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<document id="0B9BCC81BB3A1F91C395C2395E016418" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629613271721" checkinUser="felipe" docAuthor="Paquin, Pierre &amp; Dupérré, Nadine" docDate="2009" docId="03C55737FFC4FFEBFF50F888D028FCAA" docLanguage="en" docName="zt02002p067.pdf" docOrigin="Zootaxa 2002" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cicurina (Cicurella) caverna Gertsch 1992" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FFFC2F4FFFD6FFFFFFC7FFA4D12FFFC0" masterDocTitle="of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution" masterLastPageNumber="67" masterPageNumber="1" pageNumber="19" updateTime="1699083252512" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FAF44B812EAB54BBD6DC27A432029864">of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution</mods:title>
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<mods:namePart id="B24CBD5DC7F6E012855D69B54CDAB202">Paquin, Pierre</mods:namePart>
<mods:affiliation id="D51D197C92AC36BC44E161E88C5D8899">Cave and Endangered Invertebrate Research Laboratory, SWCA Environmental Consultants, 4407 Monterey Oaks Boulevard, Building 1, Suite 110, Austin, Texas, 78749, USA. E-mail: ppaquin @ swca. com &amp; This is publication no. 15 of the Karst Biosciences and Environmental Geophysics Research Laboratories, SWCA Environmental Consultants</mods:affiliation>
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<mods:namePart id="D22A5AB3CA10B8C68DE283DE7FAEB493">Dupérré, Nadine</mods:namePart>
<mods:affiliation id="EB8B1DF71EF42C3FDFEB2E116659E971">American Museum of Natural History, Division of Invertebrate Zoology, Central Park West at 79 Street, New York, New York, 10024 USA. E-mail: nduperre @ amnh. org</mods:affiliation>
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<treatment id="03C55737FFC4FFEBFF50F888D028FCAA" ID-DOI="http://doi.org/10.5281/zenodo.5243018" ID-Zenodo-Dep="5243018" LSID="urn:lsid:plazi:treatment:03C55737FFC4FFEBFF50F888D028FCAA" httpUri="http://treatment.plazi.org/id/03C55737FFC4FFEBFF50F888D028FCAA" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C376B5AAFFC4FFEDFF50F888D38EF885" box="[151,673,1835,1862]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BD3E621FFC4FFEDFF50F888D38EF885" blockId="18.[151,673,1835,1896]" box="[151,673,1835,1862]" pageId="18" pageNumber="19">
<heading id="D09B514DFFC4FFEDFF50F888D38EF885" bold="true" box="[151,673,1835,1862]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C6C9DA2FFC4FFEDFF50F888D38EF885" ID-CoL="5Z664" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[151,673,1835,1862]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="caverna" subGenus="Cicurella">
<emphasis id="B9183A33FFC4FFEDFF50F888D38EF885" bold="true" box="[151,673,1835,1862]" pageId="18" pageNumber="19">
<emphasis id="B9183A33FFC4FFEDFF50F888D0DAF886" bold="true" box="[151,501,1836,1862]" italics="true" pageId="18" pageNumber="19">Cicurina (Cicurella) caverna</emphasis>
<bibRefCitation id="EFFD9BD0FFC4FFEDFE3BF88FD38EF885" author="Gertsch, W. J." box="[508,673,1835,1861]" pageId="18" pageNumber="19" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C376B5AAFFC4FFEDFF50F8EAD047F8A8" box="[151,360,1870,1896]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BD3E621FFC4FFEDFF50F8EAD047F8A8" blockId="18.[151,673,1835,1896]" box="[151,360,1870,1896]" pageId="18" pageNumber="19">
(
<figureCitation id="1357FAA4FFC4FFEDFF58F8EAD008F8A8" box="[159,295,1870,1896]" captionStart="FIGURES 827" captionStartId="19.[151,272,1804,1828]" captionTargetBox="[275,1316,207,1785]" captionTargetId="figure-119@19.[272,1329,197,1798]" captionTargetPageId="19" captionText="FIGURES 827. Cleared epigynum of troglobitic Cicurina (Cicurella) holotypes. For each species, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. 89 Cicurina bandera Gertsch 1992. 1011 Cicurina bandida Gertsch 1992. 1213 Cicurina baronia Gertsch 1992. 1415 Cicurina barri Gertsch 1992. 1617 Cicurina browni Gertsch 1992. 1819 Cicurina brunsi" pageId="18" pageNumber="19">Figs 2627</figureCitation>
,
<figureCitation id="1357FAA4FFC4FFEDFEF5F8EAD070F8A8" box="[306,351,1870,1896]" captionStart="FIGURE 137" captionStartId="60.[151,255,1808,1832]" captionTargetBox="[151,1431,661,1785]" captionTargetId="figure-237@60.[151,1436,661,1785]" captionTargetPageId="60" captionText="FIGURE 137. Distribution of troglobitic Cicurina (subgenus Cicurella) species in West Texas. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). See figure 138 for more detailed coverage of Val Verde and Edwards Counties." pageId="18" pageNumber="19">137</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF13B6A9FFC5FFECFF50F8A8D4B3F850" pageId="19" pageNumber="20" startId="19.[151,272,1804,1828]" targetBox="[275,1316,207,1785]" targetPageId="19">
<paragraph id="8BD3E621FFC5FFECFF50F8A8D4B3F850" blockId="19.[151,1436,1804,1936]" pageId="19" pageNumber="20">
<emphasis id="B9183A33FFC5FFECFF50F8A8D079F8E4" bold="true" box="[151,342,1804,1828]" pageId="19" pageNumber="20">FIGURES 827.</emphasis>
Cleared epigynum of troglobitic
<taxonomicName id="4C6C9DA2FFC5FFECFD0CF8A9D280F8E3" authority="" authorityName="Chamberlin &amp; Ivie" authorityYear="1940" box="[715,943,1804,1828]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Cicurella">
<emphasis id="B9183A33FFC5FFECFD0CF8A9D203F8E4" box="[715,812,1805,1828]" italics="true" pageId="19" pageNumber="20">Cicurina</emphasis>
(
<emphasis id="B9183A33FFC5FFECFCF9F8A9D28AF8E4" box="[830,933,1805,1828]" italics="true" pageId="19" pageNumber="20">Cicurella</emphasis>
)
</taxonomicName>
holotypes. For each species, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view.
<emphasis id="B9183A33FFC5FFECFEA6F8F0D0A6F8AC" bold="true" box="[353,393,1876,1900]" pageId="19" pageNumber="20">89</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFE57F8F1D3C9F8AB" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[400,742,1876,1900]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bandera">
<emphasis id="B9183A33FFC5FFECFE57F8F1D362F8AC" box="[400,589,1877,1900]" italics="true" pageId="19" pageNumber="20">Cicurina bandera</emphasis>
<bibRefCitation id="EFFD9BD0FFC5FFECFD92F8F0D3C9F8AB" author="Gertsch, W. J." box="[597,742,1876,1899]" pageId="19" pageNumber="20" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9183A33FFC5FFECFD36F8F0D21DF8AC" bold="true" box="[753,818,1876,1900]" pageId="19" pageNumber="20">1011</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFCFEF8F1D5BFF8AB" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[825,1168,1876,1900]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bandida">
<emphasis id="B9183A33FFC5FFECFCFEF8F1D2D9F8AC" box="[825,1014,1877,1900]" italics="true" pageId="19" pageNumber="20">Cicurina bandida</emphasis>
<bibRefCitation id="EFFD9BD0FFC5FFECFC3AF8F0D5BFF8AB" author="Gertsch, W. J." box="[1021,1168,1876,1899]" pageId="19" pageNumber="20" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9183A33FFC5FFECFB5EF8F0D5F3F8AC" bold="true" box="[1177,1244,1876,1900]" pageId="19" pageNumber="20">1213</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFB24F8F1D01EF84F" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="baronia">
<emphasis id="B9183A33FFC5FFECFB24F8F1D4B4F8AC" box="[1251,1435,1877,1900]" italics="true" pageId="19" pageNumber="20">Cicurina baronia</emphasis>
<bibRefCitation id="EFFD9BD0FFC5FFECFF50F8DCD01EF84F" author="Gertsch, W. J." box="[151,305,1912,1935]" pageId="19" pageNumber="20" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9183A33FFC5FFECFEF9F8DCD0ACF850" bold="true" box="[318,387,1912,1936]" pageId="19" pageNumber="20">1415</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFE4BF8DDD3FAF84F" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[396,725,1912,1936]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="barri">
<emphasis id="B9183A33FFC5FFECFE4BF8DDD31DF850" box="[396,562,1913,1936]" italics="true" pageId="19" pageNumber="20">Cicurina barri</emphasis>
<bibRefCitation id="EFFD9BD0FFC5FFECFDFCF8DCD3FAF84F" author="Gertsch, W. J." box="[571,725,1912,1935]" pageId="19" pageNumber="20" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9183A33FFC5FFECFD25F8DCD208F850" bold="true" box="[738,807,1912,1936]" pageId="19" pageNumber="20">1617</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFCF7F8DDD5A2F84F" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[816,1165,1912,1936]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="B9183A33FFC5FFECFCF7F8DDD2C5F850" box="[816,1002,1913,1936]" italics="true" pageId="19" pageNumber="20">Cicurina browni</emphasis>
<bibRefCitation id="EFFD9BD0FFC5FFECFC34F8DCD5A2F84F" author="Gertsch, W. J." box="[1011,1165,1912,1935]" pageId="19" pageNumber="20" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9183A33FFC5FFECFB5DF8DCD5F0F850" bold="true" box="[1178,1247,1912,1936]" pageId="19" pageNumber="20">1819</emphasis>
<taxonomicName id="4C6C9DA2FFC5FFECFB2FF8DDD4B3F850" authorityName="Cokendolpher" authorityYear="2004" box="[1256,1436,1913,1936]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brunsi">Cicurina brunsi</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C376B5AAFFC2FFEBFF50FF33D312FF18" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BD3E621FFC2FFEBFF50FF33D312FF18" blockId="20.[151,1437,151,874]" pageId="20" pageNumber="21">
<materialsCitation id="3B04EC7CFFC2FFEBFF50FF33D312FF18" collectingDate="1964-02-21" collectionCode="AMNH" collectorName="Bill Russell &amp; Examined" location="Kimble County. Fleming's Cave" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<emphasis id="B9183A33FFC2FFEBFF50FF33D086FF71" bold="true" box="[151,425,151,177]" pageId="20" pageNumber="21">
<typeStatus id="54D75883FFC2FFEBFF50FF33D06EFF71" box="[151,321,151,177]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
female:
</emphasis>
<location id="8EB3B0FAFFC2FFEBFE7BFF3CD261FF72" LSID="urn:lsid:plazi:treatment:03C55737FFC4FFEBFF50F888D028FCAA:8EB3B0FAFFC2FFEBFE7BFF3CD261FF72" box="[444,846,152,178]" name="Kimble County. Fleming's Cave" pageId="20" pageNumber="21">Kimble County. Flemings Cave</location>
,
<locationDeviation id="9A61FF03FFC2FFEBFC9BFF3CD540FF72" box="[860,1135,152,178]" location="No. Telegraph" pageId="20" pageNumber="21">
<quantity id="4C944BC4FFC2FFEBFC9BFF3CD29BFF72" box="[860,948,152,178]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="20" pageNumber="21" unit="mi" value="4.0">4 miles</quantity>
No. Telegraph
</locationDeviation>
,
<date id="FFD2C0E1FFC2FFEBFBBBFF3CD44DFF72" box="[1148,1378,152,178]" pageId="20" pageNumber="21" value="1964-02-21">
<collectingDate id="EF963909FFC2FFEBFBBBFF3CD44DFF72" box="[1148,1378,152,178]" pageId="20" pageNumber="21" value="1964-02-21">February 21, 1964</collectingDate>
</date>
.
<collectorName id="269983F7FFC2FFEBFAA8FF3CD1C2FF18" pageId="20" pageNumber="21">Bill Russell</collectorName>
coll. ”.
<collectorName id="269983F7FFC2FFEBFE8DFF1AD0EEFF18" box="[330,449,190,216]" pageId="20" pageNumber="21">Examined</collectorName>
(
<collectionCode id="ED7D7EE4FFC2FFEBFE16FF1AD31EFF18" box="[465,561,190,216]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="20" pageNumber="21">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC2FFEBFF00FF41D1FCFD0F" pageId="20" pageNumber="21" type="description">
<paragraph id="8BD3E621FFC2FFEBFF00FF41D305FF3E" blockId="20.[151,1437,151,874]" box="[199,554,229,255]" pageId="20" pageNumber="21">
<emphasis id="B9183A33FFC2FFEBFF00FF41D070FF3F" bold="true" box="[199,351,229,255]" pageId="20" pageNumber="21">Description.</emphasis>
<emphasis id="B9183A33FFC2FFEBFEA1FF41D305FF3E" box="[358,554,229,254]" italics="true" pageId="20" pageNumber="21">
Female
<typeStatus id="54D75883FFC2FFEBFE02FF41D305FF3E" box="[453,554,229,254]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD3E621FFC2FFEBFF00FEA8D1FCFD0F" blockId="20.[151,1437,151,874]" pageId="20" pageNumber="21">Total length: 3.90; carapace length: 1.90; carapace width: 1.15; opisthosoma length: 2.00. Eyes absent. Cheliceral promargin right side with 3 teeth (2 large, 1 small), left side likewise; retromargin right side with 6 denticles (2 large, 4 small), left side 5 denticles (2 large, 3 small). Leg I: total length: 5.60; femur: 1.60; patella: 0.60; tibia: 1.25; metatarsus: 1.30; tarsus: 0.85; leg IV: total length: 6.55; femur: 1.75; patella: 0.60; tibia: 1.55; metatarsus: 1.70; tarsus: 0.95. Tarsal claw IV: 0.13. Epigynum: 0.35. Epigynal ventral plate with long transverse slit, bearing medial, very deep, inverted U-shaped indentation; atrium squared, almost completely intruded by dome-shaped dorsal plate, atrial aperture small; bursa not enlarged nor fused; copulatory ducts touching at midpoint, reaching over primary lobe apex; copulatory ducts constriction located sub-basally; primary pores not visible; stalk short, procurved joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore elongated; primary lobe elongated; secondary lobe absent; fertilization canal long, externally positioned from the dictynoid pore; fertilization duct medially attached to the primary lobe base.</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC2FFEBFF00FD7FD1FBFCDC" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BD3E621FFC2FFEBFF00FD7FD1FBFCDC" blockId="20.[151,1437,151,874]" pageId="20" pageNumber="21">
<emphasis id="B9183A33FFC2FFEBFF00FD7FD04FFD35" bold="true" box="[199,352,731,757]" pageId="20" pageNumber="21">Distribution</emphasis>
. Only reported from the
<typeStatus id="54D75883FFC2FFEBFD5CFD78D3E2FD36" box="[667,717,732,758]" pageId="20" pageNumber="21">type</typeStatus>
locality: Flemings Bat Cave (Kimble County,
<collectingRegion id="49A828C3FFC2FFEBFAD1FD78D472FD36" box="[1302,1373,732,758]" country="United States of America" name="Texas" pageId="20" pageNumber="21">Texas</collectingRegion>
,
<figureCitation id="1357FAA4FFC2FFEBFAA9FD78D1E8FCDC" captionStart="FIGURE 137" captionStartId="60.[151,255,1808,1832]" captionTargetBox="[151,1431,661,1785]" captionTargetId="figure-237@60.[151,1436,661,1785]" captionTargetPageId="60" captionText="FIGURE 137. Distribution of troglobitic Cicurina (subgenus Cicurella) species in West Texas. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). See figure 138 for more detailed coverage of Val Verde and Edwards Counties." pageId="20" pageNumber="21">Fig. 137</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC2FFEBFF50FC8CD028FCAA" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BD3E621FFC2FFEBFF50FC8CD028FCAA" blockId="20.[151,1437,151,874]" pageId="20" pageNumber="21">
<emphasis id="B9183A33FFC2FFEBFF50FC8CD03FFC82" bold="true" box="[151,272,808,834]" pageId="20" pageNumber="21">Remarks.</emphasis>
Male unknown. Data added by
<bibRefCitation id="EFFD9BD0FFC2FFEBFD43FC8DD216FC83" author="Gertsch, W. J." box="[644,825,809,835]" pageId="20" pageNumber="21" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch (1992)</bibRefCitation>
: “Flemings Bat Cave”, which is the actual name for that cave.
</paragraph>
</subSubSection>
</treatment>
</document>