treatments-xml/data/03/B4/87/03B487E9FF9B4E1791FAFF474FAAC560.xml

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<mods:title id="F91C1AB6A4A782DB433CA3772922CE39">Asiagomphus reinhardti sp. nov. (Odonata, Gomphidae) from eastern Cambodia and southern Laos</mods:title>
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<treatment id="03B487E9FF9B4E1791FAFF474FAAC560" ID-DOI="http://doi.org/10.5281/zenodo.6068461" ID-GBIF-Taxon="127640244" ID-Zenodo-Dep="6068461" LSID="urn:lsid:plazi:treatment:03B487E9FF9B4E1791FAFF474FAAC560" httpUri="http://treatment.plazi.org/id/03B487E9FF9B4E1791FAFF474FAAC560" lastPageId="6" lastPageNumber="41" pageId="1" pageNumber="36">
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<paragraph id="8BA236FFFF9B4E1091FAFF474F61C0CF" blockId="1.[151,545,151,211]" box="[151,545,151,178]" pageId="1" pageNumber="36">
<heading id="D0EA8193FF9B4E1091FAFF474F61C0CF" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="1" pageNumber="36" reason="1">
<emphasis id="B969EAEDFF9B4E1091FAFF474F61C0CF" bold="true" box="[151,545,151,178]" pageId="1" pageNumber="36">
<taxonomicName id="4C1D4D7CFF9B4E1091FAFF474CFDC0CC" ID-CoL="9X249" box="[151,445,151,177]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="reinhardti" status="sp. nov.">
<emphasis id="B969EAEDFF9B4E1091FAFF474CFDC0CC" bold="true" box="[151,445,151,177]" italics="true" pageId="1" pageNumber="36">Asiagomphus reinhardti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A5796FF9B4E1090A9FF484F61C0CF" box="[452,545,152,178]" pageId="1" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BA236FFFF9B4E1091FAFF6A4C52C0AE" blockId="1.[151,545,151,211]" box="[151,274,186,211]" pageId="1" pageNumber="36">
(
<figureCitation id="13262A7AFF9B4E1091F2FF6A4C49C0AE" box="[159,265,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1802,1824]" captionStartId-1="3.[151,250,1915,1937]" captionStartId-2="4.[151,250,1915,1937]" captionStartId-3="5.[151,250,1217,1239]" captionTargetBox-0="[151,1436,809,1777]" captionTargetBox-1="[338,1254,201,1884]" captionTargetBox-2="[156,1409,198,1883]" captionTargetBox-3="[151,1427,193,1194]" captionTargetId-0="figure@1.[151,1436,809,1777]" captionTargetId-1="figure@3.[332,1254,193,1893]" captionTargetId-2="figure@4.[156,1430,193,1893]" captionTargetId-3="figure@5.[151,1436,193,1196]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." captionText-1="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." captionText-2="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." captionText-3="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri-0="https://zenodo.org/record/271165/files/figure.png" httpUri-1="https://zenodo.org/record/271166/files/figure.png" httpUri-2="https://zenodo.org/record/271167/files/figure.png" httpUri-3="https://zenodo.org/record/271168/files/figure.png" pageId="1" pageNumber="36">Figs. 1–4</figureCitation>
)
</paragraph>
<paragraph id="8BA236FFFF9B4E1091FAFED44F06C147" blockId="1.[151,1436,259,314]" pageId="1" pageNumber="36">
<taxonomicName id="4C1D4D7CFF9B4E1091FAFED44C60C167" box="[151,288,260,282]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAEDFF9B4E1091FAFED44C60C167" box="[151,288,260,282]" italics="true" pageId="1" pageNumber="36">Asiagomphus</emphasis>
</taxonomicName>
sp.1— 
<bibRefCitation id="EF8C4B0EFF9B4E109009FED4494EC167" author="Yokoi" box="[356,1038,260,282]" pageId="1" pageNumber="36" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A List of Lao Dragonfliles. Kyoei Printing Co Ltd., Koriyama, 110 pp." type="book" year="2014" yearSuffix="p">Yokoi &amp; Souptanthong (2014: p. 15, 64; fig. 62; pl. 14, note 66)</bibRefCitation>
: ♂, 
<date id="FFA3103FFF9B4E109553FED449E7C167" box="[1086,1191,260,282]" pageId="1" pageNumber="36" value="2005-04-28">28/4/2005</date>
, Paksong, Alt. 
<quantity id="4CE59B1AFF9B4E10942BFED448D8C167" box="[1350,1432,260,282]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="36" unit="m" value="1310.0">1310 m</quantity>
, Bolaven Plt., Champsak Prov. [
<collectingCountry id="F30A766FFF9B4E10936EFEF44F7BC147" box="[515,571,292,314]" name="Laos" pageId="1" pageNumber="36">Laos</collectingCountry>
].
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF9B4E1091FAFEB6492DC272" pageId="1" pageNumber="36" type="materials_examined">
<paragraph id="8BA236FFFF9B4E1091FAFEB6492DC272" blockId="1.[151,1436,358,754]" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091FAFEB64C4DC102" bold="true" box="[151,269,358,383]" pageId="1" pageNumber="36">
<typeStatus id="54A6885DFF9B4E1091FAFEB64C48C102" box="[151,264,358,383]" pageId="1" pageNumber="36" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂ (abdomen and hind legs broken) (
<figureCitation id="13262A7AFF9B4E1093DAFEB64E64C102" box="[695,804,358,383]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1802,1824]" captionStartId-1="3.[151,250,1915,1937]" captionStartId-2="4.[151,250,1915,1937]" captionTargetBox-0="[151,1436,809,1777]" captionTargetBox-1="[338,1254,201,1884]" captionTargetBox-2="[156,1409,198,1883]" captionTargetId-0="figure@1.[151,1436,809,1777]" captionTargetId-1="figure@3.[332,1254,193,1893]" captionTargetId-2="figure@4.[156,1430,193,1893]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." captionText-1="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." captionText-2="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri-0="https://zenodo.org/record/271165/files/figure.png" httpUri-1="https://zenodo.org/record/271166/files/figure.png" httpUri-2="https://zenodo.org/record/271167/files/figure.png" pageId="1" pageNumber="36">Figs. 1–3</figureCitation>
), 
<collectingCountry id="F30A766FFF9B4E109251FEB64EF2C102" box="[828,946,358,383]" name="Cambodia" pageId="1" pageNumber="36">Cambodia</collectingCountry>
, Mondulkiri Province, 
<emphasis id="B969EAEDFF9B4E1095D6FEB64984C102" bold="true" box="[1211,1220,358,383]" pageId="1" pageNumber="36">‘</emphasis>
Loringae brook’ [a working nickname], the left tributary of the main river downstream from Buu Sraa Waterfall, 12°34'01–19'' 
<geoCoordinate id="EE295038FF9B4E1094E5FE5B4C52C1BA" direction="north" orientation="latitude" pageId="1" pageNumber="36" precision="15" value="107.41389">N 107°24'50''</geoCoordinate>
–25'03'' E, ca 
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a.s.l., 
<date id="FFA3103FFF9B4E10933EFE7E4F94C1BA" box="[595,724,430,455]" pageId="1" pageNumber="36" value="2014-06-15">15 vi 2014</date>
, 
<collectingCountry id="F30A766FFF9B4E10938EFE7F4FBAC1BA" box="[739,762,431,455]" name="Iceland" pageId="1" pageNumber="36">O</collectingCountry>
. Kosterin leg.; deposited in Naturalis Biodiversity Centre, Leiden, 
<collectingCountry id="F30A766FFF9B4E109198FE024CE8C196" box="[245,424,466,491]" name="Netherlands" pageId="1" pageNumber="36">the Netherlands</collectingCountry>
(
<collectionCode id="ED0CAE3AFF9B4E1090D7FE034F55C196" LSID="urn:lsid:biocol.org:col:34992" box="[442,533,467,491]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="36">RMNH</collectionCode>
). 
<emphasis id="B969EAEDFF9B4E109347FE024FE0C196" bold="true" box="[554,672,466,491]" pageId="1" pageNumber="36">
<typeStatus id="54A6885DFF9B4E109347FE024FDBC196" box="[554,667,466,491]" pageId="1" pageNumber="36" type="paratype">Paratype</typeStatus>
:
</emphasis>
♂ (
<figureCitation id="13262A7AFF9B4E1093BCFE024E58C196" box="[721,792,466,491]" captionStart="FIGURE 4" captionStartId="5.[151,250,1217,1239]" captionTargetBox="[151,1427,193,1194]" captionTargetId="figure@5.[151,1436,193,1196]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="1" pageNumber="36">Fig. 4</figureCitation>
), 
<collectingCountry id="F30A766FFF9B4E10925DFE034E29C197" box="[816,873,467,490]" name="Laos" pageId="1" pageNumber="36">Laos</collectingCountry>
, Champsak Province, Bolaven Plateau, Paksong, 
<quantity id="4CE59B1AFF9B4E1091FAFE274DACC273" box="[151,236,503,527]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="36" unit="m" value="1310.0">1310 m</quantity>
a.s.l., 
<date id="FFA3103FFF9B4E109058FE274CF0C272" box="[309,432,502,527]" pageId="1" pageNumber="36" value="2005-04-29">29 iv 2005</date>
, N. Yokoi leg.; deposited in the private collection of N. Yokoi.
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF9B4E1091AAFDCE49BFC2B6" pageId="1" pageNumber="36" type="etymology">
<paragraph id="8BA236FFFF9B4E1091AAFDCE482FC203" blockId="1.[151,1436,358,754]" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091AAFDCE4C0EC24A" bold="true" box="[199,334,542,567]" pageId="1" pageNumber="36">Etymology.</emphasis>
The new species is named in honour of Klaus Reinhardt, a 
<collectingCountry id="F30A766FFF9B4E109288FDCF497FC24A" box="[997,1087,543,567]" name="Germany" pageId="1" pageNumber="36">German</collectingCountry>
odonatologist who contributed greatly to our knowledge of 
<taxonomicName id="4C1D4D7CFF9B4E109089FD924F03C226" box="[484,579,578,603]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="36" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
by taking an active part in establishing the International Dragonfly Fund, including personal donations, editing of 
<emphasis id="B969EAEDFF9B4E109336FDB64E8BC203" box="[603,971,614,639]" italics="true" pageId="1" pageNumber="36">Odonatological Abstract Service</emphasis>
and numerous papers in 
<emphasis id="B969EAEDFF9B4E10958EFDB7482FC203" box="[1251,1391,615,638]" italics="true" pageId="1" pageNumber="36">IDF-Report.</emphasis>
</paragraph>
<paragraph id="8BA236FFFF9B4E1091AAFD5D49BFC2B6" blockId="1.[151,1436,358,754]" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091AAFD5D4CC9C2DB" bold="true" box="[199,393,653,678]" pageId="1" pageNumber="36">Short diagnosis.</emphasis>
<taxonomicName id="4C1D4D7CFF9B4E1090FFFD5D4F69C2DB" box="[402,553,653,678]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAEDFF9B4E1090FFFD5D4F69C2DB" box="[402,553,653,678]" italics="true" pageId="1" pageNumber="36">Asiagomphus</emphasis>
</taxonomicName>
with S10 with a large inclined caudal lobe expanding between cerci for half their length in dorsal view, and cerci expanded basally to strong median angulation of outer margins.
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF9B4E1391AAFD09481FC7E7" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36" type="materials_examined">
<paragraph id="8BA236FFFF9B4E1091AAFD094EB0C28F" blockId="1.[151,1436,358,754]" box="[199,1008,729,754]" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091AAFD094C3BC28F" bold="true" box="[199,379,729,754]" pageId="1" pageNumber="36">
<typeStatus id="54A6885DFF9B4E1091AAFD094C74C28F" box="[199,308,729,754]" pageId="1" pageNumber="36" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Body black with pale, dull yellowish markings (
<figureCitation id="13262A7AFF9B4E1092F0FD094E9EC28F" box="[925,990,729,754]" captionStart="FIGURE 1" captionStartId="1.[151,250,1802,1824]" captionTargetBox="[151,1436,809,1777]" captionTargetId="figure@1.[151,1436,809,1777]" captionTargetPageId="1" captionText="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." httpUri="https://zenodo.org/record/271165/files/figure.png" pageId="1" pageNumber="36">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF626677FF9B4E1091FAF8DA4EB5C743" httpUri="https://zenodo.org/record/271165/files/figure.png" pageId="1" pageNumber="36" targetBox="[151,1436,809,1777]" targetPageId="1">
<paragraph id="8BA236FFFF9B4E1091FAF8DA4EB5C743" blockId="1.[151,1436,1802,1854]" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091FAF8DA4C53C762" bold="true" box="[151,275,1802,1824]" pageId="1" pageNumber="36">FIGURE 1.</emphasis>
Holotype male of 
<taxonomicName id="4C1D4D7CFF9B4E1090BDF8DA4F83C75D" box="[464,707,1802,1824]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9B4E1090BDF8DA4F83C75D" box="[464,707,1802,1824]" italics="true" pageId="1" pageNumber="36">Asiagomphus reinhardti</emphasis>
</taxonomicName>
at the type locality (Cambodia, Mondulkiri Province, 
<emphasis id="B969EAEDFF9B4E109585F8DA49B0C75D" bold="true" box="[1256,1264,1802,1824]" pageId="1" pageNumber="36">‘</emphasis>
Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014).
</paragraph>
</caption>
<paragraph id="8BA236FFFF9B4E1391AAF8B64CF1C086" blockId="1.[151,1437,1894,2027]" lastBlockId="2.[151,1437,151,2018]" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
<emphasis id="B969EAEDFF9B4E1091AAF8B64C44C702" box="[199,260,1894,1919]" italics="true" pageId="1" pageNumber="36">Head</emphasis>
. Eyes green in life (
<figureCitation id="13262A7AFF9B4E109091F8B64F02C702" box="[508,578,1894,1919]" captionStart="FIGURE 1" captionStartId="1.[151,250,1802,1824]" captionTargetBox="[151,1436,809,1777]" captionTargetId="figure@1.[151,1436,809,1777]" captionTargetPageId="1" captionText="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." httpUri="https://zenodo.org/record/271165/files/figure.png" pageId="1" pageNumber="36">Fig. 1</figureCitation>
). Face black with pale pattern (
<figureCitation id="13262A7AFF9B4E1092ACF8B64947C702" box="[961,1031,1894,1919]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="1" pageNumber="36">Fig. 2</figureCitation>
d–e). Labium blackish brown with squamae yellowish and lateral lobes gradually lightening to yellowish in basal parts (
<figureCitation id="13262A7AFF9B4E109526F85A49CDC7DE" box="[1099,1165,1930,1955]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="1" pageNumber="36">Fig. 2</figureCitation>
h). Mandible bases pale. Labrum centrally with pair of indistinct slanting pale strokes at middle. Anteclypeus dark brown. Postclypeus black. Frons black with an evenly broad straight pale stripe along ridge, its ends at eyes rounded (
<figureCitation id="13262A7AFF9B4E1095BBF802485AC796" box="[1238,1306,2002,2027]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="1" pageNumber="36">Fig. 2</figureCitation>
d). Antenna brownish but 1st segment blackish basally. Vertex and occiput black (
<figureCitation id="13262A7AFF984E1392CDFF4A4EA4C0CE" box="[928,996,154,179]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">Fig. 2</figureCitation>
e–g). A paired swelling, with indistinct pale marks, behind each lateral ocellus (
<figureCitation id="13262A7AFF984E13930BFF6E4FEBC0AA" box="[614,683,190,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">Fig. 2</figureCitation>
e–f). Occiput with a small and gentle swelling at middle but hind margin straight (
<figureCitation id="13262A7AFF984E13903DFF324CD4C086" box="[336,404,226,251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">
Fig. 
<quantity id="4CE59B1AFF984E1390E9FF334CD4C086" box="[388,404,227,251]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="37" unit="g" value="2.0">2</quantity>
</figureCitation>
g).
</paragraph>
<paragraph id="8BA236FFFF984E1391AAFED6492CC267" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
<emphasis id="B969EAEDFF984E1391AAFED64C58C162" box="[199,280,262,287]" italics="true" pageId="2" pageNumber="37">Thorax</emphasis>
black with a pale pattern (
<figureCitation id="13262A7AFF984E13933EFED64FE3C162" box="[595,675,262,287]" captionStart="FIGURE 1" captionStartId="1.[151,250,1802,1824]" captionTargetBox="[151,1436,809,1777]" captionTargetId="figure@1.[151,1436,809,1777]" captionTargetPageId="1" captionText="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." httpUri="https://zenodo.org/record/271165/files/figure.png" pageId="2" pageNumber="37">Figs. 1</figureCitation>
, 
<figureCitation id="13262A7AFF984E1393D9FED64F82C162" box="[692,706,262,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">2</figureCitation>
a–b). Prothorax with a pair of lateral spots and a pair of small, nearly fused dorsal spots, its posterior margin laterally outlined with pale streak; posterior lobe black, hairy (
<figureCitation id="13262A7AFF984E139402FEFA4DE7C11A" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">Fig. 2</figureCitation>
a). Synthorax (
<figureCitation id="13262A7AFF984E139026FE9E4CCEC11A" box="[331,398,334,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">Fig. 2</figureCitation>
b) with following pale pattern: Dorsal stripes broadening anteriorly towards collar forming a pattern of two inverted 7s. Antehumeral stripes not interrupted, broadening at antealar sinus. A broad stripe occupying most of mesepimeron. Mesepisternum with a small upper spot and a tiny lower dot. Metepimeron pale except for its black margins. Inframesepisternum and inframetepisternum pale in posterior parts, gradually turning black to anterior part. Subalar ridges marked with black. Ventral side of thorax pale behind hind coxae from which two diverging stripes sprout, going along poststernum and forming a pliers-like pattern.
</paragraph>
<paragraph id="8BA236FFFF984E1391AAFDF84C17C28E" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">Legs entirely black. Mid and hind femora with numerous short spines on adaxial surface diminishing to smaller pointed knobs at sides; these spines and knobs disposed rather irregularly but middle of adaxial surface flat and free from spines, so an impression of two not well-arranged rows of larger spines (and sparse strong setae) on both sides; abaxial side smooth. Tibiae with four sharp lengthwise ridges, of which two adaxial are set with rather sparse strong spines and two abaxial with numerous, dense small spinules. Apices of extended hind femora reach posterior hamuli.</paragraph>
<paragraph id="8BA236FFFF984E1391AAFD2E4819C38F" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
Wings (
<figureCitation id="13262A7AFF984E139072FD2E4C23C36A" box="[287,355,766,791]" captionStart="FIGURE 2" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[338,1254,201,1884]" captionTargetId="figure@3.[332,1254,193,1893]" captionTargetPageId="3" captionText="FIGURE 2. Details of head and thorax of holotype male of Asiagomphus reinhardti; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale." httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="2" pageNumber="37">Fig. 2</figureCitation>
c) hyaline, very slightly tinted brownish; venation blackish brown, crossveins between principal veins at anterior edge partly brown. Antenodals (1st and 5th primary) 15 (right)–16 (left) on forewings, 10 on hindwings; postnodals 14 on forewing, 15 on hindwings. Triangles not crossed. Three crossveins between Arc and R1–R4 junction both above and below Rs on forewing, two above Rs and one below Rs on hindwing. Anal loop not defined. Anal triangle well formed, 3-celled. Tornus strongly sinuose at inner wing margin but obtuse, not pointed; membranule extremely narrow, grey. Pterostigmata brownish black, with black bordering veins, of which lengthwise ones swollen, below covering 4 cells on forewings and 4.5 (right) and 5 (left) cells on hindwings.
</paragraph>
<paragraph id="8BA236FFFF984E1391AAFC2D49F0C503" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
Abdomen rather narrow at S3–6, evenly expanding laterally at S7–9, black with pale markings as follows (
<figureCitation id="13262A7AFF984E139402FC2D4DE4C447" captionStart="FIGURE 1" captionStartId="1.[151,250,1802,1824]" captionTargetBox="[151,1436,809,1777]" captionTargetId="figure@1.[151,1436,809,1777]" captionTargetPageId="1" captionText="FIGURE 1. Holotype male of Asiagomphus reinhardti at the type locality (Cambodia, Mondulkiri Province, ‘ Loringae brook’, the left tributary of the main river downstream from Buu Sraa Waterfall, 15 vi 2014)." httpUri="https://zenodo.org/record/271165/files/figure.png" pageId="2" pageNumber="37">Fig. 1</figureCitation>
): S1 with a dorsal triangle directed anteriorly and on either side with a ventral pale area broadly extending dorsally along tergite posterior margin. S2 with a narrowly triangular dorsal spot directed posteriorly and on either side with a pale area composed of a larger anteroventral part including auricle and a smaller posteroventral part forming an irregular up-directed protuberance. S3 with broad anteroventral pale areas extending to ¼ of segment length. S4–7 with anterior semirings incised but not fully interrupted mid-dorsally, not reaching tergite ventral margins. S8 with a pair of narrow dorsolateral streaks along anterior margin and a pair of elongate, slanting, pointed-below dorsolateral spots near posterior margin. S9 with a broad apical spot at posterior margin with a broad rounded incision at middle, occupying 0.25–0.4 of tergite length (
<figureCitation id="13262A7AFF984E13929DFACD4976C54B" box="[1008,1078,1309,1334]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
a–b). Ventral margins of S3–7 tergites marked with yellowish streaks. Dorsal area along posterior margin of S1 tergite set with very dense long brown hairs; posterior margin of S1 sternite and ventral margins of S2 tergite with finer hairs.
</paragraph>
<paragraph id="8BA236FFFF984E1391AAFA5A4C36C698" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
S10 projecting between cerci as a striking large subquadrangular lobe with broadly rounded corners, directed at ca 45° down and behind; in dorsal view reaching half length of cerci (
<figureCitation id="13262A7AFF984E1392D0FA7D4EBFC5BB" box="[957,1023,1453,1478]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
a, c–d), in lateral view reaching level of lower margin of cercus base (
<figureCitation id="13262A7AFF984E13936BFA014F0AC597" box="[518,586,1489,1514]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
b, e). Cerci in dorsal view (
<figureCitation id="13262A7AFF984E1392EFFA014E86C597" box="[898,966,1489,1514]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
a, c–d) broad in basal half but tapering in apical half and nearly pointed but extreme tips slightly truncated, with evenly concave inner margins but blunt lateral prominences at middle of outer margins (which are slightly concave before and after it). In lateral view (
<figureCitation id="13262A7AFF984E139402F9C94DE5C62B" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
b, e), cercus also broad in basal half and tapering to a poined apex, with its upper margin slightly concave while its lower margin has a strong median prominence. Epiproct extends to same level as cercus, in dorsal view (
<figureCitation id="13262A7AFF984E13944BF9B14828C607" box="[1318,1384,1633,1658]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
a, c– d) with widely divaricate broad branches, with both margins concave and rounded and with slightly hooked apices; in lateral view (
<figureCitation id="13262A7AFF984E139038F9794CDCC6BF" box="[341,412,1705,1730]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
b, e) robust with broad up-hooked apices; outer surface of basal half of epiproct arms noticeably concave.
</paragraph>
<paragraph id="8BA236FFFF984E1391AAF922481FC7E7" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
Accessory genitalia black, of a structure typical for the genus and without specific features (
<figureCitation id="13262A7AFF984E139473F9214825C777" box="[1310,1381,1777,1802]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
f–h). Anterior hamulus rather slender, tapering towards apex so that broader part not seen from lateral view, posterolateral margin of apical part with six fine teeth 
<figureCitation id="13262A7AFF984E139243F8E94E35C72F" box="[814,885,1849,1874]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
h). Posterior hamulus smooth, without marked angulations but with slight anterolateral rib on anterolateral surface of its basal part, apices broadly and strongly hooked inside in ventral and subventral views (
<figureCitation id="13262A7AFF984E1393C5F8514FA9C7E7" box="[680,745,1921,1946]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
f–g). Vesica spermalis with a long flagellum (
<figureCitation id="13262A7AFF984E139584F851486AC7E7" box="[1257,1322,1921,1946]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="2" pageNumber="37">Fig. 3</figureCitation>
j–k).
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF984E1791AAF8774E94C345" lastPageId="6" lastPageNumber="41" pageId="2" pageNumber="37" type="description">
<paragraph id="8BA236FFFF984E1391AAF877497CC79F" blockId="2.[151,1437,151,2018]" pageId="2" pageNumber="37">
<emphasis id="B969EAEDFF984E1391AAF8774CFAC7C0" box="[199,442,1958,1982]" italics="true" pageId="2" pageNumber="37">Measurements (mm).</emphasis>
Hindwing 35; abdomen without anal appendages 41; total length (with head and appendages) 57; hind femur 8; fore wing pterostigma 4, hind wing pterostigma 3.5.
</paragraph>
<caption id="DF626677FF994E1291FAF8AB4C57C7B3" httpUri="https://zenodo.org/record/271166/files/figure.png" pageId="3" pageNumber="38" targetBox="[338,1254,201,1884]" targetPageId="3">
<paragraph id="8BA236FFFF994E1291FAF8AB4C57C7B3" blockId="3.[151,1437,1915,1998]" pageId="3" pageNumber="38">
<emphasis id="B969EAEDFF994E1291FAF8AB4C55C7ED" bold="true" box="[151,277,1915,1937]" pageId="3" pageNumber="38">FIGURE 2.</emphasis>
Details of head and thorax of holotype male of 
<taxonomicName id="4C1D4D7CFF994E12927BF8AB494CC7EC" box="[790,1036,1915,1937]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF994E12927BF8AB494CC7EC" box="[790,1036,1915,1937]" italics="true" pageId="3" pageNumber="38">Asiagomphus reinhardti</emphasis>
</taxonomicName>
; a—prothorax, right lateral view; bsynthorax, right lateral view; c—right wings; d—head, frontal view; e—head, dorsal view; f—vertex; g—occiput; h—labium. Not to scale.
</paragraph>
</caption>
<caption id="DF626677FF9E4E1591FAF8AB4F6DC790" httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="4" pageNumber="39" targetBox="[156,1409,198,1883]" targetPageId="4">
<paragraph id="8BA236FFFF9E4E1591FAF8AB4F6DC790" blockId="4.[151,1436,1915,2029]" pageId="4" pageNumber="39">
<emphasis id="B969EAEDFF9E4E1591FAF8AB4C55C7ED" bold="true" box="[151,277,1915,1937]" pageId="4" pageNumber="39">FIGURE 3.</emphasis>
Abdominal structures of holotype male of 
<taxonomicName id="4C1D4D7CFF9E4E1593B4F8AB4E8FC7EC" box="[729,975,1915,1937]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9E4E1593B4F8AB4E8FC7EC" box="[729,975,1915,1937]" italics="true" pageId="4" pageNumber="39">Asiagomphus reinhardti</emphasis>
</taxonomicName>
; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale.
</paragraph>
</caption>
<caption id="DF626677FF9F4E1491FAFB114C8DC569" httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="5" pageNumber="40" targetBox="[151,1427,193,1194]" targetPageId="5">
<paragraph id="8BA236FFFF9F4E1491FAFB114C8DC569" blockId="5.[151,1436,1217,1300]" pageId="5" pageNumber="40">
<emphasis id="B969EAEDFF9F4E1491FAFB114C57C4AB" bold="true" box="[151,279,1217,1239]" pageId="5" pageNumber="40">FIGURE 4.</emphasis>
Abdominal structures of paratype male of 
<taxonomicName id="4C1D4D7CFF9F4E14938AFB114E9FC4AA" box="[743,991,1217,1239]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9F4E14938AFB114E9FC4AA" box="[743,991,1217,1239]" italics="true" pageId="5" pageNumber="40">Asiagomphus reinhardti</emphasis>
</taxonomicName>
from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale.
</paragraph>
</caption>
<paragraph id="8BA236FFFF9F4E1491AAFA954CD5C523" blockId="5.[151,1437,1349,2029]" box="[199,405,1349,1374]" pageId="5" pageNumber="40">
<emphasis id="B969EAEDFF9F4E1491AAFA954C5EC523" bold="true" box="[199,286,1349,1374]" pageId="5" pageNumber="40">Female</emphasis>
unknown.
</paragraph>
<paragraph id="8BA236FFFF9F4E1491AAFABC4C0DC6FC" blockId="5.[151,1437,1349,2029]" pageId="5" pageNumber="40">
<emphasis id="B969EAEDFF9F4E1491AAFABC4C7BC5F8" bold="true" box="[199,315,1388,1413]" pageId="5" pageNumber="40">
<typeStatus id="54A6885DFF9F4E1491AAFABC4C76C5F8" box="[199,310,1388,1413]" pageId="5" pageNumber="40" type="paratype">Paratype</typeStatus>
.
</emphasis>
Similar to 
<typeStatus id="54A6885DFF9F4E1490ACFABC4F66C5F8" box="[449,550,1388,1413]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
, especially in structural characters, but differs in the following traits, mostly related to the pattern: pale elements of body pattern saturated yellow (
<figureCitation id="13262A7AFF9F4E1492A1FA414954C5D7" box="[972,1044,1425,1450]" captionStart="FIGURE 4" captionStartId="5.[151,250,1217,1239]" captionTargetBox="[151,1427,193,1194]" captionTargetId="figure@5.[151,1436,193,1196]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="5" pageNumber="40">Fig. 4</figureCitation>
b); paired yellow spots on labrum larger but diffuse; antehumeral stripe completely reduced to a tiny antealar spot (the most noticeable difference) (
<figureCitation id="13262A7AFF9F4E1491F2FA094DAAC58F" box="[159,234,1497,1522]" captionStart="FIGURE 4" captionStartId="5.[151,250,1217,1239]" captionTargetBox="[151,1427,193,1194]" captionTargetId="figure@5.[151,1436,193,1196]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="5" pageNumber="40">Fig. 4</figureCitation>
a,b); both pale spots on the metepisternum missing (
<figureCitation id="13262A7AFF9F4E14920BFA094EF1C58C" box="[870,945,1497,1522]" captionStart="FIGURE 4" captionStartId="5.[151,250,1217,1239]" captionTargetBox="[151,1427,193,1194]" captionTargetId="figure@5.[151,1436,193,1196]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="5" pageNumber="40">Fig. 4</figureCitation>
a); 12 postnodals on forewing; 13–14 on hindwing; dorsolateral spots at posterior margin of S8 very small (
<figureCitation id="13262A7AFF9F4E1492EBFA2D4E8BC668" box="[902,971,1533,1558]" captionStart="FIGURE 4" captionStartId="5.[151,250,1217,1239]" captionTargetBox="[151,1427,193,1194]" captionTargetId="figure@5.[151,1436,193,1196]" captionTargetPageId="5" captionText="FIGURE 4. Abdominal structures of paratype male of Asiagomphus reinhardti from Laos, Champsak Province, Bolaven Plateau, Paksong; a—thorax, frontal view; b—general habitus; c—anal appendages, lateral view; d—vesica spermalis distal part, lateral view. Not to scale." httpUri="https://zenodo.org/record/271168/files/figure.png" pageId="5" pageNumber="40">Fig. 4</figureCitation>
c); large dorsoposterior spot on S9 with a straight anterior margin (occupying about 1/3 of segment length). Measurements (mm): hindwing 33.5; abdomen without anal appendages 38.7; total length (with head and appendages) 54.2; hind femur 8.1; fore and hind pterostigma 3.9.
</paragraph>
<paragraph id="8BA236FFFF9F4E1491AAF9404C65C7FC" blockId="5.[151,1437,1349,2029]" pageId="5" pageNumber="40">
<emphasis id="B969EAEDFF9F4E1491AAF9404C90C6D4" bold="true" box="[199,464,1680,1705]" pageId="5" pageNumber="40">Differential diagnosis.</emphasis>
The large caudal lobe of S10 (
<figureCitation id="13262A7AFF9F4E149251F9404EC1C6D4" box="[828,897,1680,1705]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="5" pageNumber="40">Fig. 3</figureCitation>
a, c–d) is unique in the genus. Some moderate roundish expansion of the S10 posterior margin is present in 
<taxonomicName id="4C1D4D7CFF9F4E149220F9654837C6B0" authority="(Doi &amp; Okumura, 1937) (Lee 2001)" authorityName="(Doi &amp; Okumura, 1937) (Lee" authorityYear="2001" box="[845,1399,1716,1741]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="B969EAEDFF9F4E149220F9654E93C6B0" box="[845,979,1717,1741]" italics="true" pageId="5" pageNumber="40">A. coreanus</emphasis>
(Doi &amp; Okumura, 1937) (
<bibRefCitation id="EF8C4B0EFF9F4E149592F965482FC6B0" author="Lee" box="[1279,1391,1717,1741]" pageId="5" pageNumber="40" refString="Lee, S. - M. (2001) The dragonflies of Korean Peninsula (Odonata). Junghaen-Sa, Seoul." type="book" year="2001">Lee 2001</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C1D4D7CFF9F4E1494E9F9654CE7C68C" authority="Chao, 1920" authorityName="Chao" authorityYear="1920" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B969EAEDFF9F4E1494E9F9654C5DC68C" italics="true" pageId="5" pageNumber="40">A. hainanensis</emphasis>
Chao, 1920
</taxonomicName>
, 
<taxonomicName id="4C1D4D7CFF9F4E1490DCF9094FBBC68C" authority="Needham, 1930" authorityName="Needham" authorityYear="1930" box="[433,763,1752,1777]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="septimus">
<emphasis id="B969EAEDFF9F4E1490DCF9094F71C68C" box="[433,561,1753,1777]" italics="true" pageId="5" pageNumber="40">A. septimus</emphasis>
(Needham, 1930)
</taxonomicName>
and 
<taxonomicName id="4C1D4D7CFF9F4E14925DF90949A3C68C" authority="(Chao, 1953) (Chao 1990)" authorityName="(Chao, 1953) (Chao" authorityYear="1990" box="[816,1251,1752,1777]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B969EAEDFF9F4E14925DF9094EF4C68C" box="[816,948,1753,1777]" italics="true" pageId="5" pageNumber="40">A. pacificus</emphasis>
(Chao, 1953) (
<bibRefCitation id="EF8C4B0EFF9F4E149531F908499BC68C" author="Chao" box="[1116,1243,1752,1777]" pageId="5" pageNumber="40" refString="Chao [Zhao], X. - f. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou." type="book" year="1990">Chao 1990</bibRefCitation>
)
</taxonomicName>
. Another unique feature of the new species is the cercus shape (
<figureCitation id="13262A7AFF9F4E1493C9F92C4FA6C768" box="[676,742,1788,1813]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[156,1409,198,1883]" captionTargetId="figure@4.[156,1430,193,1893]" captionTargetPageId="4" captionText="FIGURE 3. Abdominal structures of holotype male of Asiagomphus reinhardti; a—end of abdomen, dorsal view; b—ditto, lateral view; c—anal appendages, dosral view; d—ditto, posterodorsal view; e—ditto, lateral view; f—secondary genitalia, lateral view; g—hamulil, oblique ventrolateral view; h—anterior hamulus apical part, lateral view; j—vesica spermalis, lateral view; k—ditto, apical part. Not to scale." httpUri="https://zenodo.org/record/271167/files/figure.png" pageId="5" pageNumber="40">Fig. 3</figureCitation>
a–e), with basal half expanded to a median angular projection of the outer margin, well seen in both dorsal and lateral view. In other species the cercus outer margin is smooth even if somewhat expanded. None of the known 
<taxonomicName id="4C1D4D7CFF9F4E1493D8F8944E0CC720" box="[693,844,1860,1885]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAEDFF9F4E1493D8F8944E0CC720" box="[693,844,1860,1885]" italics="true" pageId="5" pageNumber="40">Asiagomphus</emphasis>
</taxonomicName>
species shares either of these two characters with the new species.
</paragraph>
<paragraph id="8BA236FFFF9F4E1791AAF85C4C50C1F5" blockId="5.[151,1437,1349,2029]" lastBlockId="6.[151,1436,151,824]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
Identifying the new species (both the 
<typeStatus id="54A6885DFF9F4E14931EF85C4F95C7D8" box="[627,725,1932,1957]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54A6885DFF9F4E149263F85D4E30C7D8" box="[782,880,1933,1957]" pageId="5" pageNumber="40" type="paratype">paratype</typeStatus>
) using the key by 
<bibRefCitation id="EF8C4B0EFF9F4E14952EF85C4993C7D8" author="Chao" box="[1091,1235,1932,1957]" pageId="5" pageNumber="40" refString="Chao [Zhao], X. - f. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou." type="book" year="1990">Chao (1990)</bibRefCitation>
, based on colourpattern characters and hence not so reliable, would lead to 
<taxonomicName id="4C1D4D7CFF9F4E149225F8614EB1C7B4" box="[840,1009,1968,1993]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B969EAEDFF9F4E149225F8614EB1C7B4" box="[840,1009,1968,1993]" italics="true" pageId="5" pageNumber="40">A. hainanensis</emphasis>
</taxonomicName>
, to which the new species has some similarity of body pattern, but it has only two small pale spots at most on the metepisternum versus a broad pale stripe in 
<taxonomicName id="4C1D4D7CFF9C4E17906CFF484CEAC0CD" box="[257,426,151,176]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B969EAEDFF9C4E17906CFF484CEAC0CD" box="[257,426,151,176]" italics="true" pageId="6" pageNumber="41">A. hainanensis</emphasis>
</taxonomicName>
. Note that 
<taxonomicName id="4C1D4D7CFF9C4E17935FFF484F9BC0CD" box="[562,731,151,176]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B969EAEDFF9C4E17935FFF484F9BC0CD" box="[562,731,151,176]" italics="true" pageId="6" pageNumber="41">A. hainanensis</emphasis>
</taxonomicName>
demonstrates a variable expression of the antehumeral light stripes (
<bibRefCitation id="EF8C4B0EFF9C4E17919CFF6C4C30C0A9" author="Chao" box="[241,368,188,213]" pageId="6" pageNumber="41" refString="Chao [Zhao], X. - f. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou." type="book" year="1990">Chao 1990</bibRefCitation>
) as well as 
<taxonomicName id="4C1D4D7CFF9C4E179091FF6C4FCCC0A8" box="[508,652,188,213]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9C4E179091FF6C4FCCC0A8" box="[508,652,188,213]" italics="true" pageId="6" pageNumber="41">A. reinhardti</emphasis>
</taxonomicName>
, in which the stripes are complete in the 
<typeStatus id="54A6885DFF9C4E179500FF6C498FC0A8" box="[1133,1231,188,213]" pageId="6" pageNumber="41" type="holotype">holotype</typeStatus>
and absent in the 
<typeStatus id="54A6885DFF9C4E1791FAFF304DBBC085" box="[151,251,224,248]" pageId="6" pageNumber="41" type="paratype">paratype</typeStatus>
. Perhaps this character is environmentally modifiable; note that the 
<typeStatus id="54A6885DFF9C4E179299FF0F4916C085" box="[1012,1110,223,248]" pageId="6" pageNumber="41" type="holotype">holotype</typeStatus>
was taken at the elevation of 
<quantity id="4CE59B1AFF9C4E1791FAFED44D9FC161" box="[151,223,260,285]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="41" unit="m" value="450.0">450 m</quantity>
a.s.l. and the 
<typeStatus id="54A6885DFF9C4E179010FED54C9DC160" box="[381,477,261,285]" pageId="6" pageNumber="41" type="paratype">paratype</typeStatus>
at 
<quantity id="4CE59B1AFF9C4E17936CFED44F16C161" box="[513,598,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="6" pageNumber="41" unit="m" value="1310.0">1310 m</quantity>
a.s.l. The general shape of the hamulus of the new species also resembles that of 
<taxonomicName id="4C1D4D7CFF9C4E17918BFEF84CCAC13D" box="[230,394,295,320]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B969EAEDFF9C4E17918BFEF84CCAC13D" box="[230,394,295,320]" italics="true" pageId="6" pageNumber="41">A. hainanensis</emphasis>
</taxonomicName>
and also 
<taxonomicName id="4C1D4D7CFF9C4E179099FEF84EB9C13D" authority="Liu &amp; Chao, 1990 (Chao 1990)" authorityName="Liu &amp; Chao, 1990 (Chao" authorityYear="1990" box="[500,1017,295,320]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="motuoensis">
<emphasis id="B969EAEDFF9C4E179099FEF84FD1C13D" box="[500,657,296,320]" italics="true" pageId="6" pageNumber="41">A. motuoensis</emphasis>
Liu &amp; 
<bibRefCitation id="EF8C4B0EFF9C4E17938FFEF74E23C13D" author="Chao" box="[738,867,295,320]" pageId="6" pageNumber="41" refString="Chao [Zhao], X. - f. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou." type="book" year="1990">Chao, 1990</bibRefCitation>
(
<bibRefCitation id="EF8C4B0EFF9C4E17921CFEF74EB1C13D" author="Chao" box="[881,1009,295,320]" pageId="6" pageNumber="41" refString="Chao [Zhao], X. - f. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou." type="book" year="1990">Chao 1990</bibRefCitation>
)
</taxonomicName>
, but the cercus shape in the two latter species strongly differs from 
<taxonomicName id="4C1D4D7CFF9C4E179082FE9C4F1EC118" box="[495,606,332,357]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9C4E179082FE9C4F1EC118" box="[495,606,332,357]" italics="true" pageId="6" pageNumber="41">reinhardti</emphasis>
</taxonomicName>
in having a weak subapical expansion rather than a strong submedian expansion.
</paragraph>
<paragraph id="8BA236FFFF9C4E1791AAFE444C10C28D" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="41">
Another species which 
<taxonomicName id="4C1D4D7CFF9C4E1790BAFE444F29C1D0" box="[471,617,404,429]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9C4E1790BAFE444F29C1D0" box="[471,617,404,429]" italics="true" pageId="6" pageNumber="41">A. reinhardti</emphasis>
</taxonomicName>
resembles in general habitus is 
<taxonomicName id="4C1D4D7CFF9C4E179289FE4449CDC1D0" box="[996,1165,404,429]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="xanthenatus">
<emphasis id="B969EAEDFF9C4E179289FE4449CDC1D0" box="[996,1165,404,429]" italics="true" pageId="6" pageNumber="41">A. xanthenatus</emphasis>
</taxonomicName>
, described from Upper 
<collectingCountry id="F30A766FFF9C4E1791FAFE684C44C1AD" box="[151,260,440,464]" name="Myanmar" pageId="6" pageNumber="41">Myanmar</collectingCountry>
and reported also from the western 
<collectingCountry id="F30A766FFF9C4E1793F2FE674E43C1AD" box="[671,771,439,464]" name="Thailand" pageId="6" pageNumber="41">Thailand</collectingCountry>
and (as a melanised subspecies 
<taxonomicName id="4C1D4D7CFF9C4E17951EFE684C70C189" authority="Karube, 1990" authorityName="Karube" authorityYear="1990" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="xanthenatus" subSpecies="malayanus">
<emphasis id="B969EAEDFF9C4E17951EFE6848DCC1AD" box="[1139,1436,439,464]" italics="true" pageId="6" pageNumber="41">A. xanthenatus malayanus</emphasis>
<bibRefCitation id="EF8C4B0EFF9C4E1791FAFE0C4C70C189" author="Karube" box="[151,304,476,501]" pageId="6" pageNumber="41" refString="Karube, H. (1990) Description of a new subspecies of Asiagomphus xanthenatus (Williamson) from West Malaysia. Tombo, 33 (1 / 4), 21 - 24." type="journal article" year="1990">Karube, 1990</bibRefCitation>
</taxonomicName>
) from Fraser Hills in Peninsular 
<collectingCountry id="F30A766FFF9C4E1793CFFE0C4E48C188" box="[674,776,476,501]" name="Malaysia" pageId="6" pageNumber="41">Malaysia</collectingCountry>
(
<bibRefCitation id="EF8C4B0EFF9C4E17927AFE0C4E9CC189" author="Williamson" box="[791,988,476,501]" pageId="6" pageNumber="41" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam - II. Subfamilies Cordulegasterinae, Chlorogomphinae and Gomphinae. Proceedings of the U. S. National Museum, 33, 267 - 317. http: // dx. doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article" year="1907">Williamson 1907</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E17928AFE0C493CC189" author="Karube" box="[999,1148,476,501]" pageId="6" pageNumber="41" refString="Karube, H. (1990) Description of a new subspecies of Asiagomphus xanthenatus (Williamson) from West Malaysia. Tombo, 33 (1 / 4), 21 - 24." type="journal article" year="1990">Karube 1990</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E1795EAFE0C4D92C265" author="Hamalainen" pageId="6" pageNumber="41" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the Dragonflies of Thailand. Distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana 1999</bibRefCitation>
, Orr 2005, 
<bibRefCitation id="EF8C4B0EFF9C4E17903EFDD04CB9C265" author="Do" box="[339,505,511,536]" pageId="6" pageNumber="41" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Toan, P. Q. (2011) Anisoptera of Cuc Phuong National park, North Vietnam. International Dragonfly Fund Report, 33, 1 - 18." type="journal article" year="2011">
Do 
<emphasis id="B969EAEDFF9C4E179011FDD14CF5C265" box="[380,437,511,536]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
2011
</bibRefCitation>
). But in this species the cerci have a weak subapical ventral swelling or projection (as in most 
<taxonomicName id="4C1D4D7CFF9C4E179074FDF44CF0C240" box="[281,432,548,573]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAEDFF9C4E179074FDF44CF0C240" box="[281,432,548,573]" italics="true" pageId="6" pageNumber="41">Asiagomphus</emphasis>
</taxonomicName>
spp.) rather than a strong sumbedian projection, and the S10 posterior margin is not at all expanded. 
<taxonomicName id="4C1D4D7CFF9C4E17907CFD984F55C21D" authority="Asahina, 1996" authorityName="Asahina" authorityYear="1996" box="[273,533,583,608]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B969EAEDFF9C4E17907CFD984C27C21D" box="[273,359,584,608]" italics="true" pageId="6" pageNumber="41">A. acco</emphasis>
<bibRefCitation id="EF8C4B0EFF9C4E179002FD974F55C21D" author="Asahina" box="[367,533,583,608]" pageId="6" pageNumber="41" refString="Asahina, S. (1996) Records of the Northern Vietnamese Odonata taken by the expedition members from National Science Museum, Tokyo. 2. Gomphidae. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 22 (1), 21 - 32." type="journal article" year="1996">Asahina, 1996</bibRefCitation>
</taxonomicName>
, ranging in 
<collectingCountry id="F30A766FFF9C4E1793F0FD974FBFC21D" box="[669,767,583,608]" name="Vietnam" pageId="6" pageNumber="41">Vietnam</collectingCountry>
, 
<collectingCountry id="F30A766FFF9C4E179266FD984E01C222" box="[779,833,584,607]" name="Laos" pageId="6" pageNumber="41">Laos</collectingCountry>
and S. 
<collectingCountry id="F30A766FFF9C4E1792F4FD974E9CC21D" box="[921,988,583,608]" name="China" pageId="6" pageNumber="41">China</collectingCountry>
(
<bibRefCitation id="EF8C4B0EFF9C4E179281FD9749CDC21D" author="Asahina" box="[1004,1165,583,608]" pageId="6" pageNumber="41" refString="Asahina, S. (1996) Records of the Northern Vietnamese Odonata taken by the expedition members from National Science Museum, Tokyo. 2. Gomphidae. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 22 (1), 21 - 32." type="journal article" year="1996">Asahina 1996</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E1795F4FD97486CC21D" author="Wilson" box="[1177,1324,583,608]" pageId="6" pageNumber="41" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8 (1), 107 - 168. http: // dx. doi. org / 10.1080 / 13887890.2005.9748247" type="journal article" year="2005">Wilson 2005</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E179455FD984D91C2F9" author="Do" pageId="6" pageNumber="41" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Toan, P. Q. (2011) Anisoptera of Cuc Phuong National park, North Vietnam. International Dragonfly Fund Report, 33, 1 - 18." type="journal article" year="2011">
Do 
<emphasis id="B969EAEDFF9C4E17940EFD9948DCC21D" box="[1379,1436,583,608]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
2011
</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E17918EFDBC4F02C2F9" author="Yokoi" box="[227,578,620,645]" pageId="6" pageNumber="41" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A List of Lao Dragonfliles. Kyoei Printing Co Ltd., Koriyama, 110 pp." type="book" year="2014">Yokoi &amp; Souphanthong 2014</bibRefCitation>
), is no doubt a good species although described as a subspecies of 
<taxonomicName id="4C1D4D7CFF9C4E1794E9FDBC4C5EC2D5" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="xanthenatus">
<emphasis id="B969EAEDFF9C4E1794E9FDBC4C5EC2D5" italics="true" pageId="6" pageNumber="41">A. xanthenatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C4B0EFF9C4E17905CFD5F4C89C2D5" author="Wilson" box="[305,457,655,680]" pageId="6" pageNumber="41" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8 (1), 107 - 168. http: // dx. doi. org / 10.1080 / 13887890.2005.9748247" type="journal article" year="2005">Wilson 2005</bibRefCitation>
, 
<bibRefCitation id="EF8C4B0EFF9C4E1790B4FD404FCAC2D5" author="Do" box="[473,650,655,680]" pageId="6" pageNumber="41" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Toan, P. Q. (2011) Anisoptera of Cuc Phuong National park, North Vietnam. International Dragonfly Fund Report, 33, 1 - 18." type="journal article" year="2011">
Do 
<emphasis id="B969EAEDFF9C4E17936AFD414F04C2D5" box="[519,580,655,680]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
2011
</bibRefCitation>
). 
<taxonomicName id="4C1D4D7CFF9C4E1793CFFD404FBBC2D5" box="[674,763,656,680]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B969EAEDFF9C4E1793CFFD404FBBC2D5" box="[674,763,656,680]" italics="true" pageId="6" pageNumber="41">A. acco</emphasis>
</taxonomicName>
has the cerci with a sumbedian ventral projection in the position corresponding to the projection in 
<emphasis id="B969EAEDFF9C4E179312FD644E55C2B0" box="[639,789,692,717]" italics="true" pageId="6" pageNumber="41">
<taxonomicName id="4C1D4D7CFF9C4E179312FD644E51C2B0" box="[639,785,692,717]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">A. reinhardti</taxonomicName>
,
</emphasis>
but in 
<taxonomicName id="4C1D4D7CFF9C4E17920BFD644EFAC2B1" box="[870,954,692,716]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B969EAEDFF9C4E17920BFD644EFAC2B1" box="[870,954,692,716]" italics="true" pageId="6" pageNumber="41">A. acco</emphasis>
</taxonomicName>
it is smooth and not visible in dorsal view (
<bibRefCitation id="EF8C4B0EFF9C4E1791F2FD084C04C28D" author="Do" box="[159,324,727,752]" pageId="6" pageNumber="41" refString="Do, M. C., Bui, M. H., Nguyen, T. H. &amp; Toan, P. Q. (2011) Anisoptera of Cuc Phuong National park, North Vietnam. International Dragonfly Fund Report, 33, 1 - 18." type="journal article" year="2011">
Do 
<emphasis id="B969EAEDFF9C4E1791A4FD094C42C28D" box="[201,258,727,752]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA236FFFF9C4E1791AAFD2C4E94C345" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="41">
If a new key for 
<taxonomicName id="4C1D4D7CFF9C4E1790E3FD2C4F65C368" box="[398,549,764,789]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAEDFF9C4E1790E3FD2C4F65C368" box="[398,549,764,789]" italics="true" pageId="6" pageNumber="41">Asiagomphus</emphasis>
</taxonomicName>
species were to be produced, it would be reasonable to start it with the first couplet differentiating 
<taxonomicName id="4C1D4D7CFF9C4E1790FAFCF04F65C345" box="[407,549,799,824]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9C4E1790FAFCF04F65C345" box="[407,549,799,824]" italics="true" pageId="6" pageNumber="41">A. reinhardti</emphasis>
</taxonomicName>
from the rest of the genus, as follows:
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF9C4E1791FAFCB74E03C4D2" pageId="6" pageNumber="41" type="key">
<key id="FF8C61CEFF9C4E1791FAFCB748DDC3CA" pageId="6" pageNumber="41">
<keyStep id="30E92E5AFF9C4E1791FAFCB748DDC3CA" pageId="6" pageNumber="41">
<paragraph id="8BA236FFFF9C4E1791FAFCB748DCC3E7" blockId="6.[151,1437,871,951]" pageId="6" pageNumber="41">
<keyLead id="30EC95CAFF9C4E1791FAFCB748DCC3E7" pageId="6" pageNumber="41">
1. S10 with a large caudal lobe protruding between cerci; cerci with a blunt submedian ventrolateral prominence (
<collectingCountry id="F30A766FFF9C4E17945CFCB748D7C300" box="[1329,1431,871,893]" name="Cambodia" pageId="6" pageNumber="41">Cambodia</collectingCountry>
, 
<collectingCountry id="F30A766FFF9C4E17918CFC554C51C3E7" box="[225,273,901,922]" name="Laos" pageId="6" pageNumber="41">Laos</collectingCountry>
)...................................................................................... 
<taxonomicName id="4C1D4D7CFF9C4E17944CFC5448DCC3E7" ID-CoL="9X249" box="[1313,1436,900,922]" class="Insecta" family="Gomphidae" genus="Asiagomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B969EAEDFF9C4E17944CFC5448DCC3E7" box="[1313,1436,900,922]" italics="true" pageId="6" pageNumber="41">A. reinhardti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA236FFFF9C4E1791FAFC7248DDC3CA" blockId="6.[151,1437,871,951]" box="[151,1437,929,951]" pageId="6" pageNumber="41">
<keyLead id="30EC95CAFF9C4E1791FAFC7248DDC3CA" box="[151,1437,929,951]" pageId="6" pageNumber="41">- S10 without a large caudal lobe; cerci with smooth outlines................................. 2 (the rest of the genus).</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BA236FFFF9C4E1791AAFC344E03C4D2" blockId="6.[151,1436,996,1309]" pageId="6" pageNumber="41">
<emphasis id="B969EAEDFF9C4E1791AAFC344C6AC380" bold="true" box="[199,298,996,1021]" pageId="6" pageNumber="41">Habitat.</emphasis>
The old, worn out and not too cautious Cambodian male perched on a sunlit stone at a rocky reach of a forest brook, half shaded by tall evergreen forest, situated downstream from the popular Buu Sraa Waterfalls, at an elevation of about 
<quantity id="4CE59B1AFF9C4E1790E4FBFC4C91C439" box="[393,465,1068,1093]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="41" unit="m" value="450.0">450 m</quantity>
a.s.l. This brook descends from the left slope of the forested main river gorge crossing hilly terrain originally clad with a deciduous dipterocarp forest but presently mostly replaced by plantations. The Laotian male was found at a small mountain stream about 
<quantity id="4CE59B1AFF9C4E179221FBA44E3BC4F1" box="[844,891,1140,1164]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="41" unit="m" value="2.0">2 m</quantity>
wide, in relatively pristine environment at the elevation of 
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a.s.l. The new species is obviously rare.
</paragraph>
</subSubSection>
<subSubSection id="C3076574FF9C4E1791AAFB6C4FAAC560" pageId="6" pageNumber="41" type="distribution">
<paragraph id="8BA236FFFF9C4E1791AAFB6C4FAAC560" blockId="6.[151,1436,996,1309]" pageId="6" pageNumber="41">
<emphasis id="B969EAEDFF9C4E1791AAFB6C4C20C4A8" bold="true" box="[199,352,1212,1237]" pageId="6" pageNumber="41">Distribution.</emphasis>
The species is so far known from two localities in southern 
<collectingCountry id="F30A766FFF9C4E179570FB6D4913C4A9" box="[1053,1107,1213,1236]" name="Laos" pageId="6" pageNumber="41">Laos</collectingCountry>
and eastern 
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, both situated at the western part of the Annamense Mountains. It is expected from southern 
<collectingCountry id="F30A766FFF9C4E1795F9FB0F49B6C485" box="[1172,1270,1247,1272]" name="Vietnam" pageId="6" pageNumber="41">Vietnam</collectingCountry>
, the border of which lies just 
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east of the Cambodian locality.
</paragraph>
</subSubSection>
</treatment>
</document>