317 lines
40 KiB
XML
317 lines
40 KiB
XML
<document id="0C57BAD11FB671E55A626AF3FC1DA534" ID-CLB-Dataset="20465" ID-DOI="10.11646/zootaxa.5104.2.9" ID-GBIF-Dataset="fcb2352e-d754-4629-9277-2a5864501d13" ID-ISSN="1175-5326" ID-Zenodo-Dep="6281000" ID-ZooBank="489361F2-1C78-4C23-BD29-D80A73A71F98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645780943727" checkinUser="plazi" docAuthor="Tsvetkov, Eugene" docDate="2022" docId="03B487FE6475FFFE26D287A8FAC9FDE4" docLanguage="en" docName="zootaxa.5104.2.9.pdf" docOrigin="Zootaxa 5104 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thiodia leucocitron Tsvetkov 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="300" masterDocId="FF8DFF866474FFFD26458777FF97FF85" masterDocTitle="A new species of the genus Thiodia Hübner, [1825] (Lepidoptera: Tortricidae) from South Kazakhstan" masterLastPageNumber="300" masterPageNumber="297" pageNumber="298" updateTime="1699316520494" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4897F17364E8F69A0FC4FA9DC3822AB7">A new species of the genus Thiodia Hübner, [1825] (Lepidoptera: Tortricidae) from South Kazakhstan</mods:title>
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<mods:name id="03D16B1028E2470EBF2E36E8A6EBE00E" type="personal">
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<mods:namePart id="26B536F2CD6F9FF95A1EB1CE02552FD0">Tsvetkov, Eugene</mods:namePart>
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<mods:title id="780342D9912085380CA49C6427A9BADA">Zootaxa</mods:title>
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<mods:part id="97B27D05ACF3A17B4185B92B8A621A02">
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<mods:date id="360234523BDACA35ACC63D81B195EF03">2022</mods:date>
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<mods:number id="AE9E39A596CFEF30A79ED175D1D6AE66">5104</mods:number>
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<treatment id="03B487FE6475FFFE26D287A8FAC9FDE4" ID-DOI="http://doi.org/10.5281/zenodo.6317798" ID-GBIF-Taxon="193836212" ID-Zenodo-Dep="6317798" LSID="urn:lsid:plazi:treatment:03B487FE6475FFFE26D287A8FAC9FDE4" httpUri="http://treatment.plazi.org/id/03B487FE6475FFFE26D287A8FAC9FDE4" lastPageId="3" lastPageNumber="300" pageId="1" pageNumber="298">
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<subSubSection id="C30765636475FFFC26D287A8FDF0FF7C" box="[151,615,223,250]" pageId="1" pageNumber="298" type="nomenclature">
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<emphasis id="B969EAFA6475FFFC26D287A8FDF0FF7C" bold="true" box="[151,615,223,250]" pageId="1" pageNumber="298">
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<taxonomicName id="4C1D4D6B6475FFFC26D287A8FE6AFF7C" ID-CoL="9BDJJ" authority="Tsvetkov" authorityName="Tsvetkov" authorityYear="2022" box="[151,509,223,250]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
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<emphasis id="B969EAFA6475FFFC26D287A8FE11FF7F" bold="true" box="[151,390,223,250]" italics="true" pageId="1" pageNumber="298">Thiodia leucocitron</emphasis>
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Tsvetkov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25A57816475FFFC244C87A8FDF0FF7C" box="[521,615,223,249]" pageId="1" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30765636475FFFC26D28674FEB0FE9B" box="[151,295,259,286]" pageId="1" pageNumber="298" type="description">
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<paragraph id="8BA236E86475FFFC26D28674FEB0FE9B" blockId="1.[151,615,223,286]" box="[151,295,259,286]" pageId="1" pageNumber="298">
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(
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<figureCitation id="13262A6D6475FFFC26DA8674FF71FE98" box="[159,230,259,285]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1030,1054]" captionTargetBox="[153,1435,525,1007]" captionTargetId="figure-102@1.[151,1435,525,1007]" captionTargetPageId="1" captionText="FIGURES 1–2. Thiodia leucocitron sp. nov. and its natural habitat. 1. Imago, holotype. 2. The habitat, a mountain slope in environs of the village Baizhansai, June 29, 2019." figureDoi="http://doi.org/10.5281/zenodo.6281002" httpUri="https://zenodo.org/record/6281002/files/figure.png" pageId="1" pageNumber="298">Figs 1</figureCitation>
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,
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<figureCitation id="13262A6D6475FFFC26B18674FE89FE9B" box="[244,286,259,286]" captionStart="FIGURES 3–5" captionStartId="2.[151,264,1552,1576]" captionTargetBox="[159,1419,362,1521]" captionTargetId="figure-109@2.[151,1436,345,1529]" captionTargetPageId="2" captionText="FIGURES 3–5. Genitalia of Thiodia leucocitron sp. nov. 3. Male genitalia. 4. Aedeagus. 5. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6281004" httpUri="https://zenodo.org/record/6281004/files/figure.png" pageId="1" pageNumber="298">3–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30765636475FFFC26D2863CFCF9FE57" pageId="1" pageNumber="298" type="materials_examined">
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<paragraph id="8BA236E86475FFFC26D2863CFBD0FE0C" blockId="1.[151,1437,331,502]" pageId="1" pageNumber="298">
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<emphasis id="B969EAFA6475FFFC26D2863CFED3FEE0" bold="true" box="[151,324,331,357]" pageId="1" pageNumber="298">Type material.</emphasis>
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<materialsCitation id="3B753CB56475FFFC270F863CFBD0FE0C" ID-GBIF-Occurrence="3498808301" collectingDate="2019-06-22" collectionCode="ZISP" collectorName="E. Tsvetkov. Deposited" country="Kazakhstan" county="Tsvetkov. Deposited in Zoological Institute St. Petersburg" elevation="700" latitude="43.50861" location="Karatau Mts." longLatPrecision="19" longitude="68.862495" municipality="Ashisai" pageId="1" pageNumber="298" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54A6884A6475FFFC270F863CFE25FEE0" box="[330,434,331,357]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D1BFD616475FFFC27FD863BFE47FEE3" box="[440,464,332,358]" pageId="1" pageNumber="298" type="male">♂</specimenCount>
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.
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<collectingCountry id="F30A76786475FFFC279C863CFDF7FEE0" box="[473,608,331,357]" name="Kazakhstan" pageId="1" pageNumber="298">Kazakhstan</collectingCountry>
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:
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<location id="8EC260336475FFFC242F863BFD6AFEE3" LSID="urn:lsid:plazi:treatment:03B487FE6475FFFE26D287A8FAC9FDE4:8EC260336475FFFC242F863BFD6AFEE3" box="[618,765,332,358]" country="Kazakhstan" county="Tsvetkov. Deposited in Zoological Institute St. Petersburg" latitude="43.50861" longLatPrecision="19" longitude="68.862495" municipality="Ashisai" name="Karatau Mts." pageId="1" pageNumber="298">Karatau Mts.</location>
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,
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<quantity id="4CE59B0D6475FFFC254D863CFCA8FEE0" box="[776,831,331,358]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="298" unit="km" value="5.0">5 km</quantity>
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SW of vill.
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<collectingMunicipality id="6BC6AC926475FFFC2587863CFB8EFEE0" box="[962,1049,331,357]" pageId="1" pageNumber="298">Ashisai</collectingMunicipality>
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,
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<quantity id="4CE59B0D6475FFFC2266863CFBFCFEE3" box="[1059,1131,331,358]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="298" unit="m" value="700.0">
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<elevation id="0030D1DB6475FFFC2266863CFBFCFEE3" box="[1059,1131,331,358]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="298" unit="m" value="700.0">700 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE29502F6475FFFC2230863BFA94FEED" box="[1141,1283,332,360]" degrees="43" direction="north" minutes="30" orientation="latitude" pageId="1" pageNumber="298" precision="15" seconds="31" value="43.50861">N 43°30′31″</geoCoordinate>
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,
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<geoCoordinate id="EE29502F6475FFFC2348863BFA00FEED" box="[1293,1431,332,360]" degrees="68" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="298" precision="15" seconds="45" value="68.862495">E 68°51′45″</geoCoordinate>
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,
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<date id="FFA310286475FFFC26D28618FE85FE0C" box="[151,274,367,393]" pageId="1" pageNumber="298" value="2019-06-22">
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<collectingDate id="EFE7E9C06475FFFC26D28618FE85FE0C" box="[151,274,367,393]" pageId="1" pageNumber="298" value="2019-06-22">22.vi.2019</collectingDate>
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</date>
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, leg
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<collectorName id="26E8533E6475FFFC27038607FDDEFE0C" box="[326,585,367,394]" pageId="1" pageNumber="298">
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E.
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<collectingCounty id="62C34E646475FFFC27208618FDDEFE0C" box="[357,585,367,393]" pageId="1" pageNumber="298">Tsvetkov. Deposited</collectingCounty>
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</collectorName>
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in Zoological Institute St. Petersburg (
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<collectionCode id="ED0CAE2D6475FFFC25BE8618FBADFE0C" box="[1019,1082,367,393]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="298">ZISP</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA236E86475FFFC268286E3FCF9FE57" blockId="1.[151,1437,331,502]" pageId="1" pageNumber="298">
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<materialsCitation id="3B753CB56475FFFC268286E3FD0DFE28" ID-GBIF-Occurrence="3498808302" box="[199,666,403,430]" collectingDate="2019-06-22" collectionCode="ZISP" collectorName="E. Tsvetkov. Deposited" country="Kazakhstan" county="Tsvetkov. Deposited in Zoological Institute St. Petersburg" elevation="700" latitude="43.50861" location="Karatau Mts." longLatPrecision="19" longitude="68.862495" municipality="Ashisai" pageId="1" pageNumber="298" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="54A6884A6475FFFC268286E3FEAFFE2B" box="[199,312,404,430]" pageId="1" pageNumber="298" type="paratype">Paratypes</typeStatus>
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.
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<specimenCount id="9D1BFD616475FFFC270786E4FEFAFE2B" box="[322,365,403,430]" pageId="1" pageNumber="298" type="male">1 ♂</specimenCount>
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, same data as for holotype
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</materialsCitation>
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||
;
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<materialsCitation id="3B753CB56475FFFC24E186E4FCFDFE57" ID-GBIF-Occurrence="3498808303" collectingDate="2019-06-29" collectionCode="ZISP, ET" collectorName="E. Tsvetkov" country="Kazakhstan" latitude="43.138058" location="Karatau Mts." longLatPrecision="19" longitude="69.94139" municipality="Baizhansai" pageId="1" pageNumber="298" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="9D1BFD616475FFFC24E186E4FD47FE2B" box="[676,720,403,430]" pageId="1" pageNumber="298" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D1BFD616475FFFC249986E4FC92FE2B" box="[732,773,403,430]" pageId="1" pageNumber="298" type="female">2 ♀</specimenCount>
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,
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<collectingCountry id="F30A76786475FFFC255686E4FC0DFE28" box="[787,922,403,429]" name="Kazakhstan" pageId="1" pageNumber="298">Kazakhstan</collectingCountry>
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:
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<location id="8EC260336475FFFC25E186E3FBAFFE2B" LSID="urn:lsid:plazi:treatment:03B487FE6475FFFE26D287A8FAC9FDE4:8EC260336475FFFC25E186E3FBAFFE2B" box="[932,1080,404,430]" country="Kazakhstan" latitude="43.138058" longLatPrecision="19" longitude="69.94139" municipality="Baizhansai" name="Karatau Mts." pageId="1" pageNumber="298">Karatau Mts.</location>
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,
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<quantity id="4CE59B0D6475FFFC220686E4FBEBFE28" box="[1091,1148,403,429]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="298" unit="km" value="3.0">3 km</quantity>
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NE of vill.
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<collectingMunicipality id="6BC6AC926475FFFC234586E4FAE9FE28" box="[1280,1406,403,429]" pageId="1" pageNumber="298">Baizhansai</collectingMunicipality>
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,
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<geoCoordinate id="EE29502F6475FFFC23CD86E3FE9DFE51" degrees="43" direction="north" minutes="08" orientation="latitude" pageId="1" pageNumber="298" precision="15" seconds="17" value="43.138058">N 43°08′17″</geoCoordinate>
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,
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<geoCoordinate id="EE29502F6475FFFC275386CFFE35FE51" box="[278,418,440,468]" degrees="69" direction="east" minutes="56" orientation="longitude" pageId="1" pageNumber="298" precision="15" seconds="29" value="69.94139">E 69°56′29″</geoCoordinate>
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,
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<date id="FFA310286475FFFC27E886C0FDB0FE54" box="[429,551,439,465]" pageId="1" pageNumber="298" value="2019-06-29">
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<collectingDate id="EFE7E9C06475FFFC27E886C0FDB0FE54" box="[429,551,439,465]" pageId="1" pageNumber="298" value="2019-06-29">29.vi.2019</collectingDate>
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</date>
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, leg
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<collectorName id="26E8533E6475FFFC241986CFFD75FE54" box="[604,738,439,466]" pageId="1" pageNumber="298">E. Tsvetkov</collectorName>
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, (
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<collectionCode id="ED0CAE2D6475FFFC24B086C0FCA7FE54" box="[757,816,439,465]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="298">ZISP</collectionCode>
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,
|
||
<collectionCode id="ED0CAE2D6475FFFC257E86CFFCF5FE57" box="[827,866,440,466]" country="0" name="East Texas State University" pageId="1" pageNumber="298">ET</collectionCode>
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)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C30765636475FFFF268286ACFAA1FEC4" lastPageId="2" lastPageNumber="299" pageId="1" pageNumber="298" type="description">
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<paragraph id="8BA236E86475FFFC268286ACFA86FE70" blockId="1.[151,1437,331,502]" box="[199,1297,475,502]" pageId="1" pageNumber="298">
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<emphasis id="B969EAFA6475FFFC268286ACFECEFE70" bold="true" box="[199,345,475,501]" pageId="1" pageNumber="298">Description.</emphasis>
|
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Imago (
|
||
<figureCitation id="13262A6D6475FFFC27F386ACFE6FFE70" box="[438,504,475,501]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1030,1054]" captionTargetBox="[153,1435,525,1007]" captionTargetId="figure-102@1.[151,1435,525,1007]" captionTargetPageId="1" captionText="FIGURES 1–2. Thiodia leucocitron sp. nov. and its natural habitat. 1. Imago, holotype. 2. The habitat, a mountain slope in environs of the village Baizhansai, June 29, 2019." figureDoi="http://doi.org/10.5281/zenodo.6281002" httpUri="https://zenodo.org/record/6281002/files/figure.png" pageId="1" pageNumber="298">Fig. 1</figureCitation>
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||
). Sexual dimorphism is not expressed in habitus and in head structure.
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||
</paragraph>
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||
<caption id="DF6266606475FFFC26D28371FD06FBC6" ID-DOI="http://doi.org/10.5281/zenodo.6281002" ID-Zenodo-Dep="6281002" httpUri="https://zenodo.org/record/6281002/files/figure.png" pageId="1" pageNumber="298" startId="1.[151,264,1030,1054]" targetBox="[153,1435,525,1007]" targetPageId="1">
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<paragraph id="8BA236E86475FFFC26D28371FD06FBC6" blockId="1.[151,1436,1030,1091]" pageId="1" pageNumber="298">
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||
<emphasis id="B969EAFA6475FFFC26D28371FEABFB9B" bold="true" box="[151,316,1030,1054]" pageId="1" pageNumber="298">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C1D4D6B6475FFFC27008371FD99FB9B" authority="Tsvetkov, 2022" authorityName="Tsvetkov" authorityYear="2022" box="[325,526,1030,1054]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
|
||
<emphasis id="B969EAFA6475FFFC27008371FD99FB9B" box="[325,526,1030,1054]" italics="true" pageId="1" pageNumber="298">Thiodia leucocitron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAFA6475FFFC24538370FDFEFB9A" bold="true" box="[534,617,1031,1055]" pageId="1" pageNumber="298">
|
||
<taxonomicNameLabel id="A25A57816475FFFC24538370FDFEFB9A" box="[534,617,1031,1055]" pageId="1" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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and its natural habitat. 1. Imago, holotype. 2. The habitat, a mountain slope in environs of the village Baizhansai, June 29, 2019.
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||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236E86475FFFC26828306FCB5FB51" blockId="1.[151,1437,1137,2027]" pageId="1" pageNumber="298">
|
||
<emphasis id="B969EAFA6475FFFC26828306FE9CFB0E" box="[199,267,1137,1163]" italics="true" pageId="1" pageNumber="298">Head.</emphasis>
|
||
Frons and vertex with white hair-like scales; labial palpi about 1.1–1.2× the diameter of eye, whitish, partly brown; antennae some more than 1/2 of the forewing, white dorsally, chequered ventrally and laterally (white and dark grey areas); chaetosemata present (white scales).
|
||
</paragraph>
|
||
<paragraph id="8BA236E86475FFFC268283AAFEFBF9DA" blockId="1.[151,1437,1137,2027]" pageId="1" pageNumber="298">
|
||
<emphasis id="B969EAFA6475FFFC268283AAFEB7FB72" box="[199,288,1245,1271]" italics="true" pageId="1" pageNumber="298">Thorax.</emphasis>
|
||
Thorax white from ventral side and light yellow dorsally. Legs with white and light brown areas which form clear ring pattern on tarsus; each tarsomere brown with white ring distally. Forewing nearly triangular with sharply rounded apex and widely rounded tornus; costal margin almost straight, slightly bent distally; termen straight, dorsal margin bent basally; small rounded scaled concavity (scent organ) is present basally between Cu stalk and A2+3 vein from underside; costal fold absent. Forewing upperside general background colour white, almost entirely covered with suffusion of orange-yellow groups of scales; one of the groups of colourful scales that form a submarginal fascia. Speculum subtriangular, pale brownish with admixture of orange-yellow scales; a row of black dots along the outer side. Pairs of white costal strigulae contrasted with dividing dark brown spots and streaks. Fringe unicolorous pale brownish-yellow; basal stripe nearly 1/3 of width, motley, formed by white brown-spotted scales. Forewing underside unicolourous brown. Hindwing light brown from both sides, fringe light brown with brown basal stripe.
|
||
</paragraph>
|
||
<paragraph id="8BA236E86475FFFC2682811EFE35F922" blockId="1.[151,1437,1137,2027]" pageId="1" pageNumber="298">
|
||
<emphasis id="B969EAFA6475FFFC2682811EFEAEF906" box="[199,313,1641,1667]" italics="true" pageId="1" pageNumber="298">Abdomen.</emphasis>
|
||
Abdomen whitish, its pattern is composed of pale brownish and white rings (two differently coloured rings on each segment).
|
||
</paragraph>
|
||
<paragraph id="8BA236E86475FFFC268281C6FB49F8FA" blockId="1.[151,1437,1137,2027]" pageId="1" pageNumber="298">
|
||
<emphasis id="B969EAFA6475FFFC268281C6FEFBF94E" box="[199,364,1713,1739]" italics="true" pageId="1" pageNumber="298">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13262A6D6475FFFC273E81C6FE76F94E" box="[379,481,1713,1739]" captionStart="FIGURES 3–5" captionStartId="2.[151,264,1552,1576]" captionTargetBox="[159,1419,362,1521]" captionTargetId="figure-109@2.[151,1436,345,1529]" captionTargetPageId="2" captionText="FIGURES 3–5. Genitalia of Thiodia leucocitron sp. nov. 3. Male genitalia. 4. Aedeagus. 5. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6281004" httpUri="https://zenodo.org/record/6281004/files/figure.png" pageId="1" pageNumber="298">Figs 3, 4</figureCitation>
|
||
). Uncus small, triangular. Socii membranous, apically rounded, flat, densely covered by short bristles. Valva elongate; cucullus well defined, nearly triangular (angles rounded), separated from sacculus by neck, inner surface densely covered with long bristles except for areas along its ventral and dorsal margins. Aedeagus heavily sclerotised, bearing a long process on its left side. Process flattened dorsoventrally, strongly flattened in distal 1/3 where a jagged crest is present on the right side; distal part of aedeagus with large membranous excavation on the left side. Juxta structure typical of Eucosmini (shown ventrolaterally on
|
||
<figureCitation id="13262A6D6475FFFC22C88012FB46F8FA" box="[1165,1233,1893,1919]" captionStart="FIGURES 3–5" captionStartId="2.[151,264,1552,1576]" captionTargetBox="[159,1419,362,1521]" captionTargetId="figure-109@2.[151,1436,345,1529]" captionTargetPageId="2" captionText="FIGURES 3–5. Genitalia of Thiodia leucocitron sp. nov. 3. Male genitalia. 4. Aedeagus. 5. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6281004" httpUri="https://zenodo.org/record/6281004/files/figure.png" pageId="1" pageNumber="298">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA236E86475FFFF268280FEFAA1FEC4" blockId="1.[151,1437,1137,2027]" lastBlockId="2.[151,1436,151,321]" lastPageId="2" lastPageNumber="299" pageId="1" pageNumber="298">
|
||
<emphasis id="B969EAFA6475FFFC268280FEFE12F826" box="[199,389,1929,1955]" italics="true" pageId="1" pageNumber="298">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13262A6D6475FFFC27D680FEFE40F826" box="[403,471,1928,1955]" captionStart="FIGURES 3–5" captionStartId="2.[151,264,1552,1576]" captionTargetBox="[159,1419,362,1521]" captionTargetId="figure-109@2.[151,1436,345,1529]" captionTargetPageId="2" captionText="FIGURES 3–5. Genitalia of Thiodia leucocitron sp. nov. 3. Male genitalia. 4. Aedeagus. 5. Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6281004" httpUri="https://zenodo.org/record/6281004/files/figure.png" pageId="1" pageNumber="298">Fig. 5</figureCitation>
|
||
). Papillae anales elongate, well sclerotised, fused, almost bald, covered by sparse short bristles. Apophyses strong, well sclerotised; posterior apophyses with broadening in posterior 1/3, anterior apophyses shorter than posterior apophyses. Eighth tergum large trapezoidal, tapering posteriorly, covered by bristles in posterior 1/4; anterior margin convex. Sternum VII trapezoidal with rounded angles, ending posteriorly with very large and broad ostium. Antrum very large, asymmetrical, flattened, consisting of two large pocket-like hollows and a funnel-shaped part between them branching off from left hollow to ductus bursae; surface well sclerotised with plots of heavy sclerotisation. Ductus bursae membranous, thin, elongate sclerite present inside in junction with antrum. Bursa copulatrix pear-shaped, membranous with a tiny rounded signum convex from the inner side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6266606476FFFF26D28167FB79F9AD" ID-DOI="http://doi.org/10.5281/zenodo.6281004" ID-Zenodo-Dep="6281004" box="[151,1262,1552,1576]" httpUri="https://zenodo.org/record/6281004/files/figure.png" pageId="2" pageNumber="299" startId="2.[151,264,1552,1576]" targetBox="[159,1419,362,1521]" targetPageId="2">
|
||
<paragraph id="8BA236E86476FFFF26D28167FB79F9AD" blockId="2.[151,1262,1552,1576]" box="[151,1262,1552,1576]" pageId="2" pageNumber="299">
|
||
<emphasis id="B969EAFA6476FFFF26D28167FEACF9AD" bold="true" box="[151,315,1552,1576]" pageId="2" pageNumber="299">FIGURES 3–5.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="4C1D4D6B6476FFFF27FA8167FD10F9AD" authority="Tsvetkov, 2022" authorityName="Tsvetkov" authorityYear="2022" box="[447,647,1552,1576]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
|
||
<emphasis id="B969EAFA6476FFFF27FA8167FD10F9AD" box="[447,647,1552,1576]" italics="true" pageId="2" pageNumber="299">Thiodia leucocitron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAFA6476FFFF24C88167FD49F9AD" bold="true" box="[653,734,1552,1576]" pageId="2" pageNumber="299">
|
||
<taxonomicNameLabel id="A25A57816476FFFF24C88167FD49F9AD" box="[653,734,1552,1576]" pageId="2" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
3. Male genitalia. 4. Aedeagus. 5. Female genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30765636476FFFF26828121FA1AF8E9" pageId="2" pageNumber="299" type="diagnosis">
|
||
<paragraph id="8BA236E86476FFFF26828121FA1AF8E9" blockId="2.[151,1437,1622,2009]" pageId="2" pageNumber="299">
|
||
<emphasis id="B969EAFA6476FFFF26828121FED5F9F5" bold="true" box="[199,322,1622,1648]" pageId="2" pageNumber="299">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C1D4D6B6476FFFF27098121FDBCF9F5" authority="Tsvetkov, 2022" authorityName="Tsvetkov" authorityYear="2022" box="[332,555,1622,1648]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
|
||
<emphasis id="B969EAFA6476FFFF27098121FDBCF9F5" box="[332,555,1622,1648]" italics="true" pageId="2" pageNumber="299">Thiodia leucocitron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAFA6476FFFF24738121FD04F9F5" bold="true" box="[566,659,1622,1648]" pageId="2" pageNumber="299">
|
||
<taxonomicNameLabel id="A25A57816476FFFF24738121FD04F9F5" box="[566,659,1622,1648]" pageId="2" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C1D4D6B6476FFFF255D8121FB3DF9F4" authority="(Kennel, 1916)" baseAuthorityName="Kennel" baseAuthorityYear="1916" box="[792,1194,1622,1649]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="anatoliana">
|
||
<emphasis id="B969EAFA6476FFFF255D8121FC65F9F5" box="[792,1010,1622,1648]" italics="true" pageId="2" pageNumber="299">Thiodia anatoliana</emphasis>
|
||
(Kennel, 1916)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D6B6476FFFF22A28121FEA6F911" authority="(Mann, 1873)" baseAuthorityName="Mann" baseAuthorityYear="1873" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="fessana">
|
||
<emphasis id="B969EAFA6476FFFF22A28121FA0BF9F5" box="[1255,1436,1622,1648]" italics="true" pageId="2" pageNumber="299">Thiodia fessana</emphasis>
|
||
(Mann, 1873)
|
||
</taxonomicName>
|
||
in the forewing colouration. But in the two species mentioned above the forewing is darker, elements of the pattern are olive, olive-grey or yellowish-brown. Elements of the forewing pattern are orange-yellow in the new species. Another species,
|
||
<taxonomicName id="4C1D4D6B6476FFFF27B281B5FC03F959" authority="Staudinger, 1871" authorityName="Staudinger" authorityYear="1871" box="[503,916,1730,1756]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="placidana">
|
||
<emphasis id="B969EAFA6476FFFF27B281B5FD5FF959" box="[503,712,1730,1756]" italics="true" pageId="2" pageNumber="299">Thiodia placidana</emphasis>
|
||
<bibRefCitation id="EF8C4B196476FFFF249481B5FC03F959" author="Staudinger, O." box="[721,916,1730,1756]" pageId="2" pageNumber="299" pagination="273 - 330" refId="ref2215" refString="Staudinger, O. (1871) Beschreibung neuer Lepidopteren des europaischen Faunengebiets (Schluss). Berliner Entomologische Zeitschrift, 14, 273 - 330." type="journal article" year="1871">Staudinger, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
, has some similarity with the new species in the shape of valva in males and also in the antrum which is asymmetrical in females. But
|
||
<taxonomicName id="4C1D4D6B6476FFFF22CC8190FA82F885" authorityName="Staudinger" authorityYear="1871" box="[1161,1301,1766,1792]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="placidana">
|
||
<emphasis id="B969EAFA6476FFFF22CC8190FA82F885" box="[1161,1301,1766,1792]" italics="true" pageId="2" pageNumber="299">T. placidana</emphasis>
|
||
</taxonomicName>
|
||
well differs externally by red pattern of the forewing. Male and female genitalia are very specific in
|
||
<taxonomicName id="4C1D4D6B6476FFFF2235807CFA9DF8A1" authority="Tsvetkov, 2022" authorityName="Tsvetkov" authorityYear="2022" box="[1136,1290,1802,1828]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
|
||
<emphasis id="B969EAFA6476FFFF2235807CFA9DF8A1" box="[1136,1290,1802,1828]" italics="true" pageId="2" pageNumber="299">T. leucocitron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAFA6476FFFF2354807DFAFDF8A1" bold="true" box="[1297,1386,1802,1828]" pageId="2" pageNumber="299">
|
||
<taxonomicNameLabel id="A25A57816476FFFF2354807DFAFDF8A1" box="[1297,1386,1802,1828]" pageId="2" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The aedeagus bears a long side process in males which allows to separate the new species from other congeners. A very large ostium and the antrum consisting of two hollows are characteristic for the females of
|
||
<taxonomicName id="4C1D4D6B6476FFFF22D78024FABBF8E9" authority="Tsvetkov, 2022" authorityName="Tsvetkov" authorityYear="2022" box="[1170,1324,1874,1900]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="leucocitron" status="sp. nov.">
|
||
<emphasis id="B969EAFA6476FFFF22D78024FABBF8E9" box="[1170,1324,1874,1900]" italics="true" pageId="2" pageNumber="299">T. leucocitron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAFA6476FFFF23768025FA1AF8E9" bold="true" box="[1331,1421,1874,1900]" pageId="2" pageNumber="299">
|
||
<taxonomicNameLabel id="A25A57816476FFFF23768025FA1AF8E9" box="[1331,1421,1874,1900]" pageId="2" pageNumber="299" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765636476FFFE26828001FAFCFF50" lastPageId="3" lastPageNumber="300" pageId="2" pageNumber="299" type="etymology">
|
||
<paragraph id="8BA236E86476FFFE26828001FAFCFF50" blockId="2.[151,1437,1622,2009]" lastBlockId="3.[151,1437,151,609]" lastPageId="3" lastPageNumber="300" pageId="2" pageNumber="299">
|
||
<emphasis id="B969EAFA6476FFFF26828001FED9F815" bold="true" box="[199,334,1910,1936]" pageId="2" pageNumber="299">Etymology.</emphasis>
|
||
The name of the species is a noun derived from the Ancient Greek prefix
|
||
<emphasis id="B969EAFA6476FFFF22E18001FB7DF815" box="[1188,1258,1910,1936]" italics="true" pageId="2" pageNumber="299">leuco-</emphasis>
|
||
and the French word
|
||
<emphasis id="B969EAFA6476FFFF269D80ECFE8DF831" box="[216,282,1947,1972]" italics="true" pageId="2" pageNumber="299">citron</emphasis>
|
||
. It is associated with the forewing colouration, which combines white and orange-yellow colours. The species group name “
|
||
<taxonomicName id="4C1D4D6B6476FFFF27CC80C9FD90F85D" authorityName="Tsvetkov" authorityYear="2022" box="[393,519,1982,2008]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="leucocitron">
|
||
<emphasis id="B969EAFA6476FFFF27CC80C9FD90F85D" box="[393,519,1982,2008]" italics="true" pageId="2" pageNumber="299">leucocitron</emphasis>
|
||
</taxonomicName>
|
||
” is not a Latin or latinized word. Following the ICZN Art. 31.2.1 and 31.2.3 this compound species-group name, of which the final component word is not a Latin or latinized word (Arts. 11.2, 26) is to be treated as indeclinable, and need not agree in gender with the generic name with which it is combined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765636477FFFE268287A9FBC1FE98" pageId="3" pageNumber="300" type="biology_ecology">
|
||
<paragraph id="8BA236E86477FFFE268287A9FBC1FE98" blockId="3.[151,1437,151,609]" pageId="3" pageNumber="300">
|
||
<emphasis id="B969EAFA6477FFFE268287A9FEB0FF7D" bold="true" box="[199,295,222,248]" pageId="3" pageNumber="300">Biology.</emphasis>
|
||
The species occurs in open mountain valleys. In the area
|
||
<quantity id="4CE59B0D6477FFFE25EC87A8FC75FF7C" box="[937,994,223,249]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="300" unit="km" value="3.0">3 km</quantity>
|
||
NE of the village Baizhansai its habitat is a dry mountain valley with steppe and rocky slopes at an altitude of
|
||
<quantity id="4CE59B0D6477FFFE25E88674FC63FE98" box="[941,1012,259,285]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="300" unit="m" value="900.0">900 m</quantity>
|
||
(
|
||
<figureCitation id="13262A6D6477FFFE22468674FBDEFE98" box="[1027,1097,259,285]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1030,1054]" captionTargetBox="[153,1435,525,1007]" captionTargetId="figure-102@1.[151,1435,525,1007]" captionTargetPageId="1" captionText="FIGURES 1–2. Thiodia leucocitron sp. nov. and its natural habitat. 1. Imago, holotype. 2. The habitat, a mountain slope in environs of the village Baizhansai, June 29, 2019." figureDoi="http://doi.org/10.5281/zenodo.6281002" httpUri="https://zenodo.org/record/6281002/files/figure.png" pageId="3" pageNumber="300">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765636477FFFE26828650FAC9FDE4" pageId="3" pageNumber="300" type="discussion">
|
||
<paragraph id="8BA236E86477FFFE26828650FAC9FDE4" blockId="3.[151,1437,151,609]" pageId="3" pageNumber="300">
|
||
<emphasis id="B969EAFA6477FFFE26828650FEACFEC4" bold="true" box="[199,315,295,321]" pageId="3" pageNumber="300">Remarks.</emphasis>
|
||
Males and females of the described species have a very characteristic unusual structure of the genitalia. Such a feature as a caudally directed process of the aedeagus is functionally related to the structure of the female antrum, which is divided into two pocket-like hollows. Among the representatives of the genus
|
||
<taxonomicName id="4C1D4D6B6477FFFE23508618FAFBFE0C" authorityName="Hubner" authorityYear="1825" box="[1301,1388,367,393]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAFA6477FFFE23508618FAFBFE0C" box="[1301,1388,367,393]" italics="true" pageId="3" pageNumber="300">Thiodia</emphasis>
|
||
</taxonomicName>
|
||
, the aedeagus with a process on the left side is found only in
|
||
<taxonomicName id="4C1D4D6B6477FFFE254A86E4FB0AFE28" authority="Kuznetzov, 1972" authorityName="Kuznetzov" authorityYear="1972" box="[783,1181,403,429]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="elbursica">
|
||
<emphasis id="B969EAFA6477FFFE254A86E4FC42FE28" box="[783,981,403,429]" italics="true" pageId="3" pageNumber="300">Thiodia elbursica</emphasis>
|
||
<bibRefCitation id="EF8C4B196477FFFE259986E4FB0AFE28" author="Kuznetzov, V. I." box="[988,1181,403,429]" pageId="3" pageNumber="300" pagination="311 - 318" refId="ref2063" refString="Kuznetzov, V. I. (1972) New species of the leaf-rollers (Lepidoptera, Tortricidae) from Iran and Caucasus. Proceedings of the Zoological Institute, Academy of Sciences of the USSR, 52, 311 - 318. [in Russian]" type="journal article" year="1972">Kuznetzov, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
, a species known from
|
||
<collectingCountry id="F30A76786477FFFE26D286C0FF5FFE54" box="[151,200,439,465]" name="Iran" pageId="3" pageNumber="300">Iran</collectingCountry>
|
||
. But in this species, the process is very small and distally located. It should be noted that similar side processes occur in the other genera of the
|
||
<taxonomicName id="4C1D4D6B6477FFFE27B586ACFDFDFE70" box="[496,618,475,501]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
|
||
family, for example, in
|
||
<taxonomicName id="4C1D4D6B6477FFFE253486ACFA9CFE70" authority="Obraztsov, 1952" authorityName="Obraztsov" authorityYear="1952" box="[881,1291,474,501]" class="Insecta" family="Tortricidae" genus="Eriopsela" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="klimeschi">
|
||
<emphasis id="B969EAFA6477FFFE253486ACFBDAFE70" box="[881,1101,475,501]" italics="true" pageId="3" pageNumber="300">Eriopsela klimeschi</emphasis>
|
||
<bibRefCitation id="EF8C4B196477FFFE221786ACFA9CFE70" author="Obraztsov, N." box="[1106,1291,474,501]" pageId="3" pageNumber="300" pagination="40 - 42" refId="ref2110" refString="Obraztsov, N. (1952) Eine wenig bekannte und drei neue palaearktische Tortriciden-Arten. Zeitschrift fur Lepidopterologie, 2 (1), 40 - 42." type="journal article" year="1952">Obraztsov, 1952</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from the Alps, as well as in some species of the genus
|
||
<taxonomicName id="4C1D4D6B6477FFFE24F38688FC39FD9C" authority="Hubner, [1825]" authorityName="Hubner" authorityYear="1825" box="[694,942,510,537]" class="Insecta" family="Tortricidae" genus="Cydia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAFA6477FFFE24F38688FD6EFD9C" box="[694,761,511,537]" italics="true" pageId="3" pageNumber="300">Cydia</emphasis>
|
||
Hübner, [1825]
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C1D4D6B6477FFFE25938688FBC8FD9C" authorityName="Obraztsov" authorityYear="1952" box="[982,1119,511,537]" class="Insecta" family="Tortricidae" genus="Eriopsela" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="klimeschi">
|
||
<emphasis id="B969EAFA6477FFFE25938688FBC8FD9C" box="[982,1119,511,537]" italics="true" pageId="3" pageNumber="300">E. klimeschi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D6B6477FFFE22D18688FA9CFD9C" box="[1172,1291,511,537]" class="Insecta" family="Tortricidae" genus="Cydia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B969EAFA6477FFFE22D18688FB40FD9C" box="[1172,1239,511,537]" italics="true" pageId="3" pageNumber="300">Cydia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
the structure of the antrum of females, however, without such a characteristic feature as in
|
||
<taxonomicName id="4C1D4D6B6477FFFE22548554FB3BFDB8" authorityName="Tsvetkov" authorityYear="2022" box="[1041,1196,547,573]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="leucocitron">
|
||
<emphasis id="B969EAFA6477FFFE22548554FB3BFDB8" box="[1041,1196,547,573]" italics="true" pageId="3" pageNumber="300">T. leucocitron</emphasis>
|
||
</taxonomicName>
|
||
. The presence of the aedeagus process in the mentioned genera and in
|
||
<taxonomicName id="4C1D4D6B6477FFFE24848530FCCDFDE4" authorityName="Tsvetkov" authorityYear="2022" box="[705,858,583,609]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="leucocitron">
|
||
<emphasis id="B969EAFA6477FFFE24848530FCCDFDE4" box="[705,858,583,609]" italics="true" pageId="3" pageNumber="300">T. leucocitron</emphasis>
|
||
</taxonomicName>
|
||
, in our opinion, is an example of homoplasia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |