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<document id="270E86D88A3B8BD6F2430F40BC42F136" ID-CLB-Dataset="29023" ID-DOI="10.11646/zootaxa.4459.2.3" ID-GBIF-Dataset="d1b4add2-e5bc-4d08-9502-274bab438721" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458574" ID-ZooBank="49F5AF2B-E795-404B-A946-2C00DB6011B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288907869" checkinUser="plazi" docAuthor="Karaman, Ivo M. &amp; Horvatović, Mladen" docDate="2018" docId="404814486752FF97FF28FEC72759FBCC" docLanguage="en" docName="zootaxa.4459.2.3.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Titanethes Schioedte 1849" docType="treatment" docVersion="7" lastPageNumber="274" masterDocId="BC716C30675FFF9AFFBFFFB72642FFC6" masterDocTitle="Revision of the genera Cyphonethes Verhoeff, 1926 and Titanethes Schioedte, 1849 (Isopoda: Oniscoidea: Trichoniscidae) with a description of a new genus and three new taxa" masterLastPageNumber="284" masterPageNumber="261" pageNumber="274" updateTime="1698578336577" updateUser="ExternalLinkService">
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<mods:title id="2001882AE286A437F76A86B7BDD5C3A4">Revision of the genera Cyphonethes Verhoeff, 1926 and Titanethes Schioedte, 1849 (Isopoda: Oniscoidea: Trichoniscidae) with a description of a new genus and three new taxa</mods:title>
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<mods:namePart id="BE5E7397A88DB4FEF38108FA179EBE22">Karaman, Ivo M.</mods:namePart>
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<mods:date id="F2E0ADBD8FBB1404AA94FD02244A4ABD">2018</mods:date>
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<mods:number id="9966A64CDADDB54E6E5E373BD8C75E26">2018-08-15</mods:number>
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<treatment id="404814486752FF97FF28FEC72759FBCC" ID-DOI="http://doi.org/10.5281/zenodo.5983532" ID-GBIF-Taxon="148692698" ID-Zenodo-Dep="5983532" LSID="urn:lsid:plazi:treatment:404814486752FF97FF28FEC72759FBCC" httpUri="http://treatment.plazi.org/id/404814486752FF97FF28FEC72759FBCC" lastPageNumber="274" pageId="13" pageNumber="274">
<subSubSection id="80FBF6D56752FF97FF28FEC72477FE4C" box="[151,565,368,394]" pageId="13" pageNumber="274" type="nomenclature">
<paragraph id="C85EA55E6752FF97FF28FEC72477FE4C" blockId="13.[151,565,368,394]" box="[151,565,368,394]" pageId="13" pageNumber="274">
<heading id="931612326752FF97FF28FEC72477FE4C" bold="true" box="[151,565,368,394]" fontSize="11" level="1" pageId="13" pageNumber="274" reason="1">
<emphasis id="FA95794C6752FF97FF28FEC72477FE4C" bold="true" box="[151,565,368,394]" pageId="13" pageNumber="274">
Genus
<taxonomicName id="0FE1DEDD6752FF97FF52FEC72477FE4C" ID-CoL="7Q4JW" authority="Schioedte, 1849" authorityName="Schioedte" authorityYear="1849" box="[237,565,368,394]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="FA95794C6752FF97FF52FEC72728FE4C" bold="true" box="[237,362,368,394]" italics="true" pageId="13" pageNumber="274">Titanethes</emphasis>
Schioedte, 1849
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="80FBF6D56752FF97FF28FE01270DFCF4" pageId="13" pageNumber="274" type="materials_examined">
<paragraph id="C85EA55E6752FF97FF28FE01243BFE0A" blockId="13.[151,633,438,460]" box="[151,633,438,460]" pageId="13" pageNumber="274">
<emphasis id="FA95794C6752FF97FF28FE01275EFE0A" bold="true" box="[151,284,438,460]" pageId="13" pageNumber="274">Type species</emphasis>
.
<taxonomicName id="0FE1DEDD6752FF97FE96FE01243BFE0A" authority="(C. Koch, 1841)" authorityName="C. Koch" authorityYear="1841" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[297,633,438,460]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FE96FE01278DFE0A" box="[297,463,438,460]" italics="true" pageId="13" pageNumber="274">Titanethes albus</emphasis>
(C. Koch, 1841)
</taxonomicName>
</paragraph>
<paragraph id="C85EA55E6752FF97FF28FE4E270DFCF4" blockId="13.[151,1436,505,1034]" pageId="13" pageNumber="274">
<emphasis id="FA95794C6752FF97FF28FE4E27C4FDD4" bold="true" box="[151,390,505,530]" pageId="13" pageNumber="274">Species composition</emphasis>
.
<taxonomicName id="0FE1DEDD6752FF97FE2CFE4E250AFDD7" authority="albus (C. Koch, 1841)" authorityName="albus (C. Koch" authorityYear="1841" box="[403,840,505,530]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="subSpecies" species="albus" subSpecies="albus">
<emphasis id="FA95794C6752FF97FE2CFE4E24CFFDD4" box="[403,653,505,530]" italics="true" pageId="13" pageNumber="274">Titanethes albus albus</emphasis>
(C. Koch, 1841)
</taxonomicName>
and
<taxonomicName id="0FE1DEDD6752FF97FCC1FE4E22DEFDD7" authority="Karaman &amp; Horvatović, 2018" authorityName="Karaman &amp; Horvatović" authorityYear="2018" box="[894,1180,505,530]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="subSpecies" species="albus" status="subsp. nov." subSpecies="matovici">
<emphasis id="FA95794C6752FF97FCC1FE4E22DEFDD7" box="[894,1180,505,530]" italics="true" pageId="13" pageNumber="274">Titanethes albus matovici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1A6C4376752FF97FB1CFE4E22AEFDD4" box="[1187,1260,505,530]" pageId="13" pageNumber="274" rank="subSpecies">
<emphasis id="FA95794C6752FF97FB1CFE4E22FBFDD4" bold="true" box="[1187,1209,505,530]" pageId="13" pageNumber="274">n.</emphasis>
<taxonomicName id="0FE1DEDD6752FF97FB7FFE4E22AEFDD4" box="[1216,1260,505,530]" pageId="13" pageNumber="274" rank="subSpecies" subSpecies="described">
<emphasis id="FA95794C6752FF97FB7FFE4E22AEFDD4" bold="true" box="[1216,1260,505,530]" pageId="13" pageNumber="274">ssp.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
described here. The species
<taxonomicName id="0FE1DEDD6752FF97FE9CFDAA24CAFDF0" authority="Verhoeff, 1926" authorityName="Verhoeff" authorityYear="1926" box="[291,648,541,566]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FE9CFDAA279AFDF0" box="[291,472,541,566]" italics="true" pageId="13" pageNumber="274">Titanethes dahli</emphasis>
<bibRefCitation id="AC70D8AF6752FF97FE60FDAA24CAFDF0" box="[479,648,541,566]" pageId="13" pageNumber="274" refId="ref10131" refString="Verhoeff, K. (1926) Uber Isopoden der Balkanhalbinsel, gesammelt von Herrn Dr. Buresch. Mitteilungen der bulgarischen entomologischen Gesellschaft in Sofia, 3, 135 - 158." type="journal article">Verhoeff, 1926</bibRefCitation>
</taxonomicName>
here we treat as a junior synonym of
<taxonomicName id="0FE1DEDD6752FF97FB89FDAA22ACFDF0" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[1078,1262,541,566]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FB89FDAA22ACFDF0" box="[1078,1262,541,566]" italics="true" pageId="13" pageNumber="274">Titanethes albus</emphasis>
</taxonomicName>
although it has not yet formally been accepted. We treat the species
<taxonomicName id="0FE1DEDD6752FF97FCAEFDF52529FD9C" authorityName="Verhoeff" authorityYear="1926" box="[785,875,577,602]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FCAEFDF52529FD9C" box="[785,875,577,602]" italics="true" pageId="13" pageNumber="274">T. dahli</emphasis>
</taxonomicName>
as a &quot;
<taxonomicName id="0FE1DEDD6752FF97FC01FDF625BAFD9C" authorityName="Verhoeff" authorityYear="1926" box="[958,1016,577,602]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FC01FDF625BAFD9C" box="[958,1016,577,602]" italics="true" pageId="13" pageNumber="274">dahli</emphasis>
</taxonomicName>
&quot; form of the species
<taxonomicName id="0FE1DEDD6752FF97FABFFDF5231FFD9C" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[1280,1373,577,602]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FABFFDF5231FFD9C" box="[1280,1373,577,602]" italics="true" pageId="13" pageNumber="274">T. albus</emphasis>
</taxonomicName>
. The molecular study of
<bibRefCitation id="AC70D8AF6752FF97FECEFDD22474FDB8" box="[369,566,613,638]" pageId="13" pageNumber="274" refId="ref10012" refString="Taiti, S., Filippelli, M., Bedek, J. &amp; Baratti, M. (2011) Molecular and morphological analyses in two genera of Trichoniscidae with and without gland-piliferous Organs (Isopoda, Oniscidea). Proceedings of the 8 th International Symposium of Terrestrial Isopod Biology ISTIB, Bled, Slovenia, 19 - 23 June, 2011, 43 - 44." type="journal article">
Taiti
<emphasis id="FA95794C6752FF97FE14FDD1279EFDB8" box="[427,476,613,638]" italics="true" pageId="13" pageNumber="274">et al</emphasis>
. (2011)
</bibRefCitation>
, indicate
<taxonomicName id="0FE1DEDD6752FF97FD1DFDD124B9FDB8" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[674,763,613,638]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FD1DFDD124B9FDB8" box="[674,763,613,638]" italics="true" pageId="13" pageNumber="274">T. albus</emphasis>
</taxonomicName>
and
<taxonomicName id="0FE1DEDD6752FF97FC8EFDD125C5FDB8" authorityName="Verhoeff" authorityYear="1926" box="[817,903,613,638]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FC8EFDD125C5FDB8" box="[817,903,613,638]" italics="true" pageId="13" pageNumber="274">T. dahli</emphasis>
</taxonomicName>
as unique species with two forms, which could even appear syntopically. We hope to formally announce these findings in the near future. The &quot;
<taxonomicName id="0FE1DEDD6752FF97FB59FD3E2360FD64" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[1254,1314,649,674]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FB59FD3E2360FD64" box="[1254,1314,649,674]" italics="true" pageId="13" pageNumber="274">albus</emphasis>
</taxonomicName>
&quot; form has well-developed tergal glands and accompanying structures of males pleon tergite 4 (
<figureCitation id="50DAB9DB6752FF97FBE7FD1A22E9FD00" box="[1112,1195,685,710]" captionStart="FIGURE 35" captionStartId="12.[151,250,1121,1143]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@12.[188,1398,193,1100]" captionTargetPageId="12" captionText="FIGURE 35. Titanethes albus f. &quot;albus&quot;, male pleonite 4, lateral margin (SEM by Kostanjšek Rok &amp; Miloš Vittori)." httpUri="https://zenodo.org/record/1458596/files/figure.png" pageId="13" pageNumber="274">Fig. 35</figureCitation>
) while in the &quot;
<taxonomicName id="0FE1DEDD6752FF97FAE8FD1A23D2FD00" authorityName="Verhoeff" authorityYear="1926" box="[1367,1424,685,710]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FAE8FD1A23D2FD00" box="[1367,1424,685,710]" italics="true" pageId="13" pageNumber="274">dahli</emphasis>
</taxonomicName>
&quot; form (
<figureCitation id="50DAB9DB6752FF97FF63FD66271FFD2C" box="[220,349,721,746]" captionStart="FIGURES 36–37" captionStartId="12.[151,264,1691,1713]" captionTargetBox="[151,1436,1193,1669]" captionTargetId="figure@12.[151,1436,1193,1669]" captionTargetPageId="12" captionText="FIGURES 36–37. Titanethes albus f. &quot;dahli&quot;, male. 36, pleon and uropods. 37, pleonites 4–5, lateral margin." httpUri="https://zenodo.org/record/1458598/files/figure.png" pageId="13" pageNumber="274">Figs 36–37</figureCitation>
) they are missing. Results of
<bibRefCitation id="AC70D8AF6752FF97FD16FD66252EFD2C" box="[681,876,721,746]" pageId="13" pageNumber="274" refId="ref10012" refString="Taiti, S., Filippelli, M., Bedek, J. &amp; Baratti, M. (2011) Molecular and morphological analyses in two genera of Trichoniscidae with and without gland-piliferous Organs (Isopoda, Oniscidea). Proceedings of the 8 th International Symposium of Terrestrial Isopod Biology ISTIB, Bled, Slovenia, 19 - 23 June, 2011, 43 - 44." type="journal article">
Taiti
<emphasis id="FA95794C6752FF97FD5CFD652551FD2C" box="[739,787,721,746]" italics="true" pageId="13" pageNumber="274">et al</emphasis>
. (2011)
</bibRefCitation>
leave open the possibility that
<taxonomicName id="0FE1DEDD6752FF97FB76FD652363FD2C" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[1225,1313,721,746]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="albus">
<emphasis id="FA95794C6752FF97FB76FD652363FD2C" box="[1225,1313,721,746]" italics="true" pageId="13" pageNumber="274">T. albus</emphasis>
</taxonomicName>
(including
<taxonomicName id="0FE1DEDD6752FF97FF28FD4126AEFCC8" authorityName="Verhoeff" authorityYear="1926" box="[151,236,757,782]" class="Malacostraca" family="Trichoniscidae" genus="Titanethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="274" phylum="Arthropoda" rank="species" species="dahli">
<emphasis id="FA95794C6752FF97FF28FD4126AEFCC8" box="[151,236,757,782]" italics="true" pageId="13" pageNumber="274">T. dahli</emphasis>
</taxonomicName>
) could be a taxon complex, where the possession and structure of pleonite tergal glands are not relevant for their distinction.
</paragraph>
</subSubSection>
<subSubSection id="80FBF6D56752FF97FF78FC8A2759FBCC" pageId="13" pageNumber="274" type="diagnosis">
<paragraph id="C85EA55E6752FF97FF78FC8A2759FBCC" blockId="13.[151,1436,505,1034]" pageId="13" pageNumber="274">
<emphasis id="FA95794C6752FF97FF78FC8A2778FC90" bold="true" box="[199,314,829,854]" pageId="13" pageNumber="274">Diagnosis</emphasis>
. Large troglobite, colourless and blind animals. Distolateral part of maxilliped with rounded tip; terminal peniciles of maxilla 1 endopodite longer than endopodite diameter; exopodite of maxilla 1 with short teeth, terminal three teeth shorter than exopodite diameter; maxilla 2 carries short rod-like setae; male pleopod 1endopodite basal article longer than terminal one; male pleopod 1 exopodite protruded mediodistally; male pleopod 2 endopodite terminal article robust with sharp tip; basal article shorter than exopodite; exopodite trapezoidal.
</paragraph>
</subSubSection>
</treatment>
</document>
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