794 lines
65 KiB
XML
794 lines
65 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.67564" ID-GBIF-Dataset="57d9ade8-67e7-4d65-9581-b85a80e88893" ID-PMC="PMC8292283" ID-Pensoft-Pub="1313-2970-1048-145" ID-Pensoft-UUID="93FE96A4127A5D8FAA3A39BCB3E4B729" ID-PubMed="34316267" ID-ZooBank="578795FD41BF4807B0016F7B5AC4D4B9" ModsDocID="1313-2970-1048-145" checkinTime="1626283267302" checkinUser="pensoft" docAuthor="Lantsov, Vladimir I. & Pilipenko, Valentin E." docDate="2021" docId="E104034BF2F251DB8C145337C9D914B4" docLanguage="en" docName="ZooKeys 1048: 145-175" docOrigin="ZooKeys 1048" docPubDate="2021-07-13" docSource="http://dx.doi.org/10.3897/zookeys.1048.67564" docTitle="Tipula (Lunatipula) quadridentata Savchenko 1964" docType="treatment" docVersion="4" id="93FE96A4127A5D8FAA3A39BCB3E4B729" lastPageNumber="145" masterDocId="93FE96A4127A5D8FAA3A39BCB3E4B729" masterDocTitle="A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group" masterLastPageNumber="175" masterPageNumber="145" pageNumber="145" updateTime="1668150580932" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Lantsov, Vladimir I.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8275-496X</mods:nameIdentifier>
|
||
<mods:affiliation>Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
|
||
<mods:nameIdentifier type="email">lantsov@megalog.ru</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Pilipenko, Valentin E.</mods:namePart>
|
||
<mods:affiliation>Lomonosov Moscow State University, GSP- 1, Leninskie Gory, Moscow 119991, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-07-13</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>1048</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>145</mods:start>
|
||
<mods:end>175</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.1048.67564</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.67564</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-1048-145</mods:identifier>
|
||
<mods:identifier type="ZooBank">578795FD41BF4807B0016F7B5AC4D4B9</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">93FE96A4127A5D8FAA3A39BCB3E4B729</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="183543013" LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4" httpUri="http://treatment.plazi.org/id/E104034BF2F251DB8C145337C9D914B4" lastPageNumber="145" pageId="0" pageNumber="145">
|
||
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName LSID="E104034B-F2F2-51DB-8C14-5337C9D914B4" authority="Savchenko, 1964" authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) quadridentata" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="quadridentata" subGenus="Lunatipula">Tipula (Lunatipula) quadridentata Savchenko, 1964</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">Figs 8</figureCitation>
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">, 9</figureCitation>
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">, 12D-F</figureCitation>
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">, 13D</figureCitation>
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">, 14D,J</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName authority="quadridentata" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) quadridentata subsp. quadridentata" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="subSpecies" species="quadridentata" subGenus="Lunatipula" subSpecies="quadridentata">Tipula (Lunatipula) quadridentata quadridentata</taxonomicName>
|
||
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
|
||
: 392 (type locality: Russia, near Stavropol); Oosterbroek and Theowald 1992: 118;
|
||
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" publicationUrl="http://ccw.naturalis.nl/detail.php" refId="B8" refString="Oosterbroek, P, 2021. Catalogue of the Craneflies of the World. http://ccw.naturalis.nl/detail.php" title="Catalogue of the Craneflies of the World." url="http://ccw.naturalis.nl/detail.php" year="2021">Oosterbroek 2021</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<materialsCitation collectingDate="1954-05-21" collectingDateMax="1954-05-26" collectingDateMin="1954-05-21" collectorName="Medvedev, E. Savchenko, K. Arnoldi" country="Russia" county="Stavropolya" determinerName="E. Savchenko" elevation="850" location="Stavropolya" municipality="Staryi" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Stavropol" typeStatus="Holotype">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry name="Russia">Russia</collectingCountry>
|
||
•
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="”околицы">"okolitsy</normalizedToken>
|
||
<normalizedToken originalValue="Ставрополя">
|
||
<collectingCounty>Stavropolya</collectingCounty>
|
||
</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="байрачн.[ый">bairachn.[yi</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="лес”">les"</normalizedToken>
|
||
[near
|
||
<collectingRegion country="Russia" name="Stavropol'">Stavropol</collectingRegion>
|
||
, bairak forest] [small forest in steppe ravines];
|
||
<collectingDate value="1954-05-25">25 May 1954</collectingDate>
|
||
; [S.]
|
||
<collectorName>Medvedev</collectorName>
|
||
leg.; ZISP. /
|
||
<normalizedToken originalValue="“южн">"yuzhn</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="скл">skl</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="под">pod</normalizedToken>
|
||
<normalizedToken originalValue="пологом">pologom</normalizedToken>
|
||
[
|
||
<normalizedToken originalValue="леса]”">lesa]"</normalizedToken>
|
||
[southern slopes under the canopy forest] / "
|
||
<taxonomicName genus="T." lsidName="T." pageId="0" pageNumber="145" rank="genus">T.</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Edwards" authorityYear="1931" class="Insecta" family="Tipulidae" genus="Lunatipula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lunatipula" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Lunatipula</taxonomicName>
|
||
) quadridenta sp. n.,
|
||
<normalizedToken originalValue="опр">opr</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Е.Савченко”">
|
||
<collectorName>E.Savchenko</collectorName>
|
||
"
|
||
</normalizedToken>
|
||
[det.
|
||
<determinerName>E. Savchenko</determinerName>
|
||
]. The specimen is very badly damaged and glued together. The head is glued to the thorax; the abdomen is broken in half and glued together; two legs of uncertain position are glued to the specimen, one without coxa and trochanter, and the other missing the last four tarsal segments.
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<typeStatus>Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry name="Russia">Russia</collectingCountry>
|
||
•
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="“Старый">
|
||
"
|
||
<collectingMunicipality>Staryi</collectingMunicipality>
|
||
</normalizedToken>
|
||
<normalizedToken originalValue="лес">les</normalizedToken>
|
||
<normalizedToken originalValue="к">k</normalizedToken>
|
||
<normalizedToken originalValue="югу">yugu</normalizedToken>
|
||
<normalizedToken originalValue="от">ot</normalizedToken>
|
||
<normalizedToken originalValue="Ставрополя”">
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:69AABCC41207F67F512100E3D31EC11A" country="Russia" county="Stavropolya" municipality="Staryi" name="Stavropolya" stateProvince="Stavropol">Stavropolya</location>
|
||
"
|
||
</normalizedToken>
|
||
[Old forest south of
|
||
<collectingRegion country="Russia" name="Stavropol'">Stavropol</collectingRegion>
|
||
];
|
||
<collectingDate value="1954-05-25">25 May 1954</collectingDate>
|
||
;[S.]
|
||
<collectorName>Medvedev</collectorName>
|
||
leg.; ZISP. /
|
||
<normalizedToken originalValue="“ниж">"nizh</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="часть">chast</normalizedToken>
|
||
<normalizedToken originalValue="сев">sev</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="склона”">sklona"</normalizedToken>
|
||
[lower part of northern slope] •
|
||
<specimenCount type="female">1 female</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="“Георгиевское">"Georgievskoe</normalizedToken>
|
||
<normalizedToken originalValue="лесничество">lesnichestvo</normalizedToken>
|
||
<normalizedToken originalValue="Туапсинск">Tuapsinsk</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="р-н”">r-n"</normalizedToken>
|
||
[Georgievskoe forestry Tuapse District];
|
||
<collectingDate value="1954-05-21">21 May 1954</collectingDate>
|
||
;
|
||
<collectorName>K. Arnoldi</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
, same collection data as for preceding;
|
||
<collectingDate value="1954-05-22">22 May 1954</collectingDate>
|
||
•
|
||
<specimenCount type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount type="female">2 females</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="“г">"g</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Лысая">
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:5CFD92DB39F71A75ACE921CF6632EF3E" country="Russia" county="Stavropolya" municipality="Staryi" name="Lysaya" stateProvince="Stavropol">Lysaya</location>
|
||
</normalizedToken>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
|
||
800-900
|
||
<normalizedToken originalValue="м">m</normalizedToken>
|
||
</elevation>
|
||
</quantity>
|
||
,
|
||
<normalizedToken originalValue="Туапсинск">Tuapsinsk</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="р-н”">r-n"</normalizedToken>
|
||
[mount Lysaya, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
|
||
</quantity>
|
||
, Tuapse District];
|
||
<collectingDate value="1954-05-26">26 May 1954</collectingDate>
|
||
;
|
||
<collectorName>K. Arnoldi</collectorName>
|
||
leg.; ZISP; /
|
||
<normalizedToken originalValue="“вершинная">"vershinnaya</normalizedToken>
|
||
<normalizedToken originalValue="луговина">lugovina</normalizedToken>
|
||
<normalizedToken originalValue="и">i</normalizedToken>
|
||
<normalizedToken originalValue="опушка">opushka</normalizedToken>
|
||
<normalizedToken originalValue="леса">lesa</normalizedToken>
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
|
||
</quantity>
|
||
" [summit meadow and forest edge, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900 m</elevation>
|
||
</quantity>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="145">Additional material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<materialsCitation collectingDate="1956-05-18" collectingDateMax="2019-05-24" collectingDateMin="1956-05-18" collectorName="Gilyarov, E. N. Savchenko, Savchenko, V. Lantsov, S. Kustov, E. Hachikov., V. Gladun., V. Pilipenko" country="RUSSIA" county="Tarki-Tau Mt." elevation="135" latitude="42.941113" location="Russia" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." specimenCount="70" specimenCount-female="12" specimenCount-male="58" stateProvince="Krasnodar">
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:CA449E0163BBEACD15EEC23AC6060A0E" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Russia" stateProvince="Krasnodar">Russia</location>
|
||
•
|
||
<specimenCount type="male">9 males</specimenCount>
|
||
,
|
||
<specimenCount type="female">8 females</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="“курорт">"kurort</normalizedToken>
|
||
<normalizedToken originalValue="“Горячий">
|
||
"
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:9B26B8641EBB4CA343409FE6E24DA8AB" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Goryachii Klyuch" stateProvince="Krasnodar">
|
||
Goryachii
|
||
<normalizedToken originalValue="Ключ”">Klyuch"</normalizedToken>
|
||
</location>
|
||
</normalizedToken>
|
||
<normalizedToken originalValue="хр">khr</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Котх.[ский">Kotkh.[skii</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="Краснодар.[ский">
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
.[skii
|
||
</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="кр">kr</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="дуб.[овый">dub.[ovyi</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="лес”">les"</normalizedToken>
|
||
[Goryachiy Klyuch resort Koth[skiy] Ridge,
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
.[sky] District oak. Forest];
|
||
<collectingDate value="1956-05-18">18 May 1956</collectingDate>
|
||
;
|
||
<collectorName>Gilyarov</collectorName>
|
||
leg.; ZISP / "
|
||
<taxonomicName genus="T." lsidName="T." pageId="0" pageNumber="145" rank="genus">T.</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Edwards" authorityYear="1931" class="Insecta" family="Tipulidae" genus="Lunatipula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lunatipula" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Lunatipula</taxonomicName>
|
||
)
|
||
<taxonomicName authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula quadridentata" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="quadridentata">Tipula quadridentata</taxonomicName>
|
||
sp. n.
|
||
<normalizedToken originalValue="опр">opr</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Е.Н.Савченко”">
|
||
<collectorName>E.N.Savchenko</collectorName>
|
||
"
|
||
</normalizedToken>
|
||
•
|
||
<specimenCount type="female">1 female</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="“м">"m</normalizedToken>
|
||
. [
|
||
<normalizedToken originalValue="мыс">mys</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="Пенай">
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:EAEFD5FDD73B52323000CEC85A19FC5B" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Penai" stateProvince="Krasnodar">Penai</location>
|
||
</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="к">k</normalizedToken>
|
||
<normalizedToken originalValue="югу">yugu</normalizedToken>
|
||
<normalizedToken originalValue="от">ot</normalizedToken>
|
||
<normalizedToken originalValue="Новороссийска”">
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:0653E1A6B637B07249A2CDC775B554BB" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Novorossiiska" stateProvince="Krasnodar">Novorossiiska</location>
|
||
"
|
||
</normalizedToken>
|
||
[m. (cape) Penay, south of Novorossiysk] 24. V. [1]956;
|
||
<collectorName>Gilyarov</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount type="female">1 female</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="“окр">"okr</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="ст">st</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Смоленской">Smolenskoi</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Сиверского">Siverskogo</normalizedToken>
|
||
<normalizedToken originalValue="р-на">r-na</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Краснодар">
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="кр.”">kr."</normalizedToken>
|
||
[near
|
||
<collectingRegion country="Russia" name="Smolensk">Smolensk</collectingRegion>
|
||
station, Siversky Region,
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar District</collectingRegion>
|
||
]
|
||
<collectingDate value="1963-05-20">20 May 1963</collectingDate>
|
||
;
|
||
<collectorName>Savchenko</collectorName>
|
||
leg.; /
|
||
<normalizedToken originalValue="“опуш.[ка">"opush.[ka</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="смеш.[анного">smesh.[annogo</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="предгорн.[ого">predgorn.[ogo</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="леса”">lesa"</normalizedToken>
|
||
[edge of mixed foothill forest]; ZISP •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
;
|
||
<normalizedToken originalValue="“окр">"okr</normalizedToken>
|
||
.
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:94E8A82C5BE9F9397DEB6CBC2FDCFDDD" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="st. Krivenkovskoi" stateProvince="Krasnodar">
|
||
<normalizedToken originalValue="ст">st</normalizedToken>
|
||
.
|
||
<normalizedToken originalValue="Кривенковской">Krivenkovskoi</normalizedToken>
|
||
</location>
|
||
,
|
||
<normalizedToken originalValue="Туапс.[инского">Tuaps.[inskogo</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="р-на">r-na</normalizedToken>
|
||
<normalizedToken originalValue="Краснодар.[ского">
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
.[skogo
|
||
</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="края”">kraya"</normalizedToken>
|
||
[around village Krivenkovskaya, Tuapse Region of
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar District</collectingRegion>
|
||
];
|
||
<collectingDate value="1963-05-25">25 May 1963</collectingDate>
|
||
;
|
||
<collectorName>Savchenko</collectorName>
|
||
leg.; /
|
||
<normalizedToken originalValue="“опушка">"opushka</normalizedToken>
|
||
<normalizedToken originalValue="листв.[енного">listv.[ennogo</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="леса">lesa</normalizedToken>
|
||
<normalizedToken originalValue="у">u</normalizedToken>
|
||
<normalizedToken originalValue="реки">reki</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="вдоль">vdol</normalizedToken>
|
||
<normalizedToken originalValue="горн.[ого">gorn.[ogo</normalizedToken>
|
||
]
|
||
<normalizedToken originalValue="потока”">potoka"</normalizedToken>
|
||
[the edge of the foliage forest near the river, along the mountain stream]; ZISP •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:2DC346DB3EE5FACA545E9B50C12E0B9A" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Khosta" stateProvince="Krasnodar">Khosta</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:EF780B6AFF6E3966FB1FC0A23AAB6FF0" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Krasnodarskiy Kray" stateProvince="Krasnodar">Krasnodarskiy Kray</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:354DD67B20A9D79CDD6946A2C9F8F3B3" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Caucasian Reserve" stateProvince="Krasnodar">Caucasian Reserve</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:1DD93F305D9F517457C2C5290A8D6C57" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Tiso-samshitovaya" stateProvince="Krasnodar">Tiso-samshitovaya</location>
|
||
rosha [Yew-and-Boxwood Tree Grove];
|
||
<geoCoordinate degrees="43" direction="north" minutes="32" orientation="latitude" precision="15" seconds="014" value="43.53722">43°32'014"N</geoCoordinate>
|
||
, 39°52'621"E; alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.35" unit="m" value="135.0">135 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2018-05-10">10 May 2018</collectingDate>
|
||
;
|
||
<collectorName>V. Lantsov</collectorName>
|
||
leg.; / Fern-butcher community on the rocky slopes of the right side of the gorge of the Khosta River; IEMT •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
; Khosta, Krasnodarskiy Kray, Caucasian Reserve, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43°31'656"N, 39°52'467
|
||
<normalizedToken originalValue="’’">''</normalizedToken>
|
||
E; alt.
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">54 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2018-05-12">12 May 2018</collectingDate>
|
||
,
|
||
<collectorName>V. Lantsov</collectorName>
|
||
leg.; / collected on light; IEMT •
|
||
<specimenCount type="male">2 males</specimenCount>
|
||
(in alcohol); North Caucasus, Krasnodarskiy Kray, near village Mezmay; 44°11'291"N, 39°58'090"E; alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.08" unit="m" value="808.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.08" unit="m" value="808.0">808 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2018-05-15">15 May 2018</collectingDate>
|
||
;
|
||
<collectorName>V. Lantsov</collectorName>
|
||
leg.; / Beech (
|
||
<taxonomicName authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
|
||
</taxonomicName>
|
||
), fruit tree (
|
||
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Pyrus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pyrus" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Pyrus</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Malus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Malus" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Malus</emphasis>
|
||
</taxonomicName>
|
||
sp.) forest with
|
||
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
|
||
</taxonomicName>
|
||
in under growth,
|
||
<taxonomicName class="Dicotyledoneae" family="Caprifoliaceae" genus="Sambucus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sambucus nigra" order="Dipsacales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="nigra">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Sambucus nigra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Dicotyledoneae" family="Cornaceae" genus="Cornus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cornus mas" order="Umbelliflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="mas">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Cornus mas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Rosa canina</emphasis>
|
||
,
|
||
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Crataegus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Crataegus" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Crataegus</emphasis>
|
||
</taxonomicName>
|
||
sp., in shrub layer and sedge
|
||
<taxonomicName class="Monocotyledoneae" family="Cyperaceae" genus="Carex" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carex pendula" order="Cyperales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="pendula">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Carex pendula</emphasis>
|
||
</taxonomicName>
|
||
, cereals and herbs in ground layer [a leveled area of the light part of the forest, possibly a site of a wild pear-apple orchard 50-70 years old]; IEMT •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
;
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory, Apsheron District, environs of village Mezmay, Guam Gorge, left slope, 44°12'409"N,
|
||
<geoCoordinate degrees="39" direction="east" minutes="55" orientation="longitude" precision="15" seconds="056" value="39.932224">39°55'056"E</geoCoordinate>
|
||
, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.47" unit="m" value="547.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.47" unit="m" value="547.0">547 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2019-05-24">24 May 2019</collectingDate>
|
||
;
|
||
<collectorName>V. Lantsov</collectorName>
|
||
leg.; / Landslide foot, community with butterbur [
|
||
<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Petasites" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Petasites albus" order="Campanulales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="albus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Petasites albus</emphasis>
|
||
</taxonomicName>
|
||
] as dominant along the banks of the stream, open places at the edge of the forest; ZISP •
|
||
<specimenCount type="male">8 males</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory, Seversky District, in vicinity of village Thamaha,
|
||
<collectingDate value="2016-05-03">3 May 2016</collectingDate>
|
||
;
|
||
<collectorName>S. Kustov</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">7 males</specimenCount>
|
||
,
|
||
<specimenCount type="female">1 female</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory, Seversky District, environs of village Plancheskaya,
|
||
<collectingDate value="2016-05-03">3 May 2016</collectingDate>
|
||
.
|
||
<collectorName>S. Kustov</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">5 males</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory, environs of village Bolshoy Utrish;
|
||
<collectingDate value="2008-05-01">1 May 2008</collectingDate>
|
||
;
|
||
<collectorName>E. Hachikov.</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">3 males</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory, municipality Anapa, environs of village Sukko,
|
||
<normalizedToken originalValue="Kvashin’s">Kvashin's</normalizedToken>
|
||
Gorge, Dolgaya Niva territory;
|
||
<geoCoordinate degrees="44" direction="north" minutes="47" orientation="latitude" precision="15" seconds="20" value="44.788887">44°47'20"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="37" direction="east" minutes="28" orientation="longitude" precision="15" seconds="33" value="37.475834">37°28'33"E</geoCoordinate>
|
||
; alt.
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.7" unit="m" value="67.0">
|
||
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.7" unit="m" value="67.0">67 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2016-05-06" valueMax="2016-05-08" valueMin="2016-05-06">6-8 May 2016</collectingDate>
|
||
;
|
||
<collectorName>S. Kustov</collectorName>
|
||
,
|
||
<collectorName>V. Gladun.</collectorName>
|
||
leg.; ZISP •
|
||
<specimenCount type="male">5 males</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory,
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:39544B84DC4425ABA481E23A571574AA" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="river Shakhe Gorge" stateProvince="Krasnodar">river Shakhe Gorge</location>
|
||
;
|
||
<geoCoordinate degrees="43" direction="north" minutes="52" orientation="latitude" precision="15" seconds="46" value="43.879444">43°52'46"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="39" direction="east" minutes="50" orientation="longitude" precision="15" seconds="00" value="39.833332">39°50'00"E</geoCoordinate>
|
||
;
|
||
<collectingDate value="2012-05-02">2 May 2012</collectingDate>
|
||
;
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; VPMC •
|
||
<specimenCount type="male">3 males</specimenCount>
|
||
; (in alcohol); Khosta, Krasnodarskiy Kray,
|
||
<location LSID="urn:lsid:plazi:treatment:E104034BF2F251DB8C145337C9D914B4:605E8E70D4F8E79C295D8DEF669501B6" country="RUSSIA" county="Tarki-Tau Mt." latitude="42.941113" longLatPrecision="19" longitude="47.46889" municipality="Tarki Distr." name="Caucasian Reserve" stateProvince="Krasnodar">Caucasian Reserve</location>
|
||
, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove];
|
||
<geoCoordinate degrees="43" direction="north" minutes="32" orientation="latitude" precision="15" seconds="014" value="43.53722">43°32'014"N</geoCoordinate>
|
||
, 39°52'621"E;
|
||
<collectingDate value="2012-05-08">8 May 2012</collectingDate>
|
||
;
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; VPMC •
|
||
<specimenCount type="male">4 males</specimenCount>
|
||
(in alcohol);
|
||
<collectingRegion country="Russia" name="Krasnodar">Krasnodar</collectingRegion>
|
||
Territory,
|
||
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
|
||
to the N from Sochi, Sukhoy Canyon;
|
||
<geoCoordinate degrees="43" direction="north" minutes="32" orientation="latitude" precision="925" value="43.533333">43°32'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="39" direction="east" minutes="56" orientation="longitude" precision="925" value="39.933334">39°56'E</geoCoordinate>
|
||
;
|
||
<collectingDate value="2014-05-05">5 May 2014</collectingDate>
|
||
;
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; VPMC;
|
||
<collectingCountry name="Russia">RUSSIA</collectingCountry>
|
||
- Dagestan •
|
||
<specimenCount type="male">5 males</specimenCount>
|
||
; Makhachkala,
|
||
<collectingMunicipality>Tarki Distr.</collectingMunicipality>
|
||
;
|
||
<geoCoordinate degrees="42" direction="north" minutes="56" orientation="latitude" precision="15" seconds="57" value="42.94917">42°56'57"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="47" direction="east" minutes="29" orientation="longitude" precision="15" seconds="41" value="47.494724">47°29'41"E</geoCoordinate>
|
||
; alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2019-05-01">1 May 2019</collectingDate>
|
||
.,
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; cemetery on the hillside; VPMC •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount type="female">1 female</specimenCount>
|
||
; same locality;
|
||
<collectingDate value="2019-05-10">10 May 2019</collectingDate>
|
||
.,
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; VPMC •
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
,
|
||
<collectingCounty>Tarki-Tau Mt.</collectingCounty>
|
||
;
|
||
<geoCoordinate degrees="42" direction="north" minutes="56" orientation="latitude" precision="15" seconds="28" value="42.941113">42°56'28"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="47" direction="east" minutes="28" orientation="longitude" precision="15" seconds="08" value="47.46889">47°28'08"E</geoCoordinate>
|
||
; alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2019-05-02">2 May 2019</collectingDate>
|
||
;
|
||
<collectorName>V. Pilipenko</collectorName>
|
||
leg.; oak forest; VPMC
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Tergite 9 with four widely spaced teeth and with three rounded notches distally; caudal margin of gonocoxite with two dentate projections. Paired appendages of sternite 8 with wide base bearing thick yellow setae distally covering gap between. Cercus long and straight; hypogynial valve only slightly longer than width at base.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="redescription">
|
||
<paragraph pageId="0" pageNumber="145">Redescription.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Adult male (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8A, B</figureCitation>
|
||
). General body coloration grey. Body length 15-16 mm, wings 15.5-17 mm.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 8.</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata" subGenus="Lunatipula">T. (Lunatipula) quadridentata</taxonomicName>
|
||
, male habitus
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
general lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
head and thorax, dorsal view. Scale bars: 10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
), 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Head</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13D</figureCitation>
|
||
). Vertex dirty grey with longitudinal thin dark grey midline. Rostrum grey dorsally with long brown setae mixed with shorter white setae, but glabrous at base, laterally and ventrally dirty yellow (nearly rusty). Nasus well developed, with long, whitish procumbent setae. Frons light grey and yellow with group of 2-4 small brown setae behind each antenna. Gula grey, glabrous with narrow brown stripe. Gena light grey near eyes.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Antennae</emphasis>
|
||
.
|
||
</emphasis>
|
||
Framing around antennal sockets yellow. Scape light brown, pedicel yellow, flagellomeres brown; verticils not longer than corresponding flagellomeres.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8A, B</figureCitation>
|
||
). Dark grey with silvery pruinescence. Pronotum dark grey; katepisternum dorsally with groups of whitish setae. Scutum with 4 brownish stripes (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8B</figureCitation>
|
||
), spaces between stripes grey with long whitish bristles. Mediotergite grey with whitish bristles.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Wings</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8A</figureCitation>
|
||
). Transparent, without noticeable marble pattern, with light brown pterostigma. Longitudinal veins with macrotrichia.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<taxonomicName genus="Halteres" lsidName="Halteres" pageId="0" pageNumber="145" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Halteres</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Stem light brown to yellowish, knob brown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Femora light brown with procumbent dark brown bristles. Tarsal claws without noticeable tooth at base.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Abdomen</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. T. (Lunatipula) quadridentata, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure8" httpUri="https://binary.pensoft.net/fig/566417" pageId="0" pageNumber="145">8A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9A</figureCitation>
|
||
). Dark grey with short whitish bristles. Tergites with distal and lateral whitish edging.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 9.</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata" subGenus="Lunatipula">T. (Lunatipula) quadridentata</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
terminal segments of male abdomen, lateral view (dry)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B-L</emphasis>
|
||
male terminalia (after KOH 10% treatment)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
hypopygium, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
hypopygium, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
sternite 8, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
tergite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
tergite 9, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
inner and outer gonostylus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
|
||
inner and outer gonostylus, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
|
||
gonocoxal fragment, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
|
||
semen pump and aedeagus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Terminalia</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14D, J</figureCitation>
|
||
). Hypopygium not thickened. Tergite 9 and gonocoxite with silver pruinescence. Upper margin of gonocoxite with two dentate projections. Tergite 9 (Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9E, F</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14D</figureCitation>
|
||
) at apex with four widely spaced teeth and with three rounded notches, narrower middle notch bounded by small flat projection bearing sharp thorn beneath; small backwards projecting spine beneath tergite 9. Outer and inner gonostyli (Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9G, H</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14J</figureCitation>
|
||
) without significant features. Paired appendages of sternite 8 (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9D</figureCitation>
|
||
) large, wide base with fringe of thick yellow setae hiding gap between (apparent on pinned specimens). Paired apical appendages of sternite 9 as in Figure
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9K, L</figureCitation>
|
||
. Gonocoxal fragment (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9I</figureCitation>
|
||
) with characteristic small V-shaped fork at base. Semen pump and aedeagus as in Figure
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9J</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||
Body length 17.5-19.5 mm, wings 16-18.5 mm. Scape and pedicel brownish yellow. Terminalia (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12D-F</figureCitation>
|
||
). Cercus long and straight. Hypogynial valve (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12E</figureCitation>
|
||
) only slightly longer than wide at base. Sternite 9 and furca as in Figure
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12D, F</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="145">Comparison with closely related species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This species differs from other species of the
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
species group (in the male) by tergite 9 with four widely spaced teeth and with three small notches at the apex and a small spine beneath (Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9E, F</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14D</figureCitation>
|
||
). Paired appendages of sternite 8 is broadest, with thick yellow bristles that cover the gap between; in the other species this gap is not obscured.
|
||
<taxonomicName authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula quadridentata" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="quadridentata">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula quadridentata</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName lsidName="T. paupera" pageId="0" pageNumber="145" rank="species" species="paupera">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. paupera</emphasis>
|
||
</taxonomicName>
|
||
by having two teeth on the edge of the gonocoxite and similar coloring of the antenna. The outer gonostylus is somewhat larger than in
|
||
<taxonomicName lsidName="T. caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. caucasica</emphasis>
|
||
</taxonomicName>
|
||
. The gonocoxal fragment is similar to that of
|
||
<taxonomicName lsidName="T. talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. talyshensis</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. T. (Lunatipula) quadridentata A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure9" httpUri="https://binary.pensoft.net/fig/566418" pageId="0" pageNumber="145">9I</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11I</figureCitation>
|
||
). The female has a parallel-sided, straight cercus whereas it is upturned at the tip in
|
||
<taxonomicName lsidName="T. caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. caucasica</emphasis>
|
||
</taxonomicName>
|
||
and broader at base in
|
||
<taxonomicName lsidName="T. talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. talyshensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="elevation">
|
||
<paragraph pageId="0" pageNumber="145">Elevation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults were collected at altitudes ranging from sea level (54 m in Tiso-samshitovaya rosha) to 800-900 m (in Mount Lysaya, Tuapse district).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="flight period">
|
||
<paragraph pageId="0" pageNumber="145">Flight period.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults were collected from throughout the month of May.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="habitat">
|
||
<paragraph pageId="0" pageNumber="145">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Specimens are found in moderately humid woody deciduous communities.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Endemic to the Caucasus; currently known from the West Caucasus (northern and southern slopes; Krasnodar and Stavropol Territory) and from the East Caucasus (Dagestan; first record).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |