148 lines
17 KiB
XML
148 lines
17 KiB
XML
<document id="F6395D45375AFF055E93FD0A9BF2801C" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2019" docId="03F32A40FF90B14953679ED6F7CDFCB7" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion insularis Hyman & Köhler 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="424" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="424" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C0DC0F5F31870E46039FFEEE5FDD421B">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="1BCDC9D9558D1ACDACEF94357D33D2E5">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="9FEDE1EF0C356FBBC2280AD8960C6E9A">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="CD44334BC0DBB2D72D3B6541787C69B7">text</mods:typeOfResource>
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<mods:title id="750C2767E995D0155C26E2320A8B4A60">Contributions to Zoology</mods:title>
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<mods:part id="9DA7EB9D3787DF40729AC36CEB67C457">
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<mods:date id="E4BA8EA292ACF77DF34AD2B7E355484D">2019</mods:date>
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<mods:number id="EE0D6EB460F811FA9565DFE99E750554">2019-10-03</mods:number>
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<mods:number id="FEB2118A2BA22AE5902D7C0B6481537C">88</mods:number>
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<mods:url id="1C1BA3E7367C2D91DA83E419887706C8">http://dx.doi.org/10.1163/18759866-20191416</mods:url>
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<mods:classification id="FD796A9B52971C5C5B792FA820B7AE6F">journal article</mods:classification>
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<mods:identifier id="72C1973DADD991ABDD9722A1F0B5FA88" type="DOI">10.1163/18759866-20191416</mods:identifier>
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<treatment id="03F32A40FF90B14953679ED6F7CDFCB7" LSID="urn:lsid:plazi:treatment:03F32A40FF90B14953679ED6F7CDFCB7" httpUri="http://treatment.plazi.org/id/03F32A40FF90B14953679ED6F7CDFCB7" lastPageNumber="424" pageId="73" pageNumber="424">
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<subSubSection id="C340C8DDFF90B14953679ED6F5DDFD1B" box="[93,426,709,736]" pageId="73" pageNumber="424" type="nomenclature">
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<paragraph id="8BE59B56FF90B14953679ED6F5DDFD1B" blockId="73.[93,426,709,736]" box="[93,426,709,736]" pageId="73" pageNumber="424">
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<heading id="D0AD2C3AFF90B14953679ED6F5DDFD1B" box="[93,426,709,736]" fontSize="11" level="2" pageId="73" pageNumber="424" reason="2">
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<emphasis id="B92E4744FF90B14953679ED6F5DDFD1B" bold="true" box="[93,426,709,736]" pageId="73" pageNumber="424">
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<taxonomicName id="4C5AE0D5FF90B14953679ED6F526FD1B" ID-CoL="7RBRH" ID-ENA="2658024" authority="Hyman & Köhler, 2019" authorityName="Hyman & Köhler" authorityYear="2019" box="[93,337,709,736]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="insularis" status="sp. nov.">
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<emphasis id="B92E4744FF90B14953679ED6F526FD1B" bold="true" box="[93,337,709,736]" italics="true" pageId="73" pageNumber="424">Fastosarion insularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21DFA3FFF90B149526C9ED6F5DDFD1B" box="[342,426,710,736]" pageId="73" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340C8DDFF90B14953589EFAF7CDFCB7" pageId="73" pageNumber="424" type="description">
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<paragraph id="8BE59B56FF90B14953589EFAF571FCFE" blockId="73.[98,262,746,773]" box="[98,262,746,773]" pageId="73" pageNumber="424">
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<figureCitation id="136187D3FF90B14953589EFAF4C5FCFF" box="[98,178,746,772]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="73" pageNumber="424">
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Figs.
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<geoCoordinate id="EE6EFD91FF90B14953A09EFAF4C5FCFF" box="[154,178,746,772]" degrees="18, 19" direction="east" orientation="longitude" pageId="73" pageNumber="424" precision="555" value="18.19">18</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="136187D3FF90B14953869EFAF494FCFF" box="[188,227,746,772]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="73" pageNumber="424">19E</figureCitation>
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,
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<figureCitation id="136187D3FF90B14953D79EFAF571FCFE" box="[237,262,746,773]" captionStart="FIGURE 25" captionStartId="75.[98,171,1516,1537]" captionTargetBox="[99,1156,189,1484]" captionTargetId="figure-7@75.[98,1156,189,1484]" captionTargetPageId="75" captionText="FIGURE 25 Genitalia of F.insularis, MO85836, Palm Island. A. Reproductive system.Arrow shows the junction of the penis and epiphallus.B.Penis interior. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343111" httpUri="https://zenodo.org/record/8343111/files/figure.png" pageId="73" pageNumber="424">25</figureCitation>
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</paragraph>
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<paragraph id="8BE59B56FF90B14953669F22F4A3FCB7" blockId="73.[92,608,818,951]" box="[92,212,818,844]" pageId="73" pageNumber="424">Etymology</paragraph>
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<paragraph id="8BE59B56FF90B14953589F46F4B9FC4C" blockId="73.[92,608,818,951]" pageId="73" pageNumber="424">
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From
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<taxonomicName id="4C5AE0D5FF90B14953939F45F57FFC94" authorityName="Hyman & Köhler" authorityYear="2019" box="[169,264,853,879]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="insularis">
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<emphasis id="B92E4744FF90B14953939F45F57FFC94" box="[169,264,853,879]" italics="true" pageId="73" pageNumber="424">insularis</emphasis>
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</taxonomicName>
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(Latin = of an island), referring to the island habitat of this species; adjective.
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</paragraph>
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<paragraph id="8BE59B56FF90B14953669FF5F547FC04" blockId="73.[92,608,997,1203]" box="[92,304,997,1023]" pageId="73" pageNumber="424">Material examined</paragraph>
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<paragraph id="8BE59B56FF90B1495358981AF64AFBB3" blockId="73.[92,608,997,1203]" pageId="73" pageNumber="424">
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<emphasis id="B92E4744FF90B1495358981AF4D6FBD8" box="[98,161,1034,1059]" italics="true" pageId="73" pageNumber="424">Types</emphasis>
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:
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<materialsCitation id="3B32910BFF90B14953959819F64DFBB3" collectingDate="2001-02-22" collectionCode="QM" collectorName="G. Monteith" location="Palm Island" pageId="73" pageNumber="424" specimenCode="MO85836" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E125F4FF90B14953959819F56DFBD8" box="[175,282,1033,1059]" pageId="73" pageNumber="424">Holotype</typeStatus>
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:
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<collectionCode id="ED4B0393FF90B149521F9819F527FBD8" box="[293,336,1033,1059]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF90B149526D9819F5B4FBDF" box="[343,451,1033,1060]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">MO85836</specimenCode>
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(
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<location id="8E85CD8DFF90B14952E99819F62AFBD8" LSID="urn:lsid:plazi:treatment:03F32A40FF90B14953679ED6F7CDFCB7:8E85CD8DFF90B14952E99819F62AFBD8" box="[467,605,1033,1059]" name="Palm Island" pageId="73" pageNumber="424">Palm Island</location>
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, site 4, NEQ, coll.
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<date id="FFE4BD96FF90B1495227983DF5EAFBBC" box="[285,413,1069,1095]" pageId="73" pageNumber="424" value="2001-02-22">
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<collectingDate id="EFA0447EFF90B1495227983DF5EAFBBC" box="[285,413,1069,1095]" pageId="73" pageNumber="424" value="2001-02-22">22 Feb 2001</collectingDate>
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</date>
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,
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<collectorName id="26AFFE80FF90B149529C983DF645FBB3" box="[422,562,1069,1096]" pageId="73" pageNumber="424">G. Monteith</collectorName>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF90B14953BB9841F4A1FB6B" blockId="73.[92,608,997,1203]" pageId="73" pageNumber="424">
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<materialsCitation id="3B32910BFF90B14953BB9841F4A5FB6B" collectingDate="2001-02-22" collectionCode="QM" collectorName="G. Monteith" location="Palm Island" pageId="73" pageNumber="424" specimenCode="MO76362" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54E125F4FF90B14953BB9841F491FB90" box="[129,230,1105,1131]" pageId="73" pageNumber="424">Paratype</typeStatus>
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:
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<collectionCode id="ED4B0393FF90B14953C09841F552FB90" box="[250,293,1105,1131]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF90B149520C9841F5D7FB90" box="[310,416,1105,1131]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">MO76362</specimenCode>
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(same data as holotype)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF90B14953BB9889F5BCFB48" blockId="73.[92,608,997,1203]" box="[129,459,1177,1203]" pageId="73" pageNumber="424">
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<emphasis id="B92E4744FF90B14953BB9889F53BFB48" box="[129,332,1177,1203]" italics="true" pageId="73" pageNumber="424">
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<typeStatus id="54E125F4FF90B14953BB9889F530FB48" box="[129,327,1177,1203]" isOtherTypeStatus="true" pageId="73" pageNumber="424">Non-type material</typeStatus>
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:
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</emphasis>
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See
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<tableCitation id="C6D8AEEDFF90B14952449889F5B0FB48" box="[382,455,1177,1203]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="73" pageNumber="424" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF90B149536698F0F4BEFB01" blockId="73.[92,611,1248,1741]" box="[92,201,1248,1274]" pageId="73" pageNumber="424">Diagnosis</paragraph>
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<paragraph id="8BE59B56FF90B14953589914F55CF99A" blockId="73.[92,611,1248,1741]" pageId="73" pageNumber="424">
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<emphasis id="B92E4744FF90B14953589914F525FAE5" box="[98,338,1284,1310]" italics="true" pageId="73" pageNumber="424">External morphology</emphasis>
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: Shell (fig. 19E) small (
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<quantity id="4CA236B3FF90B14953539938F49AFAB8" box="[105,237,1320,1347]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.4" metricValueMin="9.1" pageId="73" pageNumber="424" unit="mm" value="9.25" valueMax="9.4" valueMin="9.1">9.1–9.4 mm</quantity>
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), pale amber, transparent, 2.8–3.2 whorls, flattened, thin, last whorl large. Body
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<quantity id="4CA236B3FF90B149535E9960F4A3FA70" box="[100,212,1392,1419]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="73" pageNumber="424" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
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, colour (in alcohol) brown with tiny white specks; dark bands on either side of neck; sole pale. Mantle lobes and shell lappets moderately large, covered with small white pustules, both lappets with horizontal ridge near lower edge, left lappet only with dark band below ridge.
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</paragraph>
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<paragraph id="8BE59B56FF90B14953BB9A7BF752FE66" blockId="73.[92,611,1248,1741]" lastBlockId="73.[646,1158,171,413]" pageId="73" pageNumber="424">
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<emphasis id="B92E4744FF90B14953BB9A7BF533F97E" box="[129,324,1643,1669]" italics="true" pageId="73" pageNumber="424">Genital anatomy:</emphasis>
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Genitalia (fig. 25) with long vagina; bursa copulatrix moderately short, duct distinct, bursa triangular. Penis very large, swollen proximally, internally with 3 longitudinal pilasters; internal wall smooth; 85–90% of penis contained in penial tunica. Epiphallus approx. 1.5–2 times length of penis; epiphallus 1 shorter than epiphallus 2; epiphallic caecum medium length; flagellum short, slender.
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</paragraph>
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<paragraph id="8BE59B56FF90B14951BA9DDAF695FE1F" blockId="73.[640,1158,458,844]" box="[640,738,458,484]" pageId="73" pageNumber="424">Remarks</paragraph>
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<paragraph id="8BE59B56FF90B14951BC9DFFF7CDFCB7" blockId="73.[640,1158,458,844]" pageId="73" pageNumber="424">
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<taxonomicName id="4C5AE0D5FF90B14951BC9DFFF71EFDF3" authorityName="Hyman & Köhler" authorityYear="2019" box="[646,873,494,520]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="insularis">
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<emphasis id="B92E4744FF90B14951BC9DFFF71EFDF3" box="[646,873,494,520]" italics="true" pageId="73" pageNumber="424">Fastosarion insularis</emphasis>
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</taxonomicName>
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(referred to above as ‘
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<taxonomicName id="4C5AE0D5FF90B14957619DFFF77BFDD7" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
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sp. Palm Island’) is only recorded from Palm Island in northeastern Qld (fig. 18) and is parapatric with respect to its congeners. It is most closely related to
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<taxonomicName id="4C5AE0D5FF90B14950FA9E6FF06DFD63" box="[960,1050,638,664]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="alyssa">
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<emphasis id="B92E4744FF90B14950FA9E6FF06DFD63" box="[960,1050,638,664]" italics="true" pageId="73" pageNumber="424">F. alyssa</emphasis>
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</taxonomicName>
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, which it greatly resembles, but can be distinguished by the presence of small white pustules covering the shell lappets and lobes, less distinct slime grooves, and a less transparent shell, as well as its distinct genital anatomy.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |