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<document ID-CLB-Dataset="294387" ID-DOI="http://dx.doi.org/10.3897/mycokeys.105.113591" ID-GBIF-Dataset="64ac0e11-9e76-4482-b594-a60794febc6f" ID-Pensoft-Pub="1314-4049-105-1" ID-Pensoft-UUID="5A3AED11D5C855A88B5A618B36F9843C" ModsDocID="1314-4049-105-1" checkinTime="1713442480305" checkinUser="pensoft" docAuthor="Hussain, Shah, Al-Kharousi, Moza, Al-Maqbali, Dua'a, Al-Owaisi, Arwa A., Velazhahan, Rethinasamy, Al-Yahya'ei, Mohamed N. &amp; Al-Sadi, Abdullah M." docDate="2024" docId="82BEAF0BC6A85E019615B20090130FC6" docLanguage="en" docName="MycoKeys 105: 1-19" docOrigin="MycoKeys 105" docPubDate="2024-04-17" docSource="http://dx.doi.org/10.3897/mycokeys.105.113591" docTitle="Hymenagaricus parvulus Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain 2024, sp. nov." docType="treatment" docVersion="2" id="5A3AED11D5C855A88B5A618B36F9843C" lastPageNumber="1" masterDocId="5A3AED11D5C855A88B5A618B36F9843C" masterDocTitle="Two new species of Hymenagaricus (Agaricales, Agaricaceae) from Oman, based on morphology and molecular phylogeny" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1713443100742" updateUser="ExternalLinkService">
<mods:mods id="FB0FB7F58D3556F5B884AFD8DE409552" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7167A7D8439624422E7524B0B9AFC245">
<mods:title id="D59E2EEF94349D540E6B3741F61E327B">Two new species of Hymenagaricus (Agaricales, Agaricaceae) from Oman, based on morphology and molecular phylogeny</mods:title>
</mods:titleInfo>
<mods:name id="BE1A5C64BF3529B75240930E8C0DC927" type="personal">
<mods:role id="5E876B7EDAE93EF83B65AD4AC0AF5361">
<mods:roleTerm id="2A98C60C92A5BB17C0CE2B47C34919AE">Author</mods:roleTerm>
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<mods:namePart id="CD11C137EF4E1A24EB8C1804C355DCED">Hussain, Shah</mods:namePart>
<mods:nameIdentifier id="4573B47F47C74E5F788E8A3A0EB8F0A1" type="ORCID">https://orcid.org/0000-0002-5772-7206</mods:nameIdentifier>
<mods:affiliation id="28E087E64BB4E154A4516D859FA3B7BC">Oman Animal and Plant Genetic Resources Center (Mawarid), Ministry of Higher Education, Research and Innovation, P. O. Box 515, P. C. 123, Muscat, Oman &amp; Department of Plant Sciences, College of Agricultural and Marine Sciences, Sultan Qaboos University, PO Box 34, AlKhoud 123, Oman</mods:affiliation>
</mods:name>
<mods:name id="3535BB68374B8BCD1B7D1A7E21E98453" type="personal">
<mods:role id="9F5F375EA318A198A9B34AE2DFE3BDEF">
<mods:roleTerm id="C85EEA660EF38573660BC02537303D52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E9656DC3B8CD8A2CB0DA4012DE0FFF2">Al-Kharousi, Moza</mods:namePart>
<mods:affiliation id="C347B71CA25241025E1943DE80EAB082">Oman Animal and Plant Genetic Resources Center (Mawarid), Ministry of Higher Education, Research and Innovation, P. O. Box 515, P. C. 123, Muscat, Oman</mods:affiliation>
</mods:name>
<mods:name id="A65C4CFFDF936F90488C493B0CEB0537" type="personal">
<mods:role id="ACE7C655AF1388CB8B22FA59C2919FB7">
<mods:roleTerm id="C0F15D542F8C2F5F424B5FD87036EDDD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B8181347DC038F1C88A65148A8C85B4">Al-Maqbali, Dua'a</mods:namePart>
<mods:affiliation id="53FB60755C1D8419620D2A5DC6609A5A">Oman Animal and Plant Genetic Resources Center (Mawarid), Ministry of Higher Education, Research and Innovation, P. O. Box 515, P. C. 123, Muscat, Oman</mods:affiliation>
</mods:name>
<mods:name id="C233A38388137DCA830CC96DC5C872AF" type="personal">
<mods:role id="1AE603BD92A2A3B6B0A7BDA853533E08">
<mods:roleTerm id="C067300D8DC2B87F724FFB0CA60D3E38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DBAD9D93A4A16DB1C7FB7FE6F15D01B4">Al-Owaisi, Arwa A.</mods:namePart>
<mods:affiliation id="F42A7815C0DEFEDC1F9DD6A4142353A3">Oman Animal and Plant Genetic Resources Center (Mawarid), Ministry of Higher Education, Research and Innovation, P. O. Box 515, P. C. 123, Muscat, Oman</mods:affiliation>
</mods:name>
<mods:name id="66FCE33FB342984FD2AD8E6192939B1A" type="personal">
<mods:role id="506A22D5F2A16B48CABF2A0626B19DDC">
<mods:roleTerm id="34C50D62E7CB00990A6967161B74FE55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7813621DE9D5B2097DFE068556E657B1">Velazhahan, Rethinasamy</mods:namePart>
<mods:nameIdentifier id="7E2A37F3821C3211A5D9732A6FD48E43" type="ORCID">https://orcid.org/0000-0002-9263-4371</mods:nameIdentifier>
<mods:affiliation id="1F12C0FE4BB2B0A36246BAB46BDFC596">Department of Plant Sciences, College of Agricultural and Marine Sciences, Sultan Qaboos University, PO Box 34, AlKhoud 123, Oman</mods:affiliation>
</mods:name>
<mods:name id="F5D80ADAC6387B15673B4F2ECB0370E5" type="personal">
<mods:role id="F23B3990C041EC7DA471F07455DB1DD4">
<mods:roleTerm id="7FBDF81C91C722F53C9AEE36BC45ECA3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="87187EC03B4B35AF7CAFAC5572292432">Al-Yahya'ei, Mohamed N.</mods:namePart>
<mods:nameIdentifier id="89239C9AA2A896C1A2ABF4C86081F395" type="ORCID">https://orcid.org/0000-0002-9516-5339</mods:nameIdentifier>
<mods:affiliation id="1493A74392C1E30E29182B4FF0E33E10">Oman Animal and Plant Genetic Resources Center (Mawarid), Ministry of Higher Education, Research and Innovation, P. O. Box 515, P. C. 123, Muscat, Oman</mods:affiliation>
<mods:nameIdentifier id="E2361065D779A1C9BB51E89DB23C41B3" type="email">mohamed.alyahyaei@moheri.gov.om</mods:nameIdentifier>
</mods:name>
<mods:name id="F58DA68CCBB25C5FEC60DBCB556503F6" type="personal">
<mods:role id="39E633A4F33206D248C5123308D4B303">
<mods:roleTerm id="365440E3C6BEE7CC61AEEF044A8D9599">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D304515ACB70EFCFE592D392005B6B3">Al-Sadi, Abdullah M.</mods:namePart>
<mods:nameIdentifier id="ADFC678A3510EF80B69C3FB843BC3551" type="ORCID">https://orcid.org/0000-0002-3419-8268</mods:nameIdentifier>
<mods:affiliation id="9ACD4F171375E363D7EE44C940A50429">Department of Plant Sciences, College of Agricultural and Marine Sciences, Sultan Qaboos University, PO Box 34, AlKhoud 123, Oman</mods:affiliation>
<mods:nameIdentifier id="64A129F2FEDE34859C528B2FA3A1F004" type="email">alsadi@squ.edu.om</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EC4038509204C66A6902587D63B5F70F">text</mods:typeOfResource>
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<mods:titleInfo id="C54F84E617C881C35466A74F68AB3744">
<mods:title id="6C29B9BB6276C254E14232B067B78D66">MycoKeys</mods:title>
</mods:titleInfo>
<mods:part id="2AD7154A329A9E235D98BD54B7FE91CF">
<mods:date id="63152322A378E5D4C6B67BD31F6C3848">2024</mods:date>
<mods:detail id="F20E6F97217039B7C4C5A148EB0227BE" type="pubDate">
<mods:number id="5BC8A65485FBAE29BDA19C48EAC8058A">2024-04-17</mods:number>
</mods:detail>
<mods:detail id="4923608629149238DE65BEF351FC57FC" type="volume">
<mods:number id="1A5F3BE241C77549D9C904EB7E5BDF7F">105</mods:number>
</mods:detail>
<mods:extent id="EA3E00F8B3CE97CB17846B56717F0919" unit="page">
<mods:start id="002934907473AA519738F83357BE0812">1</mods:start>
<mods:end id="59F36D0F05251D82D7AD1FBAD00C380A">19</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="A8282A161E7D4CC5A959AF7FC8B3A3D8">
<mods:url id="D203B9DE8A947C9BF178C88DACFAF0D9">http://dx.doi.org/10.3897/mycokeys.105.113591</mods:url>
</mods:location>
<mods:classification id="D86AECFF59EAFD050937BAC1500979A4">journal article</mods:classification>
<mods:identifier id="B998F6AF124BBF6F92B9C50D799B1648" type="DOI">http://dx.doi.org/10.3897/mycokeys.105.113591</mods:identifier>
<mods:identifier id="C953DC97336356F853E4D773394DEB7C" type="Pensoft-Pub">1314-4049-105-1</mods:identifier>
<mods:identifier id="CBD346D970F4A918E55974D32DA99D6E" type="Pensoft-UUID">5A3AED11D5C855A88B5A618B36F9843C</mods:identifier>
</mods:mods>
<subSection id="6A79586D05C27AD16541FD8DB6074F0F" pageId="0" pageNumber="1" type="results">
<treatment id="82BEAF0BC6A85E019615B20090130FC6" LSID="urn:lsid:plazi:treatment:82BEAF0BC6A85E019615B20090130FC6" httpUri="http://treatment.plazi.org/id/82BEAF0BC6A85E019615B20090130FC6" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="5E28767E4B643BE5E936B2DF5CCF0C63" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="7271AFC5C06EF7916E3AA77FC530A6E3" pageId="0" pageNumber="1">
<taxonomicName id="0AA07773F79927A5CAF47B02AE26757F" LSID="82BEAF0B-C6A8-5E01-9615-B20090130FC6" authority="Al‐Kharousi, Al‐Sadi, Al-Yahya’ei, &amp; S. Hussain" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus" status="sp. nov.">
Hymenagaricus parvulus Al-Kharousi, Al-Sadi,
<normalizedToken id="1D666440451E02616CEB72A5FD25581E" originalValue="Al-Yahya’ei">Al-Yahya'ei</normalizedToken>
, &amp; S. Hussain
</taxonomicName>
<taxonomicNameLabel id="82D8679650F27DE9D99A4EC02E54460C" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="30FF2B705C4FC9E6B11E9CD0559B7784" pageId="0" pageNumber="1" type="description">
<paragraph id="484A3D705EAA74B559AD49FA6F89C637" pageId="0" pageNumber="1">
<figureCitation id="597979CD08FC55A86C89D445CB0EC8A5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Basidiomata of Hymenagaricus parvulus (based on holotype collection JRZ- 22 - 004) A mature and young basidiomata B mature basidimata." figureDoi="10.3897/mycokeys.105.113591.figure5" httpUri="https://binary.pensoft.net/fig/1027429" pageId="0" pageNumber="1">Figs 5</figureCitation>
<figureCitation id="CD173618D754083D5AD462B889A7CBA3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Light microscopy of anatomical features of Hymenagaricus parvulus (based on holotype collection JRZ- 22 - 004) A basidiospores B basidia C cheilocystidia D annulus E pellicle structure F veil elements. Scale bars: 5 µm (A); 10 µm (B, C); 15 µm (D-F)." figureDoi="10.3897/mycokeys.105.113591.figure6" httpUri="https://binary.pensoft.net/fig/1027430" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation id="A34A7A7AE2511F6183E922DDD31B8F41" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Line drawings of anatomical features of Hymenagaricus parvulus (based on holotype collection JRZ- 22 - 004) A basidiospores B basidia C annulus elements D cheilocystidia E pellicle structure F veil elements. Scale bars: 5 µm (A); 10 µm (B, D); 15 µm (C, E, F)." figureDoi="10.3897/mycokeys.105.113591.figure7" httpUri="https://binary.pensoft.net/fig/1027431" pageId="0" pageNumber="1">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2CCF199AE0044043514A86BCFC71BCC5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="EDEB4DC14254CB448C2A3DA145DDEEE6" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="2BB12D7ACA5F2CB148E90DE97DFD68B6" pageId="0" pageNumber="1">
The new species
<taxonomicName id="2AE3B6AE2E8C34336E2B900D7C5205E0" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="DAF8A015F64E70968F55A14159DE63E2" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
can be differentiated from other species of the genus by its small-sized, creamy basidiomata, umbonate pileus covered with appressed pellicle.
</paragraph>
</subSubSection>
<subSubSection id="1974885222C8AD6BCF724ED9BC00A53F" pageId="0" pageNumber="1" type="holotype">
<paragraph id="5A36419AD9B6FA89832D2C741DCA2D7D" pageId="0" pageNumber="1">Holotype.</paragraph>
<paragraph id="19E98F5D6617E5401B19F9BA64D8EE80" pageId="0" pageNumber="1">
Sultanate of Oman: Dhofar, Salalah, Wadi Jarzeez, on termite mounds, under the trees of
<taxonomicName id="28D530E742AA6EB16A5C9CF5BB687BB2" class="Dicotyledoneae" family="Combretaceae" genus="Anogeissus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Anogeissus dhofarica" order="Myrtiflorae" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="dhofarica">
<emphasis id="903B5680EF3B9BFBBB38E72FBF54AC1B" italics="true" pageId="0" pageNumber="1">Anogeissus dhofarica</emphasis>
</taxonomicName>
, 8 August 2022, S. Hussain, A. Al-Owaisi,
<normalizedToken id="4D898CAF9BCF43425C46AFF561F6249D" originalValue="Al-Yahya’ei">Al-Yahya'ei</normalizedToken>
&amp; Al-Sadi, JRZ-22-004 (holotype Mawarid-JRZ-22-004), GenBank accession: ITS = OR612994, 28S = OR613017, EF-1α = OR735176.
</paragraph>
</subSubSection>
<subSubSection id="027B156F8B634D874A2A3D5E6949A6EB" pageId="0" pageNumber="1" type="etymology">
<paragraph id="DDA287FA40E7B298DD44D408231F3439" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="7696C01A890E2D1CDA64CC76F4040501" pageId="0" pageNumber="1">
The specific epithet '
<emphasis id="87110BFDF73A7F661D247B040C4BFEDC" italics="true" pageId="0" pageNumber="1">parvulus</emphasis>
' refers to the small-sized basidiomata of the new species.
</paragraph>
</subSubSection>
<subSubSection id="EECFA94714016FF6AD4ACF2A2C701286" pageId="0" pageNumber="1" type="description">
<paragraph id="83B507BEE87306250DFC876C5CE44F04" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="9623AC9EA2670D82FAE6829FA8CCF6CB" pageId="0" pageNumber="1">
<emphasis id="6AB13168B0021A9D7D033578A38E8AC5" bold="true" italics="true" pageId="0" pageNumber="1">Basidiomata</emphasis>
small-sized.
<emphasis id="082ECB1A393B4F9FE97E1222DBCF3F06" bold="true" italics="true" pageId="0" pageNumber="1">Pileus</emphasis>
15-25 mm in diam., at young stage globose to parabolic, surface floccose squamulose, squamules light pinkish to creamy, with appressed pellicle at the centre, margin appendiculate; at mature stage cap convex to hemispherical with the broadly umbonate disc, with appressed, pale brownish pellicle at the disc, surface finely floccose squamulose, squamules pale creamy to light greyish, margins striate, just exceeding the lamellae; context membranous, pinkish on cutting.
<emphasis id="8A06EB298447392794716A90820FB82D" bold="true" italics="true" pageId="0" pageNumber="1">Lamellae</emphasis>
free, pale pinkish to brownish, ventricose, sparsely crowded, with 1-2 series of lamellulae.
<emphasis id="60579B47C222BA030F769C96DBE45A8A" bold="true" italics="true" pageId="0" pageNumber="1">Stipe</emphasis>
25-35
<normalizedToken id="9B6AFC54DE274D0F39BB61D794AC3527" originalValue="×">x</normalizedToken>
2-5 mm, equal, annulus cortinate, concolorous to squamules; stem surface creamy, covered with finely floccose squamules below the annulus, smooth above the annulus, context pinkish on cutting, fistulose.
<emphasis id="B7174F90B4B445F97FD7D96531B3338F" bold="true" italics="true" pageId="0" pageNumber="1">Smell</emphasis>
pleasant.
<emphasis id="AD9076757FFA41780ACE2B5DF6AD7CC3" bold="true" italics="true" pageId="0" pageNumber="1">Taste</emphasis>
not recorded.
</paragraph>
<caption id="B545ACD13EF9126B5D3B4652E7F2DE04" doi="10.3897/mycokeys.105.113591.figure5" httpUri="https://binary.pensoft.net/fig/1027429" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="B0D722935942E7C82002CFFD1C14097C" pageId="0" pageNumber="1">
<emphasis id="9E370F69C17E60BF973636FE0CDAD85E" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Basidiomata of
<taxonomicName id="928C81F237ACEEFCD62A378C0E9941A2" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="633D2AF33D892B1C1925120CF8A0D93D" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
(based on holotype collection JRZ-22-004)
<emphasis id="E16D1886E1C129E4D1670F34B7BC8DC3" bold="true" pageId="0" pageNumber="1">A</emphasis>
mature and young basidiomata
<emphasis id="403D4A575EEE24A30CFCE9B8D3966AEB" bold="true" pageId="0" pageNumber="1">B</emphasis>
mature basidimata.
</paragraph>
</caption>
<paragraph id="B8EF1530E90167A10E6F2C8037052C33" pageId="0" pageNumber="1">
<emphasis id="D4CB99851EDBC273421A380FDE5C0243" bold="true" italics="true" pageId="0" pageNumber="1">Basidiospores</emphasis>
5.0-6.5
<normalizedToken id="41C779866312EC17E364998A956A4548" originalValue="×">x</normalizedToken>
4.0-4.5
<normalizedToken id="1A18014D45008379ACA4DFB357C541B2" originalValue="µm">µm</normalizedToken>
, average size 6.0
<normalizedToken id="9054734A38BA2A60EF333D66F2287F6D" originalValue="×">x</normalizedToken>
4.2
<normalizedToken id="FD507DFE4FEE519FAC5A5B81168CD6CD" originalValue="µm">µm</normalizedToken>
, Q = 1.3-1.5, av. Q = 1.4; ellipsoid to broadly ellipsoid, yellowish to dark brown, smooth, thick-walled, apiculus visible, germ-pore not observed.
<emphasis id="5018B57907C17A620EA6CEDA605F9884" bold="true" italics="true" pageId="0" pageNumber="1">Basidia</emphasis>
16.5-22.5
<normalizedToken id="1100E2C9287DE94752427D4337BEB7B5" originalValue="×">x</normalizedToken>
6.5-8.5
<normalizedToken id="53324943C296A7F6FB0D800F120E890C" originalValue="µm">µm</normalizedToken>
, on average 19.0
<normalizedToken id="491A810A8B134CD0F0AB4F7F42728965" originalValue="×">x</normalizedToken>
7.5
<normalizedToken id="7170136C9CE42152D7CD23682086573F" originalValue="µm">µm</normalizedToken>
, clavate to cylindrical, smooth, hyaline in KOH, tetrasporic.
<emphasis id="F6AF91E1BB805A65F3B336019B22006D" bold="true" italics="true" pageId="0" pageNumber="1">Cheilocystidia</emphasis>
19-25
<normalizedToken id="13F010891968B478236D4B7A8100FF54" originalValue="×">x</normalizedToken>
9-11
<normalizedToken id="D48215D4BD0BC4DA59C242885F65EFEC" originalValue="µm">µm</normalizedToken>
, on average 22
<normalizedToken id="155BA619828593C6429F41BCAF307825" originalValue="×">x</normalizedToken>
10
<normalizedToken id="76D10D573C9367764E969231BDAF8A92" originalValue="µm">µm</normalizedToken>
, clavate to broadly clavate, often turning to one side, with multiseptate base, smooth, thin-walled, hyaline in KOH.
<emphasis id="F0DEB13D3A0D1B66A7E0453E18654FF5" bold="true" italics="true" pageId="0" pageNumber="1">Pleurocystidia</emphasis>
absent.
<emphasis id="E0C424B5C85C771A42ACB0537C6093E2" bold="true" italics="true" pageId="0" pageNumber="1">Subhymenium</emphasis>
consisting of cylindrical to elongated cells, measuring 6-9
<normalizedToken id="2EC5352F8B08D6E2655769BE9F3D52CB" originalValue="µm">µm</normalizedToken>
diam.
<emphasis id="4D4D6FD9B76BDE5A17F55168A554E832" bold="true" italics="true" pageId="0" pageNumber="1">Pellicle</emphasis>
is a hymeniform, consisting of chains of several elements, each element measuring 14-22
<normalizedToken id="84D100AA758395D1D12035EB1BB0CDA6" originalValue="×">x</normalizedToken>
12-17
<normalizedToken id="7B9E008F4339DC322C4AD6596F13FC4C" originalValue="µm">µm</normalizedToken>
, globose to subglobose or ovoid, hyaline or pale yellowish, smooth, thin-walled; these chains of elements attached to inflated hyphae with encrusted walls.
<emphasis id="0720113B385BB902B8AF028C1575AE40" bold="true" italics="true" pageId="0" pageNumber="1">Veil</emphasis>
is a cutis to ixocutis, consisting of elongated or cylindrical elements, not easily detached, hyaline, thin-walled, with terminal element fusiform with papillate end, each element measuring 15-18
<normalizedToken id="9315939FDC982FD20D336E1FFD047A5D" originalValue="×">x</normalizedToken>
5-7
<normalizedToken id="9A4AD05E20AB4BFBAC1A21D7C65E37CA" originalValue="µm">µm</normalizedToken>
.
<emphasis id="EC89B0F5AA9B9853CDD01757E424D9AC" bold="true" italics="true" pageId="0" pageNumber="1">Annulus</emphasis>
is an intricate trichoderm, composed of hyaline hyphae, 4-7
<normalizedToken id="BB55E8461D931E9EE5AD4E6309EFD733" originalValue="µm">µm</normalizedToken>
diam., cylindrical, constituted by short elements, constricted at septa and easily disarticulated.
<emphasis id="B1513611112E04D7C269C7B0B37D8D0E" bold="true" italics="true" pageId="0" pageNumber="1">Clamp connections</emphasis>
absent in all tissues.
</paragraph>
<caption id="B6C7F800F1F64EAAF463982F33881D6A" doi="10.3897/mycokeys.105.113591.figure6" httpUri="https://binary.pensoft.net/fig/1027430" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph id="8F8843CF9BFED0C972011CC1673A7F64" pageId="0" pageNumber="1">
<emphasis id="064CDFDC72471807501230FD99ECE34B" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
Light microscopy of anatomical features of
<taxonomicName id="0B7A425B2B125681BDCCD45B0D82C8FB" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="16287AE73DA6C8BADDEDA2FD742B0BA3" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
(based on holotype collection JRZ-22-004)
<emphasis id="5B87A06994AD0CE7AE5186DAD8BFAAA6" bold="true" pageId="0" pageNumber="1">A</emphasis>
basidiospores
<emphasis id="A13385650CEB35879F31F21A2E855DC8" bold="true" pageId="0" pageNumber="1">B</emphasis>
basidia
<emphasis id="8CB19FBEC164E2506D0D1F9AF51F0C6F" bold="true" pageId="0" pageNumber="1">C</emphasis>
cheilocystidia
<emphasis id="1BF241C9989FA782E815EBBF5C2ADDD7" bold="true" pageId="0" pageNumber="1">D</emphasis>
annulus
<emphasis id="86CA45D8B7B4065940A87941469288F3" bold="true" pageId="0" pageNumber="1">E</emphasis>
pellicle structure
<emphasis id="5198D1B5CFAC8E1929F50872DB85B0E5" bold="true" pageId="0" pageNumber="1">F</emphasis>
veil elements. Scale bars: 5
<normalizedToken id="467D0E9308725701CC5A82ABFA72B9CF" originalValue="µm">µm</normalizedToken>
(
<emphasis id="A5C3E22B54E8A728F9E29DFF7F180F95" bold="true" pageId="0" pageNumber="1">A</emphasis>
); 10
<normalizedToken id="45624EF3269CCE1F71B0CCD8DD0D1240" originalValue="µm">µm</normalizedToken>
(
<emphasis id="48BE1B02A244E04ABF7C19AE560357A1" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
); 15
<normalizedToken id="8A102AD596F32530B048B9C835016FDD" originalValue="µm">µm</normalizedToken>
(
<emphasis id="3567C20DC6E5AA91687D2BD7C8FBCF50" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="AF033E2DE301CCA0D5ADEE9BB5978646" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A02BED6F85077A3B2D2FC61D1BD6B012" pageId="0" pageNumber="1">Habit, habitat and distribution.</paragraph>
<paragraph id="3B6AA7A26DECE949D102913E20901DF3" pageId="0" pageNumber="1">Fruiting body formation occurs in early August to early September, saprotrophic, scattered in small groups, found on termite mounds. Currently only known from southern Oman.</paragraph>
<caption id="ABB352F1659CC29C88C96FD0D8C068C6" doi="10.3897/mycokeys.105.113591.figure7" httpUri="https://binary.pensoft.net/fig/1027431" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph id="1E64CD03809771D41F3B229A2E800A4F" pageId="0" pageNumber="1">
<emphasis id="EAF7DAB999D278439ACAE350E2B8B5E0" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Line drawings of anatomical features of
<taxonomicName id="4280BDB3A6868CDE4CF3ADE747B3C921" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="01A080E6336871384DB5C3357B02D1B6" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
(based on holotype collection JRZ-22-004)
<emphasis id="50E972E05723AF323E707C22FF8107B7" bold="true" pageId="0" pageNumber="1">A</emphasis>
basidiospores
<emphasis id="796260F98813DB94BE2E0D7C9B040D69" bold="true" pageId="0" pageNumber="1">B</emphasis>
basidia
<emphasis id="ECCB95EBD256D11CA6684EF7E699BE90" bold="true" pageId="0" pageNumber="1">C</emphasis>
annulus elements
<emphasis id="090A800BC9E951BCDC31C5AFE56FB3A0" bold="true" pageId="0" pageNumber="1">D</emphasis>
cheilocystidia
<emphasis id="BD9A1A12141D5E2AFF5F7A2132C34936" bold="true" pageId="0" pageNumber="1">E</emphasis>
pellicle structure
<emphasis id="DBBBE976317824B6C6AF3C7A0DA30C0A" bold="true" pageId="0" pageNumber="1">F</emphasis>
veil elements. Scale bars: 5
<normalizedToken id="C11C46290FB8530A861BBBC0DA551443" originalValue="µm">µm</normalizedToken>
(
<emphasis id="0CD740D06433B4A1A1652ECCD9DB7EC5" bold="true" pageId="0" pageNumber="1">A</emphasis>
); 10
<normalizedToken id="094AE6B632407D48FE378BA2FAA70D44" originalValue="µm">µm</normalizedToken>
(
<emphasis id="35817D8A38B9E6DA21785C5C391887D2" bold="true" pageId="0" pageNumber="1">B, D</emphasis>
); 15
<normalizedToken id="B64B8B6C0B516671C02C7E250C6F7D7E" originalValue="µm">µm</normalizedToken>
(
<emphasis id="AEBFFA17AAE5595B48890B9568A3C581" bold="true" pageId="0" pageNumber="1">C, E, F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8A77786FC0040E901141E6650154BF00" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="0B47DAA6B2C93E8075CF0AF990886911" pageId="0" pageNumber="1">Additional specimens examined.</paragraph>
<paragraph id="931119C47BB12BA8C3979523C241BD75" pageId="0" pageNumber="1">
<materialsCitation id="BC4D6780CAE7E5337C138962953088A5" collectingDate="2022-08-11" collectorName="S. Hussain, A. Al-Owaisi, Al-Sadi, GenBank" country="Sultanate of Oman" location="Wadi Jarzeez" municipality="Salalah" specimenCount="1" stateProvince="Dhofar">
<collectingCountry id="EE1E2E5B5FFA94F47EFF49FEC42A66F5" name="Oman">Sultanate of Oman</collectingCountry>
:
<collectingRegion id="981820ED055E6584A9C2A45C6B8616B2" country="Oman" name="Zufar">Dhofar</collectingRegion>
,
<collectingMunicipality id="42D79603B866033A12B6572B3F3D9D79">Salalah</collectingMunicipality>
,
<location id="4D3C1D24C6F7B4115BD6F6AE0083615C" LSID="urn:lsid:plazi:treatment:82BEAF0BC6A85E019615B20090130FC6:4D3C1D24C6F7B4115BD6F6AE0083615C" country="Sultanate of Oman" municipality="Salalah" name="Wadi Jarzeez" stateProvince="Dhofar">Wadi Jarzeez</location>
, on termite mounds, under the trees of
<taxonomicName id="52E3BFB25B5758AD1C830528674FC82C" class="Dicotyledoneae" family="Combretaceae" genus="Anogeissus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Anogeissus dhofarica" order="Myrtiflorae" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="dhofarica">
<emphasis id="8F018D3F959359B8436036F2FDC3347C" italics="true" pageId="0" pageNumber="1">Anogeissus dhofarica</emphasis>
</taxonomicName>
,
<collectingDate id="E137DDC3038B0ADF29862E14AB91B83B" value="2022-08-11">11 August 2022</collectingDate>
,
<collectorName id="97D720A78044C3230DA93E6CFB74FBDE">S. Hussain</collectorName>
,
<collectorName id="B2A4D15F366DF37A82DAE3FAABB5DB5B">A. Al-Owaisi</collectorName>
,
<normalizedToken id="2F976B52B410EE8D1AB2B4CFAE50B63A" originalValue="Al-Yahya’ei">Al-Yahya'ei</normalizedToken>
&amp;
<collectorName id="2BFFC2BF15D47688E8E66C2B818D448F">Al-Sadi</collectorName>
, JRZ2-22-002 (Mawarid-NHZ-22-002),
<collectorName id="C6ED36513896B6BEE447D6FA111DDC3E">GenBank</collectorName>
accession: ITS = OR612995
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BA95CD3540C354642D43EB7A661C6A3A" pageId="0" pageNumber="1" type="notes">
<paragraph id="AFD73C795399D688E4ADA81648122527" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="9E3AA5913AA1C5C5221B01B59F553A2B" pageId="0" pageNumber="1">
<taxonomicName id="AA24B4A0C98BFA1C070FBB0490A11B8C" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="6EF2FEFF7ECFCA733B430E0B526955DE" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
is a small, cream-coloured species, differentiated from other species of the genus by its whitish to pale pinkish floccose squamules on pileus and stipe surfaces with a broadly umbonate centre.
<taxonomicName id="DAB339C34079D7B1D1A59CB71FD35392" authorityName="Al-Kharousi, Al-Sadi, Al-Yahya'ei, &amp; S. Hussain" authorityYear="2024" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="BF2D3A4256DB97DBE8FFBBE6952AF863" italics="true" pageId="0" pageNumber="1">Hymenagaricus parvulus</emphasis>
</taxonomicName>
shares basidiomata size and basidiospores morphology with
<taxonomicName id="F2904221F18D9775D77438D3F75E1A52" class="Agaricomycetes" family="Agaricaceae" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="H. pakistanicus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="pakistanicus">
<emphasis id="FAA8F286A350152B4B84395908A0477C" italics="true" pageId="0" pageNumber="1">H. pakistanicus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="A3390F223132E3AD83043D6E9EB0CC43" class="Agaricomycetes" family="Agaricaceae" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="H. pakistanicus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="pakistanicus">
<emphasis id="DA81589C356EE75290714AC43CD6464E" italics="true" pageId="0" pageNumber="1">H. pakistanicus</emphasis>
</taxonomicName>
can be differentiated from the new species by its caesptiose fruiting habit, pileus with pinkish to brownish squamulose pellicle, consisting of pseudoparenchymatous cells (
<bibRefCitation id="7182BF15C0E6ABB493EF8CD92C545F4D" DOI="https://doi.org/10.11646/phytotaxa.594.4.6" author="Syed, MF" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" pagination="292 - 300" refId="B33" refString="Syed, MF, Saba, M, Chattha, SM, Hosen, MI, 2023. Hymenagaricus pakistanicus (Agaricaceae, Agaricales), a new species from Pakistan based on morphological and molecular evidence. Phytotaxa 594 (4): 292 - 300, DOI: https://doi.org/10.11646/phytotaxa.594.4.6" title="Hymenagaricus pakistanicus (Agaricaceae, Agaricales), a new species from Pakistan based on morphological and molecular evidence." url="https://doi.org/10.11646/phytotaxa.594.4.6" volume="594" year="2023">Syed et al. 2023</bibRefCitation>
).
<taxonomicName id="51640A02E48767602EBF1D313CEE9348" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus saisamornae" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="saisamornae">
<emphasis id="C84BA3AC2E356A0542823C38F899A2E3" italics="true" pageId="0" pageNumber="1">Hymenagaricus saisamornae</emphasis>
</taxonomicName>
differs from the new species by its smaller pileus (up to 15 mm diam. Vs. 15-25 mm of
<taxonomicName id="52BD23F0C84C31246C6A18CBD28F029F" class="Agaricomycetes" family="Agaricaceae" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="H. parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="4365A2CF48C48003F846913ECFDEEE4B" italics="true" pageId="0" pageNumber="1">H. parvulus</emphasis>
</taxonomicName>
), covered with brownish pellicles and larger basidiospores (5.5-7.0
<normalizedToken id="4F793DF0CCD49FD5C65A88B54560C550" originalValue="×">x</normalizedToken>
4-4.5
<normalizedToken id="41BE4958B840057BAC4D8B4CEE67E238" originalValue="µm">µm</normalizedToken>
;
<bibRefCitation id="151827616E67E875785FD2ABB8D77FBC" author="Kumla, J" journalOrPublisher="Warasan Khana Witthayasat Maha Witthayalai Chiang Mai" pageId="0" pageNumber="1" pagination="827 - 836" refId="B20" refString="Kumla, J, Suwannarach, N, Wannathes, N, 2021. Hymenagaricus saisamornae sp. nov. (Agaricales, Basidiomycota) from Northern Thailand. Warasan Khana Witthayasat Maha Witthayalai Chiang Mai 48: 827 - 836" title="Hymenagaricus saisamornae sp. nov. (Agaricales, Basidiomycota) from Northern Thailand." volume="48" year="2021">Kumla et al. (2021)</bibRefCitation>
).
<taxonomicName id="3A023024B5D28EF670CC1B91FF32C406" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus siamensis" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="siamensis">
<emphasis id="EBAFF3B0EE28B9C7A9D349EA382C3021" italics="true" pageId="0" pageNumber="1">Hymenagaricus siamensis</emphasis>
</taxonomicName>
, another small-sized species is distinguished from the new species by its pinkish-brown cap, pellicle comprised of pseudoparenchymatous cells and larger basidiospores (6.5-8.0
<normalizedToken id="A4B95434126696B4B466FFB6FF89B58E" originalValue="×">x</normalizedToken>
4.0-5.0
<normalizedToken id="9A7BC472DCF7547AFD32BD69E8E1CE99" originalValue="µm">µm</normalizedToken>
;
<bibRefCitation id="C55C519482B82D53C8749F6D234238B9" DOI="https://doi.org/10.11646/phytotaxa.585.2.4" author="Kumla, J" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" pagination="135 - 144" refId="B21" refString="Kumla, J, Suwannarach, N, Khuna, S, Phonrob, W, Lumyong, S, 2023. Hymenagaricus siamensis (Agaricaceae, Agaricales), a novel macrofungus from northern Thailand. Phytotaxa 585 (2): 135 - 144, DOI: https://doi.org/10.11646/phytotaxa.585.2.4" title="Hymenagaricus siamensis (Agaricaceae, Agaricales), a novel macrofungus from northern Thailand." url="https://doi.org/10.11646/phytotaxa.585.2.4" volume="585" year="2023">Kumla et al. (2023)</bibRefCitation>
). Similarly,
<taxonomicName id="7FF89F7C57D01DA8D783254225115EED" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus canoruber" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="canoruber">
<emphasis id="A1A630495F62040844C5CF8C12F618B6" italics="true" pageId="0" pageNumber="1">Hymenagaricus canoruber</emphasis>
</taxonomicName>
(Berk. &amp; Br.) Heinem. &amp; Little Flower, known from India and Sri Lanka, is characterised by a small-sized pileus (15-25 mm diam.), with greyish-brown squamules, hymeniform pellicle and smaller basidiospores (4.6-5.7 3.5-4.3
<normalizedToken id="8B69E57A3EA7D4ADE79C031FBBF7BAAF" originalValue="µm">µm</normalizedToken>
;
<bibRefCitation id="3A12EE6ADE7091BA4D6284B9342B2EB2" DOI="https://doi.org/10.2307/3667871" author="Heinemann, P" journalOrPublisher="Bulletin du Jardin Botanique de l'Etat a Bruxelles" pageId="0" pageNumber="1" refId="B14" refString="Heinemann, P, Little Flower, SR, 1984. Hymenagaricus (Agaricaceae) de Kerala (Inde) et de Sri Lanka. Bulletin du Jardin Botanique National de Belgique 54(1/2): 151-182. https://doi.org/10.2307/3667871" title="Hymenagaricus (Agaricaceae) de Kerala (Inde) et de Sri Lanka. Bulletin du Jardin Botanique National de Belgique 54 (1 / 2): 151 - 182." url="https://doi.org/10.2307/3667871" year="1984">Heinemann and Little Flower (1984)</bibRefCitation>
).
<taxonomicName id="1EA7D30CC8B6E24DE1508DA74698B052" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus pallidodiscus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="pallidodiscus">
<emphasis id="AD10C6FE5F4D7326690EC7C0CD1F2AFA" italics="true" pageId="0" pageNumber="1">Hymenagaricus pallidodiscus</emphasis>
</taxonomicName>
D.A. Reid &amp; Eicker, the smallest mushroom in the genus with pileus diam. up to 11 mm, covered with brownish squamules and smaller basidiospores (4.2-5.4
<normalizedToken id="96A10B0E8150599E0F57DF56031C1A81" originalValue="×">x</normalizedToken>
3.1-3.8
<normalizedToken id="0FDA08282F8AC115A04779E1DB0FC1A9" originalValue="µm">µm</normalizedToken>
;
<bibRefCitation id="F0116EC868029D6FC97A183045D7F8DE" author="Reid, DA" journalOrPublisher="Mycotaxon" pageId="0" pageNumber="1" pagination="169 - 197" refId="B30" refString="Reid, DA, Eicker, A, 1999. South African fungi 10: New species, new records and some new observations. Mycotaxon 73: 169 - 197" title="South African fungi 10: New species, new records and some new observations." volume="73" year="1999">Reid and Eicker (1999)</bibRefCitation>
).
<taxonomicName id="A1816CF7FCA2E0F87AE78C38ACB4E0B0" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus cylindrocystis" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="cylindrocystis">
<emphasis id="7B19C8771804F107B782282F7147E7C3" italics="true" pageId="0" pageNumber="1">Hymenagaricus cylindrocystis</emphasis>
</taxonomicName>
Heinem. &amp; Little Flower another small-sized species, has been reported in Singapore and India, with a brownish cap, larger basidiospores (6.4-8.4
<normalizedToken id="F59C3B94CFA18A647875B854728A6502" originalValue="×">x</normalizedToken>
4.5-5.6
<normalizedToken id="FE01460ABF47EF1D2D5F6D412C829CA3" originalValue="µm">µm</normalizedToken>
) and a pseudoparenchymatous pellicle (
<bibRefCitation id="16FE5F584F678C305DEEC33778FE04FB" DOI="https://doi.org/10.2307/3667096" author="Heinemann, P" journalOrPublisher="Bulletin du Jardin Botanique de l'Etat a Bruxelles" pageId="0" pageNumber="1" pagination="1 - 127" refId="B11" refString="Heinemann, P, 1956. Champignons recoltes au Congo Belge par Madame M. Goossens-Fontana II. Agaricus Fries s.s. Bulletin du Jardin Botanique de l'Etat a Bruxelles 26: 1 - 127, DOI: https://doi.org/10.2307/3667096" title="Champignons recoltes au Congo Belge par Madame M. Goossens-Fontana II. Agaricus Fries s. s." url="https://doi.org/10.2307/3667096" volume="26" year="1956">Heinemann 1956</bibRefCitation>
;
<bibRefCitation id="2B6D550ADAF35CF2B24A2AAAB176BA33" DOI="https://doi.org/10.2307/3667871" author="Heinemann, P" journalOrPublisher="Bulletin du Jardin Botanique de l'Etat a Bruxelles" pageId="0" pageNumber="1" refId="B14" refString="Heinemann, P, Little Flower, SR, 1984. Hymenagaricus (Agaricaceae) de Kerala (Inde) et de Sri Lanka. Bulletin du Jardin Botanique National de Belgique 54(1/2): 151-182. https://doi.org/10.2307/3667871" title="Hymenagaricus (Agaricaceae) de Kerala (Inde) et de Sri Lanka. Bulletin du Jardin Botanique National de Belgique 54 (1 / 2): 151 - 182." url="https://doi.org/10.2307/3667871" year="1984">Heinemann and Little Flower 1984</bibRefCitation>
).
<taxonomicName id="89547C81E70C1CCBD363F8BACB288237" class="Agaricomycetes" family="Agaricaceae" genus="Hymenagaricus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenagaricus kivuensis" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="kivuensis">Hymenagaricus cf. kivuensis</taxonomicName>
and
<taxonomicName id="76129898FADE9AEBE15DBAF890A1A489" class="Agaricomycetes" family="Agaricaceae" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="H. wadijarzeezicus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="wadijarzeezicus">
<emphasis id="1676B166FF0D86D6ED9CA808E58AFBFC" italics="true" pageId="0" pageNumber="1">H. wadijarzeezicus</emphasis>
</taxonomicName>
with their medium-sized pilei can be easily differentiated from
<taxonomicName id="80C6C0EF9300EA0EA505C5ECE199EC46" class="Agaricomycetes" family="Agaricaceae" higherTaxonomySource="GBIF" kingdom="Fungi" lsidName="H. parvulus" order="Agaricales" pageId="0" pageNumber="1" phylum="Basidiomycota" rank="species" species="parvulus">
<emphasis id="C8647E32C231BBDD51DE5725FB1AF2EC" italics="true" pageId="0" pageNumber="1">H. parvulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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