudal); E. Gonarcal complex (frontoventral). c, comes; gsac, gonosaccus; g.ap., gonarcal apodeme; g.br., gonarcal bridge; h.i., hypandrium internum; mu, mediuncus; pr, unarticulated process on frontal margin of gonarcal apodeme; s.p., setose subanal plate; s.pl., paired setose plate of gonosaccus." figureDoi="http://doi.org/10.5281/zenodo.3545370" httpUri="https://zenodo.org/record/3545370/files/figure.png" pageId="5" pageNumber="92">6</figureCitation>
|
||
): Tergites, sternites, pleural regions covered with relatively dense covering of moderately long setae; microsetae present; no microtholi. T6 (lateral view) length
|
||
<quantity id="23339B6F66200225512CF91631705969" box="[1051,1142,1788,1814]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="92" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, ~7.2x height; T7 (lateral view) length
|
||
<quantity id="23339B6F662002255407F8CA348C5945" box="[304,394,1824,1850]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="92" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, ~3.8x height; T8 (lateral view) length
|
||
<quantity id="23339B6F662002255664F8CA36AB5945" box="[851,941,1824,1850]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="92" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, ~6.9x height; T9+ect (lateral view) length
|
||
<quantity id="23339B6F6620022555A0F8AE35E95921" box="[151,239,1860,1886]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="92" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, 1.5x height, ~0.6 length of T7; S6 (lateral view) length
|
||
<quantity id="23339B6F66200225565DF8AE36C45921" box="[874,962,1860,1886]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="92" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, ~2.7x height; S7 (lateral view) length
|
||
<quantity id="23339B6F66200225504EF8AE35C359FD" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="92" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, ~2.5x height. Tergites, sternites roughly rectangular, with pleural margins slightly concave. Segments A4 to A8 with large spiracles, (e.g., A7: spiracle diameter ~0.19-0.21x length of sternite). T9+ect with dorsal invagination prominent (~0.33x dorsal length of T9+ect), margins of invagination almost straight, base rounded; dorsal margin of T9+ect moderately rounded throughout, posterior margin of ectoproct convex, without knob or extension; ventral margin straight, lightly sclerotized; callus cerci large, ovate (diameter:
|
||
<quantity id="23339B6F66230226569CFF72314B5ECC" box="[939,1101,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="93" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.10-0.15 mm</quantity>
|
||
), with discrete, well separated trichobothria.
|
||
</paragraph>
|
||
<caption id="B0B466026623022655A0F95937795900" ID-DOI="http://doi.org/10.5281/zenodo.3545368" ID-Zenodo-Dep="3545368" httpUri="https://zenodo.org/record/3545368/files/figure.png" pageId="6" pageNumber="93" startId="6.[151,250,1715,1740]" targetBox="[151,1436,237,1690]" targetPageId="6">
|
||
<paragraph id="E474368A6623022655A0F95937795900" blockId="6.[151,1437,1715,1920]" pageId="6" pageNumber="93">
|
||
<emphasis id="D6BFEA986623022655A0F959341558B3" bold="true" box="[151,275,1715,1740]" pageId="6" pageNumber="93">FIGURE 5.</emphasis>
|
||
<taxonomicName id="23CB4D0966230226542DF95E370558B3" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[282,515,1716,1740]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA9866230226542DF95E370558B3" box="[282,515,1716,1740]" italics="true" pageId="6" pageNumber="93">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662302265726F95E376458B3" bold="true" box="[529,610,1716,1740]" pageId="6" pageNumber="93">
|
||
<taxonomicNameLabel id="CD8C57E3662302265726F95E376458B3" box="[529,610,1716,1740]" pageId="6" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype, cleared abdomen (male, Peru: La Libertad, FSCA). A. Abdomen (lateral); B. Abdominal terminus (lateral); C. Terminal abdominal segments and genitalia (lateral); D. Fused ninth tergite and ectoproct (dorsal); E. Fused eighth and ninth sternites (ventral, suture on dorsal integument between T8 and T9 visible). c.c., callus cerci; g.ap., gonarcal apodeme; g.br., gonarcal bridge; inv, invaginated dorsal cleft in T9+e, above anus; S3, third sternite; S8+9, fused eighth and ninth sternites, s.p., setose subanal plate; sp, spiracular opening; T4, T6, T8, T9, fourth, sixth, eighth, ninth tergites; T9+e, fused ninth tergite and ectoproct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A6623022755F0F844318F5E84" blockId="6.[151,1436,1966,2029]" lastBlockId="7.[151,1437,153,251]" lastPageId="7" lastPageNumber="94" pageId="6" pageNumber="93">S8+9 fused, with line of fusion not detectible; length (lateral view) 1.0 mm, ~4.8x height; dorsal margin with apodeme lightly sclerotized, mostly noticeable on basal two-thirds; dorsum tapering roughly from area below T9 to beyond callus cerci, then forming small ledge with upturned terminus; terminus blunt (lateral view), extending very slightly beyond distal margin of T9+ect; setae dense, slender, mostly long, simple, those along distolateral margin of similar size to other setae, without flanges. Subanal plate distinct, with ~12 small setae.</paragraph>
|
||
<caption id="B0B466026622022755A0F930341559FC" ID-DOI="http://doi.org/10.5281/zenodo.3545370" ID-Zenodo-Dep="3545370" httpUri="https://zenodo.org/record/3545370/files/figure.png" pageId="7" pageNumber="94" startId="7.[151,250,1754,1779]" targetBox="[151,1436,273,1727]" targetPageId="7">
|
||
<paragraph id="E474368A6622022755A0F930341559FC" blockId="7.[151,1436,1754,1923]" pageId="7" pageNumber="94">
|
||
<emphasis id="D6BFEA986622022755A0F9303410588C" bold="true" box="[151,278,1754,1779]" pageId="7" pageNumber="94">FIGURE 6.</emphasis>
|
||
<taxonomicName id="23CB4D09662202275417F931370B588C" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[288,525,1755,1779]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA98662202275417F931370B588C" box="[288,525,1755,1779]" italics="true" pageId="7" pageNumber="94">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA9866220227572AF9313777588C" bold="true" box="[541,625,1755,1779]" pageId="7" pageNumber="94">
|
||
<taxonomicNameLabel id="CD8C57E366220227572AF9313777588C" box="[541,625,1755,1779]" pageId="7" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype, genitalia (male, Peru: La Libertad, FSCA). A. Gonarcal complex (anterolateral); B. Gonarcus (posterodorsal); C. Gonarcal complex (frontal); D. gonarcus (caudal); E. Gonarcal complex (frontoventral). c, comes; gsac, gonosaccus; g.ap., gonarcal apodeme; g.br., gonarcal bridge; h.i., hypandrium internum; mu, mediuncus; pr, unarticulated process on frontal margin of gonarcal apodeme; s.p., setose subanal plate; s.pl., paired setose plate of gonosaccus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A6622022855F0F85B30245F18" blockId="7.[151,1436,1969,2031]" lastBlockId="8.[151,1436,153,360]" lastPageId="8" lastPageNumber="95" pageId="7" pageNumber="94">Gonarcus broadly arcuate, with bridge stout, rounded dorsally, acute interiorly; apodemes (arms) heavy, relatively straight, expanded distally, with lightly sclerotized unarticulated process on frontal margin extending distally toward gonosaccus. Mediuncus with rounded base closely attached to gonarcal bridge, narrow distally, with terminus narrow, curved downward about 90° (lateral view). Gonosaccus bilobed, with lobes closely aligned, when unexpanded forming a triangular envelope around tip of mediuncus, when expanded consisting of two lobes, rounded dorsally, mesally, laterally, but flat distally; each lobe with large, dense patch of sturdy gonosetae arising from prominent setal bases facing inward when uneverted, outward when everted; gonosetae relatively long (almost as long as length of mediuncus); hypandrium internum quadrate, with dense arms, sturdy, keel-like comes.</paragraph>
|
||
<caption id="B0B46602662D022855A0F8AC30815990" ID-DOI="http://doi.org/10.5281/zenodo.3545372" ID-Zenodo-Dep="3545372" httpUri="https://zenodo.org/record/3545372/files/figure.png" pageId="8" pageNumber="95" startId="8.[151,250,1862,1887]" targetBox="[151,1436,381,1837]" targetPageId="8">
|
||
<paragraph id="E474368A662D022855A0F8AC30815990" blockId="8.[151,1437,1862,2031]" pageId="8" pageNumber="95">
|
||
<emphasis id="D6BFEA98662D022855A0F8AC34125920" bold="true" box="[151,276,1862,1887]" pageId="8" pageNumber="95">FIGURE 7.</emphasis>
|
||
<taxonomicName id="23CB4D09662D0228542CF8AD37035920" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" box="[283,517,1863,1887]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA98662D0228542CF8AD37035920" box="[283,517,1863,1887]" italics="true" pageId="8" pageNumber="95">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662D02285724F8AD37625920" bold="true" box="[531,612,1863,1887]" pageId="8" pageNumber="95">
|
||
<taxonomicNameLabel id="CD8C57E3662D02285724F8AD37625920" box="[531,612,1863,1887]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Paratype, cleared abdomen (female, Peru: La Libertad, TRC). A. Terminal segments of abdomen (lateral); B. Fused ninth tergite and ectoproct (lateral); C, D. Spermathecal complex; E, F. Spermathecal complex and subgenitale. c.c., callus cerci; g.l. gonapophysis lateralis; inv, spermathecal invagination; pouch, pouch at base of subgenitale; sg, subgenitale; sm, spermatheca; sm.d., spermathecal duct; sm.d.(b), sm.d.(d), basal, distal sections of spermathecal duct; S7, seventh sternite; T7, T8, seventh, eighth tergites; T9+e, fused ninth tergite and ectoproct; vel, spermathecal vellum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B0B46602662C022955A0FC0F31655A16" ID-DOI="http://doi.org/10.5281/zenodo.3545374" ID-Zenodo-Dep="3545374" httpUri="https://zenodo.org/record/3545374/files/figure.png" pageId="9" pageNumber="96" startId="9.[151,250,997,1022]" targetBox="[213,1375,181,971]" targetPageId="9">
|
||
<paragraph id="E474368A662C022955A0FC0F31655A16" blockId="9.[151,1437,997,1130]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955A0FC0F34135D81" bold="true" box="[151,277,997,1022]" pageId="9" pageNumber="96">FIGURE 8.</emphasis>
|
||
Similarity in the antennal features of
|
||
<taxonomicName id="23CB4D09662C02295794FC0F368B5D82" authorityName="Tauber" authorityYear="2019" box="[675,909,997,1021]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="pseudomeleoma">
|
||
<emphasis id="D6BFEA98662C02295794FC0F368B5D82" box="[675,909,997,1021]" italics="true" pageId="9" pageNumber="96">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
with those of three species of
|
||
<taxonomicName id="23CB4D09662C022951E7FC0F30285D82" authorityName="Fitch" authorityYear="1855" box="[1232,1326,997,1021]" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662C022951E7FC0F30285D82" box="[1232,1326,997,1021]" italics="true" pageId="9" pageNumber="96">Meleoma</emphasis>
|
||
</taxonomicName>
|
||
. A.
|
||
<taxonomicName id="23CB4D09662C0229506AFC0F343C5A5E" authority="Tauber, 2019" authorityName="Tauber" authorityYear="2019" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="pseudomeleoma" status="sp. nov.">
|
||
<emphasis id="D6BFEA98662C0229506AFC0F343C5A5E" italics="true" pageId="9" pageNumber="96">Ungla pseudomeleoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D6BFEA98662C02295471FBE034905A5D" bold="true" box="[326,406,1034,1058]" pageId="9" pageNumber="96">
|
||
<taxonomicNameLabel id="CD8C57E3662C02295471FBE034905A5D" box="[326,406,1034,1058]" pageId="9" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype (Peru: La Libertad, FSCA); B.
|
||
<taxonomicName id="23CB4D09662C02295609FBE331375A5D" authority="Fitch" authorityName="Fitch" box="[830,1073,1033,1058]" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="emuncta">
|
||
<emphasis id="D6BFEA98662C02295609FBE336FE5A5E" box="[830,1016,1033,1057]" italics="true" pageId="9" pageNumber="96">Meleoma emuncta</emphasis>
|
||
Fitch
|
||
</taxonomicName>
|
||
(USA: California, TRC); C.
|
||
<taxonomicName id="23CB4D09662C02295063FBE335B05A3A" authorityName="Fitch" authorityYear="1855" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662C02295063FBE335B05A3A" italics="true" pageId="9" pageNumber="96">Meleoma</emphasis>
|
||
</taxonomicName>
|
||
undescribed sp. #1 (USA: Arizona, TRC); D.
|
||
<taxonomicName id="23CB4D09662C022957B3FBC736705A39" authority="Penny" authorityName="Penny" authorityYear="2006" box="[644,886,1069,1094]" class="Insecta" family="Chrysopidae" genus="Meleoma" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="stangei">
|
||
<emphasis id="D6BFEA98662C022957B3FBC736375A3A" box="[644,817,1069,1093]" italics="true" pageId="9" pageNumber="96">Meleoma stangei</emphasis>
|
||
Penny
|
||
</taxonomicName>
|
||
(Mexico: Baja California, San Diego County Museum of Natural History). f1, first flagellomere; h, horn extending from scape; ped, pedicel; sc, scape.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E474368A662C022955F0FB7230875B3D" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0FB72348D5ACD" box="[199,395,1176,1202]" italics="true" pageId="9" pageNumber="96">Female abdomen</emphasis>
|
||
(
|
||
<figureCitation id="7CF02A0F662C022954ABFB7234E75ACD" box="[412,481,1176,1202]" captionStart="FIGURE 7" captionStartId="8.[151,250,1862,1887]" captionTargetBox="[151,1436,381,1837]" captionTargetId="figure@8.[151,1436,381,1837]" captionTargetPageId="8" captionText="FIGURE 7. Ungla pseudomeleoma, sp. nov., Paratype, cleared abdomen (female, Peru: La Libertad, TRC). A. Terminal segments of abdomen (lateral); B. Fused ninth tergite and ectoproct (lateral); C, D. Spermathecal complex; E, F. Spermathecal complex and subgenitale. c.c., callus cerci; g.l. gonapophysis lateralis; inv, spermathecal invagination; pouch, pouch at base of subgenitale; sg, subgenitale; sm, spermatheca; sm.d., spermathecal duct; sm.d.(b), sm.d.(d), basal, distal sections of spermathecal duct; S7, seventh sternite; T7, T8, seventh, eighth tergites; T9+e, fused ninth tergite and ectoproct; vel, spermathecal vellum." figureDoi="http://doi.org/10.5281/zenodo.3545372" httpUri="https://zenodo.org/record/3545372/files/figure.png" pageId="9" pageNumber="96">Fig. 7</figureCitation>
|
||
): Tergites, sternites with uniform, moderately dense setal covering, setae of moderately long length. T6 (lateral view) length
|
||
<quantity id="23339B6F662C02295747FB5637CE5AA9" box="[624,712,1212,1238]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="96" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, ~5.9x height; T7 (lateral view) length
|
||
<quantity id="23339B6F662C02295149FB5631D05AA9" box="[1150,1238,1212,1238]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="96" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, ~7.3x height; T8 (lateral view) length
|
||
<quantity id="23339B6F662C02295449FB0A34D05A85" box="[382,470,1248,1274]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="96" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, ~4.5x height; T9+ect (lateral view) length
|
||
<quantity id="23339B6F662C02295680FB0A31085A85" box="[951,1038,1248,1274]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="96" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, 0.5x height, ~0.5x length of T7; S6 (lateral view) length
|
||
<quantity id="23339B6F662C022954B6FAEE34DF5B61" box="[385,473,1284,1310]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="96" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, ~1.6x height; S7 length 1.0 mm, ~1.8x height. Tergites, sternites roughly rectangular, with pleural margins straight. Spiracles located in center of pleural region, roughly oval externally, not enlarged.
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0FAA6367D5BD1" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
Tergite 8 (lateral view) narrower, slightly taller than T7, with pleural region not sclerotized. Tergite 9 with ectoproct fused, extending ventrally full height of segment. Gonapophysis lateralis well sclerotized, elongate, oval (length
|
||
<quantity id="23339B6F662C022955D9FA7E34405BD1" box="[238,326,1428,1454]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="96" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, width
|
||
<quantity id="23339B6F662C022954AFFA7E34F45BD1" box="[408,498,1428,1454]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="96" unit="mm" value="0.1">0.1 mm</quantity>
|
||
), with relatively long, robust setae.
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0FA52343D58D5" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0FA52371B5BAD" box="[199,541,1464,1490]" italics="true" pageId="9" pageNumber="96">Bursal-spermathecal complex.</emphasis>
|
||
Spermatheca pillbox-shaped, with vellum; vellum of approximately same height as length of base, open to bursal duct via slit on side. Spermathecal invagination truncate, extending almost full depth of spermathecal bowl. Spermathecal duct well sclerotized, wavy, with smooth surface basally, extending directly toward subgenitale, becoming straight and setose distally at base of subgenitale. Bursa copulatrix small, consisting of light membranous pouch attached to distal section of subgenitale base, with proximal section or duct extending toward spermatheca, probably with two transparent, elongate bursal glands above spermatheca and spermathecal duct.
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0F95E317B588D" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0F95E344D58B1" box="[199,331,1716,1742]" italics="true" pageId="9" pageNumber="96">Subgenitale</emphasis>
|
||
. Distal section well sclerotized, with two, distinctly separated, round lobes extending from leathery, partially sclerotized, membranous base that extends internally. Lobes with rough surface.
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0F91634345921" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0F91634A35969" box="[199,421,1788,1814]" italics="true" pageId="9" pageNumber="96">Colleterial complex</emphasis>
|
||
. Colleterial gland elongate, clear, with relatively broad duct extending to small, colleterial reservoir near gonapophyses laterales; reservoir clear, opening via membranous duct above round, striated transverse sclerite.
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0F88D319159B5" blockId="9.[151,1437,1176,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0F88D34C159FD" bold="true" box="[199,455,1895,1922]" pageId="9" pageNumber="96">Specimens examined.</emphasis>
|
||
<materialsCitation id="54A33CD7662C022954F8F882319159B5" ID-GBIF-Occurrence="2452381636" collectionCode="FSCA, TRC" pageId="9" pageNumber="96" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
|
||
<typeStatus id="3B708828662C022954F8F882373159FD" box="[463,567,1896,1922]" pageId="9" pageNumber="96" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<specimenCount id="F2CDFD03662C0229577FF882375959FD" box="[584,607,1896,1922]" pageId="9" pageNumber="96" type="male">♂</specimenCount>
|
||
,
|
||
<collectionCode id="82DAAE4F662C0229575BF88237B259FD" box="[620,692,1896,1922]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="96" type="Museum">FSCA</collectionCode>
|
||
) and three
|
||
<typeStatus id="3B708828662C02295604F882369859FD" box="[819,926,1896,1922]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
|
||
(all from same locality and date;
|
||
<specimenCount id="F2CDFD03662C0229502AF882304659FD" box="[1309,1344,1896,1922]" pageId="9" pageNumber="96" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="82DAAE4F662C02295079F882309E59FD" box="[1358,1432,1896,1922]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="96" type="Museum">FSCA</collectionCode>
|
||
;
|
||
<specimenCount id="F2CDFD03662C022955A0F86635BA59D9" box="[151,188,1932,1958]" pageId="9" pageNumber="96" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="F2CDFD03662C022955FEF86635EA59D9" box="[201,236,1932,1958]" pageId="9" pageNumber="96" type="female">1♀</specimenCount>
|
||
, Tauber Research Collection,
|
||
<collectionCode id="82DAAE4F662C02295775F866377B59D9" box="[578,637,1932,1958]" pageId="9" pageNumber="96">TRC</collectionCode>
|
||
). Both of the female specimens appear slightly teneral; their structures seem softer and more membranous than those on mature specimens of other
|
||
<taxonomicName id="23CB4D09662C022956C5F85A313E59B5" authorityName="Navas" authorityYear="1914" box="[1010,1080,1968,1994]" class="Insecta" family="Chrysopidae" genus="Ungla" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6BFEA98662C022956C5F85A313E59B5" box="[1010,1080,1968,1994]" italics="true" pageId="9" pageNumber="96">Ungla</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E474368A662C022955F0F839300C5990" blockId="9.[151,1437,1176,2031]" box="[199,1290,2003,2031]" pageId="9" pageNumber="96">
|
||
<emphasis id="D6BFEA98662C022955F0F83934E95991" bold="true" box="[199,495,2003,2030]" pageId="9" pageNumber="96">Geographic distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="3B708828662C022957D8F83E36195991" box="[751,799,2004,2030]" pageId="9" pageNumber="96">type</typeStatus>
|
||
locality (
|
||
<collectingCountry id="9CDC761A662C022956BEF83E36C45991" box="[905,962,2004,2030]" name="Peru" pageId="9" pageNumber="96">Peru</collectingCountry>
|
||
: Department of
|
||
<collectingRegion id="260FF868662C02295141F83E31F85990" box="[1142,1278,2004,2031]" country="Peru" name="La Libertad" pageId="9" pageNumber="96">La Libertad</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |