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<document id="67A0964C6A7322A1A3DCEFF61888028C" ID-DOI="10.5281/zenodo.10081494" ID-ISSN="2345-7600" ID-RefBankID="FF8FFF8D2661FFF88F4F3F0AFFDBB877" ID-Zenodo-Dep="10081494" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616096481571" checkinUser="carolina" docAuthor="Chen, Wen-Jou, Cheng, Jin-Hua, Jhy-Yun &amp; Shy" docDate="2005" docId="EC4987F9FFF3FFE60EC717B6FC0AFB5B" docLanguage="en" docName="RafflesBZool.53.1.103-110.pdf" docOrigin="Raffles Bulletin of Zoology 53 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Geothelphusa lili Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy 2005, new species" docType="treatment" docVersion="4" lastPageNumber="107" masterDocId="1070FF81FFF2FFE20D051037FFE1FFE3" masterDocTitle="On Two New Species Of Freshwater Crabs (Crustacea: Decapoda: Brachyura: Potamidae) From Southern Taiwan" masterLastPageNumber="110" masterPageNumber="103" pageNumber="104" updateTime="1700277715140" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1C78D4F14B4C46ED9B571D3DB5C74AE5">On Two New Species Of Freshwater Crabs (Crustacea: Decapoda: Brachyura: Potamidae) From Southern Taiwan</mods:title>
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<mods:namePart id="68AF00F667B0EE20F687B7F9AA5EF6D0">Chen, Wen-Jou</mods:namePart>
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<mods:namePart id="E35047AC4B89370EECE3E5BE96DEFA49">Cheng, Jin-Hua</mods:namePart>
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<mods:namePart id="029EBE048FCA9CEB243D2838D53B8BFD">Jhy-Yun</mods:namePart>
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<taxonomicName id="A3E04D6CFFF3FFE30EC717B6FB63F87A" ID-CoL="3FT42" authority="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy, 2005" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[962,1154,1921,1945]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="lili" status="sp. nov.">
<emphasis id="5694EAFDFFF3FFE30EC717B6FB63F87A" bold="true" box="[962,1154,1921,1945]" italics="true" pageId="1" pageNumber="104">Geothelphusa lili</emphasis>
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, 
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(
<figureCitation id="FCDB2A6AFFF3FFE309241796FB7AF85B" box="[1057,1179,1953,1976]" captionStart="Fig" captionStartId="2.[136,171,2065,2086]" captionTargetBox="[157,1456,1044,2021]" captionTargetPageId="2" captionText="Fig. 2, A-B: Geothelphusa shernshan, new species. Holotype male, 29.5 by 22.5 mm (NTOU F000101). C-D: G. lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A, C: dorsal view; B, D: frontal view." figureDoi="http://doi.org/10.5281/zenodo.10081498" httpUri="https://zenodo.org/record/10081498/files/figure.png" pageId="1" pageNumber="104">Figs. 2C,D</figureCitation>
, 
<figureCitation id="FCDB2A6AFFF3FFE309AD1796FB57F85B" box="[1192,1206,1953,1976]" captionStart="Fig" captionStartId="6.[136,172,2037,2058]" captionTargetBox="[193,1395,253,1956]" captionTargetId="figure-67@6.[163,1443,231,1988]" captionTargetPageId="6" captionText="Fig. 6, Geothelphusa lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A: carapace; B: ventral view of right G1; C: dorsal view of right G1; D: ventral view of right G2; d: ventral view of right G1 terminal segment. Scale: a – c = 1 mm, d = 0.1 mm, e = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10081506" httpUri="https://zenodo.org/record/10081506/files/figure.png" pageId="1" pageNumber="104">6</figureCitation>
)
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<emphasis id="5694EAFDFFF3FFE30E2917D7FC08F816" bold="true" box="[812,1001,2016,2037]" italics="true" pageId="1" pageNumber="104">Material examined</emphasis>
. 
<emphasis id="5694EAFDFFF3FFE30EF017D7FBE0F816" bold="true" box="[1013,1025,2016,2037]" pageId="1" pageNumber="104">–</emphasis>
<materialsCitation id="D4883CB2FFF3FFE3090317D7FA41F7F1" collectingDate="2000-07-26" collectionCode="NTOU" collectorName="W. J. Chen" location="Pingtung county" pageId="1" pageNumber="104" specimenCode="F000701" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="5694EAFDFFF3FFE3090317D7FB84F816" bold="true" box="[1030,1125,2016,2037]" pageId="1" pageNumber="104">
<typeStatus id="BB5B884DFFF3FFE3090317D7FB84F816" box="[1030,1125,2016,2037]" pageId="1" pageNumber="104" type="holotype">Holotype</typeStatus>
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- male (20.1 by 
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)(
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), Laii, 
<collectingRegion id="A624F80DFFF3FFE30EC917CAFB91F7F1" box="[972,1136,2045,2066]" country="Taiwan" name="Pingtung" pageId="1" pageNumber="104">Pingtung county</collectingRegion>
, coll. 
<collectorName id="C9155339FFF3FFE309A917CAFAF7F7F1" box="[1196,1302,2045,2066]" pageId="1" pageNumber="104">W.J. Chen</collectorName>
, 
<date id="105E102FFFF3FFE3082517CAFA7DF7F1" box="[1312,1436,2045,2066]" pageId="1" pageNumber="104" value="2000-07-26">
<collectingDate id="001AE9C7FFF3FFE3082517CAFA7DF7F1" box="[1312,1436,2045,2066]" pageId="1" pageNumber="104" value="2000-07-26">26 Jul. 2000</collectingDate>
</date>
.
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</paragraph>
<paragraph id="645F36EFFFF0FFE00D8D1283FF0EFCFC" blockId="2.[136,775,692,799]" pageId="2" pageNumber="105">
<materialsCitation id="D4883CB2FFF0FFE00D8D1283FE01FD05" collectingDate="2000-07-26" collectionCode="NTOU" collectorName="W. J. Chen" location="Pingtung county" pageId="2" pageNumber="105" specimenCode="F000701" specimenCount="4" specimenCount-female="4" stateProvince="Pingtung" typeStatus="paratype">
<typeStatus id="BB5B884DFFF0FFE00D8D1283FF06FD2A" box="[136,231,692,713]" pageId="2" pageNumber="105" type="paratype">Paratypes</typeStatus>
– 
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(largest female 18.7 by 
<quantity id="A3189B0AFFF0FFE00F5F1283FD55FD2A" box="[602,692,692,713]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="105" unit="mm" value="14.9">14.9 mm</quantity>
)(
<collectionCode id="02F1AE2AFFF0FFE00FC41283FCE7FD2A" box="[705,774,692,713]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="105">NTOU</collectionCode>
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), same data as holotype
</materialsCitation>
; 
<materialsCitation id="D4883CB2FFF0FFE00CEB12E6FF0AFCFC" collectingDate="2000-10-15" collectionCode="NTOU" collectorName="W. J. Chen &amp; C. F. Tseng" location="Pingtung county" pageId="2" pageNumber="105" specimenCode="F001001" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Pingtung" typeStatus="paratype">
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, 
<specimenCount id="72E6FD66FFF0FFE00F4812E6FD51FD05" box="[589,688,721,742]" pageId="2" pageNumber="105" type="female">2 females</specimenCount>
(
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), Laii, 
<collectingRegion id="A624F80DFFF0FFE00C2712D9FE23FCE0" box="[290,450,750,771]" country="Taiwan" name="Pingtung" pageId="2" pageNumber="105">Pingtung county</collectingRegion>
, coll. 
<collectorName id="C9155339FFF0FFE00CFF12D9FDBFFCE0" box="[506,606,750,771]" pageId="2" pageNumber="105">W.J. Chen</collectorName>
&amp; 
<collectorName id="C9155339FFF0FFE00F7F12D9FD07FCE0" box="[634,742,750,771]" pageId="2" pageNumber="105">C.F. Tseng</collectorName>
, 
<date id="105E102FFFF0FFE00FEB12D9FF0AFCFC" pageId="2" pageNumber="105" value="2000-10-15">
<collectingDate id="001AE9C7FFF0FFE00FEB12D9FF0AFCFC" pageId="2" pageNumber="105" value="2000-10-15">15 Oct. 2000</collectingDate>
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.
</paragraph>
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Others – 
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<specimenCount id="72E6FD66FFF0FFE00DDE1373FEF8FCBA" box="[219,281,836,857]" pageId="2" pageNumber="105" type="male">1 male</specimenCount>
(18.5 by 
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)(
<collectionCode id="02F1AE2AFFF0FFE00CD01373FDF6FCBA" box="[469,535,836,857]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="105">NTOU</collectionCode>
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), 
<location id="613F6034FFF0FFE00F841373FD48FCBA" LSID="urn:lsid:plazi:treatment:EC4987F9FFF3FFE60EC717B6FC0AFB5B:613F6034FFF0FFE00F841373FD48FCBA" box="[641,681,836,857]" name="Laii" pageId="2" pageNumber="105" stateProvince="Pingtung">Laii</location>
, 
<collectingRegion id="A624F80DFFF0FFE00FB41373FF2AFC95" country="Taiwan" name="Pingtung" pageId="2" pageNumber="105">Pingtung county</collectingRegion>
, coll. 
<collectorName id="C9155339FFF0FFE00C051356FE83FC95" box="[256,354,865,886]" pageId="2" pageNumber="105">W.J. Chen</collectorName>
&amp; 
<collectorName id="C9155339FFF0FFE00C7B1356FE06FC95" box="[382,487,865,886]" pageId="2" pageNumber="105">C.F. Tseng</collectorName>
, 
<date id="105E102FFFF0FFE00CEA1356FDBDFC95" box="[495,604,865,886]" pageId="2" pageNumber="105" value="2001-01-01">
<collectingDate id="001AE9C7FFF0FFE00CEA1356FDBDFC95" box="[495,604,865,886]" pageId="2" pageNumber="105" value="2001-01-01">1 Jan. 2001</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="D4883CB2FFF0FFE00F661356FDB4FC4C" collectingDate="2001-10-28" collectionCode="SYSU" collectorName="W. J. Chen" location="Machia" pageId="2" pageNumber="105" specimenCode="F01122802" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="72E6FD66FFF0FFE00F661356FD45FC95" box="[611,676,865,886]" pageId="2" pageNumber="105" type="male">1 male</specimenCount>
, 
<specimenCount id="72E6FD66FFF0FFE00FA91356FCE6FC95" box="[684,775,865,886]" pageId="2" pageNumber="105" type="female">3 females</specimenCount>
(largest female 20.0 by 
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) (
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), 
<location id="613F6034FFF0FFE00FB21349FCE2FC70" LSID="urn:lsid:plazi:treatment:EC4987F9FFF3FFE60EC717B6FC0AFB5B:613F6034FFF0FFE00FB21349FCE2FC70" box="[695,771,894,915]" name="Machia" pageId="2" pageNumber="105" stateProvince="Pingtung">Machia</location>
, 
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, coll. 
<collectorName id="C9155339FFF0FFE00C6613ADFE2AFC4C" box="[355,459,922,943]" pageId="2" pageNumber="105">W.J. Chen</collectorName>
, 
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<collectingDate id="001AE9C7FFF0FFE00CD113ADFDB4FC4C" box="[468,597,922,943]" pageId="2" pageNumber="105" value="2001-10-28">28 Oct. 2001</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="D4883CB2FFF0FFE00F5813ADFAEAFD2A" collectingDate="2002-01-06" collectionCode="SYSU" collectorName="W. J. Chen" location="Sandimen" pageId="2" pageNumber="105" specimenCode="F02010601" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Pingtung">
<specimenCount id="72E6FD66FFF0FFE00F5813ADFD48FC4C" box="[605,681,922,943]" pageId="2" pageNumber="105" type="male">2 males</specimenCount>
, 
<specimenCount id="72E6FD66FFF0FFE00FB613ADFCE7FC4C" box="[691,774,922,943]" pageId="2" pageNumber="105" type="female">1 female</specimenCount>
(largest male 18.3 by 
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) (
<collectionCode id="02F1AE2AFFF0FFE00CD71380FDF0FC2F" box="[466,529,951,972]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="2" pageNumber="105" type="Herbarium">SYSU</collectionCode>
<specimenCode id="34469E94FFF0FFE00F1F1380FD6CFC2F" box="[538,653,951,972]" collectionCode="SYSU" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="2" pageNumber="105" type="Herbarium">F02010601</specimenCode>
), 
<location id="613F6034FFF0FFE00F9A1380FCE2FC2F" LSID="urn:lsid:plazi:treatment:EC4987F9FFF3FFE60EC717B6FC0AFB5B:613F6034FFF0FFE00F9A1380FCE2FC2F" box="[671,771,951,972]" name="Sandimen" pageId="2" pageNumber="105" stateProvince="Pingtung">Sandimen</location>
, 
<collectingRegion id="A624F80DFFF0FFE00E461283FC06FD2A" box="[835,999,692,713]" country="Taiwan" name="Pingtung" pageId="2" pageNumber="105">Pingtung county</collectingRegion>
, coll. 
<collectorName id="C9155339FFF0FFE009261283FB6DFD2A" box="[1059,1164,692,713]" pageId="2" pageNumber="105">W.J. Chen</collectorName>
, 
<date id="105E102FFFF0FFE009931283FAEAFD2A" box="[1174,1291,692,713]" pageId="2" pageNumber="105" value="2002-01-06">
<collectingDate id="001AE9C7FFF0FFE009931283FAEAFD2A" box="[1174,1291,692,713]" pageId="2" pageNumber="105" value="2002-01-06">6 Jan. 2002</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="D4883CB2FFF0FFE008111283FAA2FCE0" collectingDate="2001-02-17" collectionCode="SYSU" collectorName="W. J. Chen &amp; C. F. Tseng" location="Taiwu" pageId="2" pageNumber="105" specimenCode="F010217" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Pingtung">
<specimenCount id="72E6FD66FFF0FFE008111283FA83FD2A" box="[1300,1378,692,713]" pageId="2" pageNumber="105" type="male">2 males</specimenCount>
, 
<specimenCount id="72E6FD66FFF0FFE008681283FA23FD2A" box="[1389,1474,692,713]" pageId="2" pageNumber="105" type="female">1 female</specimenCount>
(largest male 20.2 by 
<quantity id="A3189B0AFFF0FFE0091412E6FB83FD05" box="[1041,1122,721,742]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="2" pageNumber="105" unit="mm" value="16.1">16.1mm</quantity>
) (
<collectionCode id="02F1AE2AFFF0FFE0097112E6FB52FD05" box="[1140,1203,721,742]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="2" pageNumber="105" type="Herbarium">SYSU</collectionCode>
<specimenCode id="34469E94FFF0FFE009BC12E6FAF3FD05" box="[1209,1298,721,742]" collectionCode="SYSU" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="2" pageNumber="105" type="Herbarium">F010217</specimenCode>
), 
<location id="613F6034FFF0FFE0082712E6FA83FD05" LSID="urn:lsid:plazi:treatment:EC4987F9FFF3FFE60EC717B6FC0AFB5B:613F6034FFF0FFE0082712E6FA83FD05" box="[1314,1378,721,742]" name="Taiwu" pageId="2" pageNumber="105" stateProvince="Pingtung">Taiwu</location>
, 
<collectingRegion id="A624F80DFFF0FFE0086F12E6FC69FCE0" country="Taiwan" name="Pingtung" pageId="2" pageNumber="105">Pingtung county</collectingRegion>
, coll. 
<collectorName id="C9155339FFF0FFE00EC112D9FBCDFCE0" box="[964,1068,750,771]" pageId="2" pageNumber="105">W.J. Chen</collectorName>
&amp; 
<collectorName id="C9155339FFF0FFE0094012D9FB55FCE0" box="[1093,1204,750,771]" pageId="2" pageNumber="105">C.F. Tseng</collectorName>
, 
<date id="105E102FFFF0FFE009BA12D9FAA2FCE0" box="[1215,1347,750,771]" pageId="2" pageNumber="105" value="2001-02-17">
<collectingDate id="001AE9C7FFF0FFE009BA12D9FAA2FCE0" box="[1215,1347,750,771]" pageId="2" pageNumber="105" value="2001-02-17">17 Feb. 2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2CFA6564FFF0FFE60E46131CFAD5FD7B" lastPageId="4" lastPageNumber="107" pageId="2" pageNumber="105" type="diagnosis">
<paragraph id="645F36EFFFF0FFE60E46131CFF16FEBB" blockId="2.[835,1475,811,962]" lastBlockId="4.[136,775,193,344]" lastPageId="4" lastPageNumber="107" pageId="2" pageNumber="105">
<emphasis id="5694EAFDFFF0FFE00E46131CFC2BFCA0" bold="true" box="[835,970,811,835]" pageId="2" pageNumber="105">
<emphasis id="5694EAFDFFF0FFE00E46131CFC50FCA0" bold="true" box="[835,945,811,835]" italics="true" pageId="2" pageNumber="105">Diagnosis</emphasis>
. –
</emphasis>
Carapace with fine pits, width and length about 2.3 and 1.8 times depth; width about 1.3 times length; frontal width about 0.3 times carapace width (Table 1). Gastric region smooth, anterolateral and posterolateral regions with striae. Postorbital crista faint, orbital and frontal region gently concaved, divided into 2 lobes. Anterolateral crista distinct, epibranchial tooth absent, lined with small granules. Cervical groove shallow, faint. H-shaped gastric groove is distinct. Posterior epistomal margin sinuous, median lobe not acutely triangular.
</paragraph>
<caption id="309F6667FFF0FFE00D8D1826FC53F7A0" ID-DOI="http://doi.org/10.5281/zenodo.10081498" ID-Zenodo-Dep="10081498" httpUri="https://zenodo.org/record/10081498/files/figure.png" pageId="2" pageNumber="105" startId="2.[136,171,2065,2086]" targetBox="[157,1456,1044,2021]" targetPageId="2">
<paragraph id="645F36EFFFF0FFE00D8D1826FC53F7A0" blockId="2.[136,1474,2065,2115]" pageId="2" pageNumber="105">
Fig. 2, A-B: 
<taxonomicName id="A3E04D6CFFF0FFE00DFA1825FE0EF7C4" authority="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy, 2005" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[255,495,2066,2087]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="shernshan" status="sp. nov.">
<emphasis id="5694EAFDFFF0FFE00DFA1825FE0EF7C4" box="[255,495,2066,2087]" italics="true" pageId="2" pageNumber="105">Geothelphusa shernshan</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="4DA75786FFF0FFE00CFF1826FD91F7C5" box="[506,624,2065,2086]" pageId="2" pageNumber="105" rank="species">new species</taxonomicNameLabel>
. Holotype male, 29.5 by 22.5 mm (NTOU F000101). C-D: 
<taxonomicName id="A3E04D6CFFF0FFE009A21825FB3FF7C4" authority="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy, 2005" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[1191,1246,2066,2087]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="lili" status="sp. nov.">
<emphasis id="5694EAFDFFF0FFE009A21825FB3FF7C4" box="[1191,1246,2066,2087]" italics="true" pageId="2" pageNumber="105">G. lili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="4DA75786FFF0FFE009EC1826FA81F7C5" box="[1257,1376,2065,2086]" pageId="2" pageNumber="105" rank="species">new species</taxonomicNameLabel>
. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A, C: dorsal view; B, D: frontal view.
</paragraph>
</caption>
<caption id="309F6667FFF1FFE10D7417C2FB77F7A0" ID-DOI="http://doi.org/10.5281/zenodo.10081500" ID-Zenodo-Dep="10081500" httpUri="https://zenodo.org/record/10081500/files/figure.png" pageId="3" pageNumber="106" startId="3.[113,149,2037,2058]" targetBox="[112,1452,195,1994]" targetPageId="3">
<paragraph id="645F36EFFFF1FFE10D7417C2FB77F7A0" blockId="3.[113,1452,2037,2115]" pageId="3" pageNumber="106">
Fig. 3, A, D: 
<taxonomicName id="A3E04D6CFFF1FFE10DF017C2FE09F7E9" authority="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy, 2005" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[245,488,2037,2058]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="shernshan" status="sp. nov.">
<emphasis id="5694EAFDFFF1FFE10DF017C2FE09F7E9" box="[245,488,2037,2058]" italics="true" pageId="3" pageNumber="106">Geothelphusa shernshan</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="4DA75786FFF1FFE10CF017C2FD8EF7E9" box="[501,623,2037,2058]" pageId="3" pageNumber="106" rank="species">new species</taxonomicNameLabel>
. Paratype male, 24.3 by 18.5 mm (NTOU F011201). B, E: 
<taxonomicName id="A3E04D6CFFF1FFE109BD17C2FAEDF7E9" box="[1208,1292,2037,2058]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="neipu">
<emphasis id="5694EAFDFFF1FFE109BD17C2FAEDF7E9" box="[1208,1292,2037,2058]" italics="true" pageId="3" pageNumber="106">G. neipu</emphasis>
</taxonomicName>
. Paratype male, 34.1 by 26.4 mm (NTOU F001004). C, F: 
<taxonomicName id="A3E04D6CFFF1FFE10F0A1825FD66F7C4" authorityName="Tan &amp; Liu" authorityYear="1998" box="[527,647,2065,2087]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="pingtung">
<emphasis id="5694EAFDFFF1FFE10F0A1825FDC1F7C4" box="[527,544,2066,2087]" italics="true" pageId="3" pageNumber="106">G</emphasis>
. 
<emphasis id="5694EAFDFFF1FFE10F281825FD66F7C4" box="[557,647,2066,2087]" italics="true" pageId="3" pageNumber="106">pingtung</emphasis>
</taxonomicName>
<emphasis id="5694EAFDFFF1FFE10F821825FD6AF7C4" box="[647,651,2066,2087]" italics="true" pageId="3" pageNumber="106">.</emphasis>
Paratype male, 35.9 by 28.9 mm (NTOU F001005). All dorsal view of G1, A - C: terminal segment; D - F: synovial membrane. Scale, A = 100 m; B,C = 200 m; D,E = 1 mm; F = 2 mm.
</paragraph>
</caption>
<paragraph id="645F36EFFFF6FFE60D8D11B6FE02FDFB" blockId="4.[136,775,385,536]" pageId="4" pageNumber="107">Abdomens of male and female with 7 distinct segments (including telson); in male, length of telson 0.8 times width, longer than 6th, as in female (Table 2). Distance between tip of male abdomen and anterior margin of thoracic sternite 4 about 0.9 times of sternites 1-3.</paragraph>
<paragraph id="645F36EFFFF6FFE60D8D1276FF28FCDB" blockId="4.[136,775,577,824]" pageId="4" pageNumber="107">Chelipeds unequal, in large chela, outer surface of palm smooth, upper margin length 0.7 times length of dactylus, height 0.6 times length of chela (Table 3). Length of dactylus 0.7 times that of chela. Fingers forming obliquely triangular gape when closed, with faint cutting teeth. Carpus of large chela with 1 large spine on inner, upper surface and a subequal one below it. Inner and outer surfaces of merus with fine striae.</paragraph>
<paragraph id="645F36EFFFF6FFE60D8D1356FE64FBDB" blockId="4.[136,775,865,1080]" pageId="4" pageNumber="107">Ambulatory legs slender, with simple setae; second longest, last shortest. Second ambulatory leg about 1.8 times carapace length. In last 2 segments of right second ambulatory leg, width of propodus 0.4 times length, 0.4 times the length of dactylus; length of propodus 0.8 times dactylus (Table 4). Dactylus and propodus with a row of 3 - 9 spines on each of 4 longitudinal margins.</paragraph>
<paragraph id="645F36EFFFF6FFE60D8D1456FA23FE5B" blockId="4.[136,775,1121,1208]" lastBlockId="4.[835,1474,193,440]" pageId="4" pageNumber="107">
G1 subterminal segment straight; outer proximal margin with faint tooth and inner proximal margin dilated, both proximal margins with plumose setae, especially on inner proximal margin (
<figureCitation id="FCDB2A6AFFF6FFE60EA510F6FC05FF3B" box="[928,996,193,216]" captionStart="Fig" captionStartId="6.[136,172,2037,2058]" captionTargetBox="[193,1395,253,1956]" captionTargetId="figure-67@6.[163,1443,231,1988]" captionTargetPageId="6" captionText="Fig. 6, Geothelphusa lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A: carapace; B: ventral view of right G1; C: dorsal view of right G1; D: ventral view of right G2; d: ventral view of right G1 terminal segment. Scale: a – c = 1 mm, d = 0.1 mm, e = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10081506" httpUri="https://zenodo.org/record/10081506/files/figure.png" pageId="4" pageNumber="107">Fig. 6</figureCitation>
). Terminal segment of G1 straight, slightly curved outwards, 4-5 rings of distal scales are distributed and subterminal has sparsely small spines (
<figureCitation id="FCDB2A6AFFF6FFE609F51136FA95FEFB" box="[1264,1396,257,280]" captionStart="Fig" captionStartId="4.[136,172,2037,2058]" captionTargetBox="[137,1470,1319,1994]" captionTargetPageId="4" captionText="Fig. 4, A, B: Geothelphusa lili, new species. Paratype male, 18.5 by 15.1 mm (NTOU F010101). C, D: G. ferruginea. Paratype male, 20.8 by 16.5 mm (NTOU F020601). All dorsal view of G1; B, D: terminal segment; A, C: synovial membrane. Scale, A, C = 1 mm; B, D: 100 m." figureDoi="http://doi.org/10.5281/zenodo.10081502" httpUri="https://zenodo.org/record/10081502/files/figure.png" pageId="4" pageNumber="107">Figs. 4A, B</figureCitation>
). Total length of G1 5.5 times of terminal segment length, 3.0 times of subterminal segment width. Terminal segment length of G1 2.8 times of its width, synovial membrane length 4.6 times of its width (
<figureCitation id="FCDB2A6AFFF6FFE60ED611B6FBCDFE7B" box="[979,1068,385,408]" captionStart="Fig" captionStartId="4.[136,172,2037,2058]" captionTargetBox="[137,1470,1319,1994]" captionTargetPageId="4" captionText="Fig. 4, A, B: Geothelphusa lili, new species. Paratype male, 18.5 by 15.1 mm (NTOU F010101). C, D: G. ferruginea. Paratype male, 20.8 by 16.5 mm (NTOU F020601). All dorsal view of G1; B, D: terminal segment; A, C: synovial membrane. Scale, A, C = 1 mm; B, D: 100 m." figureDoi="http://doi.org/10.5281/zenodo.10081502" httpUri="https://zenodo.org/record/10081502/files/figure.png" pageId="4" pageNumber="107">Fig. 4A</figureCitation>
). Distal segment of G2 straight, flat, directed upwards, length 0.2 times of total length (
<figureCitation id="FCDB2A6AFFF6FFE6085A1196FA57FE5B" box="[1375,1462,417,440]" captionStart="Fig" captionStartId="6.[136,172,2037,2058]" captionTargetBox="[193,1395,253,1956]" captionTargetId="figure-67@6.[163,1443,231,1988]" captionTargetPageId="6" captionText="Fig. 6, Geothelphusa lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A: carapace; B: ventral view of right G1; C: dorsal view of right G1; D: ventral view of right G2; d: ventral view of right G1 terminal segment. Scale: a – c = 1 mm, d = 0.1 mm, e = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10081506" httpUri="https://zenodo.org/record/10081506/files/figure.png" pageId="4" pageNumber="107">Fig. 6D</figureCitation>
).
</paragraph>
<paragraph id="645F36EFFFF6FFE60E4611D6FC3AFDBB" blockId="4.[835,1475,480,600]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4611D6FC33FE1B" bold="true" box="[835,978,480,505]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4611D6FC58FE1A" bold="true" box="[835,953,481,505]" italics="true" pageId="4" pageNumber="107">Coloration</emphasis>
. –
</emphasis>
Carapace and upper surface of ambulatory legs dark brown, but posterior half of carapace slightly dark green. Dorsal part of outer and inner surfaces of cheliped purple (
<figureCitation id="FCDB2A6AFFF6FFE60E4E1276FC2FFDBB" box="[843,974,577,600]" captionStart="Fig" captionStartId="2.[136,171,2065,2086]" captionTargetBox="[157,1456,1044,2021]" captionTargetPageId="2" captionText="Fig. 2, A-B: Geothelphusa shernshan, new species. Holotype male, 29.5 by 22.5 mm (NTOU F000101). C-D: G. lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A, C: dorsal view; B, D: frontal view." figureDoi="http://doi.org/10.5281/zenodo.10081498" httpUri="https://zenodo.org/record/10081498/files/figure.png" pageId="4" pageNumber="107">Figs. 2C, D</figureCitation>
).
</paragraph>
<paragraph id="645F36EFFFF6FFE60E4612B6FAD5FD7B" blockId="4.[835,1332,640,665]" box="[835,1332,640,665]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4612B6FC76FD7A" bold="true" box="[835,919,641,665]" italics="true" pageId="4" pageNumber="107">Habitat</emphasis>
. 
<emphasis id="5694EAFDFFF6FFE60EA312B7FC52FD7B" bold="true" box="[934,947,640,664]" pageId="4" pageNumber="107">–</emphasis>
Lives under boulders near springs.
</paragraph>
</subSubSection>
<subSubSection id="2CFA6564FFF6FFE60E4612F6FAE9FD1B" pageId="4" pageNumber="107" type="etymology">
<paragraph id="645F36EFFFF6FFE60E4612F6FAE9FD1B" blockId="4.[835,1474,704,760]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4612F6FC30FD3B" bold="true" box="[835,977,704,729]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4612F6FC59FD3A" bold="true" box="[835,952,705,729]" italics="true" pageId="4" pageNumber="107">Etymology</emphasis>
. –
</emphasis>
The species is named after its 
<typeStatus id="BB5B884DFFF6FFE6081712F6FAA1FD3B" box="[1298,1344,705,728]" pageId="4" pageNumber="107">type</typeStatus>
locality and the name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="2CFA6564FFF6FFE60E461316FBD6FBBB" pageId="4" pageNumber="107" type="discussion">
<paragraph id="645F36EFFFF6FFE60E461316FBD6FBBB" blockId="4.[835,1474,800,1112]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E461316FC4CFCDB" box="[835,941,801,825]" italics="true" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E461316FC47FCDA" bold="true" box="[835,934,801,825]" italics="true" pageId="4" pageNumber="107">Remarks</emphasis>
.
</emphasis>
<emphasis id="5694EAFDFFF6FFE60EB11317FC20FCDB" bold="true" box="[948,961,800,824]" pageId="4" pageNumber="107">–</emphasis>
This species is allied to 
<taxonomicName id="A3E04D6CFFF6FFE609CB1316FC2AFCBB" authority="Shy, Ng &amp; Yu, 1994" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="5694EAFDFFF6FFE609CB1316FA83FCDB" box="[1230,1378,801,824]" italics="true" pageId="4" pageNumber="107">G. ferruginea</emphasis>
Shy, Ng &amp; Yu, 1994
</taxonomicName>
, but it can be distinguished by the following aspects: in 
<emphasis id="5694EAFDFFF6FFE60EB81356FBE4FC9B" box="[957,1029,865,888]" italics="true" pageId="4" pageNumber="107">
<taxonomicName id="A3E04D6CFFF6FFE60EB81356FC1EFC9B" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[957,1023,865,888]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="lili">G. lili</taxonomicName>
,
</emphasis>
the G1 synovial membrane length is 4.6 times its width, but 3.8 times in 
<taxonomicName id="A3E04D6CFFF6FFE609A313B6FC0CFC5B" authority="(Shy, Ng &amp; Yu, 1994: 783)" baseAuthorityName="Shy, Ng &amp; Yu" baseAuthorityPageNumber="783" baseAuthorityYear="1994" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="5694EAFDFFF6FFE609A313B6FADAFC7B" box="[1190,1339,897,920]" italics="true" pageId="4" pageNumber="107">G. ferruginea</emphasis>
(Shy, Ng &amp; Yu, 1994: 783)
</taxonomicName>
; in 
<taxonomicName id="A3E04D6CFFF6FFE609131396FBB6FC5B" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[1046,1111,929,952]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="lili">
<emphasis id="5694EAFDFFF6FFE609131396FBB6FC5B" box="[1046,1111,929,952]" italics="true" pageId="4" pageNumber="107">G. lili</emphasis>
</taxonomicName>
, terminal segment of G1 straight and slightly curves outwards but in 
<taxonomicName id="A3E04D6CFFF6FFE609CC13F6FABCFC3B" baseAuthorityName="Shy, Ng &amp; Yu" baseAuthorityYear="1994" box="[1225,1373,961,984]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="5694EAFDFFF6FFE609CC13F6FABCFC3B" box="[1225,1373,961,984]" italics="true" pageId="4" pageNumber="107">G. ferruginea</emphasis>
</taxonomicName>
it curves inwards founds the median part of the sternum (
<figureCitation id="FCDB2A6AFFF6FFE6083A13D6FA5CFC1B" box="[1343,1469,993,1016]" captionStart="Fig" captionStartId="4.[136,172,2037,2058]" captionTargetBox="[137,1470,1319,1994]" captionTargetPageId="4" captionText="Fig. 4, A, B: Geothelphusa lili, new species. Paratype male, 18.5 by 15.1 mm (NTOU F010101). C, D: G. ferruginea. Paratype male, 20.8 by 16.5 mm (NTOU F020601). All dorsal view of G1; B, D: terminal segment; A, C: synovial membrane. Scale, A, C = 1 mm; B, D: 100 m." figureDoi="http://doi.org/10.5281/zenodo.10081502" httpUri="https://zenodo.org/record/10081502/files/figure.png" pageId="4" pageNumber="107">Figs. 4A, C</figureCitation>
; 
<figureCitation id="FCDB2A6AFFF6FFE60E461436FC64FBFB" box="[835,901,1025,1048]" captionStart="Fig" captionStartId="6.[136,172,2037,2058]" captionTargetBox="[193,1395,253,1956]" captionTargetId="figure-67@6.[163,1443,231,1988]" captionTargetPageId="6" captionText="Fig. 6, Geothelphusa lili, new species. Holotype male, 20.1 by 15.5 mm (NTOU F000701). A: carapace; B: ventral view of right G1; C: dorsal view of right G1; D: ventral view of right G2; d: ventral view of right G1 terminal segment. Scale: a – c = 1 mm, d = 0.1 mm, e = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10081506" httpUri="https://zenodo.org/record/10081506/files/figure.png" pageId="4" pageNumber="107">6B, C</figureCitation>
); in 
<taxonomicName id="A3E04D6CFFF6FFE60EB31436FC17FBFB" authorityName="Chen &amp; Cheng &amp; Jhy-Yun &amp; Shy" authorityYear="2005" box="[950,1014,1025,1048]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="lili">
<emphasis id="5694EAFDFFF6FFE60EB31436FC17FBFB" box="[950,1014,1025,1048]" italics="true" pageId="4" pageNumber="107">G. lili</emphasis>
</taxonomicName>
, the total length of the G1 is 5.5 times the terminal segment length, but 6.0 times in 
<taxonomicName id="A3E04D6CFFF6FFE609F11416FBD2FBBB" authority="(Shy, Ng &amp; Yu, 1994: 783)" baseAuthorityName="Shy, Ng &amp; Yu" baseAuthorityPageNumber="783" baseAuthorityYear="1994" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="5694EAFDFFF6FFE609F11416FA62FBDB" box="[1268,1411,1057,1080]" italics="true" pageId="4" pageNumber="107">G. ferruginea</emphasis>
(Shy, Ng &amp; Yu, 1994: 783)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2CFA6564FFF6FFE60E4614B6FC0AFB5B" pageId="4" pageNumber="107" type="distribution">
<paragraph id="645F36EFFFF6FFE60E4614B6FC0AFB5B" blockId="4.[835,1474,1152,1208]" pageId="4" pageNumber="107">
<emphasis id="5694EAFDFFF6FFE60E4614B6FC2BFB7A" bold="true" box="[835,970,1153,1177]" italics="true" pageId="4" pageNumber="107">Distribution</emphasis>
. 
<emphasis id="5694EAFDFFF6FFE60ED314B7FC02FB7B" bold="true" box="[982,995,1152,1176]" pageId="4" pageNumber="107">–</emphasis>
Northern 
<collectingRegion id="A624F80DFFF6FFE6095414B6FAE5FB7B" box="[1105,1284,1153,1176]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="107">Pingtung county</collectingRegion>
, southern 
<collectingCountry id="1CF7767FFFF6FFE6087414B6FA23FB7B" box="[1393,1474,1153,1176]" name="Taiwan" pageId="4" pageNumber="107">Taiwan</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>