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<document id="8DB4A0D0899A3B2ABE2EBF9970A7A145" ID-DOI="10.11646/zootaxa.5351.5.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8392291" ID-ZooBank="C6C7769C-94F6-496A-9293-2F0E1E201330" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695993006982" checkinUser="plazi" docAuthor="Davranoglou, Leonidas-Romanos, Baňař, Petr, Kakiopoulos, George, Balázs, Attila, Avtzis, Dimitrios N. &amp; Hlaváč, Peter" docDate="2023" docId="03F487CEE47AFFC3FF2354C3FBDC4332" docLanguage="en" docName="zootaxa.5351.5.4.pdf" docOrigin="Zootaxa 5351 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.5.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afropselaphus taygetensis Davranoglou, Hlavac &amp; Banar 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="561" masterDocId="FFCDFFB6E478FFC0FFB4541CFFC84257" masterDocTitle="Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)" masterLastPageNumber="570" masterPageNumber="559" pageNumber="561" updateTime="1695993641248" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E600B31ACC27C741C0AD9DFCCDF3B0EE">Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)</mods:title>
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<mods:namePart id="D972E0B5AEC6CD0F96F12B678C2C6186">Davranoglou, Leonidas-Romanos</mods:namePart>
<mods:affiliation id="290CE6AA3758DE9FFBF280D7CA8457B9">Oxford University Museum of Natural History, University of Oxford, Parks Road, Oxford OX 1 3 PW, UK;</mods:affiliation>
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<mods:namePart id="286E885E6180589CBEDBDF4F8742EDF0">Baňař, Petr</mods:namePart>
<mods:affiliation id="76DE23E01481A6E8175F62E0DB3E6BF3">Department of Entomology, Moravian Museum, Zelný trh 6, 659 37 Brno, Czech Republic;</mods:affiliation>
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<mods:namePart id="C718282FF05F70DCF4AA31409926DF75">Kakiopoulos, George</mods:namePart>
<mods:nameIdentifier id="987CDE80742B83C64B0D44C98EB7EEDD" type="ORCID">0000-0003-3486-5099</mods:nameIdentifier>
<mods:affiliation id="BCCB71B2282AFC1A120A61720004DA8C">Ilidos 60 - 62 street, 115 27, Athens, Greece; https: // orcid. org / 0000 - 0003 - 3486 - 5099</mods:affiliation>
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<mods:namePart id="D05776D1D25EE22EC0706F7383748F52">Balázs, Attila</mods:namePart>
<mods:affiliation id="50A99ED91EE12F2500E900F7B7DECEBD">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Mendel University in Brno, Zemědělská 1, 613 00 Brno, Czech Republic;</mods:affiliation>
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<mods:namePart id="06B920C7EC59F649565E395B85DF5567">Avtzis, Dimitrios N.</mods:namePart>
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<mods:affiliation id="8E171E3FEA7DCF39F389FFF0FECCAC69">Forest Research Institute, Hellenic Agricultural Organization Demeter, 570 06 Vassilika-Thessaloniki, Greece; dimitrios. avtzis @ gmail. com; https: // orcid. org / 0000 - 0002 - 7772 - 6892</mods:affiliation>
<mods:nameIdentifier id="815D3241D86FC4DB0D581C2EA7AD74D3" type="email">dimitrios.avtzis@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5535010D60210CC69DFCB84225347AF3">Hlaváč, Peter</mods:namePart>
<mods:nameIdentifier id="1F7D1F63D4ADB139CCD87B41BCFD66F2" type="ORCID">0000-0001-5060-0811</mods:nameIdentifier>
<mods:affiliation id="C73AAB7D339B1FA5E053BA70A2429BB3">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice; peterclaviger @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
<mods:nameIdentifier id="97E93731793B937B99B7EB7AE8C97373" type="email">peterclaviger@gmail.com</mods:nameIdentifier>
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<mods:title id="8B9E2540CCA658CC364AD50B2113E7B3">Zootaxa</mods:title>
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<mods:date id="1368614642EEF21B7F52640D182DE0EC">2023</mods:date>
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<taxonomicName id="4C5D4D5BE47AFFC2FF2354C3FC9342AE" authority="Davranoglou, Hlavac &amp; Banar" authorityName="Davranoglou, Hlavac &amp; Banar" authorityYear="2023" box="[151,859,222,250]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47AFFC2FF2354C3FE1B42AE" bold="true" box="[151,467,223,250]" italics="true" pageId="2" pageNumber="561">Afropselaphus taygetensis</emphasis>
Davranoglou, Hlaváč &amp; Baňař
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,
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<paragraph id="8BE236D8E47AFFC2FF23551FFE4D434A" blockId="2.[151,938,222,285]" box="[151,389,259,285]" pageId="2" pageNumber="561">
(
<figureCitation id="13662A5DE47AFFC2FF2B551FFF32434A" box="[159,250,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="2" pageNumber="561">Figs 2A</figureCitation>
<figureCitation id="13662A5DE47AFFC2FEBE551FFEE5434A" box="[266,301,259,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="2" pageNumber="561">6A</figureCitation>
,
<figureCitation id="13662A5DE47AFFC2FE8C551FFEB3434A" box="[312,379,259,285]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="2" pageNumber="561">7A, B</figureCitation>
)
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<subSubSection id="C3476553E47AFFC2FF235557FCE243FA" pageId="2" pageNumber="561" type="materials_examined">
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<emphasis id="B929EACAE47AFFC2FF235557FEA74332" bold="true" box="[151,367,331,357]" pageId="2" pageNumber="561">Material studied.</emphasis>
<materialsCitation id="3B353C85E47AFFC2FE355557FAD943DE" collectingDate="2021-04-21" collectionCode="NMPC" collectorName="L. R. Davranoglou" country="Greece" elevation="493" latitude="37.172935" location="Mt. Taygetos" longLatPrecision="1" longitude="22.299969" pageId="2" pageNumber="561" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E6887AE47AFFC2FE355557FE384332" box="[385,496,331,357]" pageId="2" pageNumber="561">Holotype</typeStatus>
, ♁:
<collectingCountry id="F34A7648E47AFFC2FD825557FD644332" box="[566,684,331,357]" name="Greece" pageId="2" pageNumber="561">GREECE</collectingCountry>
</emphasis>
, with one label: „GREECE,
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<quantity id="4CA59B3DE47AFFC2FEEF5573FE5543DE" box="[347,413,367,393]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="2" pageNumber="561" unit="m" value="493.0">
<elevation id="0070D1EBE47AFFC2FEEF5573FE5543DE" box="[347,413,367,393]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="2" pageNumber="561" unit="m" value="493.0">493m</elevation>
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,
<date id="FFE31018E47AFFC2FE1D5573FDD743DE" box="[425,543,367,393]" pageId="2" pageNumber="561" value="2021-04-21">
<collectingDate id="EFA7E9F0E47AFFC2FE1D5573FDD743DE" box="[425,543,367,393]" pageId="2" pageNumber="561" value="2021-04-21">21.iv.2021</collectingDate>
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<taxonomicName id="4C5D4D5BE47AFFC2FD815573FD5143DE" box="[565,665,367,393]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="2" pageNumber="561" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EACAE47AFFC2FD815573FD5143DE" box="[565,665,367,393]" italics="true" pageId="2" pageNumber="561">Platanus</emphasis>
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litter /
<collectorName id="26A8530EE47AFFC2FD5F556CFC7E43DE" box="[747,950,367,394]" pageId="2" pageNumber="561">L.R. Davranoglou</collectorName>
lgt.” [white, printed] (
<collectionCode id="ED4CAE1DE47AFFC2FB075573FAC143DE" box="[1203,1289,367,393]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="561">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B353C85E47AFFC2FAAF5573FCEE43FA" collectingDate="2021-04-21" collectionCode="MMBC, PCPH" collectorName="L. R. Davranoglou" country="Greece" elevation="493" latitude="37.172935" location="Mt. Taygetos" longLatPrecision="1" longitude="22.299969" pageId="2" pageNumber="561" specimenCount="13" specimenCount-female="13" typeStatus="paratype">
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<typeStatus id="54E6887AE47AFFC2FAAF5573FA5F43DE" box="[1307,1431,367,393]" pageId="2" pageNumber="561">Paratypes</typeStatus>
:
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4 ♁♁,
<specimenCount id="9D5BFD51E47AFFC2FF50558FFEFA43F9" box="[228,306,403,430]" pageId="2" pageNumber="561" type="female">13 ♀♀</specimenCount>
, same data as for holotype (
<collectionCode id="ED4CAE1DE47AFFC2FDD8558FFD0F43FA" box="[620,711,403,429]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="2" pageNumber="561">MMBC</collectionCode>
,
<collectionCode id="ED4CAE1DE47AFFC2FD67558FFCD643FA" box="[723,798,403,429]" pageId="2" pageNumber="561">PCPH</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<paragraph id="8BE236D8E47AFFC2FF7355ABFC52404D" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF7355ABFE8A4386" bold="true" box="[199,322,439,465]" pageId="2" pageNumber="561">Diagnosis.</emphasis>
Head 1.5 times longer than wide, scape 1.8 times longer than wide, tergite 1 (IV) 1.3 times wider than long, sternite 2 (IV)1.61.7 times wider than long; metaventrite smooth; internal sac of aedeagus with median spine subdivided into three processes that lack any distinct armature.
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47AFFC2FF73563FFCA7457E" pageId="2" pageNumber="561" type="description">
<paragraph id="8BE236D8E47AFFC2FF73563FFE9C409A" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF73563FFE91406A" bold="true" box="[199,345,547,573]" pageId="2" pageNumber="561">Description.</emphasis>
Body length
<quantity id="4CA59B3DE47AFFC2FE5A563FFD8E4069" box="[494,582,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="561" unit="mm" value="1.7">1.7 mm</quantity>
. Coloration largely reddish-brown, maxillary palpi and legs lighter, posterior margin of elytra much darker, almost black (
<figureCitation id="13662A5DE47AFFC2FD3E565BFD2D4036" box="[650,741,583,609]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="2" pageNumber="561">Fig. 2A</figureCitation>
). Pilosity of body sparse; head, legs, and antennae covered by sparse, adjacent golden setae; elytra with few rows of setae, which are denser on elytral marginal line; with row of setae on distal margin of elytra; proximal margin of tergite II with very dense pilosity (
<figureCitation id="13662A5DE47AFFC2FBEE5693FB7C40FE" box="[1114,1204,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="2" pageNumber="561">Fig. 2A</figureCitation>
), remaining segment largely glabrous.
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF7356CBFDEC4646" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
Head 1.5 times longer than wide, each eye composed between 57 ommatidia (
<figureCitation id="13662A5DE47AFFC2FB8056CBFB5F40A6" box="[1076,1175,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,250,1764,1788]" captionTargetBox="[325,1263,345,1741]" captionTargetId="figure-99@4.[325,1263,345,1741]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron microscopy image of dorsal view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392301" httpUri="https://zenodo.org/record/8392301/files/figure.png" pageId="2" pageNumber="561">Figs. 3A</figureCitation>
,
<figureCitation id="13662A5DE47AFFC2FB1656CBFB0040A6" box="[1186,1224,727,753]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="2" pageNumber="561">4A</figureCitation>
); frons with pair of strongly pronounced carinae that slightly surpass proximal margin of eyes, creating a deep frontal fovea (= rostral sulcus) (
<figureCitation id="13662A5DE47AFFC2FF4C5703FE9C416E" box="[248,340,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1764,1788]" captionTargetBox="[325,1263,345,1741]" captionTargetId="figure-99@4.[325,1263,345,1741]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron microscopy image of dorsal view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392301" httpUri="https://zenodo.org/record/8392301/files/figure.png" pageId="2" pageNumber="561">Fig. 3A</figureCitation>
); gular plate with distinct tuft of setae; antenna (
<figureCitation id="13662A5DE47AFFC2FC325703FC28416D" box="[902,992,799,826]" captionStart="FIGURE 5" captionStartId="6.[151,250,1008,1032]" captionTargetBox="[361,1225,181,984]" captionTargetId="figure-26@6.[361,1225,181,984]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron microscopy image of antenna in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392305" httpUri="https://zenodo.org/record/8392305/files/figure.png" pageId="2" pageNumber="561">Fig. 5A</figureCitation>
)
<quantity id="4CA59B3DE47AFFC2FC455703FB81416D" box="[1009,1097,799,826]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="561" unit="mm" value="0.9">0.9 mm</quantity>
long, with strongly foveolate surface; scape cylindrical, 1.8 times longer than wide, and 1.8 times longer than pedicel; pedicel about as long as wide; pedicel and antennomeres 38 rounded, subequal in length, about as long as wide; antennal club composed of three antennomeres: antennomeres 910 larger than antennomeres 38, longer than wide; terminal antennomere (11) as long as 910 combined, about 1.5 times longer than wide. Maxillary palpomere 1 shortest, palpomere 2 2.5 times longer than 1; palpomere 3 extremely short and rounded; palpomere 4 1.9 times longer than 2, apex club-shaped, main stem strongly curved.
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF735000FDD146F5" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
Pronotum
<quantity id="4CA59B3DE47AFFC2FE8F5007FE564661" box="[315,414,1051,1078]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="561" unit="mm" value="0.26">0.26 mm</quantity>
wide, smooth, about 1.3 times longer than wide, slightly constricted proximally and distally, broadest medially; about 1.1 times longer than head; lacking median sulcus and carinae (
<figureCitation id="13662A5DE47AFFC2FBDA5023FB00460E" box="[1134,1224,1087,1113]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="2" pageNumber="561">Fig. 6A</figureCitation>
); median antebasal fovea weakly impressed, lateral foveae present, weakly impressed; procoxal fovea and prothoracic-mesothoracic junction with a dense tuft of setae.
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF7350B7FC4A4766" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
Elytra about 1.5 times broader than long, and 1.2 times longer than pronotum; elytral surface smooth, lacking striae or distinct punctures; posterior margin with sparse, suberect pubescence; elytra with two basal foveae, and one sutural fovea; striae 13 weakly discernible, decreasing in depth from stria 1 to stria 3 and decreasingly distinct towards posterior portion of elytra; metaventrite smooth (
<figureCitation id="13662A5DE47AFFC2FCAF510BFCBD4766" box="[795,885,1303,1329]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="2" pageNumber="561">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF735127FD8E47B2" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">Abdomen about as broad as elytra; tergite 1 (IV) (excluding paratergites) smooth, medially convex, lacking median keel, about 1.3 times wider than long, its proximal margin covered by a tuft of dense, suberect setae; basal transverse sulcus distinct, covered by elytra in natural position; proximal margin of sternite 2 (IV) also covered by tuft of setae identical to dorsal counterpart; basolateral foveae not visible in dried specimens; remaining abdominal segments extremely narrow, strip-like.</paragraph>
<paragraph id="8BE236D8E47AFFC2FF7351F3FD134479" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
Aedeagus (
<figureCitation id="13662A5DE47AFFC2FEF251F3FE77445E" box="[326,447,1519,1545]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="2" pageNumber="561">Fig. 7A, B</figureCitation>
) with internal sac comprising central spine that is dorsally subdivided into three simple branches, and second spine devoid of any armature.
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF73522BFCA7457E" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF73522BFC794406" bold="true" box="[199,945,1591,1617]" pageId="2" pageNumber="561">
Measurements of
<typeStatus id="54E6887AE47AFFC2FE17522BFDC24406" box="[419,522,1591,1617]" pageId="2" pageNumber="561">holotype</typeStatus>
(in mm)—W = width, L = length.
</emphasis>
Body, L—1.73; head, L—0.37, W—0.25; antenna, L—0.89; scapus, L—0.14, W—0.07; pedicel, L—0.07, W—0.06; antennomere III, L—0.05; antennomere IV, L—0.05; antennomere V, L—0.05; antennomere VI, L—0.05; antennomere VII, L—0.05; antennomere VIII, L—0.05; antennomere IX, L—0.07; antennomere X, L—0.09; antennomere XI, L—0.15, W—0.1; maxillary palpomere I, L—0.07; maxillary palpomere II, L—0.19; maxillary palpomere III, L—0.05; maxillary palpomere IV, L—0.36; pronotum, L—0.34, W—0.26; elytra, L—0.44, W—0.32; tergite IV, L—0.44, W—0.69; tergite IV (excluding paratergites) W—0.57; ventrite IV, L—0.42, W—0.7.
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47AFFC2FF73532FFEA445C2" pageId="2" pageNumber="561" type="etymology">
<paragraph id="8BE236D8E47AFFC2FF73532FFB92451A" blockId="2.[151,1437,331,2014]" box="[199,1114,1843,1870]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF73532FFE86451A" bold="true" box="[199,334,1843,1869]" pageId="2" pageNumber="561">Etymology.</emphasis>
The new species is named after Taygetos Mountain, the
<typeStatus id="54E6887AE47AFFC2FC7D5328FC314519" box="[969,1017,1844,1870]" pageId="2" pageNumber="561">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BE236D8E47AFFC2FF73534BFEA445C2" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF73534BFEE24526" bold="true" box="[199,298,1879,1905]" pageId="2" pageNumber="561">Habitat.</emphasis>
<taxonomicName id="4C5D4D5BE47AFFC2FE84534BFE034526" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[304,459,1879,1905]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47AFFC2FE84534BFE034526" box="[304,459,1879,1905]" italics="true" pageId="2" pageNumber="561">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47AFFC2FE66534BFDD94526" bold="true" box="[466,529,1879,1905]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A21A57B1E47AFFC2FE66534BFDD94526" box="[466,529,1879,1905]" pageId="2" pageNumber="561" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
was collected from a dense accumulation of
<taxonomicName id="4C5D4D5BE47AFFC2FBB1534BFB124526" box="[1029,1242,1879,1905]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="2" pageNumber="561" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B929EACAE47AFFC2FBB1534BFB124526" box="[1029,1242,1879,1905]" italics="true" pageId="2" pageNumber="561">Platanus orientalis</emphasis>
</taxonomicName>
leaf-litter next to a stream (
<figureCitation id="13662A5DE47AFFC2FEB15367FE9745C2" box="[261,351,1915,1941]" captionStart="FIGURE 8" captionStartId="9.[151,250,1185,1209]" captionTargetBox="[344,1243,181,1161]" captionTargetId="figure-17@9.[344,1243,181,1161]" captionTargetPageId="9" captionText="FIGURE 8. Habitat photos of examined species. A) Type locality of A. taygetensis sp. n. in Kastoreion; B) type locality of A. tymficus sp. n.; in Mt Tymfi; C) leaf litter where A. euboicus sp. n. was collected from in Mt Dirfis, the type locality; D) habitat in Mt Vitsi, where Pselaphogenius treskanus (Karaman, 1940) was collected from." figureDoi="http://doi.org/10.5281/zenodo.8392315" httpUri="https://zenodo.org/record/8392315/files/figure.png" pageId="2" pageNumber="561">Fig. 8A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47AFFC2FF735383FCE445EE" box="[199,812,1951,1978]" pageId="2" pageNumber="561" type="distribution">
<paragraph id="8BE236D8E47AFFC2FF735383FCE445EE" blockId="2.[151,1437,331,2014]" box="[199,812,1951,1978]" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF735383FEA945EE" bold="true" box="[199,353,1951,1977]" pageId="2" pageNumber="561">Distribution.</emphasis>
<collectingCountry id="F34A7648E47AFFC2FED35383FE7E45EE" box="[359,438,1951,1977]" name="Greece" pageId="2" pageNumber="561">Greece</collectingCountry>
(so far endemic to Mt Taygetos).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47AFFC3FF7353DFFBDC4332" lastPageId="3" lastPageNumber="562" pageId="2" pageNumber="561" type="discussion">
<paragraph id="8BE236D8E47AFFC3FF7353DFFBDC4332" blockId="2.[151,1437,331,2014]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="562" pageId="2" pageNumber="561">
<emphasis id="B929EACAE47AFFC2FF7353DFFEF3458A" bold="true" box="[199,315,1987,2013]" pageId="2" pageNumber="561">Remarks.</emphasis>
The aedeagus of
<taxonomicName id="4C5D4D5BE47AFFC2FDBE53DFFD6F458A" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[522,679,1987,2013]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47AFFC2FDBE53DFFD6F458A" box="[522,679,1987,2013]" italics="true" pageId="2" pageNumber="561">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47AFFC2FD0553DFFD3B458A" bold="true" box="[689,755,1987,2013]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A21A57B1E47AFFC2FD0553DFFD3B458A" box="[689,755,1987,2013]" pageId="2" pageNumber="561" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is very similar to that of
<taxonomicName id="4C5D4D5BE47AFFC2FBAA53DFFAA4458A" authority="(Reitter, 1884)" baseAuthorityName="Reitter" baseAuthorityYear="1884" box="[1054,1388,1987,2013]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="caviventris">
<emphasis id="B929EACAE47AFFC2FBAA53DFFB72458A" box="[1054,1210,1987,2013]" italics="true" pageId="2" pageNumber="561">A. caviventris</emphasis>
(Reitter, 1884)
</taxonomicName>
, the most widely distributed species in the
<collectingRegion id="4999F83AE47BFFC3FDF7548BFD1942E6" box="[579,721,151,177]" country="Greece" name="Peloponnisos" pageId="3" pageNumber="562">Peloponnese</collectingRegion>
(
<figureCitation id="13662A5DE47BFFC3FD54548BFCE942E6" box="[736,801,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,847,871]" captionTargetBox="[151,1436,179,820]" captionTargetId="figure-17@1.[151,1436,179,822]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Afropselaphus species known from Greece." figureDoi="http://doi.org/10.5281/zenodo.8392293" httpUri="https://zenodo.org/record/8392293/files/figure.png" pageId="3" pageNumber="562">Fig. 1</figureCitation>
). However, several components of the aedeagal sclerites of
<taxonomicName id="4C5D4D5BE47BFFC3FF0D54A7FE9F4283" baseAuthorityName="Reitter" baseAuthorityYear="1884" box="[185,343,187,213]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="caviventris">
<emphasis id="B929EACAE47BFFC3FF0D54A7FE9F4283" box="[185,343,187,213]" italics="true" pageId="3" pageNumber="562">A. caviventris</emphasis>
</taxonomicName>
are not found in
<taxonomicName id="4C5D4D5BE47BFFC3FD9D54A7FD0F4283" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[553,711,187,213]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47BFFC3FD9D54A7FD0F4283" box="[553,711,187,213]" italics="true" pageId="3" pageNumber="562">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47BFFC3FD6654A7FCDE4282" bold="true" box="[722,790,187,213]" pageId="3" pageNumber="562">
<taxonomicNameLabel id="A21A57B1E47BFFC3FD6654A7FCDE4282" box="[722,790,187,213]" pageId="3" pageNumber="562" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(e.g. large subtriangular sclerite behind the two central spines of the internal sac of aedeagus). Furthermore, both sexes of the specimens described here are characterised by a smooth metaventrite (
<figureCitation id="13662A5DE47BFFC3FE7D551FFDED434A" box="[457,549,259,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="3" pageNumber="562">Fig. 6A</figureCitation>
), whereas in
<taxonomicName id="4C5D4D5BE47BFFC3FD0A551FFC92434B" baseAuthorityName="Reitter" baseAuthorityYear="1884" box="[702,858,259,285]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="caviventris">
<emphasis id="B929EACAE47BFFC3FD0A551FFC92434B" box="[702,858,259,285]" italics="true" pageId="3" pageNumber="562">A. caviventris</emphasis>
</taxonomicName>
the latter is produced into a distinct protruberance (
<bibRefCitation id="EFCC4B29E47BFFC3FF2B553BFE814316" author="Besuchet, C." box="[159,329,295,321]" pageId="3" pageNumber="562" pagination="30 - 42" refId="ref6086" refString="Besuchet, C. (1961) Pselaphides palearctiques. Especes nouvelles et notes synonymiques (Coleoptera). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 34 (1), 30 - 42" type="journal article" year="1961">Besuchet 1961</bibRefCitation>
). Future molecular work is needed to test whether the morphological distinctiveness of the Taygetos population is correlated with significant genetic divergence from
<taxonomicName id="4C5D4D5BE47BFFC3FCC75557FBC54333" baseAuthorityName="Reitter" baseAuthorityYear="1884" box="[883,1037,331,357]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="caviventris">
<emphasis id="B929EACAE47BFFC3FCC75557FBC54333" box="[883,1037,331,357]" italics="true" pageId="3" pageNumber="562">A. caviventris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>