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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1003.56557" ID-GBIF-Dataset="1035018b-1a43-4681-8ba9-ca62c8bb4ee8" ID-PMC="PMC7752890" ID-Pensoft-Pub="1313-2970-1003-83" ID-Pensoft-UUID="3CD12B279B9C50768125F2697B2F0E98" ID-PubMed="33384563" ID-ZooBank="5D883EC653064AC3A0D78CA867324596" ModsDocID="1313-2970-1003-83" checkinTime="1607975014278" checkinUser="pensoft" docAuthor="Meyer, Marc De, Goergen, Georg &amp; Jordaens, Kurt" docDate="2020" docId="66EEEDF583975B93929D33E91AFC394C" docLanguage="en" docName="ZooKeys 1003: 83-160" docOrigin="ZooKeys 1003" docPubDate="2020-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1003.56557" docTitle="Senaspis nigrita" docType="treatment" docVersion="4" id="3CD12B279B9C50768125F2697B2F0E98" lastPageNumber="83" masterDocId="3CD12B279B9C50768125F2697B2F0E98" masterDocTitle="Taxonomic revision of the Afrotropical hover fly genus Senaspis Macquart (Diptera, Syrphidae)" masterLastPageNumber="160" masterPageNumber="83" pageNumber="83" updateTime="1668146589315" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Afrotropical hover fly genus Senaspis Macquart (Diptera, Syrphidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meyer, Marc De</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0755-2898</mods:nameIdentifier>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B 3080 Tervuren, Belgium</mods:affiliation>
<mods:nameIdentifier type="email">marc.de.meyer@africamuseum.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goergen, Georg</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4496-0495</mods:nameIdentifier>
<mods:affiliation>International Institute of Tropical Agriculture, Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jordaens, Kurt</mods:namePart>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B 3080 Tervuren, Belgium</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1003</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>160</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1003.56557</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1003.56557</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1003-83</mods:identifier>
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<treatment ID-GBIF-Taxon="174795557" LSID="urn:lsid:plazi:treatment:66EEEDF583975B93929D33E91AFC394C" httpUri="http://treatment.plazi.org/id/66EEEDF583975B93929D33E91AFC394C" lastPageNumber="83" pageId="0" pageNumber="83">
<subSubSection pageId="0" pageNumber="83" type="nomenclature">
<paragraph pageId="0" pageNumber="83">
<taxonomicName LSID="66EEEDF5-8397-5B93-929D-33E91AFC394C" authority="(Bigot, 1859)" baseAuthorityName="Bigot" baseAuthorityYear="1859" class="Insecta" family="Syrphidae" genus="Senaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Senaspis nigrita" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nigrita">Senaspis nigrita (Bigot, 1859)</taxonomicName>
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Senaspis species, habitus, lateral view 7 S. nigrita (Bigot) (♂) 8 S. xanthorrhoea (Bezzi) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures7-8" httpUri="https://binary.pensoft.net/fig/486700" pageId="0" pageNumber="83">Figs 7</figureCitation>
<figureCitation captionStart="Figures 3639" captionStartId="F13" captionText="Figures 36 - 39. Head, frontal and lateral view S. nigrita (Bigot) (36, 37 ♂ 38, 39 ♀)." figureDoi="10.3897/zookeys.1003.56557.figures36-39" httpUri="https://binary.pensoft.net/fig/486709" pageId="0" pageNumber="83">, 36-39</figureCitation>
<figureCitation captionStart="Figures 5659" captionStartId="F18" captionText="Figures 56 - 59. Senaspis species, thorax, dorsal view 56 S. flaviceps Macquart (♂) 57 S. haemorrhoa (Gerstaecker) (♂) 58 S. melanthysana (Speiser) (♂) 59 S. nigrita (Bigot) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures56-59" httpUri="https://binary.pensoft.net/fig/486714" pageId="0" pageNumber="83">, 59</figureCitation>
<figureCitation captionStart="Figures 7476" captionStartId="F22" captionText="Figures 74 - 76. Senaspis species, right wing 74 S. melanthysana (Speiser) (♂) 75 S. nigrita (Bigot) (♂) 76 S. nr umbrifera (Walker) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures74-76" httpUri="https://binary.pensoft.net/fig/486718" pageId="0" pageNumber="83">, 75</figureCitation>
<figureCitation captionStart="Figures 8690" captionStartId="F25" captionText="Figures 86 - 90. Senaspis species, hind leg, anterior view 86 S. melanthysana (Speiser) (♂) 87 S. nigrita (Bigot) (♂) 88 S. nr umbrifera (Walker) (♂) 89 S. xanthorrhoea (Bezzi) (♂) 90 S. pennata (Herve-Bazin) (♀)." figureDoi="10.3897/zookeys.1003.56557.figures86-90" httpUri="https://binary.pensoft.net/fig/486721" pageId="0" pageNumber="83">, 87</figureCitation>
<figureCitation captionStart="Figures 97101" captionStartId="F27" captionText="Figures 97 - 101. Senaspis species, abdomen, dorsal view 97 S. nigrita (Bigot) (♂) 98 S. umbrifera (Walker) (♂ holotype) 99 S. nr umbrifera (Walker) (♂) 100 S. xanthorrhoea (Bezzi) (♀) 101 S. pennata (Herve-Bazin) (♀)." figureDoi="10.3897/zookeys.1003.56557.figures97-99" httpUri="https://binary.pensoft.net/fig/486723" pageId="0" pageNumber="83">, 97</figureCitation>
<figureCitation captionStart="Figures 102113" captionStartId="F28" captionText="Figures 102 - 113. Senaspis species, male terminalia 102 - 108, 110, 112 dorsal view 109, 111, 113 lateral view 102 S. dentipes (Macquart) 103 S. dibapha (Walker) 104 S. elliotii Austen 105 S. flaviceps Macquart 106 S. haemorrhoa (Gerstaecker) 107 S. nigrita (Bigot) 108, 109 S. melanthysana (Speiser) 110, 111 S. umbrifera (Walker) (holotype) 112, 113 S. nr umbrifera (Walker)." figureDoi="10.3897/zookeys.1003.56557.figures102-113" httpUri="https://binary.pensoft.net/fig/486724" pageId="0" pageNumber="83">, 107</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="reference_group">
<paragraph pageId="0" pageNumber="83">
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus nigritus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nigritus">Dolichomerus nigritus</taxonomicName>
Bigot, 1859: 431.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="differential diagnosis">
<paragraph pageId="0" pageNumber="83">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="83">
A brown to black-brown species (Fig.
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Senaspis species, habitus, lateral view 7 S. nigrita (Bigot) (♂) 8 S. xanthorrhoea (Bezzi) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures7-8" httpUri="https://binary.pensoft.net/fig/486700" pageId="0" pageNumber="83">7</figureCitation>
) without distinct macula on wing but with darker medial area (Fig.
<figureCitation captionStart="Figures 7476" captionStartId="F22" captionText="Figures 74 - 76. Senaspis species, right wing 74 S. melanthysana (Speiser) (♂) 75 S. nigrita (Bigot) (♂) 76 S. nr umbrifera (Walker) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures74-76" httpUri="https://binary.pensoft.net/fig/486718" pageId="0" pageNumber="83">75</figureCitation>
). It can be diferentiated from other
<taxonomicName authorityName="Macquart" authorityYear="1850" class="Insecta" family="Syrphidae" genus="Senaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Senaspis" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="83">Senaspis</emphasis>
</taxonomicName>
species with dark scutum and similar wing marking (
<taxonomicName lsidName="S. melanthysana" pageId="0" pageNumber="83" rank="species" species="melanthysana">
<emphasis italics="true" pageId="0" pageNumber="83">S. melanthysana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="S. umbrifera" pageId="0" pageNumber="83" rank="species" species="umbrifera">
<emphasis italics="true" pageId="0" pageNumber="83">S. umbrifera</emphasis>
</taxonomicName>
) by the contrasting yellow scutellum (Fig.
<figureCitation captionStart="Figures 5659" captionStartId="F18" captionText="Figures 56 - 59. Senaspis species, thorax, dorsal view 56 S. flaviceps Macquart (♂) 57 S. haemorrhoa (Gerstaecker) (♂) 58 S. melanthysana (Speiser) (♂) 59 S. nigrita (Bigot) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures56-59" httpUri="https://binary.pensoft.net/fig/486714" pageId="0" pageNumber="83">59</figureCitation>
) (scutellum largely concolourous with scutum, at least along basal half in other species (Figs
<figureCitation captionStart="Figures 5659" captionStartId="F18" captionText="Figures 56 - 59. Senaspis species, thorax, dorsal view 56 S. flaviceps Macquart (♂) 57 S. haemorrhoa (Gerstaecker) (♂) 58 S. melanthysana (Speiser) (♂) 59 S. nigrita (Bigot) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures56-59" httpUri="https://binary.pensoft.net/fig/486714" pageId="0" pageNumber="83">58</figureCitation>
,
<figureCitation captionStart="Figures 6063" captionStartId="F19" captionText="Figures 60 - 63. Senaspis species, thorax, dorsal view 60 S. nr umbrifera (Walker) (♂) 61 S. umbrifera (Walker) (♂ holotype) 62 S. xanthorrhoea (Bezzi) (♂) 63 S. pennata (Herve-Bazin) (♀)." figureDoi="10.3897/zookeys.1003.56557.figures60-63" httpUri="https://binary.pensoft.net/fig/486715" pageId="0" pageNumber="83">61</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="materials_examined">
<paragraph pageId="0" pageNumber="83">Examined material.</paragraph>
<paragraph pageId="0" pageNumber="83">
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus nigritus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nigritus">
<emphasis italics="true" pageId="0" pageNumber="83">Dolichomerus nigritus</emphasis>
</taxonomicName>
Bigot:
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Lectotype</emphasis>
(hereby designated), female, &quot;Syn- // type&quot; &quot;ex. coll. Bigot. // Prs. by // G.H. Verrall. // B.M. 1901-14.&quot; &quot;SYNTYPE ♀ of //
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">Dolichomerus</taxonomicName>
//
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus nigritus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nigritus">Dolichomerus nigritus</taxonomicName>
Bigot // MADAGASCAR&quot; &quot;BMNH(E) # // 230785&quot;
<normalizedToken originalValue="“NHMUK010369878”">&quot;NHMUK010369878&quot;</normalizedToken>
<normalizedToken originalValue="“LECTOTYPUS”">&quot;LECTOTYPUS&quot;</normalizedToken>
[NHMUK].
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Paralectotype</emphasis>
, female, &quot;Syn- / /type&quot; &quot;SYNTYPE ♀ of //
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">Dolichomerus</taxonomicName>
//
<taxonomicName class="Insecta" family="Syrphidae" genus="Dolichomerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichomerus nigritus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="nigritus">Dolichomerus nigritus</taxonomicName>
Bigot // MADAGASCAR&quot; &quot;ex. coll. Bigot. // Pres. by // G.H. Verrall. // B.M.1901-14.&quot; &quot;BMNH(E) # // 230786&quot; &quot;PARA- // LECTOTYPUS&quot; [NHMUK].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="materials_examined">
<paragraph pageId="0" pageNumber="83">Other material.</paragraph>
<paragraph pageId="0" pageNumber="83">
Madagascar • 2♂♂ 1♀; 5 km N Ampotaka, Beloha; 5-15 Mar. 2009; M. Irwin and R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
leg.; CDFA • 1♀; Ambatobevandza; 1919; R. Decary leg.; MNHN • 1♀; Ambatolampy; 1931;
<normalizedToken originalValue="Lasère">Lasere</normalizedToken>
leg.; MNHN • 1♀; Ambatosoratra; Nov. 1960; P. Soga leg.; MNHN • 1♀; Ambodivoangy; Jan. 1967; J. Vadon leg.; KMMA • 1♀; Ambre; Oct. 1902; MNHN • 1♀; Ampassamadinika, Tamatave; 26 Jan. 1992; A. Pauly leg.; KMMA • 1♀; Analmalotra; 1 Nov. 1993; C. Kassebeer leg.; CDFA • 1♀; Andasibe National Park; 17 Dec. 2017; Schmid-Egger leg.; CDFA • 1♀; Andranobe Forest, route
<normalizedToken originalValue="dAndriamena">d'Andriamena</normalizedToken>
; MNHN • 1♀; Anja Reserve, 10 km SSW Ambalavao; 24 Dec. 2017; Schmid-Egger leg.; CDFA • 1♀; Ankaratra Massif, Manjakatompo; Dec. 1951; Benoist leg.; KMMA • 1♂; Ankaratra Massif, Manjakatompo; Jan. 1958; B. Stuckenberg leg.; NMSA • 1♂;
<normalizedToken originalValue="Anosibé">Anosibe</normalizedToken>
; Breuning leg.; KMMA • 1♀; Bekily; Mar. 1930; A. Seyrig leg.; MNHN • 1♂; same collection data as for preceding; May 1933 • 1♂; same collection data as for preceding; Jun. 1933 • 1♂ 5♀♀; Belambo, 20 km NW Boriziny, Mahajanga; 4 Jan. 2007; A.H. Kirk-Spriggs leg.; AMGS • 1♀; Betafo, Ambositra and Miandrivazo regions, Central; 1905; Bouet leg.; MNHN • 1♀; Betsileo; NHMUK • 1♀; Brickaville; Jul. 1959; Sigwalt leg.; MNHN • 2♂♂ 3♀♀; Bevilany, Tulear, 12 Apr. 1968; K.M.G. and P.D. leg.; NHMUK • 1♀; same collection data as for preceding; 14-18 Apr. 1968 • 1♂; Didy; 12 Apr. 1992; A. Pauly leg.; KMMA • 1♂; same collection data as for preceding; 16 Apr. 1992 • 1♀; Fampanambo; 1962; J. Vadon leg.; KMMA • 1♀; Fianarantsoa; A. Bigneux leg.; CNC • 2♀♀; Forest Belt; S. Hutchins leg.; NHMUK • 1♀; Fort Dauphin [= Taolagnaro]; NHMUK • 1♀; Foulpointe [= Mahavelona]; 12 Nov. 1993; A. Pauly leg.; KMMA • 1♂ 2♀♀; same collection data as for preceding; May 1995 • 1♂ 5♀♀; Imerina, Andrangoloaka Forest; A. Grandidier leg.; MNHN • 1♀; Ivoloina; Mar. 1960; Sigwalt leg.; MNHN • 1♂; Lac Froid, Ambatolampy; 11-15 Dec. 1957; B. Stuckenberg leg.; NMSA • 1♀; 7 km N Joffreville; 20 Mar.-7 Apr. 2001; R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
leg.; CDFA • 1♂ 2♀♀; Mananjara Province; 1910; Goissaud leg.; MNHN • 1♀; Maroantsetra; J. Vadon leg.; MNHN • 1♀; same collection data as for preceding; Nov. 1959; KMMA • 1♀; Montagne
<normalizedToken originalValue="dAmbre">d'Ambre</normalizedToken>
; L. Chopard leg.; MNHN • 1♀; Ranomafana National Park; 21 Dec. 2017; Schmid-Egger leg.; CDFA • 1♀; Ranomafana National Park, Radio Tower; 16 Nov. 2001; M.E. Irwin; F.D. Parker and R.
<normalizedToken originalValue="Harinhala">Harin'hala</normalizedToken>
leg.; CDFA • 3♀♀; Ranomafana National Park, ValBio Research Centre; 15-17 Jan. 2007; A.H. Kirk-Spriggs leg.; AMGS • 4♀♀; same collection data as for preceding; 13 Jan. 2014; M. Hauser leg.; CDFA • 3♀♀; Ranomafana National Park, Talatakely; 11 June 2014; A.H. Kirk-Spriggs and R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
leg.; BMSA • 1♀; Sakalava Beach; 7-25 Jun. 2001; R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
leg.; CDFA • 1♀; Tananarive; 1897; E. Dorr leg.; MNHN • 1♂ 5♀♀; same collection data as for preceding; 1914; G. Waterlot leg.; • 1♂ 1♀; same collection data as for preceding; 1916 • 3♀♀; same collection data as for preceding; 1919 • 1♀; same collection data as for preceding; 22 Dec. 1920; R. Decary leg. • 1♂; same collection data as for preceding; 29 Dec. 1920 • 1♀; same collection data as for preceding; 6 Jan. 1921 • 1♀; same collection data as for preceding; 17 Jan. 1921 • 1♀; same collection data as for preceding; 1921 • 1♀; same collection data as for preceding; Jan. 1922 • 1♀; same collection data as for preceding; 1931;
<normalizedToken originalValue="Lasère">Lasere</normalizedToken>
leg. • 2♂♂; Tananarive; Nov. 1952; E.S. Brown leg.; NHMUK • 1♂ 1♀; Tananarive; Jan. 1958; B. Stuckenberg leg.; NMSA • 1♀; Tananarive; 8 Jan. 1971; H May Daly leg.; NHMUK • 1♀; Tananarive; [no date]; MNHN • 1♀; same collection data as for preceding; KMMA • 1♀; Tananarive; [no date]; Le Moult leg.; MNHN • 1♀; same collection data as for preceding; Leray leg. • 1♀; Tsimbazaza, Antanan. Park; 16-22 Oct. 1993; C. Kassebeer leg.; CNC • 2♂♂; Vitambani Forest, 5 km N Ampotaka, Beloha; 12 Aug.-29 Sep. 2009; M. Irwin and
<normalizedToken originalValue="Rinha">Rin'ha</normalizedToken>
leg.; CDFA • 1♂; [no locality]; 1894; P.
<normalizedToken originalValue="Camboué">Camboue</normalizedToken>
leg.; MNHN• 1♀; 1898; Delcroix leg.; MNHN • 1♀; 1906; MNHN • F. Geay leg.; MNHN • 2♂♂ 13♀♀; P.
<normalizedToken originalValue="Camboué">Camboue</normalizedToken>
leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="description">
<paragraph pageId="0" pageNumber="83">Description.</paragraph>
<paragraph pageId="0" pageNumber="83">Body length: 14.2-18.2 mm. Wing length: 8.7-13.5 mm.</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">Male</emphasis>
(Fig.
<figureCitation captionStart="Figures 7, 8" captionStartId="F4" captionText="Figures 7, 8. Senaspis species, habitus, lateral view 7 S. nigrita (Bigot) (♂) 8 S. xanthorrhoea (Bezzi) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures7-8" httpUri="https://binary.pensoft.net/fig/486700" pageId="0" pageNumber="83">7</figureCitation>
). Head (Figs
<figureCitation captionStart="Figures 3639" captionStartId="F13" captionText="Figures 36 - 39. Head, frontal and lateral view S. nigrita (Bigot) (36, 37 ♂ 38, 39 ♀)." figureDoi="10.3897/zookeys.1003.56557.figures36-39" httpUri="https://binary.pensoft.net/fig/486709" pageId="0" pageNumber="83">36</figureCitation>
,
<figureCitation captionStart="Figures 3639" captionStartId="F13" captionText="Figures 36 - 39. Head, frontal and lateral view S. nigrita (Bigot) (36, 37 ♂ 38, 39 ♀)." figureDoi="10.3897/zookeys.1003.56557.figures36-39" httpUri="https://binary.pensoft.net/fig/486709" pageId="0" pageNumber="83">37</figureCitation>
). Eye bare; holoptic, eye contiguity for distance equal to length of ocellar triangle, facets dorsally slightly larger, approx. three times as large in diameter as ventral ones. Frons black-brown; subshiny with brownish pollinosity; long pale yellow pile, especially dorsally and along eye margin. Face black-brown; subshiny with pale brownish to greyish pollinosity, in parts more densely so, ventral lateral margins largely devoid of pollinosity; in parts with dispersed long pale pile; facial tubercle strongly pronounced. Gena colour and pollinosity as ventral lateral margins of face; with short to long pale pile. Occiput black-brown, covered with grey pollinosity; with dispersed pale pile except dorsally where sometimes darker yellow. Antennal segments black-brown, basoflagellomere apically more brown, arista orange-brown.
</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Thorax</emphasis>
(Fig.
<figureCitation captionStart="Figures 5659" captionStartId="F18" captionText="Figures 56 - 59. Senaspis species, thorax, dorsal view 56 S. flaviceps Macquart (♂) 57 S. haemorrhoa (Gerstaecker) (♂) 58 S. melanthysana (Speiser) (♂) 59 S. nigrita (Bigot) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures56-59" httpUri="https://binary.pensoft.net/fig/486714" pageId="0" pageNumber="83">59</figureCitation>
). Scutum subshiny black, weak brownish pollinose; with short pale brown pile in anterior half, posterior part more black pile; postpronotum and notopleuron with longer mixed pale and dark pile. Scutellum apical margin weakly rounded, distinctly marginated, three times as wide as long; pale yellow to straw yellow, anterior margin narrowly black-brown; with short pale pile, along posterior margin somewhat longer. Pleura ground colour black-brown, covered with dispersed long pale or dark pile except on meron, dorsomedial anepimeron, ventral part of katepimeron, anterior part of katepisternum and anterior anepisternum.
</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Legs</emphasis>
. Brown to black-brown, sometimes more rufous; with short black pile, along posterior margin of pro- and mesofemora with longer pale pile, along ventral margin of metafemur with longer dark pile. Metaleg (Fig.
<figureCitation captionStart="Figures 8690" captionStartId="F25" captionText="Figures 86 - 90. Senaspis species, hind leg, anterior view 86 S. melanthysana (Speiser) (♂) 87 S. nigrita (Bigot) (♂) 88 S. nr umbrifera (Walker) (♂) 89 S. xanthorrhoea (Bezzi) (♂) 90 S. pennata (Herve-Bazin) (♀)." figureDoi="10.3897/zookeys.1003.56557.figures86-90" httpUri="https://binary.pensoft.net/fig/486721" pageId="0" pageNumber="83">87</figureCitation>
), femur slightly thickened, with two distinct ventral swellings in apical third, swellings with dispersed short thick setae; tibia bent at base, thickened and slightly curved, pile along ventral margin in apical half to two-thirds longer and more dense.
</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Wing</emphasis>
(Fig.
<figureCitation captionStart="Figures 7476" captionStartId="F22" captionText="Figures 74 - 76. Senaspis species, right wing 74 S. melanthysana (Speiser) (♂) 75 S. nigrita (Bigot) (♂) 76 S. nr umbrifera (Walker) (♂)." figureDoi="10.3897/zookeys.1003.56557.figures74-76" httpUri="https://binary.pensoft.net/fig/486718" pageId="0" pageNumber="83">75</figureCitation>
). Faint yellowish tinge, near cross-veins r-m and bm-cu more yellow-brown but no distinct medial macula; distal part of cells r2+3 and r4+5 more hyaline, as well as along posterior margin, paler areas not distinctly demarcated; alula largely hyaline. Calypters dull straw yellow with fringe of silvery white pile. Cell r1 closed, petiole very short, at most half height of base of stigma, rarely longer. Vein R4+5 sinuate, appendix absent or present.
</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figures 97101" captionStartId="F27" captionText="Figures 97 - 101. Senaspis species, abdomen, dorsal view 97 S. nigrita (Bigot) (♂) 98 S. umbrifera (Walker) (♂ holotype) 99 S. nr umbrifera (Walker) (♂) 100 S. xanthorrhoea (Bezzi) (♀) 101 S. pennata (Herve-Bazin) (♀)." figureDoi="10.3897/zookeys.1003.56557.figures97-99" httpUri="https://binary.pensoft.net/fig/486723" pageId="0" pageNumber="83">97</figureCitation>
). Uniformly subshiny brown to black-brown, sometimes more rufous; with short dark pile except tergum I where more densely silver-grey pile; lateral margins intermixed with short pale pile, especially near postabdomen. Terga with dispersed long pale pile. Male genitalia as in Fig.
<figureCitation captionStart="Figures 102113" captionStartId="F28" captionText="Figures 102 - 113. Senaspis species, male terminalia 102 - 108, 110, 112 dorsal view 109, 111, 113 lateral view 102 S. dentipes (Macquart) 103 S. dibapha (Walker) 104 S. elliotii Austen 105 S. flaviceps Macquart 106 S. haemorrhoa (Gerstaecker) 107 S. nigrita (Bigot) 108, 109 S. melanthysana (Speiser) 110, 111 S. umbrifera (Walker) (holotype) 112, 113 S. nr umbrifera (Walker)." figureDoi="10.3897/zookeys.1003.56557.figures102-113" httpUri="https://binary.pensoft.net/fig/486724" pageId="0" pageNumber="83">107</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">Female.</emphasis>
As male except for the following character states: Eye dichoptic (Figs
<figureCitation captionStart="Figures 3639" captionStartId="F13" captionText="Figures 36 - 39. Head, frontal and lateral view S. nigrita (Bigot) (36, 37 ♂ 38, 39 ♀)." figureDoi="10.3897/zookeys.1003.56557.figures36-39" httpUri="https://binary.pensoft.net/fig/486709" pageId="0" pageNumber="83">38</figureCitation>
,
<figureCitation captionStart="Figures 3639" captionStartId="F13" captionText="Figures 36 - 39. Head, frontal and lateral view S. nigrita (Bigot) (36, 37 ♂ 38, 39 ♀)." figureDoi="10.3897/zookeys.1003.56557.figures36-39" httpUri="https://binary.pensoft.net/fig/486709" pageId="0" pageNumber="83">39</figureCitation>
), facets subequal in size. Frons and face pollinosity predominantly greyish; frons with short dark pile dorsally, greyish pile ventrally; with narrow pale fascia dorsally of antennal implant. Thorax dark pile sometimes more extensively so. Leg pilosity sometimes more brownish.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="distribution">
<paragraph pageId="0" pageNumber="83">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="83">
Madagascar. Records from Tanzania (
<bibRefCitation DOI="https://doi.org/10.1002/mmnd.18880310225" author="Karsch, FA" journalOrPublisher="Entomologische Zeitschrift" pageId="0" pageNumber="83" pagination="367 - 382" refId="B31" refString="Karsch, FA, 1888. Bericht ueber die durch Herrn Lieutenant Dr Carl Wilhelm Schmidt in Ost-Afrika gesammelten und von der zoologischen Abtheilung des Koeniglichen Museums fuer Naturkunde in Berlin erworbenen Dipteren. Entomologische Zeitschrift 31: 367 - 382, DOI: https://doi.org/10.1002/mmnd.18880310225" title="Bericht ueber die durch Herrn Lieutenant Dr Carl Wilhelm Schmidt in Ost-Afrika gesammelten und von der zoologischen Abtheilung des Koeniglichen Museums fuer Naturkunde in Berlin erworbenen Dipteren." url="https://doi.org/10.1002/mmnd.18880310225" volume="31" year="1888">Karsch 1888</bibRefCitation>
;
<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="83" pagination="488 - 510" refId="B46" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth 1980</bibRefCitation>
) and Uganda (
<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="83" pagination="488 - 510" refId="B46" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth 1980</bibRefCitation>
) could not be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>