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<document id="F22E9B949D2276DFCA537DB2D99CA7F4" ID-DOI="10.11865/zs.201807" ID-ISSN="2095-6827" ID-Zenodo-Dep="5366525" ID-ZooBank="0FD73836-E490-433F-BD64-C305D6826F11" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630257160038" checkinUser="diego" docAuthor="Pan, Zhao &amp; Ren, Guodong" docDate="2018" docId="03F393673614FFB3F0D0FC01FECDA58E" docLanguage="en" docName="Taxonomic revision of the subfamily Meloinae (Coleoptera_ Meloidae) from Xizang, China, with description of a new species.pdf" docOrigin="Zoological Systematics 43 (1)" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Meloe (Meloe) scabrus Pan &amp; Ren 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="78" masterDocId="FFCAEB1F361FFFBFF058FFC1FFB2A618" masterDocTitle="Taxonomic revision of the subfamily Meloinae (Coleoptera: Meloidae) from Xizang, China, with description of a new species" masterLastPageNumber="88" masterPageNumber="66" pageNumber="77" updateTime="1699199682714" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="47666F6026419344384540FCF982963B">Taxonomic revision of the subfamily Meloinae (Coleoptera: Meloidae) from Xizang, China, with description of a new species</mods:title>
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<paragraph id="8BE522713614FFB4F0D0FC01FD15A5C0" blockId="11.[136,679,960,984]" box="[136,679,960,984]" pageId="11" pageNumber="77">
<heading id="D0AD951D3614FFB4F0D0FC01FD15A5C0" box="[136,679,960,984]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="77" reason="2">
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<taxonomicName id="4C5A59F23614FFB4F0D0FC01FE36A5C0" authority="Pan &amp; Ren, 2018" authorityName="Pan &amp; Ren" authorityYear="2018" box="[136,388,960,984]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="scabrus" status="sp. nov." subGenus="Meloe">
<emphasis id="B92EFE633614FFB4F0D0FC01FF7EA5C0" bold="true" box="[136,204,960,984]" italics="true" pageId="11" pageNumber="77">Meloe</emphasis>
(
<emphasis id="B92EFE633614FFB4F084FC01FE92A5C0" bold="true" box="[220,288,960,984]" italics="true" pageId="11" pageNumber="77">Meloe</emphasis>
) 
<emphasis id="B92EFE633614FFB4F168FC01FE36A5C0" bold="true" box="[304,388,960,984]" italics="true" pageId="11" pageNumber="77">scabrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D43183614FFB4F1D3FC01FE52A5C0" box="[395,480,960,984]" pageId="11" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13613EF43614FFB4F1A5FC01FDE2A5C0" box="[509,592,960,984]" captionStart="Figures 13–25" captionStartId="9.[155,227,1962,1984]" captionTargetBox="[206,1394,373,1904]" captionTargetId="figure-111@9.[189,1399,370,1910]" captionTargetPageId="9" captionText="Figures 13–25. Habitus, dorsal view. 13. Lytta taliana Pic. 14. Meloe servulus Bates. 15. Meloe proscarabaeus proscarabaeus Linnaeus. 16. Meloe scabrus sp. nov. 17. Hycleus cichorii (Linnaeus). 18. Hycleus dorsetiferus Pan, Ren &amp; Wang. 19. Hycleus medioinsignatus (Pic). 20. Hycleus phaleratus phaleratus (Pallas). 21. Pseudabris hingstoni (Blair). 22. Pseudabris latimaculata Pan &amp; Bologna. 23. Pseudabris longiventris (Blair). 24. Pseudabris przewalskyi (Dokhtouroff). 25. Pseudabris regularis Pan &amp; Bologna." figureDoi="http://doi.org/10.5281/zenodo.5366529" httpUri="https://zenodo.org/record/5366529/files/figure.png" pageId="11" pageNumber="77">Figs 16</figureCitation>
, 
<figureCitation id="13613EF43614FFB4F203FC01FD2DA5C0" box="[603,671,960,984]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">26–33</figureCitation>
)
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</paragraph>
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<subSubSection id="C34071FA3614FFB3F0E5FC39FECDA58E" lastPageId="12" lastPageNumber="78" pageId="11" pageNumber="77" type="diagnosis">
<paragraph id="8BE522713614FFB4F0E5FC39FC4AA282" blockId="11.[136,1458,1016,2079]" pageId="11" pageNumber="77">
Diagnosis. Body is unicolor black, almost without metallic blue. Head and pronotum with irregular large shallow punctures, but the frons. Male antennomeres V–VII are modified, as in 
<figureCitation id="13613EF43614FFB4F3D2FBDBFBB6A22A" box="[906,1028,1050,1074]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Figs 26–27</figureCitation>
. Pronotum is slightly longer than wide, with a shallow large depression at center, sides sinuate. Mesonotum is triangularly stretch out, clearly visible (
<figureCitation id="13613EF43614FFB4F54BFBFCFAD0A24D" box="[1299,1378,1085,1109]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 29</figureCitation>
). Male parameres with three sub-rounded yellow spots on ventral surface. Penial distal hook is positioned almost in the middle between apex and the proximal hook, and slightly smaller than the proximal hook.
</paragraph>
<paragraph id="8BE522713614FFB4F0E5FB64FAD9A3A8" blockId="11.[136,1458,1016,2079]" pageId="11" pageNumber="77">
The new species has similar pronotal punctures with four species/subspecies from the Himalayan region, 
<emphasis id="B92EFE633614FFB4F5D6FB67FE93A2C7" italics="true" pageId="11" pageNumber="77">M. semicoriaceus</emphasis>
Fairmaire, 
<taxonomicName id="4C5A59F23614FFB4F1C5FB09FDECA2C7" authorityName=": MacSwain" authorityYear="1956" box="[413,606,1223,1247]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="proscarabaeus">
<emphasis id="B92EFE633614FFB4F1C5FB09FDECA2C7" box="[413,606,1223,1247]" italics="true" pageId="11" pageNumber="77">M. proscarabaeus</emphasis>
</taxonomicName>
<taxonomicName id="4C5A59F23614FFB4F23DFB06FCDFA2C7" authority="Linnaeus" authorityName="Linnaeus" box="[613,877,1223,1247]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="proscarabaeus">
<emphasis id="B92EFE633614FFB4F23DFB06FCB1A2C7" box="[613,771,1223,1247]" italics="true" pageId="11" pageNumber="77">proscarabaeus</emphasis>
Linnaeus
</taxonomicName>
, 
<emphasis id="B92EFE633614FFB4F321FB09FB51A2C7" box="[889,1251,1223,1247]" italics="true" pageId="11" pageNumber="77">
<taxonomicName id="4C5A59F23614FFB4F321FB09FB88A2C7" box="[889,1082,1223,1247]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="proscarabaeus">M. proscarabaeus</taxonomicName>
sericeorugosus
</emphasis>
Aksentjev, and 
<emphasis id="B92EFE633614FFB4F5D7FB09FEBDA31A" italics="true" pageId="11" pageNumber="77">M. arunachalae</emphasis>
<collectingRegion id="499EEC933614FFB4F14EFB2BFEFFA31A" box="[278,333,1258,1282]" country="Russia" name="Sakha" pageId="11" pageNumber="77">Saha</collectingRegion>
. However, 
<taxonomicName id="4C5A59F23614FFB4F19EFB2AFD8FA31A" authorityName="Pan &amp; Ren" authorityYear="2018" box="[454,573,1258,1282]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="B92EFE633614FFB4F19EFB2AFD8FA31A" box="[454,573,1258,1282]" italics="true" pageId="11" pageNumber="77">M. scabrus</emphasis>
</taxonomicName>
is distinguishable from them by the following features: 1) antennomere VII of 
<emphasis id="B92EFE633614FFB4F5D6FB2AFE92A33C" italics="true" pageId="11" pageNumber="77">M. semicoriaceus</emphasis>
is triangluar; antennomere VII of 
<taxonomicName id="4C5A59F23614FFB4F2E8FACFFC9FA33D" authorityName="Pan &amp; Ren" authorityYear="2018" box="[688,813,1293,1317]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="B92EFE633614FFB4F2E8FACFFC9FA33D" box="[688,813,1293,1317]" italics="true" pageId="11" pageNumber="77">M. scabrus</emphasis>
</taxonomicName>
is sub-quadrangluar (
<figureCitation id="13613EF43614FFB4F47EFACCFBC9A33D" box="[1062,1147,1293,1317]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 27</figureCitation>
); 2) antennomere V of 
<taxonomicName id="4C5A59F23614FFB4F5D6FACFFE7BA35F" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="proscarabaeus">
<emphasis id="B92EFE633614FFB4F5D6FACFFE7BA35F" italics="true" pageId="11" pageNumber="77">M. proscarabaeus proscarabaeus</emphasis>
</taxonomicName>
and 
<emphasis id="B92EFE633614FFB4F1A1FAF1FCEDA35F" box="[505,863,1327,1351]" italics="true" pageId="11" pageNumber="77">
<taxonomicName id="4C5A59F23614FFB4F1A1FAF1FD0BA35F" box="[505,697,1327,1351]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="proscarabaeus">M. proscarabaeus</taxonomicName>
sericeorugosus
</emphasis>
are not transverse widened (fig. 132c in Bolonga, 1991), and their parameres have big yellow spot on ventral surface (fig. 
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Aksentjev, 1987); but antennomere V of 
<taxonomicName id="4C5A59F23614FFB4F545FA92FA21A372" authorityName="Pan &amp; Ren" authorityYear="2018" box="[1309,1427,1362,1386]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="B92EFE633614FFB4F545FA92FA21A372" box="[1309,1427,1362,1386]" italics="true" pageId="11" pageNumber="77">M. scabrus</emphasis>
</taxonomicName>
is widened (
<figureCitation id="13613EF43614FFB4F0AEFAB4FEFBA395" box="[246,329,1397,1421]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 27</figureCitation>
), and parameres with three small yellow spots (
<figureCitation id="13613EF43614FFB4F333FAB4FC72A395" box="[875,960,1397,1421]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 30</figureCitation>
); 3) antennomere VII of 
<emphasis id="B92EFE633614FFB4F487FAB4FA3DA395" box="[1247,1423,1397,1421]" italics="true" pageId="11" pageNumber="77">M. arunachalae</emphasis>
is conspicuously wider than VI (fig. 
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<bibRefCitation id="EFCB5F803614FFB4F27DFA56FD12A3A8" author="Saha" box="[549,672,1431,1456]" pageId="11" pageNumber="77" pagination="1 - 146" refId="ref20188" refString="Saha, G. N. 1979. Revision of Indian blister beetles (Coleoptera: Meloidae: Meloinae). Records of the Zoological Survey of India, 74: 1 - 146." type="journal article" year="1979">
<collectingRegion id="499EEC933614FFB4F27DFA56FDE9A3B7" box="[549,603,1431,1455]" country="Russia" name="Sakha" pageId="11" pageNumber="77">Saha</collectingRegion>
, 1979
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); but antennomere VII of 
<taxonomicName id="4C5A59F23614FFB4F3ECFA59FB98A3B7" authorityName="Pan &amp; Ren" authorityYear="2018" box="[948,1066,1431,1455]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="B92EFE633614FFB4F3ECFA59FB98A3B7" box="[948,1066,1431,1455]" italics="true" pageId="11" pageNumber="77">M. scabrus</emphasis>
</taxonomicName>
is narrower than VI (
<figureCitation id="13613EF43614FFB4F557FA56FAEDA3A8" box="[1295,1375,1431,1456]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BE522713614FFB4F0E5FA7BFC29A000" blockId="11.[136,1458,1016,2079]" pageId="11" pageNumber="77">
Description. Body unicolor black with, at most, antennae and legs slightly metallic blue; surface opaque. Body with sparse and very short black setae, also on ventral side, only with golden setae on tarsal pads. Body length (apex of mandiblesapex of abdomen) 
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(
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, male), 
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(
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, female).
</paragraph>
<paragraph id="8BE522713614FFB4F0E5F9E3FB18A1C2" blockId="11.[136,1458,1016,2079]" pageId="11" pageNumber="77">
Head subquadrate, approximately 0.8 as long as wide, subparallel on sides. Punctures large, irregular, and dense (its diameter conspicuously larger than the distance between punctures), but inconspicuous at center. Frons rugulose, with a finely impressed median line and an inconspicuous depression in middle, between eyes. Eye subreniform, weakly narrowed in ventral, with antero-dorsal margin slightly sinuate, just behind antennal insertion. Temple subparallel, only slightly curved posteriad and conspicuously longer than longitudinal diameter of eye. Clypeus posteriorly with large punctures and anteriorly almost smooth, posterior margin (frontoclypeal suture) obtuse-angled curved in middle. Labrum entire, anterior margin conspicuously sinuate; maxillary palpomeres slightly enlarged apically, palpomere IV slightly shorter than II; labial palpomere III conspicuously widening; mandibles curved and progressively narrowed on apical half. Antennae modified in male, but almost non modified in female (
<figureCitation id="13613EF43614FFB4F212F8F6FD7AA157" box="[586,712,1847,1871]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Figs 26–28</figureCitation>
): male antennomere III subequel in length to I, IV subequel in length to II, V–VII conspicuously modified (
<figureCitation id="13613EF43614FFB4F278F89BFD2FA16A" box="[544,669,1882,1906]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Figs 26–27</figureCitation>
), VI positioned almost in middle of V in dorsal view, VII slightly longer but narrower than VI in exteral lateral view, VIII slightly shorter than III, decreasing in width from VIII to IX, IX to XI similar in width, XI subfusiformed, nearly 1.8 as long as X; female antennomeres similar to male but slightly stronger, antennomeres V–VII not modified, only slightly wider than others, and XI subcylindrical (
<figureCitation id="13613EF43614FFB4F416F803FB2CA1C2" box="[1102,1182,1986,2010]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8BE522713614FFB3F0E5F824FA94A797" blockId="11.[136,1458,1016,2079]" lastBlockId="12.[136,1452,236,919]" lastPageId="12" lastPageNumber="78" pageId="11" pageNumber="77">
Pronotum (
<figureCitation id="13613EF43614FFB4F16DF824FE30A1E5" box="[309,386,2021,2045]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="11" pageNumber="77">Fig. 29</figureCitation>
) slightly longer than wide (length/width approximately equal to 1.05), sides sinuate, widest at apical third, and subparallel on sides on basal third; disk with a median shallow depression in basal half; punctures similar but slightly denser than those of head; posterior margin conspicuously sinuate in middle, fully visible from above. Mesonotum to triangularly stretch out, clearly visible (
<figureCitation id="13613EF43613FFB3F215FECEFD2FA73F" box="[589,669,271,295]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="12" pageNumber="78">Fig. 29</figureCitation>
). Elytra obsolescently rugose, its length more than two times (ca. 2.5) of pronotal length. All tibiae with two spurs in both sexes; inner protibial spur longer than external spur, both slender; mesotibial spurs both slender either, in similar length; external metatibial spur widened, spoon-shaped at apex, longer and wider than inner one. Tarsal pads moderately developed on all legs in both sexes but absent on metatarsomere I of female.
</paragraph>
<paragraph id="8BE522713613FFB3F0E5FE5BFB54A7AA" blockId="12.[136,1452,236,919]" box="[189,1254,410,434]" pageId="12" pageNumber="78">Posterior margin of last visible sternite shallowly emarginate in male, and almost straight in female.</paragraph>
<paragraph id="8BE522713613FFB3F0E5FE7CFD13A424" blockId="12.[136,1452,236,919]" pageId="12" pageNumber="78">
Parameres slightly shorter than phallobase, with three sub-rounded yellow spots in middle of parameres in ventral view (
<figureCitation id="13613EF43613FFB3F0C8FE1EFF6DA7EF" box="[144,223,479,503]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="12" pageNumber="78">Fig. 30</figureCitation>
); apical lobe ca. 0.25 times of total length of parameres in lateral view (
<figureCitation id="13613EF43613FFB3F380FE1EFB95A7EF" box="[984,1063,479,503]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="12" pageNumber="78">Fig. 31</figureCitation>
). Penis (
<figureCitation id="13613EF43613FFB3F4DAFE1EFB7DA7EF" box="[1154,1231,479,503]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="12" pageNumber="78">Fig. 32</figureCitation>
) slender, distal hook positioned almost in middle between apex and proximal hook, and slightly smaller than proximal hook; endophallic hook small and slender. 
<emphasis id="B92EFE633613FFB3F117FDE5FDA2A424" box="[335,528,548,572]" italics="true" pageId="12" pageNumber="78">
<taxonomicName id="4C5A59F23613FFB3F117FDE5FE00A424" authorityName="Brauckmann &amp; Herd" authorityYear="2005" box="[335,434,548,572]" class="Insecta" family="Hapalopteridae" genus="Spiculum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caloneurodea" pageId="12" pageNumber="78" phylum="Arthropoda" rank="genus">Spiculum</taxonomicName>
gastrale
</emphasis>
as in 
<figureCitation id="13613EF43613FFB3F217FDE5FD2CA424" box="[591,670,548,572]" captionStart="Figure 26–33" captionStartId="12.[155,218,1706,1728]" captionTargetBox="[189,1398,978,1666]" captionTargetId="figure-456@12.[189,1398,978,1666]" captionTargetPageId="12" captionText="Figure 26–33. Meloe (Meloe) scabrus sp. nov. 26. Antenna, dorsal view, male. 27. Antennomeres V– VII, external lateral view. 28. Antenna, dorsal view, female. 29. Pronotum and mesonotum, dorsal view. 30. Tegmen, ventral view. 31. Tegmen, lateral view. 32. Penis, lateral view. 33. Spiculum gastrale, dorsal view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5366531" httpUri="https://zenodo.org/record/5366531/files/figure.png" pageId="12" pageNumber="78">Fig. 33</figureCitation>
.
</paragraph>
<paragraph id="8BE522713613FFB3F0E5FD86FC56A537" blockId="12.[136,1452,236,919]" pageId="12" pageNumber="78">
<materialsCitation id="3B32282C3613FFB3F0E5FD86FC56A537" collectingDate="2010-08-14" collectingDateMax="2014-08-08" collectingDateMin="2010-08-14" collectionCode="CWNU, MHBU" collectorName="Material &amp; Prof. Aimin Shi &amp; Yunchun Li &amp; Chinese &amp; Junsheng Shan" country="China" elevation="4262" latitude="28.4599" location="Chinese" longLatPrecision="1" longitude="91.42738" pageId="12" pageNumber="78" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Xizang" typeStatus="holotype">
<collectorName id="26AF47A73613FFB3F0E5FD86FEA5A447" box="[189,279,583,607]" pageId="12" pageNumber="78">Material</collectorName>
examined. 
<typeStatus id="54E19CD33613FFB3F1CBFD86FE44A447" box="[403,502,583,607]" pageId="12" pageNumber="78" type="holotype">Holotype</typeStatus>
<specimenCount id="9D5CE9F83613FFB3F1A5FD86FDA3A446" box="[509,529,583,606]" pageId="12" pageNumber="78" type="male">♂</specimenCount>
, 
<specimenCount id="9D5CE9F83613FFB3F246FD86FDF4A446" box="[542,582,583,607]" pageId="12" pageNumber="78" type="female">1 ♀</specimenCount>
<typeStatus id="54E19CD33613FFB3F214FD86FD1AA447" box="[588,680,583,607]" pageId="12" pageNumber="78" type="paratype">paratype</typeStatus>
(presented by 
<collectorName id="26AF47A73613FFB3F31DFD86FC5FA447" box="[837,1005,583,607]" pageId="12" pageNumber="78">Prof. Aimin Shi</collectorName>
of 
<collectionCode id="ED4BBAB43613FFB3F448FD86FBD6A447" box="[1040,1124,583,607]" pageId="12" pageNumber="78">CWNU</collectionCode>
, deposited in 
<collectionCode id="ED4BBAB43613FFB3F4AEFD86FAFAA447" box="[1270,1352,583,607]" pageId="12" pageNumber="78">MHBU</collectionCode>
) with the following labels: “ 
<date id="FFE404B13613FFB3F108FDABFE6FA49A" box="[336,477,618,642]" pageId="12" pageNumber="78" value="2010-08-14">
<collectingDate id="EFA0FD593613FFB3F108FDABFE6FA49A" box="[336,477,618,642]" pageId="12" pageNumber="78" value="2010-08-14">2010.VIII.14</collectingDate>
</date>
// Comai, 
<collectingRegion id="499EEC933613FFB3F209FDABFD2CA49A" box="[593,670,618,642]" country="China" name="Xizang" pageId="12" pageNumber="78">Xizang</collectingRegion>
// Yongsheng Pan &amp; 
<collectorName id="26AF47A73613FFB3F3DEFDABFBB7A49A" box="[902,1029,618,642]" pageId="12" pageNumber="78">Yunchun Li</collectorName>
leg. // Museum of 
<collectingCountry id="F34D62E13613FFB3F48EFDABFAA4A49A" box="[1238,1302,618,642]" name="China" pageId="12" pageNumber="78">China</collectingCountry>
West Normal University” (white, printed, in 
<collectorName id="26AF47A73613FFB3F196FD4DFD9AA4BC" box="[462,552,652,676]" pageId="12" pageNumber="78">Chinese</collectorName>
), “ 
<typeStatus id="54E19CD33613FFB3F21BFD4DFD60A4BC" box="[579,722,652,676]" pageId="12" pageNumber="78" type="holotype">HOLOTYPE</typeStatus>
(and 
<typeStatus id="54E19CD33613FFB3F355FD4DFC20A4BC" box="[781,914,652,676]" pageId="12" pageNumber="78" type="paratype">PARATYPE</typeStatus>
respectively) // 
<taxonomicName id="4C5A59F23613FFB3F464FD4DFA19A4BC" authority="Pan &amp; Ren" authorityName="Pan &amp; Ren" authorityYear="2018" box="[1084,1451,652,676]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="78" phylum="Arthropoda" rank="species" species="scabrus" subGenus="Meloe">
<emphasis id="B92EFE633613FFB3F464FD4DFBCDA4BC" box="[1084,1151,652,676]" italics="true" pageId="12" pageNumber="78">Meloe</emphasis>
(
<emphasis id="B92EFE633613FFB3F4D6FD4DFB63A4BC" box="[1166,1233,652,676]" italics="true" pageId="12" pageNumber="78">Meloe</emphasis>
) 
<emphasis id="B92EFE633613FFB3F487FD4DFA80A4BC" box="[1247,1330,652,676]" italics="true" pageId="12" pageNumber="78">scabrus</emphasis>
Pan &amp; Ren
</taxonomicName>
det. 2016” (red or yellow, printed and handwritten). 
<specimenCount id="9D5CE9F83613FFB3F2F7FD6EFD64A4DE" box="[687,726,687,711]" pageId="12" pageNumber="78" type="male">1 ♂</specimenCount>
<typeStatus id="54E19CD33613FFB3F284FD6EFC8AA4DF" box="[732,824,687,711]" pageId="12" pageNumber="78" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED4BBAB43613FFB3F31DFD6EFC2AA4DF" box="[837,920,687,711]" pageId="12" pageNumber="78">MHBU</collectionCode>
) with the following labels: “ 
<date id="FFE404B13613FFB3F49CFD6EFAF6A4DF" box="[1220,1348,687,711]" pageId="12" pageNumber="78" value="2014-08-08">
<collectingDate id="EFA0FD593613FFB3F49CFD6EFAF6A4DF" box="[1220,1348,687,711]" pageId="12" pageNumber="78" value="2014-08-08">2014.VIII.8</collectingDate>
</date>
// Comai, 
<collectingRegion id="499EEC933613FFB3F0D0FD10FF67A4F1" box="[136,213,721,745]" country="China" name="Xizang" pageId="12" pageNumber="78">Xizang</collectingRegion>
// Guodong Ren, Xinglong Bai &amp; 
<collectorName id="26AF47A73613FFB3F210FD10FD5BA4F1" box="[584,745,721,745]" pageId="12" pageNumber="78">Junsheng Shan</collectorName>
leg. // Museum of 
<collectingRegion id="499EEC933613FFB3F3EDFD10FC47A4F1" box="[949,1013,721,745]" country="China" name="Hebei" pageId="12" pageNumber="78">Hebei</collectingRegion>
University” (white, printed, in 
<collectorName id="26AF47A73613FFB3F51EFD10FA12A4F1" box="[1350,1440,721,745]" pageId="12" pageNumber="78">Chinese</collectorName>
), “ 
<geoCoordinate id="EE6E44B63613FFB3F0CAFD35FEA8A514" box="[146,282,756,780]" degrees="28" direction="north" minutes="27.594" orientation="latitude" pageId="12" pageNumber="78" precision="1" value="28.4599">28°27.594'N</geoCoordinate>
// 
<geoCoordinate id="EE6E44B63613FFB3F162FD35FE0CA514" box="[314,446,756,780]" degrees="91" direction="east" minutes="25.643" orientation="longitude" pageId="12" pageNumber="78" precision="1" value="91.42738">91°25.643'E</geoCoordinate>
// Alt. 
<quantity id="4CA28F943613FFB3F255FD35FDE9A514" box="[525,603,756,780]" metricMagnitude="3" metricUnit="m" metricValue="4.262" pageId="12" pageNumber="78" unit="m" value="4262.0">
<elevation id="0077C5423613FFB3F255FD35FDE9A514" box="[525,603,756,780]" metricMagnitude="3" metricUnit="m" metricValue="4.262" pageId="12" pageNumber="78" unit="m" value="4262.0">4262 m</elevation>
</quantity>
// Museum of 
<collectingRegion id="499EEC933613FFB3F2A7FD35FC8DA514" box="[767,831,756,780]" country="China" name="Hebei" pageId="12" pageNumber="78">Hebei</collectingRegion>
University” (white, printed, in 
<location id="8E8574AA3613FFB3F4CFFD35FB43A514" LSID="urn:lsid:plazi:treatment:03F393673614FFB3F0D0FC01FECDA58E:8E8574AA3613FFB3F4CFFD35FB43A514" box="[1175,1265,756,780]" country="China" latitude="28.4599" longLatPrecision="1" longitude="91.42738" name="Chinese" pageId="12" pageNumber="78" stateProvince="Xizang">Chinese</location>
), “ 
<typeStatus id="54E19CD33613FFB3F555FD34FA26A515" box="[1293,1428,757,781]" pageId="12" pageNumber="78" type="paratype">PARATYPE</typeStatus>
// 
<taxonomicName id="4C5A59F23613FFB3F0D0FCD6FE49A536" authority="Pan &amp; Ren" authorityName="Pan &amp; Ren" authorityYear="2018" box="[136,507,791,815]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="78" phylum="Arthropoda" rank="species" species="scabrus" subGenus="Meloe">
<emphasis id="B92EFE633613FFB3F0D0FCD6FF79A537" box="[136,203,791,815]" italics="true" pageId="12" pageNumber="78">Meloe</emphasis>
(
<emphasis id="B92EFE633613FFB3F082FCD6FEAFA537" box="[218,285,791,815]" italics="true" pageId="12" pageNumber="78">Meloe</emphasis>
) 
<emphasis id="B92EFE633613FFB3F174FCD6FECDA537" box="[300,383,791,815]" italics="true" pageId="12" pageNumber="78">scabrus</emphasis>
Pan &amp; Ren
</taxonomicName>
det. 2016” (yellow, printed and handwritten).
</materialsCitation>
</paragraph>
<paragraph id="8BE522713613FFB3F0E5FCFBFD04A54A" blockId="12.[136,1452,236,919]" box="[189,694,826,850]" pageId="12" pageNumber="78">
Distribution. 
<collectingCountry id="F34D62E13613FFB3F114FCFBFE22A54A" box="[332,400,826,850]" name="China" pageId="12" pageNumber="78">China</collectingCountry>
: 
<collectingRegion id="499EEC933613FFB3F1C1FCFBFE54A54A" box="[409,486,826,850]" country="China" name="Xizang" pageId="12" pageNumber="78">Xizang</collectingRegion>
(Comai (MHBU)).
</paragraph>
<paragraph id="8BE522713613FFB3F0E5FC9DFECDA58E" blockId="12.[136,1452,236,919]" pageId="12" pageNumber="78">
Etymology. The specific name “ 
<taxonomicName id="4C5A59F23613FFB3F24AFC9DFDD7A56C" authorityName="Pan &amp; Ren" authorityYear="2018" box="[530,613,860,884]" class="Insecta" family="Meloidae" genus="Meloe" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="78" phylum="Arthropoda" rank="species" species="scabrus">
<emphasis id="B92EFE633613FFB3F24AFC9DFDD7A56C" box="[530,613,860,884]" italics="true" pageId="12" pageNumber="78">scabrus</emphasis>
</taxonomicName>
” is a Latin adjective (= scabrous), referring to its large and irregular punctation on head and pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>