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<document id="D22A87B5AA9D854912607E36890BBB77" ID-CLB-Dataset="256481" ID-DOI="10.11646/zootaxa.5240.1.1" ID-GBIF-Dataset="4d650293-3f9a-4bbe-9530-204a3b09ef8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7639547" ID-ZooBank="2FE467C1-EAC2-4E90-B673-CCE2CCA93C1D" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1676380923589" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="03B487C0FFC1251CFF47C1B0F839FCEB" docLanguage="en" docName="zootaxa.5240.1.1.pdf" docOrigin="Zootaxa 5240 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5240.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) bella Fernandes &amp; Silva 2015" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FF8DFFB8FFCA2511FFD0C712FB1CFFB6" masterDocTitle="Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="12" updateTime="1699475138964" updateUser="ExternalLinkService">
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<mods:title id="0C949835EDA145B310FB0A6875C63AA5">Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae)</mods:title>
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<mods:namePart id="3065FD59D9948E418C10AF835E0E6B65">Mendonça, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="27C70784164D379A6CBD6FDDBA64B386">Silva, Valéria Juliete Da</mods:namePart>
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<mods:namePart id="AF97C39B5A10DC824733A894B439DDD4">Fernandes, José Antônio Marin</mods:namePart>
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<treatment id="03B487C0FFC1251CFF47C1B0F839FCEB" ID-DOI="http://doi.org/10.5281/zenodo.7639868" ID-GBIF-Taxon="209578830" ID-Zenodo-Dep="7639868" LSID="urn:lsid:plazi:treatment:03B487C0FFC1251CFF47C1B0F839FCEB" httpUri="http://treatment.plazi.org/id/03B487C0FFC1251CFF47C1B0F839FCEB" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
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<paragraph id="8BA236D6FFC1251AFF47C1B0F982F90A" blockId="11.[151,670,1698,1760]" box="[151,670,1698,1725]" pageId="11" pageNumber="12">
<heading id="D0EA81BAFFC1251AFF47C1B0F982F90A" bold="true" box="[151,670,1698,1725]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<taxonomicName id="4C1D4D55FFC1251AFF47C1B0F982F90A" authority="Fernandes &amp; Silva, 2015" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[151,670,1698,1725]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bella" subGenus="Edessa">
<emphasis id="B969EAC4FFC1251AFF47C1B0F982F90A" bold="true" box="[151,670,1698,1725]" pageId="11" pageNumber="12">
<emphasis id="B969EAC4FFC1251AFF47C1B0FA78F90B" bold="true" box="[151,356,1698,1725]" italics="true" pageId="11" pageNumber="12">Edessa (E.) bella</emphasis>
<bibRefCitation id="EF8C4B27FFC1251AFEBBC1B0F982F90A" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[363,670,1698,1724]" pageId="11" pageNumber="12" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">Fernandes &amp; Silva, 2015</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C307655DFFC1251AFF47C1D4FA85F956" box="[151,409,1734,1760]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BA236D6FFC1251AFF47C1D4FA85F956" blockId="11.[151,670,1698,1760]" box="[151,409,1734,1760]" pageId="11" pageNumber="12">
(
<figureCitation id="13262A53FFC1251AFF4FC1D4FBF3F956" box="[159,239,1734,1760]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="11" pageNumber="12">Figs. 2</figureCitation>
,
<figureCitation id="13262A53FFC1251AFF2BC1D4FA52F956" box="[251,334,1734,1760]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="11" pageNumber="12">24 C,D</figureCitation>
,
<figureCitation id="13262A53FFC1251AFE8AC1D4FA8DF956" box="[346,401,1734,1760]" captionStart="FIGURE 31" captionStartId="59.[152,255,1448,1472]" captionTargetBox="[164,1423,181,1414]" captionTargetId="figure-19@59.[164,1423,181,1425]" captionTargetPageId="59" captionText="FIGURE 31. A. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639677" httpUri="https://zenodo.org/record/7639677/files/figure.png" pageId="11" pageNumber="12">31 A</figureCitation>
)
</paragraph>
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<subSubSection id="C307655DFFC1251AFF47C017F925F8AB" box="[151,569,1797,1821]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BA236D6FFC1251AFF47C017F925F8AB" blockId="11.[151,569,1797,1821]" box="[151,569,1797,1821]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB0D11F8FFC1251AFF47C017F925F8AB" box="[151,569,1797,1821]" pageId="11" pageNumber="12">
<treatmentCitation id="0ABC10C7FFC1251AFF47C017F929F8AB" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[151,565,1797,1821]" page="513" pageId="11" pageNumber="12" year="2015">
<taxonomicName id="4C1D4D55FFC1251AFF47C017F929F8AB" authority="Fernandes et al., 2015: 513" authorityName="Fernandes" authorityPageNumber="513" authorityYear="2015" box="[151,565,1797,1821]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B969EAC4FFC1251AFF47C017FA04F8AB" box="[151,280,1797,1821]" italics="true" pageId="11" pageNumber="12">Edessa bella</emphasis>
<bibRefCitation id="EF8C4B27FFC1251AFECEC017F929F8AB" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[286,565,1797,1821]" pageId="11" pageNumber="12" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="B969EAC4FFC1251AFE5DC014FAA5F8AB" box="[397,441,1797,1821]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 2015: 513
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C307655DFFC1251CFF47C054F961FEAB" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BA236D6FFC1251AFF47C054F9CFF8D7" blockId="11.[151,1436,1862,1996]" box="[151,723,1862,1889]" pageId="11" pageNumber="12">
<typeStatus id="54A68874FFC1251AFF47C054FBE3F8D6" box="[151,255,1862,1888]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54A68874FFC1251AFEE6C055FABAF8D7" box="[310,422,1863,1889]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
: see
<bibRefCitation id="EF8C4B27FFC1251AFE0BC054F9CCF8D7" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[475,720,1862,1889]" pageId="11" pageNumber="12" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="B969EAC4FFC1251AFD85C055F991F8D6" box="[597,653,1862,1888]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2015
</bibRefCitation>
.
</paragraph>
<paragraph id="8BA236D6FFC1251AFF47C078FAEBF87A" blockId="11.[151,1436,1862,1996]" pageId="11" pageNumber="12">
The original description and comments were presented in
<bibRefCitation id="EF8C4B27FFC1251AFCEAC078FF56F833" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[826,1098,1898,1925]" pageId="11" pageNumber="12" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="B969EAC4FFC1251AFC67C079F8EEF832" box="[951,1010,1898,1924]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2015)
</bibRefCitation>
. Here are photographs of the
<typeStatus id="54A68874FFC1251AFF47C09CFBE5F81E" box="[151,249,1934,1960]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
and a plate with photographs of the male and female external genitalia, as the original description only includes hand-drawn drawings.
</paragraph>
<caption id="DF62665EFFC6251DFF48C085FF09F865" ID-DOI="http://doi.org/10.5281/zenodo.7639555" ID-Zenodo-Dep="7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="12" pageNumber="13" startId="12.[152,255,1943,1967]" targetBox="[186,1401,193,1905]" targetPageId="12">
<paragraph id="8BA236D6FFC6251DFF48C085FF09F865" blockId="12.[151,1437,1943,2004]" pageId="12" pageNumber="13">
<emphasis id="B969EAC4FFC6251DFF48C085FA06F819" bold="true" box="[152,282,1943,1967]" pageId="12" pageNumber="13">FIGURE 2.</emphasis>
A–G.
<taxonomicName id="4C1D4D55FFC6251DFE8FC08AF807F819" authority="Fernandes &amp; Silva, 2015" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[351,795,1943,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bella" subGenus="Edessa">
<emphasis id="B969EAC4FFC6251DFE8FC08AFABBF806" box="[351,423,1944,1968]" italics="true" pageId="12" pageNumber="13">Edessa</emphasis>
(
<emphasis id="B969EAC4FFC6251DFE67C08AFAD1F819" box="[439,461,1944,1967]" italics="true" pageId="12" pageNumber="13">E.</emphasis>
)
<emphasis id="B969EAC4FFC6251DFE0DC08AF90CF806" box="[477,528,1944,1968]" italics="true" pageId="12" pageNumber="13">bella</emphasis>
<bibRefCitation id="EF8C4B27FFC6251DFDC7C08AF807F819" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[535,795,1943,1968]" pageId="12" pageNumber="13" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">Fernandes &amp; Silva, 2015</bibRefCitation>
</taxonomicName>
: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BA236D6FFC7251CFF47C784FAC7FF4E" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="14">
<emphasis id="B969EAC4FFC7251CFF47C784FA91FF06" bold="true" box="[151,397,150,176]" pageId="13" pageNumber="14">Additional material.</emphasis>
<materialsCitation id="3B753C8BFFC7251CFE46C784FBC6FF62" ID-GBIF-Occurrence="4097197418" collectingDate="1998-08-31" collectionCode="JEE" collectorName="C. W. &amp; L. B. O'Brien" country="Costa Rica" location="Biol. Sta. La Selva" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Heredia">
<emphasis id="B969EAC4FFC7251CFE46C784F9A3FF06" bold="true" box="[406,703,150,176]" pageId="13" pageNumber="14">
<collectingCountry id="F30A7646FFC7251CFE46C784F959FF06" box="[406,581,150,176]" name="Costa Rica" pageId="13" pageNumber="14">COSTA RICA</collectingCountry>
,
<collectingRegion id="49D9F834FFC7251CFD86C784F9A6FF06" box="[598,698,150,176]" country="Costa Rica" name="Heredia" pageId="13" pageNumber="14">Heredia</collectingRegion>
:
</emphasis>
2♁,
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,
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<collectingDate id="EFE7E9FEFFC7251CFC38C784FF9EFF06" box="[1000,1154,150,176]" pageId="13" pageNumber="14" value="1998-08-31">31-VIII-1998</collectingDate>
</date>
,
<collectorName id="26E85300FFC7251CFB40C784FE80FF06" box="[1168,1436,150,177]" pageId="13" pageNumber="14">C. W. &amp; L. B. O’Brien</collectorName>
(
<collectionCode id="ED0CAE13FFC7251CFF4FC7A8FBCEFF62" box="[159,210,186,212]" pageId="13" pageNumber="14">JEE</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B753C8BFFC7251CFF0AC7A8F8C6FF62" ID-GBIF-Occurrence="4097197364" box="[218,986,186,213]" collectingDate="2009-10-09" collectionCode="NMPC" country="Costa Rica" location="Puerto Viejo La Selva Biol. Station" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Heredia">
;
<specimenCount id="9D1BFD5FFFC7251CFF35C7A8FA14FF62" box="[229,264,186,212]" pageId="13" pageNumber="14" type="female">1♀</specimenCount>
,
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of
<location id="8EC2600DFFC7251CFEBCC7A9F9EAFF62" LSID="urn:lsid:plazi:treatment:03B487C0FFC1251CFF47C1B0F839FCEB:8EC2600DFFC7251CFEBCC7A9F9EAFF62" box="[364,758,186,213]" country="Costa Rica" name="Puerto Viejo La Selva Biol. Station" pageId="13" pageNumber="14" stateProvince="Heredia">Puerto Viejo La Selva Biol. Station</location>
,
<date id="FFA31016FFC7251CFCD1C7A8F871FF62" box="[769,877,186,212]" pageId="13" pageNumber="14" value="2009-10-09">
<collectingDate id="EFE7E9FEFFC7251CFCD1C7A8F871FF62" box="[769,877,186,212]" pageId="13" pageNumber="14" value="2009-10-09">9-X-2009</collectingDate>
</date>
(
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)
</materialsCitation>
<materialsCitation id="3B753C8BFFC7251CFC0AC7A8FACBFF4E" ID-GBIF-Occurrence="4097197437" collectingDate="2004-11-02" collectionCode="NMPC" country="Costa Rica" location="Puerto Viejo La Selva Biol. Station" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Heredia">
;
<specimenCount id="9D1BFD5FFFC7251CFC34C7A8FF1BFF62" box="[996,1031,186,212]" pageId="13" pageNumber="14" type="female">1♀</specimenCount>
,
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of
<location id="8EC2600DFFC7251CFBBAC7A9FBF5FF4E" LSID="urn:lsid:plazi:treatment:03B487C0FFC1251CFF47C1B0F839FCEB:8EC2600DFFC7251CFBBAC7A9FBF5FF4E" country="Costa Rica" name="Puerto Viejo La Selva Biol. Station" pageId="13" pageNumber="14" stateProvince="Heredia">Puerto Viejo La Selva Biol. Station</location>
,
<date id="FFA31016FFC7251CFF25C7CCFA77FF4E" box="[245,363,222,248]" pageId="13" pageNumber="14" value="2004-11-02">
<collectingDate id="EFE7E9FEFFC7251CFF25C7CCFA77FF4E" box="[245,363,222,248]" pageId="13" pageNumber="14" value="2004-11-02">2-XI-2004</collectingDate>
</date>
(
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)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA236D6FFC7251CFF17C610F961FEAB" blockId="13.[151,1437,150,861]" box="[199,637,258,285]" pageId="13" pageNumber="14">
<emphasis id="B969EAC4FFC7251CFF17C610FA27FEAA" bold="true" box="[199,315,258,284]" pageId="13" pageNumber="14">Diagnose.</emphasis>
See
<bibRefCitation id="EF8C4B27FFC7251CFEA2C610F964FEAB" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[370,632,258,285]" pageId="13" pageNumber="14" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="B969EAC4FFC7251CFE3CC611F938FEAA" box="[492,548,258,284]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFC7251CFF17C634FAAAFD32" pageId="13" pageNumber="14" type="description">
<paragraph id="8BA236D6FFC7251CFF17C634FAAAFD32" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="14">
<emphasis id="B969EAC4FFC7251CFF17C634F903FEF6" bold="true" box="[199,543,294,320]" pageId="13" pageNumber="14">Complementary description:</emphasis>
Humeral angles as long as wide (
<figureCitation id="13262A53FFC7251CFC46C634FF1EFEF6" box="[918,1026,294,320]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 C</figureCitation>
); humeral angle black spot restricted to the distal part of the angle in dorsal and ventral views (
<figureCitation id="13262A53FFC7251CFC90C658F8D2FED2" box="[832,974,330,356]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 C,D</figureCitation>
). Connexival segments with concavities partially covered by black spots; rounded spots in ventral view (
<figureCitation id="13262A53FFC7251CFCA6C67CFF1EFE3E" box="[886,1026,366,392]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 C,D</figureCitation>
). Ventral surface. Dark stripe of the propleuron covering 2/3 of the width of the sclerite (
<figureCitation id="13262A53FFC7251CFD33C680F84EFE1A" box="[739,850,402,428]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 D</figureCitation>
). Legs with tibiae and tarsi darker than the rest (
<figureCitation id="13262A53FFC7251CFABFC680FBCFFE66" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 D</figureCitation>
). Abdomen with spine of third segment rounded (
<figureCitation id="13262A53FFC7251CFCD7C6A4F87AFE66" box="[775,870,438,464]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="13" pageNumber="14">Fig. 2 G</figureCitation>
). Intersegmental areas black, not reaching ventral spots of connexivum (
<figureCitation id="13262A53FFC7251CFE5DC6C8FAE7FE42" box="[397,507,474,500]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 D</figureCitation>
). Pseudosutures concolorous with surface (
<figureCitation id="13262A53FFC7251CFC0AC6C8FF5BFE42" box="[986,1095,474,500]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 D</figureCitation>
). Male genitalia, dorsal side of the pygophore with a suffused brown area occupying 2/3 of the surface (
<figureCitation id="13262A53FFC7251CFC18C6ECFF46FDAE" box="[968,1114,510,536]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="13" pageNumber="14">Fig. 2 A,B,E</figureCitation>
). Superior process of genital cup continuing ventrally in a high crenulated carina, ending in a small dentiform projection (
<figureCitation id="13262A53FFC7251CFB61C530FE36FD8A" box="[1201,1322,546,572]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="13" pageNumber="14">Fig. 2 B,E</figureCitation>
). Anterior half of proctiger brown (
<figureCitation id="13262A53FFC7251CFE60C554F97CFDD6" box="[432,608,582,608]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="13" pageNumber="14">Fig. 2 A,B,D,E</figureCitation>
). Female genitalia, laterotergites VIII with two small dark spots on outer lateral margins (
<figureCitation id="13262A53FFC7251CFE9DC578FAB6FD32" box="[333,426,618,644]" captionStart="FIGURE 2" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[186,1401,193,1905]" captionTargetId="figure-31@12.[186,1401,179,1917]" captionTargetPageId="12" captionText="FIGURE 2. A–G. Edessa (E.) bella Fernandes &amp; Silva, 2015: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7639555" httpUri="https://zenodo.org/record/7639555/files/figure.png" pageId="13" pageNumber="14">Fig. 2 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFC7251CFF17C59CFE75FC8E" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BA236D6FFC7251CFF17C59CFE75FC8E" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="14">
<emphasis id="B969EAC4FFC7251CFF17C59CFA53FD1E" bold="true" box="[199,335,654,680]" pageId="13" pageNumber="14">Comments.</emphasis>
This species is unique in the subgenus due to color pattern of the humeral angle which is orange and black (
<figureCitation id="13262A53FFC7251CFEC3C5A0FA80FD7A" box="[275,412,690,716]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 C,D</figureCitation>
). The other species of the subgenus that resembles
<taxonomicName id="4C1D4D55FFC7251CFC0DC5A1FF7BFD7A" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[989,1127,690,717]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bella" subGenus="Edessa">
<emphasis id="B969EAC4FFC7251CFC0DC5A1FF7BFD7A" box="[989,1127,690,717]" italics="true" pageId="13" pageNumber="14">E. (E.) bella</emphasis>
</taxonomicName>
is
<taxonomicName id="4C1D4D55FFC7251CFB58C5A1FE07FD7A" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[1160,1307,690,717]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lewisi" subGenus="Edessa">
<emphasis id="B969EAC4FFC7251CFB58C5A1FE07FD7A" box="[1160,1307,690,717]" italics="true" pageId="13" pageNumber="14">E. (E.) lewisi</emphasis>
</taxonomicName>
. Both have a similar distribution (
<figureCitation id="13262A53FFC7251CFE40C5C4F914FD46" box="[400,520,726,752]" captionStart="FIGURE 31" captionStartId="59.[152,255,1448,1472]" captionTargetBox="[164,1423,181,1414]" captionTargetId="figure-19@59.[164,1423,181,1425]" captionTargetPageId="59" captionText="FIGURE 31. A. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639677" httpUri="https://zenodo.org/record/7639677/files/figure.png" pageId="13" pageNumber="14">Figs. 31 A</figureCitation>
,
<figureCitation id="13262A53FFC7251CFDC4C5C4F978FD46" box="[532,612,726,752]" captionStart="FIGURE 32" captionStartId="60.[152,255,1984,2008]" captionTargetBox="[189,1399,181,1954]" captionTargetId="figure-31@60.[189,1399,181,1960]" captionTargetPageId="60" captionText="FIGURE 32. A–B. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639683" httpUri="https://zenodo.org/record/7639683/files/figure.png" pageId="13" pageNumber="14">32 A,B</figureCitation>
) and size of the body (
<figureCitation id="13262A53FFC7251CFCB6C5C4F8E5FD46" box="[870,1017,726,752]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Figs. 24 C,D</figureCitation>
,
<figureCitation id="13262A53FFC7251CFBD4C5C4FF44FD46" box="[1028,1112,726,752]" captionStart="FIGURE 27" captionStartId="55.[152,255,1943,1967]" captionTargetBox="[269,1321,188,1918]" captionTargetId="figure-19@55.[250,1337,181,1919]" captionTargetPageId="55" captionText="FIGURE 27. A–F. Dorsal and ventral surface: A–B, Edessa (E.) leucocera Dallas, 1851; C–D, Edessa (E.) lewisi Fernandes &amp; Silva, 2015; E–F, Edessa (E.) miniata Westwood, 1837. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639663" httpUri="https://zenodo.org/record/7639663/files/figure.png" pageId="13" pageNumber="14">27 C,D</figureCitation>
), but the former has the connexival segments with dark spots and corial veins concolorous (
<figureCitation id="13262A53FFC7251CFC8AC5E8F8DFFCA2" box="[858,963,762,788]" captionStart="FIGURE 24" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[269,1331,190,1904]" captionTargetId="figure-31@52.[251,1336,181,1919]" captionTargetPageId="52" captionText="FIGURE 24. A–F. Dorsal and ventral surface: A–B, Edessa (E.) alces Erichson, 1848; C–D, Edessa (E.) bella Fernandes &amp; Silva, 2015; E–F, Edessa (E.) congrua Walker, 1868. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639650" httpUri="https://zenodo.org/record/7639650/files/figure.png" pageId="13" pageNumber="14">Fig. 24 C</figureCitation>
) and the later has the connexival segments without spots and corial veins creamy (
<figureCitation id="13262A53FFC7251CFD9EC40CF9A7FC8E" box="[590,699,798,824]" captionStart="FIGURE 27" captionStartId="55.[152,255,1943,1967]" captionTargetBox="[269,1321,188,1918]" captionTargetId="figure-19@55.[250,1337,181,1919]" captionTargetPageId="55" captionText="FIGURE 27. A–F. Dorsal and ventral surface: A–B, Edessa (E.) leucocera Dallas, 1851; C–D, Edessa (E.) lewisi Fernandes &amp; Silva, 2015; E–F, Edessa (E.) miniata Westwood, 1837. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639663" httpUri="https://zenodo.org/record/7639663/files/figure.png" pageId="13" pageNumber="14">Fig. 27 C</figureCitation>
), clearly contrasting with the color of the corium (
<figureCitation id="13262A53FFC7251CFB3FC40CFE40FC8E" box="[1263,1372,798,824]" captionStart="FIGURE 27" captionStartId="55.[152,255,1943,1967]" captionTargetBox="[269,1321,188,1918]" captionTargetId="figure-19@55.[250,1337,181,1919]" captionTargetPageId="55" captionText="FIGURE 27. A–F. Dorsal and ventral surface: A–B, Edessa (E.) leucocera Dallas, 1851; C–D, Edessa (E.) lewisi Fernandes &amp; Silva, 2015; E–F, Edessa (E.) miniata Westwood, 1837. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7639663" httpUri="https://zenodo.org/record/7639663/files/figure.png" pageId="13" pageNumber="14">Fig. 27 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C307655DFFC7251CFF17C450F839FCEB" box="[199,805,834,861]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BA236D6FFC7251CFF17C450F839FCEB" blockId="13.[151,1437,150,861]" box="[199,805,834,861]" pageId="13" pageNumber="14">
<emphasis id="B969EAC4FFC7251CFF17C450FA45FCEA" bold="true" box="[199,345,834,860]" pageId="13" pageNumber="14">Distribution</emphasis>
(
<figureCitation id="13262A53FFC7251CFEB9C450FAC9FCEA" box="[361,469,834,860]" captionStart="FIGURE 31" captionStartId="59.[152,255,1448,1472]" captionTargetBox="[164,1423,181,1414]" captionTargetId="figure-19@59.[164,1423,181,1425]" captionTargetPageId="59" captionText="FIGURE 31. A. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.7639677" httpUri="https://zenodo.org/record/7639677/files/figure.png" pageId="13" pageNumber="14">Fig. 31 A</figureCitation>
). See
<bibRefCitation id="EF8C4B27FFC7251CFDCAC450F83CFCEB" author="Fernandes, J. A. M. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[538,800,834,861]" pageId="13" pageNumber="14" pagination="511 - 536" refId="ref37486" refString="Fernandes, J. A. M., Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="B969EAC4FFC7251CFD44C451F9D0FCEA" box="[660,716,834,860]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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