526 lines
52 KiB
XML
526 lines
52 KiB
XML
<document id="A070F63A4B86D703C4A1530C8D57DBF7" ENCODING="UTF-8" ID-GBIF-Dataset="21d21894-7003-4d6d-882b-5322ea07968d" ModsDocAuthor="Longino, J. T." ModsDocID="21311" checkinTime="1244213549108" checkinUser="thomas" docAuthor="Longino, J. T." docDate="2007" docId="431358CAECCE0833C6AB79289BD0B271" docLanguage="en" docName="21311_gg2.xml" docOrigin="Zootaxa 1491" docSource="http://www.antbase.org/ants/publications/21311/21311.pdf" docTitle="Azteca sericeasur Longino, 2007, new species" docType="treatment" docVersion="12" lastPageNumber="48" masterDocId="A75589B0B733EFAA7DECE020EC76B43D" masterDocTitle="A taxonomic review of the genus Azteca (Hymenoptera: Formicidae) in Costa Rica and a global revision of the aurita group." masterLastPageNumber="63" masterPageNumber="1" pageNumber="46" updateTime="1701311503130" updateUser="plazi">
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<mods:mods id="E4CE25079519214F697C6E6CA9CBF3CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EBF9B3EEBD28624D7BC0A1290DA8DAA4">
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<mods:title id="A785A0093818E7353D51B6DE19588EE2">A taxonomic review of the genus Azteca (Hymenoptera: Formicidae) in Costa Rica and a global revision of the aurita group.</mods:title>
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<mods:name id="0589B3BB6B163B8A383547845D19E89E" type="personal">
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<mods:roleTerm id="BA0F8F5132E04F20B6FDE2573B7289BA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1E5946F6FABE2F729BD702473DF4ECE3">Longino, J. T.</mods:namePart>
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<mods:typeOfResource id="6734B01C5915D1C21CD442B3A30F68A5">text</mods:typeOfResource>
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<mods:title id="08F3EAAC6F330D132BC058285F1A2BB7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A453E5DA1D36DAD141BAA1021D7BF6E7">
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<mods:date id="CAB8E43C77F3C5B4082C61351EC84AED">2007</mods:date>
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<mods:detail id="FBC41F027A7362036DE756029991F39B" type="volume">
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<mods:number id="BE790FA0960AA1BAB910C191D73CDB0A">1491</mods:number>
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</mods:detail>
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<mods:extent id="5B055122EF83ADC248B546A11CA70289" unit="page">
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<mods:start id="AB50B295F4BC48BF6A45C91FAE588CCA">1</mods:start>
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<mods:end id="FEAB6300498480015484FAD5D9260D71">63</mods:end>
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<mods:location id="6955E6B660BAA87E39C23690627CDF9B">
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<mods:url id="7CD2F51DF4E5C6E30B756F69458C39F9">http://www.antbase.org/ants/publications/21311/21311.pdf</mods:url>
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</mods:location>
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<mods:classification id="5231751FF9D78B24F97645242D2A1491">journal article</mods:classification>
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<mods:identifier id="03CE8DD78F0DF919EF84C1C8AC1BB951" type="HNS-Pub">21311</mods:identifier>
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<mods:identifier id="302C284CF0DBB10353DBDFD5D00905F1" type="ZooBank">C31A1226-724D-4D1A-8471-E6BB441EE3EF</mods:identifier>
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</mods:mods>
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<treatment id="431358CAECCE0833C6AB79289BD0B271" ID-DOI="http://doi.org/10.5281/zenodo.6246540" ID-GBIF-Taxon="100125939" ID-Zenodo-Dep="6246540" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271" httpUri="http://treatment.plazi.org/id/431358CAECCE0833C6AB79289BD0B271" lastPageNumber="48" pageNumber="46">
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<subSubSection id="0DCA1BBD66F6A86FD187CA778B78F984" type="nomenclature">
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<paragraph id="590B1412A91E98477674519C71A88AED" pageNumber="46">
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<taxonomicName id="CF0B8A70BC204AEAF08F434197F9177E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239265" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericeasur" order="Hymenoptera" pageNumber="46" phylum="Arthropoda" rank="species" species="sericeasur">Azteca sericeasur</taxonomicName>
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<taxonomicNameLabel id="6B92450BD6270325693670EE6078D01B" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="F8BF8657A1F044689FC458F934F37A22" type="description">
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<paragraph id="8DFA54760666D4D733EA53BB1F2BB15A" pageNumber="46">Figures 3,4A,5,6E,6F,7.</paragraph>
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</subSubSection>
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<subSubSection id="E1FDC2724F9D995783B7B70A0AE2C067" type="nomenclature">
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<paragraph id="81103436DA41D49F4E3C08F1F2D7B8B8" pageNumber="46">
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<materialsCitation id="385AD64DA25592548BD39AB6799DB309" ID-GBIF-Occurrence="923890755" collectingDate="1925-07-11" collectionCode="USNM" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000005642" stateProvince="Prov. Limon" typeStatus="Holotype">
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<typeStatus id="4761C263566847232D11251AAF5B615B">Holotype</typeStatus>
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alate queen:
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<collectingCountry id="28AA898ED020EEE0B1573EE705725B79">Costa Rica</collectingCountry>
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,
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<collectingRegion id="027B1D58133531A7C608C6B7B101CEFA">
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Prov.
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<normalizedToken id="10C5508B33C3590A5F609905FD4F7BCE" originalValue="Limón">Limon</normalizedToken>
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</collectingRegion>
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,
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<location id="992F68C8C50E1E37FBF92B884F20264B" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:992F68C8C50E1E37FBF92B884F20264B" country="Costa Rica" latitude="10.25" longitude="-83.45" name="Hamburg Farm" stateProvince="Prov. Limon">Hamburg Farm</location>
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[
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<geoCoordinate id="3A4679DAB19F9A2A03E6E6C3D9E0E7BB" direction="north" orientation="latitude" value="10.25">10°15'N</geoCoordinate>
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,
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<geoCoordinate id="A6D183D29DCBE0DF747D56E747C00676" direction="west" orientation="longitude" value="-83.45">83°27'W</geoCoordinate>
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],
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<location id="1E4C8CE43599108D49C08CF1D5B03E7B" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:1E4C8CE43599108D49C08CF1D5B03E7B" country="Costa Rica" latitude="10.25" longitude="-83.45" name="50 m" stateProvince="Prov. Limon">50m</location>
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,
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<date id="337F300D0713DDDB56694913A43BA38A" day="11" month="07" value="1925-07-11" year="1925">
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<collectingDate id="B2415977C9F4E6277BA2C7553E4E85C8" day="11" month="07" value="1925-07-11" year="1925">11 Jul 1925</collectingDate>
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</date>
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(
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<collectorName id="B55C29A135C050C77155478796DD01E8">F. Nevermann</collectorName>
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) [
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<collectionCode id="3D963855C73A884B9028A973FE6D25CC" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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, specimen (pin) code
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<specimenCode id="DADAB052164314D34E1DB64BD188D2BC">JTLC000005642</specimenCode>
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].
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</materialsCitation>
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</paragraph>
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<paragraph id="388BFCCB9D1AE0555ECFF067E109C611" country="Costa Rica" pageNumber="47">
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<materialsCitation id="9DF5019912026DF6A8600241D557716D" ID-GBIF-Occurrence="923890780" collectingDate="1925-07-11" collectionCode="USNM" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000005642" stateProvince="Prov. Limon" typeStatus="Paratypes">
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<typeStatus id="B3EA1740B44BCD617DFA25B30D828B74">Paratypes</typeStatus>
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: same data as holotype; 2 workers [
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<specimenCode id="207943AF2DBD5320D1061C7EBB3D820B">JTLC000005642</specimenCode>
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,
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<collectionCode id="C9D168C7E96E5EFC49F693123CC63718" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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];
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</materialsCitation>
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<materialsCitation id="D14461A500F1B96679C1069AF5AAB0C8" ID-GBIF-Occurrence="923890757" collectingDate="1925-07-11" collectionCode="MCZC" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000005643" stateProvince="Prov. Limon" typeStatus="Paratypes">
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alate queen, worker, male [
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<specimenCode id="976E47B120F250D694FA874C2770A658">JTLC000005643</specimenCode>
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,
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<collectionCode id="5A8358730634603E698EEFFE8B2A1263" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
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];
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</materialsCitation>
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<materialsCitation id="3FC1352273F1477380286C01ACE833B6" ID-GBIF-Occurrence="923890769" collectingDate="1925-07-11" collectionCode="INBC" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000009416" stateProvince="Prov. Limon" typeStatus="Paratypes">
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alate queen, 2 workers [
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<specimenCode id="D8FD750AC49F9EA89A36643B4A0BA5A0">JTLC000009416</specimenCode>
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,
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<collectionCode id="7763C39E76194DDFFD932C86338778D1" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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];
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</materialsCitation>
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<materialsCitation id="DC4D0367522BB1673ED40A5974FB75B5" ID-GBIF-Occurrence="923890761" collectingDate="1925-07-11" collectionCode="LACM" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000009415" stateProvince="Prov. Limon" typeStatus="Paratypes">
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2 workers, 1 male [
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<specimenCode id="19442E9F1486145C7B3C6DAEC1B8F156">JTLC000009415</specimenCode>
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,
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<collectionCode id="B026EEA4AF120A2C3351752C68AADDC0" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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];
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</materialsCitation>
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<materialsCitation id="446C23238DAA701B3717169E501EFEEF" ID-GBIF-Occurrence="923890770" collectingDate="1925-07-11" collectionCode="USNM" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000009412, JTLC 000009413, JTLC 000009414" stateProvince="Prov. Limon" typeStatus="Paratypes">
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8 workers [
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<specimenCode id="6E98D20C7A371FCF54ED84A1BA726D37">JTLC000009412</specimenCode>
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,
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<specimenCode id="82C2394502309BE5BC0EAAA2D4ADCE39">JTLC000009413</specimenCode>
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,
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<specimenCode id="5B4FFB297E813F7D43BAD8154FF1ED59">JTLC000009414</specimenCode>
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,
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<collectionCode id="78C89EC51ED5B9D607FEF3FC86D79CB4" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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];
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</materialsCitation>
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<materialsCitation id="07D6DCDA9941DEEA0D24D1A5EFB007A2" ID-GBIF-Occurrence="923890776" collectingDate="1925-07-11" collectionCode="JTLC" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" specimenCode="JTLC 000009405" stateProvince="Prov. Limon" typeStatus="Paratypes">
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3 workers [
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<specimenCode id="9E6372A45F561609DAC17FB8DCA6DFB4">JTLC000009405</specimenCode>
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,
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<collectionCode id="C0DA73657995EF833F6FAA7FCAC40D3C" collectionName="John T. Longino">JTLC</collectionCode>
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]
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="809651B16A49AA4BAE266A7BC0E1CC56" type="description">
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<paragraph id="C4681984B8C4B2A3E63B18367F6E5F73" pageNumber="47">Measurements of holotype. HLA 2.14, HW 2.13, SL 1.335, EL 0.518, OC 0.102, MTSC 30.</paragraph>
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</subSubSection>
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<subSubSection id="0C4B9418060262CFD6FEFE5B822DC070" type="diagnosis">
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<paragraph id="C1344E7DCBA98EC2F005031314C1F924" pageNumber="47">
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Diagnosis. The shape and size of the queen head distinguishes this species from all others except
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<taxonomicName id="6210BBFB2677A3DD4F14A1A7FB300C81" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericea">A. sericea</taxonomicName>
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. It differs from
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<taxonomicName id="1EDAED324D0C65A67CABCD656F3F992E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericea">A. sericea</taxonomicName>
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in the presence of erect setae on the posterior margin of the head. It is very similar to
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<taxonomicName id="B2DD9500941E9089A83696C573B59EA9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26035" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca velox Forel" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="velox">A. velox</taxonomicName>
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, differing only in the larger size.
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<taxonomicName id="8F1C5430210BEB9A94E8ABD9BE810DEB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25993" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca gnava Forel" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="gnava">Azteca gnava</taxonomicName>
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and
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<taxonomicName id="A5D384E3ABD4925FA8320CFBBAC80E94" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25999" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca instabilis (Smith)" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="instabilis">A. instabilis</taxonomicName>
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are similar but larger.
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<taxonomicName id="6828FEC6F1787F99CAE18EC463D22199" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25993" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca gnava Forel" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="gnava">Azteca gnava</taxonomicName>
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has the head solid dark brown;
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<taxonomicName id="C6FC8D07B7B9A2180DDE0244F9DD8A71" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239265" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericeasur" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericeasur">A. sericeasur</taxonomicName>
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has much of the anterior and lateral head light orange brown.
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<taxonomicName id="47DB0F13833D048743991C7BD1D6A7D2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25999" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca instabilis (Smith)" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="instabilis">Azteca instabilis</taxonomicName>
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has large ocelli (OCW> 0.20mm);
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<taxonomicName id="7F157D0D6117D14CFF127622C0F224E3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239265" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericeasur" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericeasur">A. sericeasur</taxonomicName>
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has small ocelli (OCW about 0.10mm).
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</paragraph>
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</subSubSection>
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<subSubSection id="7FFD491A64BA3B5EA9813CA4E52D2BFD" type="description">
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<paragraph id="1447D65FBB4295269FB16D70CBAE03EF" pageNumber="47">Queen characters. Measurements (n=10): HLA 2.00 (1.94-2.14), HW 2.00 (1.94-2.17), SL 1.31 (1.28- 1.39), CI 100 (99-104), SI 66 (62-67).</paragraph>
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<paragraph id="CB8BFED3752F1E6387F0B484360FC0BE" pageNumber="47">Palpal formula 6,4; ocelli small; middle and hind tibia with prominent pectinate apical spur; dorsal surface of mandible with small piligerous puncta, setae in puncta short, subdecumbent, interspaces between puncta smooth and shiny on apical half of mandible, gradually becoming faintly microareolate at base; medial and lateral clypeal lobes at about same level; head broadly cordate with rounded sides; petiolar node tall, strongly compressed into thin scale at apex; posteroventral petiolar lobe evenly convex from front to back; scape with abundant erect setae, about as long as one half maximum width of scape; middle and hind tibia with abundant erect setae, longest of these about as long as one third to one half maximum width of tibia (MTSC 20-30); sides of head without erect setae; posterior margin of head with abundant erect setae; pronotum with erect setae on posterior margin; mesoscutum, scutellum, and propodeum with abundant erect setae; petiolar node with rim of dense erect whitish pubescence, sparse longer setae, one pair extending above apex in profile, posterior border of sternal lobe of petiole with dense layer of long erect pubescence and several erect setae that are about twice as long as pubescence; gastral terga with sparse long erect setae; light orange brown coloration on clypeus, malar area, antennal fossa, and side of head, variable extent of infuscation on medial and posterior vertex.</paragraph>
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<paragraph id="33EFE0C86779521151A94737FF85F0F4" pageNumber="47">Worker characters. Measurements (n=9): HLA 1.32 (1.10-1.69), HW 1.31 (1.08-1.76), SL 1.09 (0.99- 1.26), CI 99 (96-104), SI 83 (75-90).</paragraph>
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<paragraph id="CCF26D40C515BD558FA85456E2F3706A" pageNumber="47">Palpal formula 6,4; middle and hind tibia with prominent pectinate apical spur; dorsal surface of mandible smooth and shining, with moderately abundant small piligerous puncta, setae in puncta short, erect, larger puncta with long setae near masticatory margin; medial and lateral clypeal lobes at about same level; head with convex sides, strongly cordate posterior margin; in lateral profile promesonotum forming single convexity; scape with abundant erect setae, length of setae about one half maximum width of scape; mid and hind tibia with abundant erect setae, longest setae about one half maximum width of tibia; sides of head without erect setae; posterior margin of head with erect setae; pronotum and mesonotum with abundant long erect setae, somewhat sparser erect setae on dorsal face of propodeum; color light to dark brown, if somewhat bicolored gaster is darker then mesosoma.</paragraph>
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</subSubSection>
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<subSubSection id="855C700251CBA86C1DF3E27822DAC99B" type="etymology">
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<paragraph id="D5540A784F861AA14459359B5431CE78" pageNumber="47">
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Etymology. The name refers to this species being a southern version of the Mexican species
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<taxonomicName id="C595DA4A86F50241C6971506CD2151C6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericea">A. sericea</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="74CC44C20C9E899793258A6C78A129F4" type="distribution">
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<paragraph id="EC179FF2A348E1EE01ACF3560FFFE684" pageNumber="47">Range. Guatemala, Costa Rica, Panama.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="53AB32A0DCDA0D94D21B7DCCFB40E0EE" type="biology_ecology">
|
|
<paragraph id="D89A43F25A8971E05D5E694D596F9206" pageNumber="47">
|
|
Biology.
|
|
<taxonomicName id="D4C0B3E4CD7D3A3DA8C0A4FAF1CE54F0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239265" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericeasur" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="species" species="sericeasur">Azteca sericeasur</taxonomicName>
|
|
occurs in moist to wet forest habitats. Queens establish in live stems of understory trees, near the base. Mature colonies are polydomous and dispersed. Foraging columns extend across low vegetation and on the forest floor, connecting multiple small pavilions made of carton (paper-like material constructed by the ants from masticated plant fibers). Clusters of workers and brood occur in the pavilions, and the pavilions may cover membracids and pink scale insects, which the ants tend. Thus the queen is located in a permanent and well-protected site, while the colony is widely spread in a large number of small ephemeral nests.
|
|
</paragraph>
|
|
<paragraph id="09D3D57B98D1D53D4CEAA4160EE4008D" pageNumber="48">
|
|
Along the
|
|
<normalizedToken id="009583C235F314A9618E81C184760070" originalValue="Río">Rio</normalizedToken>
|
|
San Luis near Monteverde, I found workers coming and going from a fissure at the base of a live
|
|
<taxonomicName id="DD31AA87BC73B99CDEA8CCF66165D65B" genus="Xanthoxylem" pageNumber="48" rank="genus">Xanthoxylem</taxonomicName>
|
|
(Rutaceae) at a pasture edge. A colony of
|
|
<taxonomicName id="39B3BBE02E2E759A247695D00E6A2D3C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26991" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus novogranadensis Mayr" order="Hymenoptera" pageNumber="48" phylum="Arthropoda" rank="species" species="novogranadensis">Camponotus novogranadensis</taxonomicName>
|
|
was using the same entrance, in an apparent case of parabiosis. I have several observations of colonies from the Osa Peninsula. One colony was spread over an area of several square meters, in the dead, hollow core of a live
|
|
<taxonomicName id="D7A8CB6F68DDA9B49B049516DABAE6AB" class="Magnoliopsida" family="Rubiaceae" genus="Chimarrhis" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageNumber="48" phylum="Tracheophyta" rank="species" species="parviflora">Chimarrhis parviflora</taxonomicName>
|
|
(Rubiaceae), in nearby dead sticks, and filling hollow stems of an adjacent
|
|
<taxonomicName id="39E4486CDE0D6CAA9A638705378ACBF4" class="Magnoliopsida" family="Salicaceae" genus="Tetrathylacium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageNumber="48" phylum="Tracheophyta" rank="species" species="costaricensis">Tetrathylacium costaricensis</taxonomicName>
|
|
plant (Flacourtiaceae). The
|
|
<taxonomicName id="776A8F787276A1100C86125F2381CB9D" class="Magnoliopsida" family="Salicaceae" genus="Tetrathylacium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageNumber="48" phylum="Tracheophyta" rank="genus">Tetrathylacium</taxonomicName>
|
|
contained workers, small brood, and mealybugs, but no sexuals or large brood. Small carton nests covered some of the stems and nest entrances. The dead core of the
|
|
<taxonomicName id="D875CD2925BB247E79BBD4C38702EC76" class="Magnoliopsida" family="Rubiaceae" genus="Chimarrhis" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageNumber="48" phylum="Tracheophyta" rank="genus">Chimarrhis</taxonomicName>
|
|
contained a small pocket of carton, in which I found the grossly distended colony queen and abundant small brood. In another case I found workers streaming through low vegetation. A nest was in a small sapling; the lowest part of the nest was 4m above the ground. There were numerous small cavities and knotholes in the trunk; these were all filled with workers and brood. The lowermost knothole was covered with a carton nest, and the bulk of the workers were in the carton portion. I looked in the carton and in all the knotholes, but I never found a queen. Leanne Tennant, in her study of
|
|
<taxonomicName id="0EAEEA69638760BD70D36D43825D5F2B" class="Magnoliopsida" family="Salicaceae" genus="Tetrathylacium" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageNumber="48" phylum="Tracheophyta" rank="species" species="costaricensis">Tetrathylacium costaricensis</taxonomicName>
|
|
in Corcovado, found at least two of her study plants inhabited by
|
|
<taxonomicName id="19797B861F15C7FB7BE29E42EEED5755" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239265" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericeasur" order="Hymenoptera" pageNumber="48" phylum="Arthropoda" rank="species" species="sericeasur">A. sericeasur</taxonomicName>
|
|
. Males may be produced in large numbers in the small satellite nests. In August, 1982, I observed a series of carton shelters on vine stems at the base of a buttressed tree, and they were packed with hundreds of adult males. Alate queens are few in collections; two were collected in October and one in July. Nest series with alate queens have been taken twice, both in July. I collected a founding queen in the internode of a
|
|
<taxonomicName id="2771EB1B1A17050212ADDF94940C9519" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" higherTaxonomySource="CoL" kingdom="Plantae" order="Rosales" pageNumber="48" phylum="Tracheophyta" rank="genus">Cecropia</taxonomicName>
|
|
sapling at Estacion Pitilla in the Guanacaste Conservation Area, and Joel Dunn found an incipient colony in a
|
|
<taxonomicName id="060759ACF805E8753BB9F9E4D0F9C844" class="Magnoliopsida" family="Cordiaceae" genus="Cordia" higherTaxonomySource="CoL" kingdom="Plantae" order="Boraginales" pageNumber="48" phylum="Tracheophyta" rank="species" species="bicolor">Cordia bicolor</taxonomicName>
|
|
plant near Monteverde.
|
|
</paragraph>
|
|
<paragraph id="F03AEB0DFECE5C0DCCF98458BC9335EA" pageNumber="48">
|
|
The queens are generally setose ants, with abundant fine flexuous setae on most body parts. An exception was the queen from the
|
|
<taxonomicName id="CD817A90A5EBE4E17D275D83FC084990" class="Magnoliopsida" family="Rubiaceae" genus="Chimarrhis" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageNumber="48" phylum="Tracheophyta" rank="genus">Chimarrhis</taxonomicName>
|
|
plant described above. This queen was almost completely devoid of erect setae anywhere on the body, including the tibiae, and the pubescence was sparse and patchy. When collecting from this colony, it took me more than an hour to locate the colony center, chop into the
|
|
<taxonomicName id="31D5EEF254DB7DA736BC7894D159FAD0" class="Magnoliopsida" family="Rubiaceae" genus="Chimarrhis" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageNumber="48" phylum="Tracheophyta" rank="genus">Chimarrhis</taxonomicName>
|
|
tree, and extract the physogastric queen. This is probably the oldest queen I have examined, the others being alates or foundresses in incipient colonies (and thus more easy to locate and collect). It may be that once queens establish and become sedentary in the middle of a large colony, they gradually become depilated, perhaps through constant grooming by workers.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AFD5A70AF1FCD2075D9197C722D0308C" type="discussion">
|
|
<paragraph id="4E1EB84722D023E5F81FCD247C4D8A9D" pageNumber="48">
|
|
Comments. This species is very similar to
|
|
<taxonomicName id="30BCF7E15EA263B8ED65DA744B813B08" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="48" phylum="Arthropoda" rank="species" species="sericea">A. sericea</taxonomicName>
|
|
, differing in the presence of erect setae on the posterior margin of the head.
|
|
<taxonomicName id="C7BF07E3348A44DDC874C98AD806D7B8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="48" phylum="Arthropoda" rank="species" species="sericea">Azteca sericea</taxonomicName>
|
|
is known from Mexico and Guatemala, where it is associated with myrmecophytic orchids. See further discussion of
|
|
<taxonomicName id="13FDE0E165E256F9B29855C0B8624DBD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26022" class="Insecta" family="Formicidae" genus="Azteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Azteca sericea (Mayr)" order="Hymenoptera" pageNumber="48" phylum="Arthropoda" rank="species" species="sericea">A. sericea</taxonomicName>
|
|
in the Addendum.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D2EE918AB6106D549532FFC3C4E55F0C" type="materials_examined">
|
|
<paragraph id="A2B32CEA90BDDD3C9427E4A1C97753C4" pageNumber="48">
|
|
Additional material examined.
|
|
<materialsCitation id="6F38207EE7715C7151FEB116773AC7FB" ID-GBIF-Occurrence="923890772" collectingDate="1963-07-06" collectionCode="USNM" collectorName="D. H. Janzen" country="COSTA RICA" latitude="10.516666" location="Florencia" locationDeviation="11 mi N" longitude="-84.48333" stateProvince="Alajuela">
|
|
<collectingCountry id="B9886B05E78F888CAE9C4A225721BF06">COSTA RICA</collectingCountry>
|
|
:
|
|
<collectingRegion id="8A3A6D1369B06072F1A703E5BE606551">Alajuela</collectingRegion>
|
|
:
|
|
<locationDeviation id="20004C733B7338C4918D7FB505C329AF">11mi N</locationDeviation>
|
|
<location id="AB68233DA49DF45A082B8A8384E71736" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:AB68233DA49DF45A082B8A8384E71736" country="COSTA RICA" latitude="10.516666" longitude="-84.48333" name="Florencia" stateProvince="Alajuela">Florencia</location>
|
|
,
|
|
<geoCoordinate id="A8B073C1C8A889BD661F9481125E989D" direction="north" orientation="latitude" value="10.516666">10°31'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="A8CE9A27C5B91EA695B89B0609AD0E1E" direction="west" orientation="longitude" value="-84.48333">84°29'W</geoCoordinate>
|
|
,
|
|
<date id="9A8B34DCB1EE42827E8B9DBC1839DED7" day="06" month="07" value="1963-07-06" year="1963">
|
|
<collectingDate id="E4D3BCC0D81C10FC7AB0F8125209EEBC" day="06" month="07" value="1963-07-06" year="1963">6 Jul 1963</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="E31CC61CD4E94584CE9B37F24F80A1FC">D. H. Janzen</collectorName>
|
|
) - workers [
|
|
<collectionCode id="94F8A352755188E214A64AB554E6D6F6" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="91CB625D778781C8CD301962C4E67FB2" ID-GBIF-Occurrence="923890766" collectingDate="1988" collectionCode="MUCR" collectorName="unknown" country="Costa Rica" latitude="10.983334" location="Cerro El Hacha, Guanacaste Conservation Area, 300 m" longitude="-85.55" stateProvince="Guanacaste">
|
|
<collectingRegion id="919E5E0F8F4071698468ED305EE8C3D4">Guanacaste</collectingRegion>
|
|
:
|
|
<location id="1C20DC114E5E65193EC035490A4AD41F" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:1C20DC114E5E65193EC035490A4AD41F" country="Costa Rica" latitude="10.983334" longitude="-85.55" name="Cerro El Hacha, Guanacaste Conservation Area" stateProvince="Guanacaste">Cerro El Hacha, Guanacaste Conservation Area</location>
|
|
,
|
|
<geoCoordinate id="2EC370A9D30545E51CF886D02F8DD873" direction="north" orientation="latitude" value="10.983334">10°59'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EB03214A5C0D4F3F68DCF035E054A155" direction="west" orientation="longitude" value="-85.55">85°33'W</geoCoordinate>
|
|
,
|
|
<location id="9DCA7EDC24DDB59D7B96F8BC4F5205F8" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:9DCA7EDC24DDB59D7B96F8BC4F5205F8" country="Costa Rica" latitude="10.983334" longitude="-85.55" name="300 m" stateProvince="Guanacaste">300m</location>
|
|
,
|
|
<collectingDate id="B10030D529A4862C2EB2E55A591A17C8">1988</collectingDate>
|
|
(
|
|
<collectorName id="5838AA09B501087AC8BC1BC71ED19481">unknown</collectorName>
|
|
) - alate queen [
|
|
<collectionCode id="09860049D8002B30968ABCE1F3C43FBD" collectionName="Costa Rica, Ciudad Universitaria, Universidad de Costa Rica, Museo de Insectos">MUCR</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="13781FC12D89C4C0A223CF451EE40D17" ID-GBIF-Occurrence="923890758" collectingDate="1991-01-24" collectorName="J. Longino" country="Costa Rica" latitude="10.983334" location="Estacion Pitilla, Guanacaste Cons. Area, 700 m" longitude="-85.433334" stateProvince="Guanacaste">
|
|
<location id="E12ED3178A2EB12B384E933B312843BF" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:E12ED3178A2EB12B384E933B312843BF" country="Costa Rica" latitude="10.983334" longitude="-85.433334" name="Estacion Pitilla, Guanacaste Cons. Area" stateProvince="Guanacaste">Estacion Pitilla, Guanacaste Cons. Area</location>
|
|
,
|
|
<geoCoordinate id="BD9E5958795315B92DC4D8D9F1CA04A1" direction="north" orientation="latitude" value="10.983334">10°59'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="26F6829EA8D053BF150871732173DC0B" direction="west" orientation="longitude" value="-85.433334">85°26'W</geoCoordinate>
|
|
,
|
|
<location id="03268C7DA3DE7369D5DD154A954347D5" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:03268C7DA3DE7369D5DD154A954347D5" country="Costa Rica" latitude="10.983334" longitude="-85.433334" name="700 m" stateProvince="Guanacaste">700m</location>
|
|
,
|
|
<date id="4689373C7E7C0F81576656806AC55B0C" day="24" month="01" value="1991-01-24" year="1991">
|
|
<collectingDate id="DDBACCC4522F0B432DEC7A5455BAE829" day="24" month="01" value="1991-01-24" year="1991">24 Jan 1991</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="EB801261AD229B2D7D3C519848D7DAC3">J. Longino</collectorName>
|
|
) - dealate queen
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="ABF05E3D54B469D880B8697AFA5EA9C9" ID-GBIF-Occurrence="923890762" collectingDate="2003-04-17" collectorName="ALAS" country="Costa Rica" latitude="10.333333" location="La Virgen, 500 m" locationDeviation="11 km SE" longitude="-84.066666" stateProvince="Heredia">
|
|
<collectingRegion id="8DA149A2FC66E1D56A032D80059C0B3F">Heredia</collectingRegion>
|
|
:
|
|
<locationDeviation id="5210B903D227E58A8751D3D8AA4D5283">11km SE</locationDeviation>
|
|
<location id="462332C6010C7E16E2612CDF6AE2220D" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:462332C6010C7E16E2612CDF6AE2220D" country="Costa Rica" latitude="10.333333" longitude="-84.066666" name="La Virgen" stateProvince="Heredia">La Virgen</location>
|
|
,
|
|
<geoCoordinate id="E3D2BDD18B6B4A3CC911AEF50AAF59BF" direction="north" orientation="latitude" value="10.333333">10°20'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="6DF1C0DE1D53B649F0A35790ACF56AAB" direction="west" orientation="longitude" value="-84.066666">84°04'W</geoCoordinate>
|
|
,
|
|
<location id="57229B92B8724608249C3A9415DB796C" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:57229B92B8724608249C3A9415DB796C" country="Costa Rica" latitude="10.333333" longitude="-84.066666" name="500 m" stateProvince="Heredia">500m</location>
|
|
,
|
|
<date id="84EAF2555A5879B4B248D8F44E40F7F7" day="17" month="04" value="2003-04-17" year="2003">
|
|
<collectingDate id="BFE4E5F4FE1F16EE01E8978A2B799FF7" day="17" month="04" value="2003-04-17" year="2003">17 Apr 2003</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="E224B0009A7BDEBC0E7CEF457BF0F6A6">ALAS</collectorName>
|
|
) - alate queen
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="94B15CF9FD27F986AD392F8AD9A4C337" ID-GBIF-Occurrence="923890763" collectingDate="1925-07-11" collectorName="F. Nevermann" country="Costa Rica" latitude="10.25" location="Hamburg Farm, 50 m" longitude="-83.45" stateProvince="Limon">
|
|
<collectingRegion id="AD2E9D33A4BFA21BD59CEBE6639520B2">Limon</collectingRegion>
|
|
:
|
|
<location id="7C9EE589F9F3D8B55B256DFBDF0B1465" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:7C9EE589F9F3D8B55B256DFBDF0B1465" country="Costa Rica" latitude="10.25" longitude="-83.45" name="Hamburg Farm" stateProvince="Limon">Hamburg Farm</location>
|
|
,
|
|
<geoCoordinate id="A05358CF892B270AC9B8C4B701C7D1CD" direction="north" orientation="latitude" value="10.25">10°15'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="25FB33061D82BE59559A49C9E615150A" direction="west" orientation="longitude" value="-83.45">83°27'W</geoCoordinate>
|
|
,
|
|
<location id="13D9AF59DE2F787C1018B664E9C33CD3" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:13D9AF59DE2F787C1018B664E9C33CD3" country="Costa Rica" latitude="10.25" longitude="-83.45" name="50 m" stateProvince="Limon">50m</location>
|
|
,
|
|
<date id="A616A1DACF4B8503D1D7203878FE0CF2" day="11" month="07" value="1925-07-11" year="1925">
|
|
<collectingDate id="D2453E085702CB4A8BBA77BB8610423E" day="11" month="07" value="1925-07-11" year="1925">11 Jul 1925</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="E1FDC310A81F4AA1EF7860B2AF01611F">F. Nevermann</collectorName>
|
|
) - workers, alate queens, males
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="359F06DA33070167A0D37DFB52226794" ID-GBIF-Occurrence="923890756" collectorName="J. Longino" country="Costa Rica" latitude="8.483334" location="Sirena, Corcovado National Park, 5 m" longitude="-83.6" stateProvince="Puntarenas">
|
|
<collectingRegion id="803F18DC7E0D2437607189A012744BBA">Puntarenas</collectingRegion>
|
|
:
|
|
<location id="0DB075A4062FAE11E158D2298CF96FDD" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:0DB075A4062FAE11E158D2298CF96FDD" country="Costa Rica" latitude="8.483334" longitude="-83.6" name="Sirena, Corcovado National Park" stateProvince="Puntarenas">Sirena, Corcovado National Park</location>
|
|
,
|
|
<geoCoordinate id="E3B18436093E928B3A13F6353FEA7506" direction="north" orientation="latitude" value="8.483334">8°29'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="CF4656235A591B042277F75CE4C71513" direction="west" orientation="longitude" value="-83.6">83°36'W</geoCoordinate>
|
|
,
|
|
<location id="AF2CE368A1DF37DC6F0EFD45BE59ED4F" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:AF2CE368A1DF37DC6F0EFD45BE59ED4F" country="Costa Rica" latitude="8.483334" longitude="-83.6" name="5 m" stateProvince="Puntarenas">5m</location>
|
|
, numerous collections and dates (
|
|
<collectorName id="8C773B11A9B6CEAD7B2162169B8D6293">J. Longino</collectorName>
|
|
) - workers, males;
|
|
</materialsCitation>
|
|
<materialsCitation id="525261A096FF4E907270E114903907E2" ID-GBIF-Occurrence="923890797" collectingDate="1990-10" collectionCode="INBC" collectorName="J. Longino" country="Costa Rica" latitude="8.483334" location="Sirena, Corcovado National Park, 5 m" longitude="-83.6" stateProvince="Puntarenas">
|
|
same locality,
|
|
<date id="35E1D6363229135AAB850094A4C13092" month="10" value="1990-10" year="1990">
|
|
<collectingDate id="1F4DF7839E6E60E983AE1029FDA696F7" month="10" value="1990-10" year="1990">Oct 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="A4EDC17821ACFF6C6D99541C3BEF408C">Saborio</collectorName>
|
|
) - alate queen [
|
|
<collectionCode id="69969CBDDDA6C1B5239C3720178308AA" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="EF1138EF9BAD7E8B2E959B5374597AB8" ID-GBIF-Occurrence="923890785" collectingDate="1981-12-04" collectorName="J. Longino" country="Costa Rica" latitude="8.633333" location="Los Planes, Corcovado National Park, 100 m" longitude="-83.666664" stateProvince="Puntarenas">
|
|
<location id="2DB1AA980BA646023278A37C739BE4E9" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:2DB1AA980BA646023278A37C739BE4E9" country="Costa Rica" latitude="8.633333" longitude="-83.666664" name="Los Planes, Corcovado National Park" stateProvince="Puntarenas">Los Planes, Corcovado National Park</location>
|
|
,
|
|
<geoCoordinate id="821245F71777DBD6B17A8D90419DCC78" direction="north" orientation="latitude" value="8.633333">8°38'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C1F4EF872BD9BFCC36E7777AA1C52C69" direction="west" orientation="longitude" value="-83.666664">83°40'W</geoCoordinate>
|
|
,
|
|
<location id="8945DB43978D4D161A942729795227CF" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:8945DB43978D4D161A942729795227CF" country="Costa Rica" latitude="8.633333" longitude="-83.666664" name="100 m" stateProvince="Puntarenas">100m</location>
|
|
,
|
|
<date id="1045F3BC61F5B9B64DFA2EE0DA051895" day="04" month="12" value="1981-12-04" year="1981">
|
|
<collectingDate id="1BD361E93A11FEEFE3F606CA4DAF23CD" day="04" month="12" value="1981-12-04" year="1981">4 Dec 1981</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="D8DFEEBC5EDFF3B4442BF4D463AABC81">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="B3FA377DA22322153B05DBC6C12ABE07" ID-GBIF-Occurrence="923890804" collectingDate="1981-03-22" collectorName="J. Longino" country="Costa Rica" latitude="8.616667" location="San Pedrillo, Corcovado National Park, 5 m" longitude="-83.73333" stateProvince="Puntarenas">
|
|
<location id="7E13EDEBE68D20C257664AA8357806DC" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:7E13EDEBE68D20C257664AA8357806DC" country="Costa Rica" latitude="8.616667" longitude="-83.73333" name="San Pedrillo, Corcovado National Park" stateProvince="Puntarenas">San Pedrillo, Corcovado National Park</location>
|
|
,
|
|
<geoCoordinate id="6F4E9D1CBA46797A46F365649318BD03" direction="north" orientation="latitude" value="8.616667">8°37'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="D909AC25EEE4659522067FB04EC4A4E4" direction="west" orientation="longitude" value="-83.73333">83°44'W</geoCoordinate>
|
|
,
|
|
<location id="97FC97824FBDE981D8448339255733B2" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:97FC97824FBDE981D8448339255733B2" country="Costa Rica" latitude="8.616667" longitude="-83.73333" name="5 m" stateProvince="Puntarenas">5m</location>
|
|
,
|
|
<date id="97859A4AD7A1184D2ED87A1449974655" day="22" month="03" value="1981-03-22" year="1981">
|
|
<collectingDate id="F70E401A06F48833014FD674DAE5A5D8" day="22" month="03" value="1981-03-22" year="1981">22 Mar 1981</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="A2A498D33B344127B61D50A0A5D0C25C">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="6FA7071028DAAE6D17C074336F8B4D88" ID-GBIF-Occurrence="923890788" collectingDate="1982-01-02" collectorName="J. Longino" country="Costa Rica" latitude="8.583333" location="Llorona, Corcovado National Park, 5 m" longitude="-83.7" stateProvince="Puntarenas">
|
|
<location id="0EA2E265D8A65C2D038E35984D4A9D8A" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:0EA2E265D8A65C2D038E35984D4A9D8A" country="Costa Rica" latitude="8.583333" longitude="-83.7" name="Llorona, Corcovado National Park" stateProvince="Puntarenas">Llorona, Corcovado National Park</location>
|
|
,
|
|
<geoCoordinate id="634B6663C94B8A149F1D28D4C87C018E" direction="north" orientation="latitude" value="8.583333">8°35'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="139DA75F167DDBEC055A3288D0F670C4" direction="west" orientation="longitude" value="-83.7">83°42'W</geoCoordinate>
|
|
,
|
|
<location id="5F2185FAC66BB1479C41CC45BDF43CE5" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:5F2185FAC66BB1479C41CC45BDF43CE5" country="Costa Rica" latitude="8.583333" longitude="-83.7" name="5 m" stateProvince="Puntarenas">5m</location>
|
|
,
|
|
<date id="8F52DA729E96ED22842E3517DF06B139" day="02" month="01" value="1982-01-02" year="1982">
|
|
<collectingDate id="922E50B1210D3692D1E6275B5DDFC72A" day="02" month="01" value="1982-01-02" year="1982">2 Jan 1982</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="8FE3AE2B9310FB738430ECD97208594F">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="8CA17A6F2E13BE1D6622EA1F27777E09" ID-GBIF-Occurrence="923890799" collectingDate="1990-06-04" collectorName="J. Longino" country="Costa Rica" latitude="10.283334" location="Rio San Luis, 850 m" longitude="-84.8" stateProvince="Puntarenas">
|
|
<location id="6797438DDC2977194F06FA22390BF297" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:6797438DDC2977194F06FA22390BF297" country="Costa Rica" latitude="10.283334" longitude="-84.8" name="Rio San Luis" stateProvince="Puntarenas">Rio San Luis</location>
|
|
,
|
|
<geoCoordinate id="C80CF73620D013E070EB3323BFE9BFB6" direction="north" orientation="latitude" value="10.283334">10°17'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C99F099F0F215B97A6D5EDD7B8C180C1" direction="west" orientation="longitude" value="-84.8">84°48'W</geoCoordinate>
|
|
,
|
|
<location id="78AD63346E69C2D5F39727D39FF5F658" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:78AD63346E69C2D5F39727D39FF5F658" country="Costa Rica" latitude="10.283334" longitude="-84.8" name="850 m" stateProvince="Puntarenas">850m</location>
|
|
,
|
|
<date id="6FD54DFE6E1DF76BA2479A3EA5A2CD8A" day="04" month="06" value="1990-06-04" year="1990">
|
|
<collectingDate id="27A9DBDA8A3A2D2CB9621DC63A90C481" day="04" month="06" value="1990-06-04" year="1990">4 Jun 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="C97D7E6E810674C0B18834D5F70995B3">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="9656C9E77B34A576A0EAD5121C89529A" ID-GBIF-Occurrence="923890787" collectingDate="1990-03-23" collectorName="J. Longino" country="Costa Rica" latitude="8.783334" location="Wilson Botanical Garden, 4 km S San Vito, 1200 m" longitude="-82.96667" stateProvince="Puntarenas">
|
|
<location id="330B9B71C76FA31E4C4BC91E335D16EE" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:330B9B71C76FA31E4C4BC91E335D16EE" country="Costa Rica" latitude="8.783334" longitude="-82.96667" name="Wilson Botanical Garden, 4 km S San Vito" stateProvince="Puntarenas">Wilson Botanical Garden, 4km S San Vito</location>
|
|
,
|
|
<geoCoordinate id="6A771C7FCA4408E0CEB7439C85253D5D" direction="north" orientation="latitude" value="8.783334">8°47'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="36E35BD203758220C8CD7C76752F7ED3" direction="west" orientation="longitude" value="-82.96667">82°58'W</geoCoordinate>
|
|
,
|
|
<location id="2380753D49AF5A5F48B9716DF7C7A53D" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:2380753D49AF5A5F48B9716DF7C7A53D" country="Costa Rica" latitude="8.783334" longitude="-82.96667" name="1200 m" stateProvince="Puntarenas">1200m</location>
|
|
,
|
|
<date id="5719813C9DA10D24DBD122E5A95CBAD2" day="23" month="03" value="1990-03-23" year="1990">
|
|
<collectingDate id="D8D7C541357ADEE47ACDE5FF4690DEE1" day="23" month="03" value="1990-03-23" year="1990">23 Mar 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="E037DDCCA1CAC7F941ADA4EF862F2923">J. Longino</collectorName>
|
|
) - workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="4119DB462CC73A8E243C269696004491" ID-GBIF-Occurrence="923890795" collectingDate="1990-11-30" collectionCode="INBC" collectorName="B. Apu" country="Costa Rica" latitude="8.7" location="Rancho Quemado, Osa Peninsula, 200 m" longitude="-83.55" stateProvince="Puntarenas">
|
|
<location id="44FC8A9FD4F68C9A103FB667BB85AABB" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:44FC8A9FD4F68C9A103FB667BB85AABB" country="Costa Rica" latitude="8.7" longitude="-83.55" name="Rancho Quemado, Osa Peninsula" stateProvince="Puntarenas">Rancho Quemado, Osa Peninsula</location>
|
|
,
|
|
<geoCoordinate id="825762B1843A0EDB2E3EC58AD998A8A9" direction="north" orientation="latitude" value="8.7">8°42'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="831ECF67644019D631F95E43DBED963A" direction="west" orientation="longitude" value="-83.55">83°33'W</geoCoordinate>
|
|
,
|
|
<location id="4B5FD3A7715A99DBEB7D82A0497DE90B" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:4B5FD3A7715A99DBEB7D82A0497DE90B" country="Costa Rica" latitude="8.7" longitude="-83.55" name="200 m" stateProvince="Puntarenas">200m</location>
|
|
,
|
|
<date id="110F3AFFACB4E30B89DFCF697ECB0BE1" day="01" month="10">
|
|
<collectingDate id="B5F5D41A8EF3D6634098FC57511A2393" day="01" month="10">1 Oct-</collectingDate>
|
|
</date>
|
|
<date id="D4BA6CF69C36A0D0408912871B792A81" day="30" month="11" value="1990-11-30" year="1990">
|
|
<collectingDate id="721E89DF3A799D3C35A0723C76C10B86" day="30" month="11" value="1990-11-30" year="1990">30 Nov 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="A34C102B2B6222ADBA2400DF1D6BA570">B. Apu</collectorName>
|
|
) - alate queen [
|
|
<collectionCode id="BA5B5DBF4A69475970A1183FBC4FE004" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="C9ED8655706DE1CAB579101EB14D3B78" ID-GBIF-Occurrence="923890786" collectingDate="1990-12-15" collectionCode="INBC" collectorName="J. Longino" country="Costa Rica" latitude="8.7" location="Rancho Quemado, Osa Peninsula, 200 m" longitude="-83.55" stateProvince="Puntarenas">
|
|
same data,
|
|
<date id="90F0AF869DA85103191D7AFEF8CC837D" day="15" month="12" value="1990-12-15" year="1990">
|
|
<collectingDate id="1F8AC49BCB09D4A313E76F2DCB2A8ECA" day="15" month="12" value="1990-12-15" year="1990">15 Dec 1990</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="C623FB1B03EE7E13A494751EAB0B3422">J. Longino</collectorName>
|
|
) - queen and workers
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="1F94E125A94703B7CA6CF3FFE58BDDF0" ID-GBIF-Occurrence="923890791" collectingDate="1957-07-23" collectionCode="LACM" collectorName="Truxal, Menke" country="Costa Rica" latitude="8.65" location="Golfito, 50 m" longitude="-83.15" stateProvince="Puntarenas">
|
|
<location id="477FEE8BFF1DFF1E0C023598C5546555" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:477FEE8BFF1DFF1E0C023598C5546555" country="Costa Rica" latitude="8.65" longitude="-83.15" name="Golfito" stateProvince="Puntarenas">Golfito</location>
|
|
,
|
|
<geoCoordinate id="8DC1D1AE65775F3D7BA45A68F99A1DC2" direction="north" orientation="latitude" value="8.65">8°39'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="3815F122DDAA17A7EA64D70EF1FC3A1B" direction="west" orientation="longitude" value="-83.15">83°09'W</geoCoordinate>
|
|
,
|
|
<location id="22CEDB14BD0B55D33ACC53EE3043F453" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:22CEDB14BD0B55D33ACC53EE3043F453" country="Costa Rica" latitude="8.65" longitude="-83.15" name="50 m" stateProvince="Puntarenas">50m</location>
|
|
,
|
|
<date id="51C09B76ADFFC5560436081DD10E2092" day="23" month="07" value="1957-07-23" year="1957">
|
|
<collectingDate id="C24431EB6079AC34F7C2E896CC5714E4" day="23" month="07" value="1957-07-23" year="1957">23 Jul 1957</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="0F66A07E8AD7E825B7F944ACAEE90E5F">Truxal & Menke</collectorName>
|
|
) - alate queen [
|
|
<collectionCode id="B9364C131C1F15E71409A8BFCDAD7CDA" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="2039A7A5C0A86B93D7B3204033E5DE20" ID-GBIF-Occurrence="923890778" collectingDate="1999-02-07" collectorName="J. Dunn" country="Costa Rica" latitude="10.3" location="Monteverde, 1400 m" longitude="-84.8" stateProvince="Puntarenas">
|
|
<location id="860663431A210EF2A0CB2FECCD4511AB" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:860663431A210EF2A0CB2FECCD4511AB" country="Costa Rica" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
|
|
,
|
|
<geoCoordinate id="A3FB97D64BFFF0D96B7121518CF44DE1" direction="north" orientation="latitude" value="10.3">10°18'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="A925A8A0BD4D768AC5DA789A926B1FD5" direction="west" orientation="longitude" value="-84.8">84°48'W</geoCoordinate>
|
|
,
|
|
<location id="B03ED39230E47A17DCCAAD968D411E8C" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:B03ED39230E47A17DCCAAD968D411E8C" country="Costa Rica" latitude="10.3" longitude="-84.8" name="1400 m" stateProvince="Puntarenas">1400m</location>
|
|
,
|
|
<date id="FED7A94E95006AC9D1B18110E0D53ACA" day="07" month="02" value="1999-02-07" year="1999">
|
|
<collectingDate id="9EA7E6032B3F968C0B61875B48D9D45C" day="07" month="02" value="1999-02-07" year="1999">7 Feb 1999</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="21F463FFEA02E55D7279C133FFC9FDE9">J. Dunn</collectorName>
|
|
) - queen and workers
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="A42F210C38338FC1CA655BD2A844984B" ID-GBIF-Occurrence="923890775" collectionCode="USNM" country="GUATEMALA" location="no specific locality">
|
|
<collectingCountry id="57A88F8B646D993A980AF1577A3F2E68">GUATEMALA</collectingCountry>
|
|
:
|
|
<location id="3169AC5B788717C34714E265752EBA8B" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:3169AC5B788717C34714E265752EBA8B" country="GUA- TEMALA" name="no specific locality">no specific locality</location>
|
|
- dealate queen [
|
|
<collectionCode id="8439927C4A8B914E7EEAA2109800BE4D" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="9BBD857CEC05F84CD8400D08600855F3" ID-GBIF-Occurrence="923890768" collectingDate="1924-07-12" collectorName="G C. Wheeler" country="PANAMA" location="Las Cascadas, cacao plantation" stateProvince="Canal Zone">
|
|
<collectingCountry id="A8C5ABA4D8BE1031645425070FCA305D">PANAMA</collectingCountry>
|
|
:
|
|
<collectingRegion id="8FB1F0EB85DA42FFB6CBC98B7EA69472">Canal Zone</collectingRegion>
|
|
:
|
|
<location id="6A08CBAC325A30D8633546C87270DD0E" LSID="urn:lsid:plazi:treatment:431358CAECCE0833C6AB79289BD0B271:6A08CBAC325A30D8633546C87270DD0E" country="PANAMA" name="Las Cascadas, cacao plantation" stateProvince="Canal Zone">Las Cascadas, cacao plantation</location>
|
|
,
|
|
<date id="CC26DC2419FD82901F30D3ECC3E41480" day="12" month="07" value="1924-07-12" year="1924">
|
|
<collectingDate id="201A9FA96B8EC846FA335998CF019879" day="12" month="07" value="1924-07-12" year="1924">12 Jul 1924</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="A69DB0B7A38C6CFE5BFF19A9F04AEF94">G C. Wheeler</collectorName>
|
|
) - alate queen, workers
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |