411 lines
53 KiB
XML
411 lines
53 KiB
XML
<document id="776841EE09381EB0AD98BCAD8D529022" ID-CLB-Dataset="45690" ID-DOI="10.5281/zenodo.279614" ID-GBIF-Dataset="b1782435-1df3-4d5d-a446-16efa7283c37" ID-ISSN="1175-5326" ID-Zenodo-Dep="279614" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460425584293" checkinUser="plazi" docAuthor="Last, Peter R. & Séret, Bernard" docDate="2012" docId="03B487D6FFE3FFF3E2EA57D6FB37F8F9" docLanguage="en" docName="zt03155p064.pdf" docOrigin="Zootaxa 3155" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Brochiraja vittacauda Last & Séret, 2012, sp. nov." docType="treatment" docVersion="11" lastPageNumber="61" masterDocId="FF8DFFAEFFEAFFFDE27D5405FF9FFFA9" masterDocTitle="Two new softnose skates of the genus Brochiraja (Rajoidei: Arhynchobatidae) from the deepwater slopes and banks of the Norfolk Ridge (South-West Pacific)" masterLastPageNumber="64" masterPageNumber="47" pageNumber="56" updateTime="1698330627290" updateUser="ExternalLinkService">
|
||
<mods:mods id="1A077D16A58BD58E0B023FDA29FAFB42" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="55C859F92443CA57AA694540F9A4B465">
|
||
<mods:title id="E1264B1B573343F883ED342049B44FFE">Two new softnose skates of the genus Brochiraja (Rajoidei: Arhynchobatidae) from the deepwater slopes and banks of the Norfolk Ridge (South-West Pacific)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="342F434F313597E194B5F2030E6DC5FA" type="personal">
|
||
<mods:role id="B1E056767ECF824E275003100BB536C0">
|
||
<mods:roleTerm id="416DB0574320B244314B1AFDE578A855">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CAE422F97E496F4EB5EAA4682549919B">Last, Peter R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="11F6775544BC1D183C70047DE966D7C3" type="personal">
|
||
<mods:role id="2B0A735FD20E034FFDAB6D21A846689C">
|
||
<mods:roleTerm id="6FE918D12D526A6069CC6418D352AA62">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6AA6F8300804CFFE0CBAC1C82A919024">Séret, Bernard</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A69D93F8BC11B095A45CCEA642483FF4">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B89CB7E6B08224FB7194525E12DADE6D" type="host">
|
||
<mods:titleInfo id="188F58C5F7DA5E1B69F0DC446A7C414C">
|
||
<mods:title id="98570951AE30DA5FF53348B5BF7C0126">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="5C9550E978800B154F498B533129A068">
|
||
<mods:date id="3D8C394989E1021F7C53125B7CA44442">2012</mods:date>
|
||
<mods:detail id="5F73387ABE79AA9BF9C3AB824A2CCA27" type="volume">
|
||
<mods:number id="A2D0349DAF66015BFCCAAF027EA0DAE4">3155</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="46B9BF661E07BC5C1B9259DA2C51FAAA" unit="page">
|
||
<mods:start id="B80ACF45B847D28CB3EFAC6955647B69">47</mods:start>
|
||
<mods:end id="11FE2CAA02D55F9A3F99D6CDF151A558">64</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="45AF81DD297E71950B1638D48E485FA3">journal article</mods:classification>
|
||
<mods:identifier id="37697C4E9102689571D13F4A7688217F" type="CLB-Dataset">45690</mods:identifier>
|
||
<mods:identifier id="61DAB662491DB2BDD726F4FE78F1FA2C" type="DOI">10.5281/zenodo.279614</mods:identifier>
|
||
<mods:identifier id="B88E311FE8A8D94C1145F1D1535A0CA0" type="GBIF-Dataset">b1782435-1df3-4d5d-a446-16efa7283c37</mods:identifier>
|
||
<mods:identifier id="9434FD31F0F10588E03E6E934478F1E4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="6CA4533690D25EB458A0A8967A57135F" type="Zenodo-Dep">279614</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B487D6FFE3FFF3E2EA57D6FB37F8F9" ID-DOI="http://doi.org/10.5281/zenodo.5679989" ID-GBIF-Taxon="119428290" ID-Zenodo-Dep="5679989" LSID="urn:lsid:plazi:treatment:03B487D6FFE3FFF3E2EA57D6FB37F8F9" httpUri="http://treatment.plazi.org/id/03B487D6FFE3FFF3E2EA57D6FB37F8F9" lastPageId="14" lastPageNumber="61" pageId="9" pageNumber="56">
|
||
<subSubSection id="C307654BFFE3FFF4E2EA57D6FEAAFBDC" pageId="9" pageNumber="56" type="nomenclature">
|
||
<paragraph id="8BA236C0FFE3FFF4E2EA57D6FE6BFC45" blockId="9.[151,500,979,1039]" box="[151,500,979,1004]" pageId="9" pageNumber="56">
|
||
<heading id="D0EA81ACFFE3FFF4E2EA57D6FE6BFC45" bold="true" box="[151,500,979,1004]" fontSize="10" level="2" pageId="9" pageNumber="56" reason="2">
|
||
<emphasis id="B969EAD2FFE3FFF4E2EA57D6FE6BFC45" bold="true" box="[151,500,979,1004]" pageId="9" pageNumber="56">
|
||
<taxonomicName id="4C1D4D43FFE3FFF4E2EA57D6FE0CFC45" ID-CoL="NC7J" box="[151,403,979,1004]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE3FFF4E2EA57D6FE0CFC45" bold="true" box="[151,403,979,1004]" italics="true" pageId="9" pageNumber="56">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57A9FFE3FFF4E3E757D1FE6BFC45" box="[410,500,980,1004]" pageId="9" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE3FFF4E2EA57F2FEE5FBA6" blockId="9.[151,500,979,1039]" box="[151,378,1015,1039]" pageId="9" pageNumber="56">
|
||
(
|
||
<figureCitation id="13262A45FFE3FFF4E2E257F2FE8DFBA6" box="[159,274,1015,1039]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStart-4="FIGURE 12" captionStart-5="FIGURE 13" captionStart-6="FIGURE 14" captionStartId-0="10.[151,250,1900,1923]" captionStartId-1="11.[151,250,1472,1495]" captionStartId-2="12.[151,250,1002,1025]" captionStartId-3="13.[151,250,2013,2036]" captionStartId-4="14.[151,250,768,791]" captionStartId-5="14.[151,250,1174,1197]" captionStartId-6="15.[151,250,1980,2003]" captionTargetBox-0="[242,1286,709,1821]" captionTargetBox-1="[434,1152,211,1451]" captionTargetBox-2="[368,1218,445,981]" captionTargetBox-3="[226,1360,824,1992]" captionTargetBox-4="[188,1403,193,728]" captionTargetBox-5="[208,1383,870,1138]" captionTargetBox-6="[207,1390,231,1959]" captionTargetId-0="figure@10.[151,1436,661,1879]" captionTargetId-1="figure@11.[434,1152,113,1451]" captionTargetId-2="figure@12.[368,1219,445,981]" captionTargetId-3="figure@13.[226,1360,803,1992]" captionTargetId-4="figure@14.[188,1403,193,729]" captionTargetId-5="figure@14.[208,1383,870,1138]" captionTargetId-6="figure@15.[151,1436,193,2007]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="14" captionTargetPageId-5="14" captionTargetPageId-6="15" captionText-0="FIGURE 8. Brochiraja vittacauda sp. nov., adult male holotype (NMNZ P 39563, 719 mm TL, fresh): A, dorsal surface; B, ventral surface." captionText-1="FIGURE 9. Brochiraja vittacauda sp. nov., adult male holotype (NMNZ P 39563, 719 mm TL, preserved) showing snout, orbito-spiracular and nuchal regions." captionText-2="FIGURE 10. Ventral view of the head of Brochiraja vittacauda sp. nov.: adult male holotype (NMNZ P 39563, 719 mm TL, preserved) showing the oronasal region and tooth bands." captionText-3="FIGURE 11. Radiograph of the head of Brochiraja vittacauda sp. nov., adult male holotype (NMNZ P 39563, 719 mm TL)." captionText-4="FIGURE 12. Thorns at tail midlength of Brochiraja vittacauda sp. nov.: A, dorsal and B, lateral views of adult male holotype (NMNZ P 39563, 719 mm TL, preserved)." captionText-5="FIGURE 13. Lateral view of the dorsal and caudal fins of Brochiraja vittacauda sp. nov., adult male holotype (NMNZ P 39563, 719 mm TL, preserved)." captionText-6="FIGURE 14. Dorsal view of bifurcated rostral thorns of A, Brochiraja heuresa sp. nov. (female paratype, CSIRO H 6091 – 01, 364 mm TL, preserved) and B, B. vittacauda sp. nov. (adult male holotype, NMNZ P 39563, 719 mm TL, preserved)." httpUri-0="https://zenodo.org/record/279622/files/figure.png" httpUri-1="https://zenodo.org/record/279623/files/figure.png" httpUri-2="https://zenodo.org/record/279624/files/figure.png" httpUri-3="https://zenodo.org/record/279625/files/figure.png" httpUri-4="https://zenodo.org/record/279626/files/figure.png" httpUri-5="https://zenodo.org/record/279627/files/figure.png" httpUri-6="https://zenodo.org/record/279628/files/figure.png" pageId="9" pageNumber="56">Figs 8–14</figureCitation>
|
||
;
|
||
<tableCitation id="C69F037BFFE3FFF4E36357F2FEF0FBA6" box="[286,367,1015,1039]" captionStart="TABLE 1" captionStartId="6.[151,242,151,175]" captionTargetBox="[151,1440,273,1948]" captionTargetPageId="6" captionText="TABLE 1. Morphometrics for the adult male holotype (CSIRO H 6073 – 01) and ranges for 9 paratypes (see methods) of Brochiraja heuresa sp. nov., and the adult male holotype (NMNZ P 39563) and single paratype (NMNZ P 42691) of B. vittacauda sp. nov. Values are expressed as a percentage of total length (TL)." httpUri="http://table.plazi.org/id/DF626648FFECFFFBE2EA5492FC53FF5C" pageId="9" pageNumber="56" tableUuid="DF626648FFECFFFBE2EA5492FC53FF5C">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE3FFF4E2EA5045FEAAFBDC" blockId="9.[151,1436,1088,1141]" pageId="9" pageNumber="56">
|
||
<taxonomicName id="4C1D4D43FFE3FFF4E2EA5045FF7AFBFF" box="[151,229,1088,1110]" class="Elasmobranchii" family="Rajidae" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="56" phylum="Chordata" rank="family">Rajidae</taxonomicName>
|
||
n. gen. sp. E: Last in
|
||
<bibRefCitation id="EF8C4B31FFE3FFF4E3BC5045FD08FBFF" author="Williams" box="[449,663,1088,1110]" pageId="9" pageNumber="56" refString="Williams A., Gowlett - Holmes, K. & Althaus, F. (eds) (2006) Explore lost world of the deep. NORFANZ Voyage. 10 May to 8 June 2003. Final report to the Department of the Environment and Heritage (National Oceans Office). June 2006. Biodiversity survey of seamounts & slopes of the Norfolk Ridge and Lord Howe Rise. NIWA & CSIRO, 591 pp." type="book" year="2006">
|
||
Williams
|
||
<emphasis id="B969EAD2FFE3FFF4E05E5044FDD0FBFF" box="[547,591,1088,1110]" italics="true" pageId="9" pageNumber="56">et al</emphasis>
|
||
., 2006
|
||
</bibRefCitation>
|
||
, Appendix 6, p. 1 (field identification, listed); Appendix 8: p. 6 (2 photos n° 37031808)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307654BFFE3FFF4E2EA50A7FA8AFA8C" pageId="9" pageNumber="56" type="materials_examined">
|
||
<paragraph id="8BA236C0FFE3FFF4E2EA50A7FC7FFB74" blockId="9.[151,1437,1185,2037]" pageId="9" pageNumber="56">
|
||
<emphasis id="B969EAD2FFE3FFF4E2EA50A7FE94FB13" bold="true" box="[151,267,1186,1210]" pageId="9" pageNumber="56">
|
||
<typeStatus id="54A68862FFE3FFF4E2EA50A7FE99FB13" box="[151,262,1186,1210]" pageId="9" pageNumber="56" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED0CAE05FFE3FFF4E36B50A7FEF7FB10" box="[278,360,1186,1209]" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="9" pageNumber="56">NMNZ</collectionCode>
|
||
P. 39563, adult male
|
||
<quantity id="4CE59B25FFE3FFF4E01350A4FD50FB10" box="[622,719,1185,1209]" metricMagnitude="-1" metricUnit="m" metricValue="7.19" pageId="9" pageNumber="56" unit="mm" value="719.0">719 mm</quantity>
|
||
<collectionCode id="ED0CAE05FFE3FFF4E0A450A7FD61FB10" box="[729,766,1186,1209]" pageId="9" pageNumber="56">TL</collectionCode>
|
||
, West
|
||
<collectingCountry id="F30A7650FFE3FFF4E12D50A4FC36FB10" box="[848,937,1185,1209]" name="Norfolk Island" pageId="9" pageNumber="56">Norfolk</collectingCountry>
|
||
Ridge,
|
||
<geoCoordinate id="EE295007FFE3FFF4E67550A4FBE8FB10" box="[1032,1143,1185,1209]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="92" value="-34.315">34°18.9'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE295007FFE3FFF4E6FA50A4FA95FB10" box="[1159,1290,1185,1209]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="92" value="168.405">168°24.3’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CE59B25FFE3FFF4E76450A4FA03FB10" box="[1305,1436,1185,1209]" metricMagnitude="2" metricUnit="m" metricValue="8.33" metricValueMax="8.57" metricValueMin="8.09" pageId="9" pageNumber="56" unit="m" value="833.0" valueMax="857.0" valueMin="809.0">809–857 m</quantity>
|
||
depth, RV
|
||
<taxonomicName id="4C1D4D43FFE3FFF4E37250C3FEE5FB74" box="[271,378,1222,1245]" class="Arachnida" family="Uloboridae" genus="Tangaroa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAD2FFE3FFF4E37250C3FEE5FB74" box="[271,378,1222,1245]" italics="true" pageId="9" pageNumber="56">Tangaroa</emphasis>
|
||
</taxonomicName>
|
||
NORFANZ TAN 0308/147, bottom trawl,
|
||
<date id="FFA31000FFE3FFF4E12050C0FC43FB74" box="[861,988,1221,1245]" pageId="9" pageNumber="56" value="2003-06-03">3 Jun 2003</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE3FFF4E2BA50EFFA8AFA8C" blockId="9.[151,1437,1185,2037]" pageId="9" pageNumber="56">
|
||
<emphasis id="B969EAD2FFE3FFF4E2BA50EFFEA4FAAB" bold="true" box="[199,315,1258,1282]" pageId="9" pageNumber="56">
|
||
<typeStatus id="54A68862FFE3FFF4E2BA50EFFEA9FAAB" box="[199,310,1258,1282]" pageId="9" pageNumber="56" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Single specimen from the Tasman Sea.
|
||
<collectionCode id="ED0CAE05FFE3FFF4E17F50EFFCCAFAA8" box="[770,853,1258,1281]" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="9" pageNumber="56">NMNZ</collectionCode>
|
||
P. 42691, juvenile male
|
||
<quantity id="4CE59B25FFE3FFF4E61050ECFB54FAA8" box="[1133,1227,1257,1281]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="56" unit="mm" value="360.0">360 mm</quantity>
|
||
<collectionCode id="ED0CAE05FFE3FFF4E6AE50EFFB67FAA8" box="[1235,1272,1258,1281]" pageId="9" pageNumber="56">TL</collectionCode>
|
||
, West
|
||
<collectingCountry id="F30A7650FFE3FFF4E73E50ECFA03FAA8" box="[1347,1436,1257,1281]" name="Norfolk Island" pageId="9" pageNumber="56">Norfolk</collectingCountry>
|
||
Ridge,
|
||
<geoCoordinate id="EE295007FFE3FFF4E2955108FECBFA8C" box="[232,340,1293,1317]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="92" value="-33.68">33º40.8'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE295007FFE3FFF4E31D5108FE75FA8C" box="[352,490,1293,1317]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="92" value="167.84166">167º 50.5' E</geoCoordinate>
|
||
,
|
||
<quantity id="4CE59B25FFE3FFF4E38B5108FDEAFA8C" box="[502,629,1293,1317]" metricMagnitude="2" metricUnit="m" metricValue="8.01" metricValueMax="9.73" metricValueMin="6.29" pageId="9" pageNumber="56" unit="m" value="801.0" valueMax="973.0" valueMin="629.0">629–973 m</quantity>
|
||
depth, F.V.
|
||
<emphasis id="B969EAD2FFE3FFF4E086510BFC4EFA8C" box="[763,977,1293,1317]" italics="true" pageId="9" pageNumber="56">Seamount Explorer</emphasis>
|
||
, bottom trawl,
|
||
<date id="FFA31000FFE3FFF4E6005108FA8EFA8C" box="[1149,1297,1293,1317]" pageId="9" pageNumber="56" value="2006-03-13">13 Mar 2006</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307654BFFE3FFF4E2B85134FF5AF8B4" pageId="9" pageNumber="56" type="diagnosis">
|
||
<paragraph id="8BA236C0FFE3FFF4E2B85134FF5AF8B4" blockId="9.[151,1437,1185,2037]" pageId="9" pageNumber="56">
|
||
<emphasis id="B969EAD2FFE3FFF4E2B85134FEA0FAE3" bold="true" box="[197,319,1329,1354]" pageId="9" pageNumber="56">Diagnosis.</emphasis>
|
||
A large species of
|
||
<taxonomicName id="4C1D4D43FFE3FFF4E0605134FD08FAE0" box="[541,663,1329,1353]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="B969EAD2FFE3FFF4E0605134FD08FAE0" box="[541,663,1329,1353]" italics="true" pageId="9" pageNumber="56">Brochiraja</emphasis>
|
||
</taxonomicName>
|
||
distinguished by the following combination of characters: long tail, 1.2–1.4 times disc length, 2.3–2.8 times ventral head length, 4.7–5.8 times snout length; orbit moderately large,
|
||
<quantity id="4CE59B25FFE3FFF4E2EA517CFE97FA38" box="[151,264,1401,1425]" metricMagnitude="-2" metricUnit="m" metricValue="7.493" metricValueMax="7.874" metricValueMin="7.111999999999999" pageId="9" pageNumber="56" unit="in" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 in</quantity>
|
||
preorbital snout length; tail slender but with relatively broad conical base, evenly tapering to mid-length of tail, width at pelvic-fin insertions 2.0–2.5 times width at midlength; anterior lobe of pelvic fin broadly spatulate; central dorsal disc with fine denticles, otherwise naked apart from enlarged, thorn-like denticles around margin of snout and near alar patch in adult males; tail with mediodorsal row of about 36–51 hooked thorns and small spiny denticles concentrated on its dorsolateral surface; single preorbital thorn, 0–1 midorbital thorns, 1–3 postorbital thorns; snout length 2.6–3.7 times distance from snout tip to bifurcate rostral thorn; calcified rostral shaft short; ventral disc and tail naked; alar thorns in a broad patch, flanked by smaller malar thorns; additional separate patch of malar thorns on snout forward of mid-eye; lateral skin fold very broad posteriorly, originating well forward, beside pelvic-fin inner margins in adult male; dorsal surface dark violet, ventral surface brownish blue, darker than dorsal surface; no beard-like patch beneath mouth; rows of faint pale-edged pores on ventral surface; tail tip white; nuchal pore patch distinct; total pectoral radials 76–77; predorsal vertebrae 112–114; 40–44 tooth rows in upper jaw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307654BFFE3FFF3E2BA532CFB81FAB9" lastPageId="14" lastPageNumber="61" pageId="9" pageNumber="56" type="description">
|
||
<paragraph id="8BA236C0FFE3FFF7E2BA532CFB1DFDF6" blockId="9.[151,1437,1185,2037]" lastBlockId="10.[151,1437,151,607]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
|
||
<emphasis id="B969EAD2FFE3FFF4E2BA532CFEC6F8EB" bold="true" box="[199,345,1833,1858]" pageId="9" pageNumber="56">Description.</emphasis>
|
||
Disc heart shaped to suboval,
|
||
<quantity id="4CE59B25FFE3FFF4E0CF532CFC9EF8E8" box="[690,769,1833,1857]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" pageId="9" pageNumber="56" unit="in" value="1.03">1.03 in</quantity>
|
||
adult male
|
||
<typeStatus id="54A68862FFE3FFF4E1FB532CFC77F8E8" box="[902,1000,1833,1857]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
(
|
||
<quantity id="4CE59B25FFE3FFF4E18A532CFBD8F8E8" box="[1015,1095,1833,1857]" metricMagnitude="-2" metricUnit="m" metricValue="2.7686" pageId="9" pageNumber="56" unit="in" value="1.09">1.09 in</quantity>
|
||
juvenile male
|
||
<typeStatus id="54A68862FFE3FFF4E693532CFACFF8E8" box="[1262,1360,1833,1857]" pageId="9" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
) times as broad as long; maximum angle in front of spiracles 89° (112°); anterior margin weakly double concave, almost straight opposite spiracles in
|
||
<typeStatus id="54A68862FFE3FFF4E3A05377FDDFF823" box="[477,576,1906,1930]" pageId="9" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
, apex and rear tip broadly rounded, posterior margin convex; axis of greatest width about 56% (53%) of disc length. Head moderately elongate, 18.8% (16.7%) TL; snout angular, broader in
|
||
<typeStatus id="54A68862FFE3FFF4E2EA53BFFF65F87B" box="[151,250,1978,2002]" pageId="9" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
, preorbital snout length 3.12 (2.78) times orbit diameter, 3.68 (3.15) times interorbital distance; preoral snout length 1.96 (1.69) times internarial distance. Snout tip lobe like, forming small, subtriangular, fleshy process in
|
||
<typeStatus id="54A68862FFE0FFF7E2C9549DFE87FF19" box="[180,280,152,176]" pageId="10" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
; process distinctly demarcated from anterior profile of disc. Orbit large, elevated (more so in
|
||
<typeStatus id="54A68862FFE0FFF7E7565492FA0FFF06" box="[1323,1424,151,175]" pageId="10" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
), diameter 1.18 (1.13) times interorbital distance. Spiracle of moderate size; main pore covering most of spiracular depression, its length of 2.09 (2.25) in orbit diameter; its anteriormost extension slightly forward of hind margin of eye. Nostril suboval (distorted in both
|
||
<typeStatus id="54A68862FFE0FFF7E0155501FD38FEB5" box="[616,679,260,284]" pageId="10" pageNumber="57">types</typeStatus>
|
||
), small, with about 20 lamellae in
|
||
<typeStatus id="54A68862FFE0FFF7E62D5501FB29FEB5" box="[1104,1206,260,284]" pageId="10" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
; anterior nasal flap expanded laterally, lobe like, partly concealed by nasal curtain anteriorly, weakly tubular along its lateral margin, posterior margin well concealed beneath nasal curtain. Nasal curtain well developed, broader than long; its lateral margin concave anteriorly (wrapping around anterior lobe), expanded medially, then broadly rounded posterolaterally; labial margin poorly fringed; internarial distance 1.60 (1.78) in distance between first gill slits, 0.94 (1.03) in distance between fifth gill slits. Upper and lower jaws strongly arched at symphysis, much less so in
|
||
<typeStatus id="54A68862FFE0FFF7E74955BDFA07FE79" box="[1332,1432,440,464]" pageId="10" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
. Teeth small with long, pointed cusps; cusps longest near middle of jaws; cusps at symphysis of upper jaw slightly smaller than those adjacent; cusps short, knob like near jaw angle; in upper jaw, cusps directly posterolaterally, in lower jaw directed anterolaterally. In
|
||
<typeStatus id="54A68862FFE0FFF7E0355620FD33FD94" box="[584,684,549,573]" pageId="10" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
. quincuncial with elevated, crowns; crowns plate-like, suboval to quadrangular, with short blunt cusps; cusps posteriorly directed, best developed lingually.
|
||
</paragraph>
|
||
<caption id="DF626648FFE0FFF7E2EA5369FEABF808" httpUri="https://zenodo.org/record/279622/files/figure.png" pageId="10" pageNumber="57" targetBox="[242,1286,709,1821]" targetPageId="10">
|
||
<paragraph id="8BA236C0FFE0FFF7E2EA5369FEABF808" blockId="10.[151,1435,1900,1953]" pageId="10" pageNumber="57">
|
||
<emphasis id="B969EAD2FFE0FFF7E2EA5369FE8BF828" bold="true" box="[151,276,1900,1923]" pageId="10" pageNumber="57">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C1D4D43FFE0FFF7E3665369FE63F82B" box="[283,508,1900,1922]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="57" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE0FFF7E3665369FE63F82B" box="[283,508,1900,1922]" italics="true" pageId="10" pageNumber="57">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAD2FFE0FFF7E0795368FDCAF82B" bold="true" box="[516,597,1901,1922]" pageId="10" pageNumber="57">
|
||
<taxonomicNameLabel id="A25A57A9FFE0FFF7E0795368FDCAF82B" box="[516,597,1901,1922]" pageId="10" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult male holotype (NMNZ P 39563, 719 mm TL, fresh): A, dorsal surface; B, ventral surface.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF626648FFE1FFF6E2EA51C5FE75FA5C" httpUri="https://zenodo.org/record/279623/files/figure.png" pageId="11" pageNumber="58" targetBox="[434,1152,211,1451]" targetPageId="11">
|
||
<paragraph id="8BA236C0FFE1FFF6E2EA51C5FE75FA5C" blockId="11.[151,1436,1472,1525]" pageId="11" pageNumber="58">
|
||
<emphasis id="B969EAD2FFE1FFF6E2EA51C5FE8DFA7F" bold="true" box="[151,274,1472,1495]" pageId="11" pageNumber="58">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C1D4D43FFE1FFF6E36551C4FE68FA7E" box="[280,503,1473,1495]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="58" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE1FFF6E36551C4FE68FA7E" box="[280,503,1473,1495]" italics="true" pageId="11" pageNumber="58">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAD2FFE1FFF6E38151C4FDD3FA7F" bold="true" box="[508,588,1473,1494]" pageId="11" pageNumber="58">
|
||
<taxonomicNameLabel id="A25A57A9FFE1FFF6E38151C4FDD3FA7F" box="[508,588,1473,1494]" pageId="11" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult male holotype (NMNZ P 39563, 719 mm TL, preserved) showing snout, orbito-spiracular and nuchal regions.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236C0FFE1FFF1E2B85227FB13FE21" blockId="11.[151,1436,1570,2027]" lastBlockId="12.[151,1436,151,392]" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
|
||
Pelvic fin deeply incised; anterior lobe fleshy, relatively elongate, spatulate, broad along its entire length with a broadly rounded apex, inner margin not incised; posterior lobe relatively short, with weakly incised and weakly convex lateral margin, free rear tip forming a narrower angle than fin apex, inner margin almost straight, slightly convex in
|
||
<typeStatus id="54A68862FFE1FFF6E371528AFEEFF90E" box="[268,368,1679,1703]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
; length of anterior lobe 0.78 (0.87) times length of posterior lobe. Tail slender at base, tapering evenly and very gradually posteriorly, pointed at its tip in
|
||
<typeStatus id="54A68862FFE1FFF6E15A52B6FC14F962" box="[807,907,1715,1739]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
; width to height ratios variable, width at insertions of pelvic fins 2.01 (2.46) times width at midlength of tail and 1.97 (3.21) times width at first dorsal fin origin respectively; length from rear of cloaca 1.37 (1.62) times distance from tip of snout to rear of cloaca; in cross section,
|
||
<typeStatus id="54A68862FFE1FFF6E2AE531BFEAAF89F" box="[211,309,1822,1846]" pageId="11" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
relatively more depressed than
|
||
<typeStatus id="54A68862FFE1FFF6E0D8531AFC97F89E" box="[677,776,1823,1847]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
, dorsal surface convex, ventral surface convex anteriorly, becoming almost flat posteriorly; weak median groove along tail on ventral surface (less obvious in
|
||
<typeStatus id="54A68862FFE1FFF6E7575347FA0FF8F3" box="[1322,1424,1858,1882]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
); width 1.34 (1.59) times height at insertion of pelvic fin, 2.02 (1.46) times height at first dorsal fin origin. Lateral skin fold very well developed; originating beside inner margin of pelvic fin; terminating beyond tip of tail in
|
||
<typeStatus id="54A68862FFE1FFF6E71C538FFF55F86E" pageId="11" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
, but abruptly near mid-base of epichordal lobe in
|
||
<typeStatus id="54A68862FFE1FFF6E08553AAFCC3F86E" box="[760,860,1967,1991]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
; very broad distally, particularly in
|
||
<typeStatus id="54A68862FFE1FFF6E69753ABFAD0F86F" box="[1258,1359,1966,1990]" pageId="11" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
, maximum width on each side almost equivalent to height of first dorsal fin, width on each side of
|
||
<typeStatus id="54A68862FFE1FFF6E6D953D6FA9BF842" box="[1188,1284,2003,2027]" pageId="11" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
equivalent to height of epichordal lobe of caudal fin. Dorsal fins small, short-based, low, located near tail tip; first dorsal fin slightly smaller than second, strongly raked, its apex almost over its insertion, first dorsal-fin height 2.11 (2.30) times in its base length; second dorsal fin similar in shape, also strongly raked, posterior margin subequal in length to inner margin; fin bases clearly separated, rear tip of first dorsal fin overlapping second dorsal-fin origin, interdorsal distance 2.26 (1.73) times in length of first dorsal-fin base. Epichordal caudal-fin lobe small in
|
||
<typeStatus id="54A68862FFE6FFF1E7785522FAF5FE96" box="[1285,1386,295,319]" pageId="12" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
; relatively well developed in
|
||
<typeStatus id="54A68862FFE6FFF1E3C35548FDBEFECC" box="[446,545,333,357]" pageId="12" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
, more than half height of first dorsal fin; distance from second dorsal-fin base subequal to interdorsal distance; hypochordal caudal lobe absent, rudimentary in
|
||
<typeStatus id="54A68862FFE6FFF1E6595575FB17FE21" box="[1060,1160,368,392]" pageId="12" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF626648FFE6FFF1E2EA57EFFD4BFBB6" httpUri="https://zenodo.org/record/279624/files/figure.png" pageId="12" pageNumber="59" targetBox="[368,1218,445,981]" targetPageId="12">
|
||
<paragraph id="8BA236C0FFE6FFF1E2EA57EFFD4BFBB6" blockId="12.[151,1436,1002,1055]" pageId="12" pageNumber="59">
|
||
<emphasis id="B969EAD2FFE6FFF1E2EA57EFFEBFFBA9" bold="true" box="[151,288,1002,1025]" pageId="12" pageNumber="59">FIGURE 10.</emphasis>
|
||
Ventral view of the head of
|
||
<taxonomicName id="4C1D4D43FFE6FFF1E03857EEFCB9FBA8" box="[581,806,1003,1025]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="59" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE6FFF1E03857EEFCB9FBA8" box="[581,806,1003,1025]" italics="true" pageId="12" pageNumber="59">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAD2FFE6FFF1E15057EEFCE1FBA9" bold="true" box="[813,894,1003,1024]" pageId="12" pageNumber="59">
|
||
<taxonomicNameLabel id="A25A57A9FFE6FFF1E15057EEFCE1FBA9" box="[813,894,1003,1024]" pageId="12" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: adult male holotype (NMNZ P 39563, 719 mm TL, preserved) showing the oronasal region and tooth bands.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236C0FFE6FFF1E2BA5042FE28FAD6" blockId="12.[151,1436,1095,2019]" pageId="12" pageNumber="59">
|
||
Dorsal surface of disc and pelvic fins mostly naked, except for minor granulations and orbital, rostral, alar, malar and tail thorns; widely spaced, smaller denticles on mid disc extending into tail, best developed dorsolaterally and continuing to tail tip in about 3 rows below dorsal fins; ventral surface entirely naked. In
|
||
<typeStatus id="54A68862FFE6FFF1E6945095FAD3FB01" box="[1257,1356,1168,1192]" pageId="12" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
, dorsal surface naked, except for a few thorns and a band of minute, widely spaced denticles extending along mid-disc from nuchal region to tail; band broadest over scapular region; band divided above cloaca into two narrower bands, each extending along lateral and dorsolateral surfaces of tail to second dorsal fin; dorsal tail beside median thorns naked; sparse covering of denticles on dorsal fins and epichordal lobe of caudal fin; ventral surfaces of disc and tail naked. Denticles unicuspid, with partly embedded, mostly quadrate bases; cusps extremely short, slender, pungent, mostly strongly recurved.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE6FFF1E2BA518EFEA4F8DE" blockId="12.[151,1436,1095,2019]" pageId="12" pageNumber="59">
|
||
Thorns confined to mid-snout, orbits, midline of tail, and as well-developed alar and malar patches in adult male
|
||
<typeStatus id="54A68862FFE6FFF1E2A951AAFEA5FA6E" box="[212,314,1455,1479]" pageId="12" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
. Bifurcated rostral thorn enlarged in juvenile
|
||
<typeStatus id="54A68862FFE6FFF1E14B51B5FC09FA61" box="[822,918,1456,1480]" pageId="12" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
(anterior length ca
|
||
<quantity id="4CE59B25FFE6FFF1E60D51AAFB55FA6E" box="[1136,1226,1455,1479]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="59" unit="mm" value="3.3">3.3 mm</quantity>
|
||
), broad based with prominent, strongly oblique, bifurcated crown; less obvious in
|
||
<typeStatus id="54A68862FFE6FFF1E12151D6FC5EFA42" box="[860,961,1491,1515]" pageId="12" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
; forward on snout, its distance from snout tip about 2.57 (3.68) in preorbital snout length.
|
||
<typeStatus id="54A68862FFE6FFF1E0CD51FDFC8DF9B9" box="[688,786,1528,1552]" pageId="12" pageNumber="59" type="paratype">Paratype</typeStatus>
|
||
with two large and prominent, broad-based orbital thorns with recurved tips; one preorbital and one postorbital thorn aligned with anterior and posterior orbital margins respectively;
|
||
<typeStatus id="54A68862FFE6FFF1E353523AFE0FF9FE" box="[302,400,1599,1623]" pageId="12" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
with additional 1–2 postorbital thorns and one smaller median thorn, all positioned around orbital margin; 36 thorns on dorsal midline of tail in
|
||
<typeStatus id="54A68862FFE6FFF1E0955261FCD4F9D5" box="[744,843,1636,1660]" pageId="12" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
, enlarged (up to
|
||
<quantity id="4CE59B25FFE6FFF1E6755266FBC0F9D2" box="[1032,1119,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="59" unit="mm" value="4.3">4.3 mm</quantity>
|
||
anterior length), in a single, closely spaced, regular row; bases of thorns compressed and subtriangular, distal half slender, recurved; integument of ventral surface relatively thick, fleshy; median series persistent in adult, 51 thorns in an almost regular row, size variable, notably smaller near dorsal fins; no interdorsal thorns. Alar thorn patch of mature male well developed; non-retractable, posteromedially directed thorns in 2–4 longitudinal rows on mid-outer pectoral fin; thorns flanked laterally by a broad band of slightly smaller, more posteriorly directed malar thorns that extend forward in a narrow band along disc margin to spiracle; additional pair of broad patches of malar thorns on snout forward of orbits.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE6FFF0E2B85386FD87FEB5" blockId="12.[151,1436,1095,2019]" lastBlockId="13.[151,1437,151,788]" lastPageId="13" lastPageNumber="60" pageId="12" pageNumber="59">
|
||
Rostral cartilage (from radiograph) narrow basally, calcified component elongate, extending anteriorly beyond rostral thorn; rostral appendices well developed, visible on radiograph; anterior fontanelle broadly flask shaped, extending onto rostral cartilage well anterior to leading edge of nasal capsules; propterygium of pectoral girdle not reaching rostral node; lateral prepelvic processes of pelvic girdle moderately elongate. Claspers slender, elongate, postcloacal length 20.9% TL when mature; glans little expanded, bluntly pointed distally, distal aperture moderately large; configuration similar to other members of the genus as in
|
||
<bibRefCitation id="EF8C4B31FFE7FFF0E1E354E5FB58FF5E" author="Last" box="[926,1223,223,247]" pageId="13" pageNumber="60" refString="Last P. R. & McEachran, J. D. (2006) New softnose skate genus Brochiraja from New Zealand (Rajidae: Arhynchobatinae) with description of four new species. New Zealand Journal of Marine and Fresh water Research, 40, 65 - 90." type="journal article" year="2006">Last & McEachran (2006)</bibRefCitation>
|
||
; tip of ventral marginal cartilage recurved, spatulate.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE7FFF0E2BA5522FB26FE2E" blockId="13.[151,1437,151,788]" pageId="13" pageNumber="60">Tooth rows in upper jaw 44 (40, n=1); lower jaw 40 (38). Pectoral-fin propterygial radials 35 (35, n=1); mesopterygial radials 14 (13–14); metapterygial radials 28 (28); total radials 77 (76). Monospondylous centra 28 (28, n=1); predorsal caudal centra 84 (86); predorsal centra 112 (114); total centra about 142 (139).</paragraph>
|
||
<paragraph id="8BA236C0FFE7FFF0E2BA5596FA9CFCBD" blockId="13.[151,1437,151,788]" pageId="13" pageNumber="60">
|
||
<emphasis id="B969EAD2FFE7FFF0E2BA5596FED0FE05" bold="true" box="[199,335,403,428]" pageId="13" pageNumber="60">Coloration.</emphasis>
|
||
In
|
||
<typeStatus id="54A68862FFE7FFF0E30E5591FE4AFE05" box="[371,469,404,428]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
(when fresh), dorsal surface mostly dark violet, and outer disc, pelvic fins and tail reddish brown, orbital membranes similar to adjacent disc; ventral surface slightly darker brownish blue, sensory pores paler, rows evident; jaws, most of second dorsal fin, caudal fin, half of dorsal clasper, distal ventral surface of anterior pelvic-fin lobe, ventral tip of tail, margin of first dorsal fin, and clasper white; outer half of both anterior pelvic lobes covered with unusually thick, white mucous; preserved coloration similar, bluish dorsally, darker brownish ventrally, whitish areas remaining obvious; enlarged denticles and thorns white, strongly contrasted with rest of disc. In juvenile
|
||
<typeStatus id="54A68862FFE7FFF0E3D85668FD9AFD2C" box="[421,517,621,645]" pageId="13" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
(when fresh), dorsal surface of disc and tail almost uniformly dark blue; iris and thorns pale, strongly contrasted with base colour; first dorsal fin white basally, dark distally; second dorsal fin and epichordal lob of caudal almost uniformly white; ventral surface of disc bluish brown (slightly darker than dorsal surface); ventral pores not obvious, although some pale pores on anterior head and on anterior disc; anterior nasal flap, mouth, gill-slit anterior membranes, clasper tips, and origin and subdorsal length of tail whitish.
|
||
</paragraph>
|
||
<caption id="DF626648FFE7FFF0E2EA53D8FA18F85A" box="[151,1415,2013,2036]" httpUri="https://zenodo.org/record/279625/files/figure.png" pageId="13" pageNumber="60" targetBox="[226,1360,824,1992]" targetPageId="13">
|
||
<paragraph id="8BA236C0FFE7FFF0E2EA53D8FA18F85A" blockId="13.[151,1415,2013,2036]" box="[151,1415,2013,2036]" pageId="13" pageNumber="60">
|
||
<emphasis id="B969EAD2FFE7FFF0E2EA53D8FE81F85A" bold="true" box="[151,286,2013,2036]" pageId="13" pageNumber="60">FIGURE 11.</emphasis>
|
||
Radiograph of the head of
|
||
<taxonomicName id="4C1D4D43FFE7FFF0E04E53DBFC8DF85D" box="[563,786,2014,2036]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="60" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE7FFF0E04E53DBFC8DF85D" box="[563,786,2014,2036]" italics="true" pageId="13" pageNumber="60">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57A9FFE7FFF0E16553DBFCF6F85A" box="[792,873,2014,2035]" pageId="13" pageNumber="60" rank="species">
|
||
<emphasis id="B969EAD2FFE7FFF0E16553DBFCF6F85A" bold="true" box="[792,873,2014,2035]" pageId="13" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male holotype (NMNZ P 39563, 719 mm TL).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF626648FFE4FFF3E2EA5705FDD7FC9C" httpUri="https://zenodo.org/record/279626/files/figure.png" pageId="14" pageNumber="61" targetBox="[188,1403,193,728]" targetPageId="14">
|
||
<paragraph id="8BA236C0FFE4FFF3E2EA5705FDD7FC9C" blockId="14.[151,1436,768,821]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2EA5705FEBFFCBC" bold="true" box="[151,288,768,791]" pageId="14" pageNumber="61">FIGURE 12.</emphasis>
|
||
Thorns at tail midlength of
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E0405705FC82FCBF" box="[573,797,768,790]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE4FFF3E0405705FC82FCBF" box="[573,797,768,790]" italics="true" pageId="14" pageNumber="61">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAD2FFE4FFF3E1595704FCEBFCBF" bold="true" box="[804,884,769,790]" pageId="14" pageNumber="61">
|
||
<taxonomicNameLabel id="A25A57A9FFE4FFF3E1595704FCEBFCBF" box="[804,884,769,790]" pageId="14" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A, dorsal and B, lateral views of adult male holotype (NMNZ P 39563, 719 mm TL, preserved).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF626648FFE4FFF3E2EA5093FE44FB62" httpUri="https://zenodo.org/record/279627/files/figure.png" pageId="14" pageNumber="61" targetBox="[208,1383,870,1138]" targetPageId="14">
|
||
<paragraph id="8BA236C0FFE4FFF3E2EA5093FE44FB62" blockId="14.[151,1436,1174,1227]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2EA5093FEBDFB05" bold="true" box="[151,290,1174,1197]" pageId="14" pageNumber="61">FIGURE 13.</emphasis>
|
||
Lateral view of the dorsal and caudal fins of
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E1765092FC72FB04" box="[779,1005,1175,1197]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="vittacauda" status="sp. nov.">
|
||
<emphasis id="B969EAD2FFE4FFF3E1765092FC72FB04" box="[779,1005,1175,1197]" italics="true" pageId="14" pageNumber="61">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B969EAD2FFE4FFF3E18B5092FBD6FB05" bold="true" box="[1014,1097,1175,1196]" pageId="14" pageNumber="61">
|
||
<taxonomicNameLabel id="A25A57A9FFE4FFF3E18B5092FBD6FB05" box="[1014,1097,1175,1196]" pageId="14" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult male holotype (NMNZ P 39563, 719 mm TL, preserved).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236C0FFE4FFF3E2BA50F2FB81FAB9" blockId="14.[151,1437,1271,1872]" box="[199,1054,1271,1296]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2BA50F2FF61FAB9" bold="true" box="[199,254,1271,1296]" pageId="14" pageNumber="61">Size.</emphasis>
|
||
Males reach at least
|
||
<quantity id="4CE59B25FFE4FFF3E39450FDFDD9FAB9" box="[489,582,1272,1296]" metricMagnitude="-1" metricUnit="m" metricValue="7.19" pageId="14" pageNumber="61" unit="mm" value="719.0">719 mm</quantity>
|
||
TL; no information available for females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307654BFFE4FFF3E2BA511EFE56FAF1" pageId="14" pageNumber="61" type="distribution">
|
||
<paragraph id="8BA236C0FFE4FFF3E2BA511EFE56FAF1" blockId="14.[151,1437,1271,1872]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2BA511EFEFFFA9D" bold="true" box="[199,352,1307,1332]" pageId="14" pageNumber="61">Distribution.</emphasis>
|
||
Known from small section of the West
|
||
<collectingCountry id="F30A7650FFE4FFF3E1655119FCEEFA9D" box="[792,881,1308,1332]" name="Norfolk Island" pageId="14" pageNumber="61">Norfolk</collectingCountry>
|
||
Ridge (ca 33º40–
|
||
<geoCoordinate id="EE295007FFE4FFF3E6405119FB08FA9D" box="[1085,1175,1308,1332]" direction="south" orientation="latitude" pageId="14" pageNumber="61" precision="925" value="-34.3">34°18' S</geoCoordinate>
|
||
, 167º 50–
|
||
<geoCoordinate id="EE295007FFE4FFF3E7775119FAAAFA9D" box="[1290,1333,1308,1332]" direction="east" orientation="longitude" pageId="14" pageNumber="61" precision="925" value="168.4">168</geoCoordinate>
|
||
°24’ E) at
|
||
<quantity id="4CE59B25FFE4FFF3E2EA5145FE89FAF1" box="[151,278,1344,1368]" metricMagnitude="2" metricUnit="m" metricValue="8.01" metricValueMax="9.73" metricValueMin="6.29" pageId="14" pageNumber="61" unit="m" value="801.0" valueMax="973.0" valueMin="629.0">629–973 m</quantity>
|
||
depth (
|
||
<figureCitation id="13262A45FFE4FFF3E3175145FE24FAF1" box="[362,443,1344,1368]" captionStart="FIGURE 11" captionStartId="13.[151,250,2013,2036]" captionTargetBox="[226,1360,824,1992]" captionTargetId="figure@13.[226,1360,803,1992]" captionTargetPageId="13" captionText="FIGURE 11. Radiograph of the head of Brochiraja vittacauda sp. nov., adult male holotype (NMNZ P 39563, 719 mm TL)." httpUri="https://zenodo.org/record/279625/files/figure.png" pageId="14" pageNumber="61">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307654BFFE4FFF3E2BA5161FB37F8F9" pageId="14" pageNumber="61" type="etymology">
|
||
<paragraph id="8BA236C0FFE4FFF3E2BA5161FAE7FA09" blockId="14.[151,1437,1271,1872]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2BA5161FED1FAD5" bold="true" box="[199,334,1380,1404]" pageId="14" pageNumber="61">Etymology.</emphasis>
|
||
A combination of the Latin
|
||
<emphasis id="B969EAD2FFE4FFF3E0F25160FD5EFAD5" box="[655,705,1381,1404]" italics="true" pageId="14" pageNumber="61">vitta</emphasis>
|
||
(ribbon) and
|
||
<emphasis id="B969EAD2FFE4FFF3E1215161FC3EFAD5" box="[860,929,1380,1404]" italics="true" pageId="14" pageNumber="61">cauda</emphasis>
|
||
(tail) with reference to the appearance of the tail due to its unusually broad lateral skin folds that terminating near its tip. Vernacular name: Ribbontail Skate.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE4FFF3E2BA51AEFCFDF969" blockId="14.[151,1437,1271,1872]" pageId="14" pageNumber="61">
|
||
<emphasis id="B969EAD2FFE4FFF3E2BA51AEFEF3FA6D" bold="true" box="[199,364,1451,1476]" pageId="14" pageNumber="61">Comparisons.</emphasis>
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E30951A9FDF3FA6D" box="[372,620,1452,1476]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="vittacauda">
|
||
<emphasis id="B969EAD2FFE4FFF3E30951A9FDF3FA6D" box="[372,620,1452,1476]" italics="true" pageId="14" pageNumber="61">Brochiraja vittacauda</emphasis>
|
||
</taxonomicName>
|
||
is either sympatric or parapatric with
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E66051A8FB01FA6D" box="[1053,1182,1453,1476]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="aenigma">
|
||
<emphasis id="B969EAD2FFE4FFF3E66051A8FB01FA6D" box="[1053,1182,1453,1476]" italics="true" pageId="14" pageNumber="61">B. aenigma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E6AB51A8FAD2FA6D" box="[1238,1357,1452,1476]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="heuresa">
|
||
<emphasis id="B969EAD2FFE4FFF3E6AB51A8FB78FA6D" box="[1238,1255,1453,1476]" italics="true" pageId="14" pageNumber="61">B</emphasis>
|
||
.
|
||
<emphasis id="B969EAD2FFE4FFF3E68B51A9FAD2FA6D" box="[1270,1357,1452,1476]" italics="true" pageId="14" pageNumber="61">heuresa</emphasis>
|
||
</taxonomicName>
|
||
on the
|
||
<collectingCountry id="F30A7650FFE4FFF3E2EA51D5FF6FFA41" box="[151,240,1488,1512]" name="Norfolk Island" pageId="14" pageNumber="61">Norfolk</collectingCountry>
|
||
Ridge. It is a much larger skate than
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E0F351D4FC9AFA41" box="[654,773,1488,1512]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="heuresa">
|
||
<emphasis id="B969EAD2FFE4FFF3E0F351D4FD00FA41" box="[654,671,1489,1512]" italics="true" pageId="14" pageNumber="61">B</emphasis>
|
||
.
|
||
<emphasis id="B969EAD2FFE4FFF3E0D051D5FC9AFA41" box="[685,773,1488,1512]" italics="true" pageId="14" pageNumber="61">heuresa</emphasis>
|
||
</taxonomicName>
|
||
(adult males to about
|
||
<quantity id="4CE59B25FFE4FFF3E18051D5FBDEFA41" box="[1021,1089,1488,1512]" metricMagnitude="-1" metricUnit="m" metricValue="7.2" pageId="14" pageNumber="61" unit="cm" value="72.0">72 cm</quantity>
|
||
vs.
|
||
<quantity id="4CE59B25FFE4FFF3E61251D5FB41FA41" box="[1135,1246,1488,1512]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="14" pageNumber="61" unit="cm" value="37.0" valueMax="38.0" valueMin="36.0">36–38 cm</quantity>
|
||
TL) and is probably larger than
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E32D51F0FE4EF9A5" box="[336,465,1525,1548]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="aenigma">
|
||
<emphasis id="B969EAD2FFE4FFF3E32D51F0FE4EF9A5" box="[336,465,1525,1548]" italics="true" pageId="14" pageNumber="61">B. aenigma</emphasis>
|
||
</taxonomicName>
|
||
. The dorsal surface is less spinulose than in either of these species (denticles concentrated near mid-disc rather than being distributed all over), and well developed thorns are present on in pre and postorbit (rather than being reduced or absent). The pectoral-fin radial and vertebral counts are high as in
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E7F95238FF67F9D1" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="aenigma">
|
||
<emphasis id="B969EAD2FFE4FFF3E7F95238FF67F9D1" italics="true" pageId="14" pageNumber="61">B. aenigma</emphasis>
|
||
</taxonomicName>
|
||
(total pectoral radials 76–77 vs. 73–74 and predorsal centra 112–114 vs.
|
||
<quantity id="4CE59B25FFE4FFF3E6505265FBECF9D1" box="[1069,1139,1632,1656]" metricMagnitude="0" metricUnit="m" metricValue="2.921" pageId="14" pageNumber="61" unit="in" value="115.0">115 in</quantity>
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E6075264FB66F9D1" box="[1146,1273,1633,1656]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="aenigma">
|
||
<emphasis id="B969EAD2FFE4FFF3E6075264FB66F9D1" box="[1146,1273,1633,1656]" italics="true" pageId="14" pageNumber="61">B. aenigma</emphasis>
|
||
</taxonomicName>
|
||
; total pectoral radials 58–63 and predorsal centra
|
||
<quantity id="4CE59B25FFE4FFF3E0515281FD00F935" box="[556,671,1668,1692]" metricMagnitude="0" metricUnit="m" metricValue="2.3876" metricValueMax="2.54" metricValueMin="2.2352" pageId="14" pageNumber="61" unit="in" value="94.0" valueMax="100.0" valueMin="88.0">88–100 in</quantity>
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E0D45280FCBDF935" box="[681,802,1668,1692]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="heuresa">
|
||
<emphasis id="B969EAD2FFE4FFF3E0D45280FD25F935" box="[681,698,1669,1692]" italics="true" pageId="14" pageNumber="61">B</emphasis>
|
||
.
|
||
<emphasis id="B969EAD2FFE4FFF3E0B75281FCBDF935" box="[714,802,1668,1692]" italics="true" pageId="14" pageNumber="61">heuresa</emphasis>
|
||
</taxonomicName>
|
||
), but the disc is more angular and the tooth row count higher (40–44 vs. about 30 rows in upper jaw) than
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E0A152ACFCC4F969" box="[732,859,1705,1728]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="aenigma">
|
||
<emphasis id="B969EAD2FFE4FFF3E0A152ACFCC4F969" box="[732,859,1705,1728]" italics="true" pageId="14" pageNumber="61">B. aenigma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236C0FFE4FFF3E2B852C9FB37F8F9" blockId="14.[151,1437,1271,1872]" pageId="14" pageNumber="61">
|
||
Of the
|
||
<collectingCountry id="F30A7650FFE4FFF3E36852C8FE33F94D" box="[277,428,1740,1764]" name="New Zealand" pageId="14" pageNumber="61">New Zealand</collectingCountry>
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E3C952C9FDB1F94D" box="[436,558,1740,1764]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="genus">
|
||
<emphasis id="B969EAD2FFE4FFF3E3C952C9FDB1F94D" box="[436,558,1740,1764]" italics="true" pageId="14" pageNumber="61">Brochiraja</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E04052C8FD4BF94D" box="[573,724,1740,1764]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="vittacauda">
|
||
<emphasis id="B969EAD2FFE4FFF3E04052C8FD4BF94D" box="[573,724,1740,1764]" italics="true" pageId="14" pageNumber="61">B. vittacauda</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C1D4D43FFE4FFF3E1D752C8FBA3F94D" box="[938,1084,1740,1764]" class="Elasmobranchii" family="Arhynchobatidae" genus="Brochiraja" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="61" phylum="Chordata" rank="species" species="leviveneta">
|
||
<emphasis id="B969EAD2FFE4FFF3E1D752C8FBA3F94D" box="[938,1084,1740,1764]" italics="true" pageId="14" pageNumber="61">B. leviveneta</emphasis>
|
||
</taxonomicName>
|
||
but has a broader, more spatulate, anterior pelvic-fin lobe, much broader lateral skin fold, higher pectoral-fin radial and vertebral counts (total pectoral radials 76–77 vs. 64–69 and predorsal centra 112–114 vs. 95–100), better developed alar and malar thorns in adult males, and reaches a larger adult size (adult males to about
|
||
<quantity id="4CE59B25FFE4FFF3E1F1533DFC4EF8F9" box="[908,977,1848,1872]" metricMagnitude="-1" metricUnit="m" metricValue="7.2" pageId="14" pageNumber="61" unit="cm" value="72.0">72 cm</quantity>
|
||
vs.
|
||
<quantity id="4CE59B25FFE4FFF3E182533DFBF1F8F9" box="[1023,1134,1848,1872]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="14" pageNumber="61" unit="cm" value="37.0" valueMax="38.0" valueMin="36.0">36–38 cm</quantity>
|
||
TL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |