276 lines
31 KiB
XML
276 lines
31 KiB
XML
<document id="2128FF8C653E5668EB9E517E585780D0" ID-DOI="10.5281/zenodo.188147" ID-GBIF-Dataset="14731d1d-b0be-4109-b6bc-e678b89dc740" ID-ISSN="1175-5326" ID-Zenodo-Dep="188147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461084195760" checkinUser="plazi" docAuthor="Meegaskumbura, Madhava, Manamendra-Arachchi, Kelum & Pethiyagoda, Rohan" docDate="2009" docId="03B40459643EFFF426A2FF3CFABEFB18" docLanguage="en" docName="zt02122p068.pdf" docOrigin="Zootaxa 2122" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Philautus singu Meegaskumbura, Manamendra-Arachchi & Pethiyagoda, 2009, new species" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FF8D7C216436FFFF2635FFABFF9EFFF5" masterDocTitle="Two new species of shrub frogs (Rhacophoridae: Philautus) from the lowlands of Sri Lanka" masterLastPageNumber="68" masterPageNumber="51" pageNumber="59" updateTime="1698587828333" updateUser="plazi">
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<mods:titleInfo id="9035EEF327E68135B6677306EF4DBE1E">
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<mods:title id="AF2D18268C2EA43093DFD3966341D464">Two new species of shrub frogs (Rhacophoridae: Philautus) from the lowlands of Sri Lanka</mods:title>
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</mods:titleInfo>
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<mods:name id="2A79EB5225A2862564B9B470343555CB" type="personal">
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<mods:roleTerm id="85A9DC30CC43EB2B3DA652B1AEC5DE42">Author</mods:roleTerm>
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<mods:namePart id="11418187D6E3E688F0F87C733EF6F370">Meegaskumbura, Madhava</mods:namePart>
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<mods:roleTerm id="220517AC271DA9D9EDB8964B7C285A67">Author</mods:roleTerm>
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<mods:namePart id="1E70D5DF30CEDBD954275DDCEEA8C434">Manamendra-Arachchi, Kelum</mods:namePart>
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<mods:name id="AF9E3C9FF83E0DC8EBB1B4E4836108F8" type="personal">
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<mods:roleTerm id="DBB14688D3967B25BB2C1965212AE073">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="014B024B9F780404458B0C2C36EBF816">Pethiyagoda, Rohan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A222C742C0B986FAB9F77E096CACAC84">text</mods:typeOfResource>
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<mods:title id="4E5A0DF597FCA153572D759190CEE9B5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="700CCC2F1B980E4783745F21DF57E2FC">
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<mods:date id="32BEA82C99D6BF9654EB63AB216BF0D8">2009</mods:date>
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<mods:detail id="F0EDF8680AD749497BF2353D65678532" type="volume">
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<mods:number id="977F1A3999871A6C45088251B1FAEBFE">2122</mods:number>
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<mods:classification id="FC631BC58CCB3D40E217FE0B29C88A38">journal article</mods:classification>
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<mods:identifier id="F1DC859A0D0D39C67724FB8E63F22AD2" type="DOI">10.5281/zenodo.188147</mods:identifier>
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<mods:identifier id="D71BF24FBAEE9BEBB85EA15E8E45DFAD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B40459643EFFF426A2FF3CFABEFB18" ID-DOI="http://doi.org/10.5281/zenodo.5624837" ID-GBIF-Taxon="119618558" ID-Zenodo-Dep="5624837" LSID="urn:lsid:plazi:treatment:03B40459643EFFF426A2FF3CFABEFB18" httpUri="http://treatment.plazi.org/id/03B40459643EFFF426A2FF3CFABEFB18" lastPageId="11" lastPageNumber="62" pageId="8" pageNumber="59">
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<subSubSection id="C307E6C4643EFFF726A2FF3CFE89FF21" pageId="8" pageNumber="59" type="nomenclature">
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<paragraph id="8BA2B54F643EFFF726A2FF3CFE6BFF44" blockId="8.[151,501,151,212]" box="[151,501,151,177]" pageId="8" pageNumber="59">
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<heading id="D0EA0223643EFFF726A2FF3CFE6BFF44" bold="true" box="[151,501,151,177]" fontSize="11" level="1" pageId="8" pageNumber="59" reason="1">
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<emphasis id="B969695D643EFFF726A2FF3CFE6BFF44" bold="true" box="[151,501,151,177]" pageId="8" pageNumber="59">
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<taxonomicName id="4C1DCECC643EFFF726A2FF3CFEC8FF44" ID-CoL="4G5L8" box="[151,342,151,177]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="singu" status="sp. nov.">
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<emphasis id="B969695D643EFFF726A2FF3CFEC8FF44" bold="true" box="[151,342,151,177]" italics="true" pageId="8" pageNumber="59">Philautus singu</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25AD426643EFFF72750FF3CFE6BFF44" box="[357,501,151,177]" pageId="8" pageNumber="59" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA2B54F643EFFF726A2FF11FE89FF21" blockId="8.[151,501,151,212]" box="[151,279,186,212]" pageId="8" pageNumber="59">
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(
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<figureCitation id="1326A9CA643EFFF726AAFF11FE91FF21" box="[159,271,186,212]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="8.[151,255,1552,1576]" captionStartId-1="9.[151,255,1954,1978]" captionStartId-2="10.[151,257,1742,1766]" captionStartId-3="12.[151,256,1599,1623]" captionTargetBox-0="[151,1434,679,1526]" captionTargetBox-1="[176,1416,470,1882]" captionTargetBox-2="[158,1420,208,1695]" captionTargetBox-3="[379,1273,583,1577]" captionTargetId-0="figure@8.[151,1436,669,1526]" captionTargetId-1="figure@9.[151,1436,435,1931]" captionTargetId-2="figure@10.[151,1436,194,1718]" captionTargetId-3="figure@12.[303,1275,583,1578]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="12" captionText-0="FIGURE 4. Philautus singu (WHT 6340), in life, Kottawa Forest Reserve, Galle." captionText-1="FIGURE 5. Philautus singu: a, lateral; b, dorsal; and c, ventral aspects, respectively, of head of holotype, male, WHT 6034, 16.2 mm SVL. Scale bar: 1 mm." captionText-2="FIGURE 6. Philautus singu: a, ventral aspect of left manus; b, ventral aspect of left pes; and c, semi-diagrammatic representation of left-pes webbing pattern of the holotype, male, WHT 6034, 16.2 mm SVL. Scale bar: 1 mm." captionText-3="FIGURE 7. South-western Sri Lanka, showing the distribution of Philautus singu (squares) and P. t a n u (circles), and place-names cited in the text." httpUri-0="https://zenodo.org/record/188151/files/figure.png" httpUri-1="https://zenodo.org/record/188152/files/figure.png" httpUri-2="https://zenodo.org/record/188153/files/figure.png" httpUri-3="https://zenodo.org/record/188154/files/figure.png" pageId="8" pageNumber="59">Figs. 4–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C307E6C4643EFFF726A2FEACFD19FE16" pageId="8" pageNumber="59" type="materials_examined">
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<paragraph id="8BA2B54F643EFFF726A2FEACFD81FEBD" blockId="8.[151,1437,263,638]" pageId="8" pageNumber="59">
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<emphasis id="B969695D643EFFF726A2FEACFE07FED4" bold="true" box="[151,409,263,289]" pageId="8" pageNumber="59">Material examined.</emphasis>
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<typeStatus id="54A60BED643EFFF72797FEA3FD84FED7" box="[418,538,264,290]" pageId="8" pageNumber="59" type="holotype">Holotype</typeStatus>
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: mature male,
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<quantity id="4CE518AA643EFFF724E7FEA3FCDEFED7" box="[722,832,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="8" pageNumber="59" unit="mm" value="16.2">16.2 mm</quantity>
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SVL, WHT6034, Kitulgala, alt.
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<quantity id="4CE518AA643EFFF722E9FEA3FAB4FED7" box="[1244,1322,264,290]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="59" unit="m" value="101.0">101 m</quantity>
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(
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<geoCoordinate id="EE29D388643EFFF72308FEA3FA09FED7" box="[1341,1431,264,290]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="925" value="6.9833336">6°59'N</geoCoordinate>
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,
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<geoCoordinate id="EE29D388643EFFF726A2FE85FF66FEBD" box="[151,248,302,328]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="925" value="80.333336">80°20'E</geoCoordinate>
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), coll.
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<date id="FFA3938F643EFFF72772FE85FD85FEBD" box="[327,539,302,328]" pageId="8" pageNumber="59" value="1999-02-22">22 February 1999</date>
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.
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</paragraph>
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<paragraph id="8BA2B54F643EFFF726F0FEFEFD19FE16" blockId="8.[151,1437,263,638]" pageId="8" pageNumber="59">
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<typeStatus id="54A60BED643EFFF726F0FEFEFEA5FE9A" box="[197,315,341,367]" pageId="8" pageNumber="59" type="paratype">Paratypes</typeStatus>
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: mature males,
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<quantity id="4CE518AA643EFFF727C7FEFEFDC4FE9A" box="[498,602,341,367]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="8" pageNumber="59" unit="mm" value="16.6">16.6 mm</quantity>
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SVL, WHT6340, Kottawa Forest Reserve (Galle), alt.
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<quantity id="4CE518AA643EFFF722D7FEFEFA80FE9A" box="[1250,1310,341,367]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="59" unit="m" value="60.0">60 m</quantity>
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(
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<geoCoordinate id="EE29D388643EFFF7231BFEFEFA09FE9A" box="[1326,1431,341,367]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="925" value="6.1">06°06’N</geoCoordinate>
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,
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||
<geoCoordinate id="EE29D388643EFFF726A2FED7FF62FE63" box="[151,252,380,406]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="925" value="80.333336">80°20’E</geoCoordinate>
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), coll.
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<date id="FFA3938F643EFFF7277EFED7FE44FE63" box="[331,474,380,406]" pageId="8" pageNumber="59" value="1999-06-03">3 June 1999</date>
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;
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<quantity id="4CE518AA643EFFF727D0FED7FDD3FE63" box="[485,589,380,406]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="8" pageNumber="59" unit="mm" value="16.1">16.1 mm</quantity>
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SVL, WHT5977, Sinharaja World Heritage Site (near Kudawa), alt.
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<quantity id="4CE518AA643EFFF72345FED7FF30FE49" metricMagnitude="2" metricUnit="m" metricValue="5.13" pageId="8" pageNumber="59" unit="m" value="513.0">513 m</quantity>
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(
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<geoCoordinate id="EE29D388643EFFF7268AFE09FEB9FE49" box="[191,295,418,444]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="925" value="6.4166665">06°25’N</geoCoordinate>
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, 80°25E), coll.
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<date id="FFA3938F643EFFF727D7FE09FD36FE49" box="[482,680,418,444]" pageId="8" pageNumber="59" value="1999-01-25">25 January 1999</date>
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;
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<quantity id="4CE518AA643EFFF72480FE09FC81FE49" box="[693,799,418,444]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="8" pageNumber="59" unit="mm" value="16.1">16.1 mm</quantity>
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SVL, WHT6341, Kottawa Forest reserve (Galle), alt.
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<quantity id="4CE518AA643EFFF726A2FE62FF4CFE16" box="[151,210,457,483]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="59" unit="m" value="60.0">60 m</quantity>
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(
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<geoCoordinate id="EE29D388643EFFF726D6FE62FEAEFE16" box="[227,304,457,483]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="925" value="6.0833335">6° 5'N</geoCoordinate>
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,
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<geoCoordinate id="EE29D388643EFFF72708FE62FE3FFE16" box="[317,417,457,483]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="925" value="80.3">80°18’E</geoCoordinate>
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), coll.
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<date id="FFA3938F643EFFF727C7FE62FD1DFE16" box="[498,643,457,483]" pageId="8" pageNumber="59" value="1999-06-03">3 June 1999</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C307E6C4643EFFF726F0FE44FD97FD8B" pageId="8" pageNumber="59" type="diagnosis">
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<paragraph id="8BA2B54F643EFFF726F0FE44FD97FD8B" blockId="8.[151,1437,263,638]" pageId="8" pageNumber="59">
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<emphasis id="B969695D643EFFF726F0FE44FED7FDFC" bold="true" box="[197,329,495,521]" pageId="8" pageNumber="59">Diagnosis</emphasis>
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.
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<taxonomicName id="4C1DCECC643EFFF7276EFE5BFDB8FDFC" box="[347,550,496,521]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="singu">
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<emphasis id="B969695D643EFFF7276EFE5BFDB8FDFC" box="[347,550,496,521]" italics="true" pageId="8" pageNumber="59">Philautus singu</emphasis>
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</taxonomicName>
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is distinguished from all its Sri Lankan congeners by the following combination of characters: size small, mature males
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<quantity id="4CE518AA643EFFF7252FFDBDFC58FDC5" box="[794,966,534,560]" metricMagnitude="-2" metricUnit="m" metricValue="1.635" metricValueMax="1.66" metricValueMin="1.61" pageId="8" pageNumber="59" unit="mm" value="16.35" valueMax="16.6" valueMin="16.1">16.1-16.6 mm</quantity>
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SVL; a prominent tubercle present on upper eyelid; tympanum distinct; supratympanic fold distinct; canthal edges rounded; vomerine ridge absent; throat, chest and belly granular.
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</paragraph>
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</subSubSection>
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<caption id="DF62E5C7643EFFF726A2F9BBFB9EF9D2" box="[151,1024,1552,1576]" httpUri="https://zenodo.org/record/188151/files/figure.png" pageId="8" pageNumber="59" targetBox="[151,1434,679,1526]" targetPageId="8">
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<paragraph id="8BA2B54F643EFFF726A2F9BBFB9EF9D2" blockId="8.[151,1024,1552,1576]" box="[151,1024,1552,1576]" pageId="8" pageNumber="59">
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<emphasis id="B969695D643EFFF726A2F9BBFE87F9DD" bold="true" box="[151,281,1552,1576]" pageId="8" pageNumber="59">FIGURE 4.</emphasis>
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<taxonomicName id="4C1DCECC643EFFF72715F9BBFE58F9D2" box="[288,454,1552,1575]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="singu">
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<emphasis id="B969695D643EFFF72715F9BBFE58F9D2" box="[288,454,1552,1575]" italics="true" pageId="8" pageNumber="59">Philautus singu</emphasis>
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</taxonomicName>
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(WHT 6340), in life, Kottawa Forest Reserve, Galle.
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</paragraph>
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</caption>
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<subSubSection id="C307E6C4643EFFF426F9F9FEFBE7FCE9" lastPageId="11" lastPageNumber="62" pageId="8" pageNumber="59" type="description">
|
||
<paragraph id="8BA2B54F643EFFF626F9F9FEFC61FE6F" blockId="8.[151,1437,1621,2034]" lastBlockId="9.[151,1437,152,410]" lastPageId="9" lastPageNumber="60" pageId="8" pageNumber="59">
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||
<emphasis id="B969695D643EFFF726F9F9FEFEFAF99A" bold="true" box="[204,356,1621,1647]" pageId="8" pageNumber="59">Description.</emphasis>
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Body slender. Head laterally convex above. Snout obtusely pointed in dorsal view, rounded in lateral view. Canthal edges rounded. Loreal region concave. Interorbital space convex. Internasal space concave. Nostrils oval. Pupil oval, horizontal. Tympanum distinct, oval, oblique, its outer rim distinct. Pineal ocellus absent. Vomerine ridge absent. Tongue moderate, emarginate, sometimes (n = 1) bearing a rounded lingual papilla. Supratympanic fold distinct. Cephalic ridges absent. Co-ossified skin on head absent. Upper and lower arms short, thin. Fingers slender. Relative length of fingers, 1 <2 <4 <3. Tips of fingers with discs bearing circum-marginal grooves. Webbing and lateral dermal fringe absent on fingers. Subarticular tubercles on fingers prominent, oval, single, sometimes absent on fingers III (n = 2) and IV (n = 2). Prepollex oval, distinct. Two palmar tubercles, oval, distinct; outer tubercle bifid. Supernumerary tubercles present. Thigh, shank slender. Toes thin, relative length 1 <2 <3 = 5 <4 (
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<typeStatus id="54A60BED643EFFF72573F81AFC2DF83E" box="[838,947,1969,1995]" pageId="8" pageNumber="59" type="holotype">holotype</typeStatus>
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), (1 <2 <5 <3 <4, WHT6340), (1 <2 <3 <5 <4, WHT5977, WHT6341). Tips of toes with discs, with circum-marginal groves. Webbing present on toes. Subarticular tubercles on toes prominent, oval, single, sometimes absent on toe V (n = 2). Inner metatarsal tubercle distinct, oval. Outer metatarsal tubercle absent. Tarsal fold absent; some dermal tubercles on outer edge of foot. Supernumerary tubercles on toes absent. Tarsal tubercle absent. Dorsal and lateral parts of snout, between eyes, side of head, anterior part of back, posterior part of back, both upper and lower flanks with scattered, glandular tubercles. Dorsolateral fold absent. Dorsal and lateral parts of upper arm, lower arm, thigh, shank and foot with scattered glandular tubercles. Throat, chest, belly and ventral side of thigh granular; underside of thigh smooth. Nuptial pad absent. Internal vocal slits present.
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</paragraph>
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<caption id="DF62E5C7643FFFF626A2F809FDAAF829" httpUri="https://zenodo.org/record/188152/files/figure.png" pageId="9" pageNumber="60" targetBox="[176,1416,470,1882]" targetPageId="9">
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<paragraph id="8BA2B54F643FFFF626A2F809FDAAF829" blockId="9.[151,1436,1954,2012]" pageId="9" pageNumber="60">
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<emphasis id="B969695D643FFFF626A2F809FE85F84F" bold="true" box="[151,283,1954,1978]" pageId="9" pageNumber="60">FIGURE 5.</emphasis>
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<taxonomicName id="4C1DCECC643FFFF62717F808FE55F84F" box="[290,459,1955,1978]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="singu">
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<emphasis id="B969695D643FFFF62717F808FE55F84F" box="[290,459,1955,1978]" italics="true" pageId="9" pageNumber="60">Philautus singu</emphasis>
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</taxonomicName>
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:
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<emphasis id="B969695D643FFFF627EFF808FE71F84F" box="[474,495,1955,1978]" italics="true" pageId="9" pageNumber="60">a,</emphasis>
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lateral;
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<emphasis id="B969695D643FFFF6247DF808FDCBF84F" box="[584,597,1955,1978]" italics="true" pageId="9" pageNumber="60">b</emphasis>
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, dorsal; and
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<emphasis id="B969695D643FFFF624D6F808FD71F84F" box="[739,751,1955,1978]" italics="true" pageId="9" pageNumber="60">c</emphasis>
|
||
, ventral aspects, respectively, of head of holotype, male, WHT 6034, 16.2 mm SVL. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF62E5C7643CFFF526A2F965FABBF8FD" httpUri="https://zenodo.org/record/188153/files/figure.png" pageId="10" pageNumber="61" targetBox="[158,1420,208,1695]" targetPageId="10">
|
||
<paragraph id="8BA2B54F643CFFF526A2F965FABBF8FD" blockId="10.[151,1436,1742,1800]" pageId="10" pageNumber="61">
|
||
<emphasis id="B969695D643CFFF526A2F965FE83F913" bold="true" box="[151,285,1742,1766]" pageId="10" pageNumber="61">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C1DCECC643CFFF52710F964FE4CF913" box="[293,466,1743,1766]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="61" phylum="Chordata" rank="species" species="singu">
|
||
<emphasis id="B969695D643CFFF52710F964FE4CF913" box="[293,466,1743,1766]" italics="true" pageId="10" pageNumber="61">Philautus singu</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B969695D643CFFF527D4F964FE70F913" box="[481,494,1743,1766]" italics="true" pageId="10" pageNumber="61">a</emphasis>
|
||
, ventral aspect of left manus;
|
||
<emphasis id="B969695D643CFFF52574F964FCD0F913" box="[833,846,1743,1766]" italics="true" pageId="10" pageNumber="61">b</emphasis>
|
||
, ventral aspect of left pes; and
|
||
<emphasis id="B969695D643CFFF52298F964FB27F913" box="[1197,1209,1743,1766]" italics="true" pageId="10" pageNumber="61">c</emphasis>
|
||
, semi-diagrammatic representation of left-pes webbing pattern of the holotype, male,WHT 6034, 16.2 mm SVL. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA2B54F643CFFF426F0F89CFF69FED3" blockId="10.[151,1437,1847,2028]" lastBlockId="11.[151,1437,152,1261]" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61">
|
||
<emphasis id="B969695D643CFFF526F0F89CFE3FF8A4" bold="true" box="[197,417,1847,1873]" pageId="10" pageNumber="61">Coloration in life.</emphasis>
|
||
Dorsal and lateral parts of head and body brown (
|
||
<figureCitation id="1326A9CA643CFFF525D8F89CFBA8F8A4" box="[1005,1078,1847,1873]" captionStart="FIGURE 4" captionStartId="8.[151,255,1552,1576]" captionTargetBox="[151,1434,679,1526]" captionTargetId="figure@8.[151,1436,669,1526]" captionTargetPageId="8" captionText="FIGURE 4. Philautus singu (WHT 6340), in life, Kottawa Forest Reserve, Galle." httpUri="https://zenodo.org/record/188151/files/figure.png" pageId="10" pageNumber="61">Fig. 4</figureCitation>
|
||
). Interorbital area dark brown. A dark-brown “W”-shaped marking on mid-back. Upper flank brown, lower flank yellow with brown pigments. Inguinal zone pale yellowish brown. Loreal region dark brown. Tympanic region blackish brown. Upper half of tympanum dark brown, lower half pale yellowish light brown. Upper lip brown with paleyellow patches. Dorsal and lateral parts of forelimb pale yellow with brown pigments, lacking distinct crossbars. Thigh and shank pale brown with three dark-brown crossbars. Dorsal side of foot pale brown with darker patches. Posterior thigh pale brown. Throat and margin of throat with dark-brown pigments on a paleyellow background. Chest, belly, thigh and webbing on toes with dark-brown pigments and pale-yellow patches.
|
||
</paragraph>
|
||
<paragraph id="8BA2B54F643DFFF426F0FE99FDB3FD77" blockId="11.[151,1437,152,1261]" pageId="11" pageNumber="62">
|
||
<emphasis id="B969695D643DFFF426F0FE99FE53FEB9" bold="true" box="[197,461,306,332]" pageId="11" pageNumber="62">Coloration in alcohol</emphasis>
|
||
(based on
|
||
<typeStatus id="54A60BED643DFFF4247AFE99FD27FEB9" box="[591,697,306,332]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
, WHT6034). Dorsal and lateral parts of head and body brown. Interorbital area dark brown. A dark-brown “W”-shaped marking on mid-back. Upper flank brown, lower flank yellow with brown pigments. Inguinal zone pale yellowish brown. Loreal region brown. Tympanic region dark brown. Upper half of tympanum dark brown, lower half pale yellowish light brown. Upper lip brown with pale-yellow patches. Dorsal and lateral parts of forelimb pale yellow with brown pigments; no distinct crossbars. Thigh and shank pale brown with three dark-brown crossbars. Dorsal side of foot pale brown with dark-brown patches. Posterior side of thigh pale brown. Throat and margin of throat with darkbrown pigments on a pale-yellow background. Chest, belly, thigh and webbing on toes with dark-brown pigments and pale-yellow patches.
|
||
</paragraph>
|
||
<paragraph id="8BA2B54F643DFFF426F0FD25FBE7FCE9" blockId="11.[151,1437,152,1261]" pageId="11" pageNumber="62">
|
||
Measurements of
|
||
<typeStatus id="54A60BED643DFFF427A3FD25FE63FD5D" box="[406,509,654,680]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
(WHT6034, in mm): DBE, 6.3; DFE, 3.6; DL, 0.7; DW, 1.0; ED, 2.5; EN, 2.0; ES, 2.3; FEL, 7.7; FL I, 1.2; FL II, 1.7; FL III, 3.0; FL IV, 2.2; FOL, 11.1; HL, 6.4; HW, 6.2; IML, 0.9; IN, 1.8; IO, 1.6; LAL, 3.2; MBE, 2.2; MFE, 4.0; MN, 5.7; NS, 1.2; PAL, 5.1; SVL, 16.2; TBL, 8.2; TL I, 1.3; TL II, 1.7; TL III, 2.8; TL IV, 3.9; TL V, 2.8;
|
||
<collectingCountry id="F30AF5DF643DFFF424B5FCA9FD21FCE9" box="[640,703,770,796]" name="Chad" pageId="11" pageNumber="62">TYD</collectingCountry>
|
||
, 0.7; TYE, 0.7; UAW, 3.6; UEW, 2.2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307E6C4643DFFF426F0FC83FD68FC9F" pageId="11" pageNumber="62" type="etymology">
|
||
<paragraph id="8BA2B54F643DFFF426F0FC83FD68FC9F" blockId="11.[151,1437,152,1261]" pageId="11" pageNumber="62">
|
||
<emphasis id="B969695D643DFFF426F0FC83FED1FCB7" bold="true" box="[197,335,808,834]" pageId="11" pageNumber="62">Etymology</emphasis>
|
||
. The species name is Sinhala for horn, an allusion to the horn-like tubercles on the upper eyelids of this frog; applied as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307E6C4643DFFF426F0FCDDFE7BFB8D" pageId="11" pageNumber="62" type="discussion">
|
||
<paragraph id="8BA2B54F643DFFF426F0FCDDFE7BFB8D" blockId="11.[151,1437,152,1261]" pageId="11" pageNumber="62">
|
||
<emphasis id="B969695D643DFFF426F0FCDDFEA9FC65" bold="true" box="[197,311,886,912]" pageId="11" pageNumber="62">Remarks</emphasis>
|
||
. Morphologically,
|
||
<taxonomicName id="4C1DCECC643DFFF42422FCDCFD51FC65" box="[535,719,887,912]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="singu">
|
||
<emphasis id="B969695D643DFFF42422FCDCFD51FC65" box="[535,719,887,912]" italics="true" pageId="11" pageNumber="62">Philautus singu</emphasis>
|
||
</taxonomicName>
|
||
is reminiscent of
|
||
<emphasis id="B969695D643DFFF42597FCDCFB50FC65" box="[930,1230,887,912]" italics="true" pageId="11" pageNumber="62">P. d e c o r i s, P. m i t t e r m e i e r i</emphasis>
|
||
and
|
||
<emphasis id="B969695D643DFFF4233EFCDCFAC6FC65" box="[1291,1368,887,912]" italics="true" pageId="11" pageNumber="62">P. t a n u</emphasis>
|
||
, new species. It is distinguished from
|
||
<taxonomicName id="4C1DCECC643DFFF4241DFC36FD3CFC43" box="[552,674,925,950]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="decoris">
|
||
<emphasis id="B969695D643DFFF4241DFC36FD3CFC43" box="[552,674,925,950]" italics="true" pageId="11" pageNumber="62">P. decoris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1DCECC643DFFF424EAFC36FC0BFC43" box="[735,917,925,950]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="mittermeieri">
|
||
<emphasis id="B969695D643DFFF424EAFC36FC0BFC43" box="[735,917,925,950]" italics="true" pageId="11" pageNumber="62">P. mittermeieri</emphasis>
|
||
</taxonomicName>
|
||
, however, by the presence of a prominent tubercle on the upper eyelid (absent in
|
||
<taxonomicName id="4C1DCECC643DFFF4246AFC6FFC93FC28" box="[607,781,964,989]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="mittermeieri">
|
||
<emphasis id="B969695D643DFFF4246AFC6FFC93FC28" box="[607,781,964,989]" italics="true" pageId="11" pageNumber="62">P. mittermeieri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1DCECC643DFFF4252EFC6FFC13FC28" box="[795,909,964,989]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="decoris">
|
||
<emphasis id="B969695D643DFFF4252EFC6FFC13FC28" box="[795,909,964,989]" italics="true" pageId="11" pageNumber="62">P. decoris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B969695D643DFFF425F3FC6FFB8DFC28" box="[966,1043,964,989]" italics="true" pageId="11" pageNumber="62">P. t a n u</emphasis>
|
||
); by having the snout rounded in lateral aspect (pointed in
|
||
<emphasis id="B969695D643DFFF427F4FC41FDE8FBF6" box="[449,630,1002,1027]" italics="true" pageId="11" pageNumber="62">
|
||
<taxonomicName id="4C1DCECC643DFFF427F4FC41FDECFBF6" box="[449,626,1002,1027]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="mittermeieri">P. mittermeieri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and obtusely pointed in both
|
||
<taxonomicName id="4C1DCECC643DFFF425E3FC41FBD7FBF6" box="[982,1097,1002,1027]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="decoris">
|
||
<emphasis id="B969695D643DFFF425E3FC41FBD7FBF6" box="[982,1097,1002,1027]" italics="true" pageId="11" pageNumber="62">P. decoris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1DCECC643DFFF422B6FC41FB4BFBF6" box="[1155,1237,1002,1027]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="tanu">
|
||
<emphasis id="B969695D643DFFF422B6FC41FB4BFBF6" box="[1155,1237,1002,1027]" italics="true" pageId="11" pageNumber="62">P. tanu</emphasis>
|
||
</taxonomicName>
|
||
); by the absence of a tarsal tubercle (present in
|
||
<taxonomicName id="4C1DCECC643DFFF427CBFBBAFD32FBDF" box="[510,684,1041,1066]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="mittermeieri">
|
||
<emphasis id="B969695D643DFFF427CBFBBAFD32FBDF" box="[510,684,1041,1066]" italics="true" pageId="11" pageNumber="62">P. mittermeieri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B969695D643DFFF424D3FBBAFCC8FBDF" box="[742,854,1041,1066]" italics="true" pageId="11" pageNumber="62">P. d e c o r i s</emphasis>
|
||
); absence of a lateral dermal fringe and webbing on fingers (present in
|
||
<emphasis id="B969695D643DFFF427A8FB93FDD7FBA4" box="[413,585,1080,1105]" italics="true" pageId="11" pageNumber="62">P. m i t t e r m e i e r i</emphasis>
|
||
and
|
||
<taxonomicName id="4C1DCECC643DFFF424B2FB93FD62FBA4" box="[647,764,1080,1105]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="decoris">
|
||
<emphasis id="B969695D643DFFF424B2FB93FD62FBA4" box="[647,764,1080,1105]" italics="true" pageId="11" pageNumber="62">P. decoris</emphasis>
|
||
</taxonomicName>
|
||
); and by the absence of a tarsal fold (presenct in both
|
||
<taxonomicName id="4C1DCECC643DFFF423B3FB93FEB6FB8D" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="mittermeieri">
|
||
<emphasis id="B969695D643DFFF423B3FB93FEB6FB8D" italics="true" pageId="11" pageNumber="62">P. mittermeieri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B969695D643DFFF42754FBF4FE4EFB8D" box="[353,464,1119,1144]" italics="true" pageId="11" pageNumber="62">P. d e c o r i s</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307E6C4643DFFF426F0FB2EFABEFB18" pageId="11" pageNumber="62" type="distribution">
|
||
<paragraph id="8BA2B54F643DFFF426F0FB2EFABEFB18" blockId="11.[151,1437,152,1261]" pageId="11" pageNumber="62">
|
||
<emphasis id="B969695D643DFFF426F0FB2EFEC0FB6A" bold="true" box="[197,350,1157,1183]" pageId="11" pageNumber="62">Distribution</emphasis>
|
||
. We observed males of
|
||
<taxonomicName id="4C1DCECC643DFFF4244EFB2EFD46FB6B" box="[635,728,1157,1182]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="singu">
|
||
<emphasis id="B969695D643DFFF4244EFB2EFD46FB6B" box="[635,728,1157,1182]" italics="true" pageId="11" pageNumber="62">P. singu</emphasis>
|
||
</taxonomicName>
|
||
perched on leaves of shrubs,
|
||
<quantity id="4CE518AA643DFFF4220DFB2EFB34FB6A" box="[1080,1194,1157,1183]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="11" pageNumber="62" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5-1.5 m</quantity>
|
||
above ground level, in the rainforest understory. Although we recorded the species only from the Kottawa and Kitulgala Forest Reserves, it probably occurs also in other rainforest patches in the wet-zone lowlands of
|
||
<collectingCountry id="F30AF5DF643DFFF42293FB78FA82FB18" box="[1190,1308,1235,1261]" name="Sri Lanka" pageId="11" pageNumber="62">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |