206 lines
35 KiB
XML
206 lines
35 KiB
XML
<document id="8378407692A4F62EC39981FFA9CA0ABC" ID-CLB-Dataset="7717" ID-DOI="10.5252/g2011n2a1" ID-GBIF-Dataset="601437bd-40a5-40dc-8fda-72b0aa119041" ID-ISSN="1638-9395" ID-Zenodo-Dep="4597045" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615470577137" checkinUser="felipe" docAuthor="Derycke, Claire & Goujet, Daniel" docDate="2011" docId="CA2B87AB6C41FFA9F8B6E909918EC99E" docLanguage="en" docName="Geodiversitas.33.2.209-226.pdf" docOrigin="Geodiversitas 33 (2)" docSource="http://dx.doi.org/10.5252/g2011n2a1" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Milesacanthus ancestralis Burrow, Lelievre & Janjou 2006" docType="treatment" docVersion="8" lastPageNumber="222" masterDocId="3612FFD36C4CFFA4F855EB1D950DCA55" masterDocTitle="Multicuspidate shark teeth associated with chondrichthyan and acanthodian scales from the Emsian (Devonian) of southern Algeria" masterLastPageNumber="226" masterPageNumber="209" pageNumber="222" updateTime="1698933075774" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="709CEEB849D63318AD88D572E8786EAA">Multicuspidate shark teeth associated with chondrichthyan and acanthodian scales from the Emsian (Devonian) of southern Algeria</mods:title>
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<mods:name id="4446C737ACB3ECB9E49B7B4A1B09C83B" type="personal">
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<mods:namePart id="D819872DA8F25B6879EC99D1A2CF6BC0">Derycke, Claire</mods:namePart>
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<mods:namePart id="BC0BCDDB4E1CAAE086C59996D13A00B5">Goujet, Daniel</mods:namePart>
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<mods:typeOfResource id="BECB8407594BB0B0D9A2302D67658C08">text</mods:typeOfResource>
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<mods:title id="304F22942F9C7C2884362AB14596C6E8">Geodiversitas</mods:title>
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<mods:part id="D3C951E3EF9FA8800F3393D76833842D">
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<mods:date id="F20A30A33830901F0DAD0E5ED8082DB2">2011</mods:date>
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<mods:number id="6F8F30EECE8E59C1D1BE2D9ECE312F03">2011-06-30</mods:number>
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<mods:number id="B334541FAB3EF8D639864640DE604E0E">33</mods:number>
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<treatment id="CA2B87AB6C41FFA9F8B6E909918EC99E" ID-DOI="http://doi.org/10.5281/zenodo.4596946" ID-GBIF-Taxon="180584331" ID-Zenodo-Dep="4596946" LSID="urn:lsid:plazi:treatment:CA2B87AB6C41FFA9F8B6E909918EC99E" httpUri="http://treatment.plazi.org/id/CA2B87AB6C41FFA9F8B6E909918EC99E" lastPageNumber="222" pageId="13" pageNumber="222">
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<subSubSection id="0A9865366C41FFA9F8B6E90994A9C81A" pageId="13" pageNumber="222" type="nomenclature">
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<paragraph id="423D36BD6C41FFA9F8B6E90994A9C81A" blockId="13.[227,533,532,591]" pageId="13" pageNumber="222">
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<heading id="197581D16C41FFA9F8B6E90994A9C81A" centered="true" fontSize="11" level="2" pageId="13" pageNumber="222" reason="2">
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<taxonomicName id="85824D3E6C41FFA9F8B6E9099718C87A" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" box="[227,533,532,560]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
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<emphasis id="70F6EAAF6C41FFA9F8B6E9099470C87A" bold="true" box="[227,381,532,559]" italics="true" pageId="13" pageNumber="222">Milesacanthus</emphasis>
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cf.
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<emphasis id="70F6EAAF6C41FFA9F9F0E9099718C87A" bold="true" box="[421,533,532,559]" italics="true" pageId="13" pageNumber="222">ancestralis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="DAB92A386C41FFA9F909E9299496C81A" box="[348,411,564,591]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9865366C41FFA9F824E9739436C8EF" pageId="13" pageNumber="222" type="materials_examined">
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<paragraph id="423D36BD6C41FFA9F824E9739436C8EF" blockId="13.[113,645,621,698]" pageId="13" pageNumber="222">
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<collectionCode id="2493AE786C41FFA9F824E97395D1C8D1" box="[113,220,622,645]" pageId="13" pageNumber="222">MATERIAL</collectionCode>
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. — Nine scales, one of which is lost (M N H N. F.A L D -4 0), a n d o n e t h i n s e c t i o n
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<collectingRegion id="8046F85F6C41FFA9F824E9BF95CBC8EF" box="[113,198,674,698]" country="Ireland" name="Monaghan" pageId="13" pageNumber="222">MNHN</collectingRegion>
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.F.ALD-41.
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9865366C41FFA9F827E9CE96DCC845" pageId="13" pageNumber="222" type="description">
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<paragraph id="423D36BD6C41FFA9F827E9CE940FC8BE" blockId="13.[111,647,723,1700]" box="[114,258,723,749]" pageId="13" pageNumber="222">DESCRIPTION</paragraph>
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<paragraph id="423D36BD6C41FFA9F824E9EF94FDC9D9" blockId="13.[111,647,723,1700]" pageId="13" pageNumber="222">
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The crown of these four scales is diamond-shaped and its anterior part may be more rounded as in
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<collectingRegion id="8046F85F6C41FFA9F826E82F95C7C918" box="[115,202,818,845]" country="Ireland" name="Monaghan" pageId="13" pageNumber="222">MNHN</collectingRegion>
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.F.ALD-38 (
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<figureCitation id="DAB92A386C41FFA9F91DE82F9499C918" box="[328,404,818,845]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8C</figureCitation>
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<figureCitation id="DAB92A386C41FFA9F9C6E8229491C905" box="[403,412,831,848]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F9C6E8229491C905" attach="left" box="[403,412,831,848]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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). The convex base, also diamond-shaped, and with anterior and rounded posterior edges is located anteriorly.
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</paragraph>
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<paragraph id="423D36BD6C41FFA9F8DDE88C94BBCF5D" blockId="13.[111,647,723,1700]" pageId="13" pageNumber="222">
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The scales with a flat crown show sub-parallel grooves (18 for
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<figureCitation id="DAB92A386C41FFA9F977E8AD947CC99E" box="[290,369,944,971]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8A</figureCitation>
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<figureCitation id="DAB92A386C41FFA9F924E8A09476C99B" box="[369,379,957,974]" captionStart="FIG" captionStartId="2.[113,124,1635,1652]" captionTargetBox="[187,1139,214,1498]" captionTargetId="figure-19@2.[146,1177,183,1520]" captionTargetPageId="2" captionText="FIG. 1. — Geological map of North-West Africa after Fabre (1976:13). Numbers correspond to localities:1, Bechar; 2, Béni Abbès; 3, In Salah (Gour Bedda); 4, Reganne; 5, Ahnet; 6, Mouydir; 7, Illizi (Fort Polignac); 8, Ajjers; 9, Tin Seririne Syncline (Anou In Ateï-Taberia); 10, Oued Felaou; 11, In Debirène." figureDoi="http://doi.org/10.5281/zenodo.4597047" httpUri="https://zenodo.org/record/4597047/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F924E8A09476C99B" attach="left" box="[369,379,957,974]" fontSize="7" pageId="13" pageNumber="222">1</subScript>
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</figureCitation>
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, B
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<figureCitation id="DAB92A386C41FFA9F9C9E8A094ABC99B" box="[412,422,957,974]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F9C9E8A094ABC99B" attach="left" box="[412,422,957,974]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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</figureCitation>
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and possibly more), diverging in the front part (
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<figureCitation id="DAB92A386C41FFA9F9F6E8CD94F9C9BE" box="[419,500,976,1003]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8C</figureCitation>
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<figureCitation id="DAB92A386C41FFA9F9A0E8C094F2C9BB" box="[501,511,989,1006]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F9A0E8C094F2C9BB" attach="right" box="[501,511,989,1006]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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</figureCitation>
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), like in the material of
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<taxonomicName id="85824D3E6C41FFA9F8A1E8ED94F2CE5F" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" box="[244,511,1008,1034]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
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<emphasis id="70F6EAAF6C41FFA9F8A1E8ED94F2CE5F" box="[244,511,1008,1034]" italics="true" pageId="13" pageNumber="222">Milesacanthus ancestralis</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="3A95762D6C41FFA9FA1DE8ED95B6CE7E" name="Saudi Arabia" pageId="13" pageNumber="222">Saudi Arabia</collectingCountry>
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.The grooves are not really deep but narrower than the ridges. Both grooves and ridges run along the entire crown length of the scale. In the scale MNHN.F.ALD-36 (
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<figureCitation id="DAB92A386C41FFA9F907EF7294ADCEDF" box="[338,416,1135,1162]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8A</figureCitation>
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<figureCitation id="DAB92A386C41FFA9F9F5EF6694A7CED9" box="[416,426,1147,1164]" captionStart="FIG" captionStartId="2.[113,124,1635,1652]" captionTargetBox="[187,1139,214,1498]" captionTargetId="figure-19@2.[146,1177,183,1520]" captionTargetPageId="2" captionText="FIG. 1. — Geological map of North-West Africa after Fabre (1976:13). Numbers correspond to localities:1, Bechar; 2, Béni Abbès; 3, In Salah (Gour Bedda); 4, Reganne; 5, Ahnet; 6, Mouydir; 7, Illizi (Fort Polignac); 8, Ajjers; 9, Tin Seririne Syncline (Anou In Ateï-Taberia); 10, Oued Felaou; 11, In Debirène." figureDoi="http://doi.org/10.5281/zenodo.4597047" httpUri="https://zenodo.org/record/4597047/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F9F5EF6694A7CED9" attach="left" box="[416,426,1147,1164]" fontSize="7" pageId="13" pageNumber="222">1</subScript>
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</figureCitation>
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), grooves and ridges start just at the front edge of the scale, whereas in the scale MNHN.F.ALD-37 (
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<figureCitation id="DAB92A386C41FFA9F9E6EFB3970ECE9C" box="[435,515,1198,1225]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8B</figureCitation>
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<figureCitation id="DAB92A386C41FFA9FA56EFA69700CE99" box="[515,525,1211,1228]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9FA56EFA69700CE99" attach="left" box="[515,525,1211,1228]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
|
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</figureCitation>
|
|
) the ridges start at the vertical edge of the crown and the posterior edge is more crenulated.
|
|
</paragraph>
|
|
<paragraph id="423D36BD6C41FFA9F8DDEE1094F4CCF1" blockId="13.[111,647,723,1700]" pageId="13" pageNumber="222">
|
|
One scale (ALD-38) shows 13 larger grooves (
|
|
<figureCitation id="DAB92A386C41FFA9F82EEE3095C7CF1D" box="[123,202,1325,1352]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8C</figureCitation>
|
|
<figureCitation id="DAB92A386C41FFA9F89FEE2795D9CF1E" box="[202,212,1338,1355]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F89FEE2795D9CF1E" attach="left" box="[202,212,1338,1355]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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</figureCitation>
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), but only in its anterior part. Pore openings, located mainly at the beginning of the grooves, connect to an ascending canal system. The grooves begin after the anterior edge of the crown and most lateral grooves undulate, slightly converging in the beginning, then parallel to the diagonal, and stop in the posterior third of the crown. The neck is well marked with a row of foramina posteriorly and anteriorly (
|
|
<figureCitation id="DAB92A386C41FFA9F8E0ED37940DCC10" box="[181,256,1578,1605]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8A</figureCitation>
|
|
<figureCitation id="DAB92A386C41FFA9F955ED2A945BCC1D" box="[256,342,1578,1608]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
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<subScript id="DE0634F86C41FFA9F955ED2A9404CC1D" attach="left" box="[256,265,1591,1608]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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, B
|
|
<subScript id="DE0634F86C41FFA9F973ED2A9422CC1D" attach="right" box="[294,303,1591,1608]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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, C
|
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<subScript id="DE0634F86C41FFA9F918ED2A945BCC1D" attach="left" box="[333,342,1591,1608]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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</figureCitation>
|
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). Furthermore, wart-like protuberances are visible on the posterior neck (
|
|
<figureCitation id="DAB92A386C41FFA9FA7EED57977BCC30" box="[555,630,1610,1637]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8A</figureCitation>
|
|
<figureCitation id="DAB92A386C41FFA9FA23ED4A9772CC3D" box="[630,639,1623,1640]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
|
|
<subScript id="DE0634F86C41FFA9FA23ED4A9772CC3D" attach="left" box="[630,639,1623,1640]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
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</figureCitation>
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; more evident on
|
|
<figureCitation id="DAB92A386C41FFA9F971ED74947ECCD1" box="[292,371,1641,1668]" captionStart="FIG" captionStartId="12.[113,124,1498,1515]" captionTargetBox="[168,1156,214,1437]" captionTargetId="figure-19@12.[160,1163,207,1454]" captionTargetPageId="12" captionText="FIG. 7. — Histology and corresponding morphology of chondrichthyan scales from the Emsian (Devonian) of southern Algeria: A-E, H, Tassiliodus lessardi n. gen., n. sp.; F, G, Milesacanthus cf. ancestralis; identical letters refer to the same scale in morphology and in thin section (except for D, corresponding to an internal view of a broken scale photographed by SEM, and for F, which was lost in the SEM); A, crown view and sagittal thin section 1 of a scale with converging ridges posteriorly and high-neck (MNHN.F.ALD-30); B, thin section 2 and crown view of a dumpy scale with radiating ridges only fringing the crown (MNHN.F.ALD-31); C, lateral view of a high-neck scale and thin section 3 (MNHN.F.ALD-32); D, SEM micrograph of internal view of a high-necked scale (MNHN.F.ALD-33); E, crown view of a dumpy scale with radiating ridges reaching the middle of the scale, and thin section 4 (MNHN.F.ALD-34); H, lateral view of a scale with converging ridges in the middle of the scale and a high neck, and thin section 5 (MNHN.F.ALD-35); F, posterolateral view (MNHN.F.ALD-40); G, thin section 9 (MNHN.F.ALD-41). Scale bars: A-E, H, 500 μm; F, 300 μm; G, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4597061" httpUri="https://zenodo.org/record/4597061/files/figure.png" pageId="13" pageNumber="222">Fig. 7F</figureCitation>
|
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), visible on “young” scales according
|
|
<bibRefCitation id="26134B4C6C41FFA9F8B5ED9494F8CCF1" author="BURROW C. J. & LELIEVRE H. & JANJOU D." box="[224,501,1673,1700]" pageId="13" pageNumber="222" pagination="537 - 560" refId="ref9231" refString="BURROW C. J., LELIEVRE H. & JANJOU D. 2006. - Gnathostome microremains from the Lower Devonian Jawf Formation, Saudi Arabia. Journal of Paleontology 80 (3): 537 - 560." type="journal article" year="2006">
|
|
Burrow
|
|
<emphasis id="70F6EAAF6C41FFA9F962ED979466CCF6" box="[311,363,1673,1699]" italics="true" pageId="13" pageNumber="222">et al.</emphasis>
|
|
(2006: 547)
|
|
</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="423D36BD6C41FFA9FAF0EBC5962FCAA5" blockId="13.[676,1212,216,528]" box="[677,802,216,242]" pageId="13" pageNumber="222">HISTOLOGY</paragraph>
|
|
<paragraph id="423D36BD6C41FFA9FAF0EBE596DCC845" blockId="13.[676,1212,216,528]" pageId="13" pageNumber="222">
|
|
The thin section 9
|
|
<collectingRegion id="8046F85F6C41FFA9FB2DEBE596DECB46" box="[888,979,248,275]" country="Ireland" name="Monaghan" pageId="13" pageNumber="222">MNHN</collectingRegion>
|
|
.F.ALD-41 (
|
|
<figureCitation id="DAB92A386C41FFA9FC0FEBE591BCCB46" box="[1114,1201,248,275]" captionStart="FIG" captionStartId="12.[113,124,1498,1515]" captionTargetBox="[168,1156,214,1437]" captionTargetId="figure-19@12.[160,1163,207,1454]" captionTargetPageId="12" captionText="FIG. 7. — Histology and corresponding morphology of chondrichthyan scales from the Emsian (Devonian) of southern Algeria: A-E, H, Tassiliodus lessardi n. gen., n. sp.; F, G, Milesacanthus cf. ancestralis; identical letters refer to the same scale in morphology and in thin section (except for D, corresponding to an internal view of a broken scale photographed by SEM, and for F, which was lost in the SEM); A, crown view and sagittal thin section 1 of a scale with converging ridges posteriorly and high-neck (MNHN.F.ALD-30); B, thin section 2 and crown view of a dumpy scale with radiating ridges only fringing the crown (MNHN.F.ALD-31); C, lateral view of a high-neck scale and thin section 3 (MNHN.F.ALD-32); D, SEM micrograph of internal view of a high-necked scale (MNHN.F.ALD-33); E, crown view of a dumpy scale with radiating ridges reaching the middle of the scale, and thin section 4 (MNHN.F.ALD-34); H, lateral view of a scale with converging ridges in the middle of the scale and a high neck, and thin section 5 (MNHN.F.ALD-35); F, posterolateral view (MNHN.F.ALD-40); G, thin section 9 (MNHN.F.ALD-41). Scale bars: A-E, H, 500 μm; F, 300 μm; G, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4597061" httpUri="https://zenodo.org/record/4597061/files/figure.png" pageId="13" pageNumber="222">Fig. 7G</figureCitation>
|
|
) shows a classical pattern for an acanthodian, with a primordium in the middle of the crown that is made of dentine covered with thin box-in-box growth zones continuing from the crown into the acellular bone base. Contrary to
|
|
<taxonomicName id="85824D3E6C41FFA9FC75EA8B961CCB9A" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
|
|
<emphasis id="70F6EAAF6C41FFA9FC75EA8B961CCB9A" italics="true" pageId="13" pageNumber="222">Milesacanthus ancestralis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="85824D3E6C41FFA9FB18EAAB97E7CBA5" authority="(Young & Burrow 2004)" baseAuthorityName="Young & Burrow" baseAuthorityYear="2004" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="antarctica">
|
|
<emphasis id="70F6EAAF6C41FFA9FB18EAAB96EFCB9A" box="[845,994,438,463]" italics="true" pageId="13" pageNumber="222">M. antarctica</emphasis>
|
|
(
|
|
<bibRefCitation id="26134B4C6C41FFA9FBA6EAAB97EECBA5" author="YOUNG G. C. & BURROW C. J." pageId="13" pageNumber="222" pagination="23 - 43" refId="ref11020" refString="YOUNG G. C. & BURROW C. J. 2004. - Diplacanthid acanthodians from the Aztec Siltstone (late Middle Devonian) of southern Victoria Land, Antarctica. Fossils and Strata 50: 23 - 43." type="journal article" year="2004">Young & Burrow 2004</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, no vascular canal was found, but only one thin section has been made.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0A9865366C41FFA9FAF0E928918EC99E" pageId="13" pageNumber="222" type="discussion">
|
|
<paragraph id="423D36BD6C41FFA9FAF0E928962BC818" blockId="13.[675,1212,565,971]" box="[677,806,565,591]" pageId="13" pageNumber="222">DISCUSSION</paragraph>
|
|
<paragraph id="423D36BD6C41FFA9FAF0E949918EC99E" blockId="13.[675,1212,565,971]" pageId="13" pageNumber="222">
|
|
The number and the development of ridges on the crown recalls the scales of
|
|
<taxonomicName id="85824D3E6C41FFA9FB9CE9699154C8D8" baseAuthorityName="Young & Burrow" baseAuthorityYear="2004" box="[969,1113,628,653]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="antarctica">
|
|
<emphasis id="70F6EAAF6C41FFA9FB9CE9699154C8D8" box="[969,1113,628,653]" italics="true" pageId="13" pageNumber="222">M. antarctica</emphasis>
|
|
</taxonomicName>
|
|
, but the presence of wart-like bumps in the posterior part of the neck is more suggestive of
|
|
<taxonomicName id="85824D3E6C41FFA9FC4BE9A991BFC898" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" box="[1054,1202,691,717]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
|
|
<emphasis id="70F6EAAF6C41FFA9FC4BE9A991BFC898" box="[1054,1202,691,717]" italics="true" pageId="13" pageNumber="222">M. ancestralis</emphasis>
|
|
</taxonomicName>
|
|
. Although wide canals were not detected here, they are present in both
|
|
<taxonomicName id="85824D3E6C41FFA9FB39E9EE96FBC959" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" box="[876,1014,754,780]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
|
|
<emphasis id="70F6EAAF6C41FFA9FB39E9EE96FBC959" box="[876,1014,754,780]" italics="true" pageId="13" pageNumber="222">M. ancestralis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="85824D3E6C41FFA9FC7CE9EE91BEC959" baseAuthorityName="Young & Burrow" baseAuthorityYear="2004" box="[1065,1203,755,780]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="antarctica">
|
|
<emphasis id="70F6EAAF6C41FFA9FC7CE9EE91BEC959" box="[1065,1203,755,780]" italics="true" pageId="13" pageNumber="222">M. antarctica</emphasis>
|
|
</taxonomicName>
|
|
, and pore openings are visible at the surface of the scale
|
|
<collectingRegion id="8046F85F6C41FFA9FA8AE82F9634C918" box="[735,825,818,845]" country="Ireland" name="Monaghan" pageId="13" pageNumber="222">MNHN</collectingRegion>
|
|
.F.ALD-38 (
|
|
<figureCitation id="DAB92A386C41FFA9FB95E82F9119C918" box="[960,1044,818,845]" captionStart="FIG" captionStartId="14.[113,124,963,980]" captionTargetBox="[113,1196,229,891]" captionTargetId="figure-357@14.[387,662,392,569]" captionTargetPageId="14" captionText="FIG. 8. — Acanthodian scales of Milesacanthus cf. ancestralis from the Emsian (Devonian) of southern Algeria: A, MNHN.F.ALD-36; A1, crown view; A2, lateral view; B, MNHN.F.ALD-37;B1, basal view; B2, front view; C, MNHN.F.ALD-38; C1, basal view; C2, front view; D, MNHN.F.ALD-39; D1, crown view; D2, basal view. Scale bars: A-B, C2, 400 μm; C1, D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4597063" httpUri="https://zenodo.org/record/4597063/files/figure.png" pageId="13" pageNumber="222">Fig. 8C</figureCitation>
|
|
<figureCitation id="DAB92A386C41FFA9FC41E8229113C905" box="[1044,1054,831,848]" captionStart="FIG" captionStartId="3.[113,124,1332,1349]" captionTargetBox="[136,1187,212,1291]" captionTargetId="figure-136@3.[126,1197,205,1307]" captionTargetPageId="3" captionText="FIG. 2. — Geological map of the Devonian and Carboniferous in southern Ahaggar after Fabre (1976: 154). Localities: 1, In Debirène; 2, Tin Rerhoh; 3, Oued Felaou; 4, Anou Izileg; 5, Touaret-Tim Mersoï; 6, Talak." figureDoi="http://doi.org/10.5281/zenodo.4597049" httpUri="https://zenodo.org/record/4597049/files/figure.png" pageId="13" pageNumber="222">
|
|
<subScript id="DE0634F86C41FFA9FC41E8229113C905" attach="left" box="[1044,1054,831,848]" fontSize="7" pageId="13" pageNumber="222">2</subScript>
|
|
</figureCitation>
|
|
). The cone of the base inserted in the crown is, in our material, more developed. In conclusion, some acanthodian scales are closer to
|
|
<taxonomicName id="85824D3E6C41FFA9FB3BE88F96F6C9FE" baseAuthorityName="Young & Burrow" baseAuthorityYear="2004" box="[878,1019,914,939]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="antarctica">
|
|
<emphasis id="70F6EAAF6C41FFA9FB3BE88F96F6C9FE" box="[878,1019,914,939]" italics="true" pageId="13" pageNumber="222">M. antarctica</emphasis>
|
|
</taxonomicName>
|
|
and the others to
|
|
<taxonomicName id="85824D3E6C41FFA9FAF0E8AC963EC99F" authorityName="Burrow, Lelievre & Janjou" authorityYear="2006" box="[677,819,944,970]" class="Chondrichthyes" family="Diplacanthidae" genus="Milesacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acanthodei" pageId="13" pageNumber="222" phylum="Chordata" rank="species" species="ancestralis">
|
|
<emphasis id="70F6EAAF6C41FFA9FAF0E8AC963EC99F" box="[677,819,944,970]" italics="true" pageId="13" pageNumber="222">M. ancestralis</emphasis>
|
|
</taxonomicName>
|
|
but their histology is different.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |