treatments-xml/data/55/0B/87/550B87C3C15AFF83FF3C745DFE0D03CD.xml
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<document id="C921361A7697FC5E42D7337BB2EE5B75" ID-DOI="10.5281/zenodo.4651080" ID-ISSN="1638-9395" ID-Zenodo-Dep="4651080" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617195131926" checkinUser="felipe" docAuthor="Bianucci, Giovanni, Lambert, Olivier &amp; Post, Klaas" docDate="2007" docId="550B87C3C15AFF83FF3C745DFE0D03CD" docLanguage="en" docName="Geodiversitas.29.4.561-618.pdf" docOrigin="Geodiversitas 29 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Pterocetus benguelae Bianucci &amp; Lambert &amp; Post 2007" docType="treatment" docVersion="3" lastPageNumber="598" masterDocId="A932FFBBC17EFFA5FFC47277FFBE056E" masterDocTitle="A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa" masterLastPageNumber="618" masterPageNumber="561" pageNumber="597" updateTime="1698897450673" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="1B77965EB04CC7A081253AF70E43D761">A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa</mods:title>
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<mods:namePart id="C38617B2C438D63BB50A8F62F1F67912">Bianucci, Giovanni</mods:namePart>
<mods:affiliation id="584A5F5573B3ECF410B04BE1C54DD9B2">Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria, 53, I- 56126 Pisa (Italy) bianucci @ dst. unipi. it</mods:affiliation>
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<mods:namePart id="DBAA5C6E40390BC0DD8964D43A6AD0E5">Lambert, Olivier</mods:namePart>
<mods:affiliation id="C2618BD69D94D4847E8905363A3269CA">Institut royal des Sciences naturelles de Belgique, Département de Paléontologie, rue Vautier, 29, B- 1000 Brussels (Belgium) olivier. lambert @ naturalsciences. be</mods:affiliation>
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<mods:namePart id="E082DC8AAD16AAF5347EA64F014032EF">Post, Klaas</mods:namePart>
<mods:affiliation id="783469964375EFB349C362FB32A72D6B">Natuurhistorisch Museum Rotterdam, P. O. Box 23452, NL- 3001 Rotterdam (The Netherlands) klaas @ fiskano. nl</mods:affiliation>
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<mods:title id="46F3B7F65EBEB22C33F1246F498A1F8D">Geodiversitas</mods:title>
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<mods:date id="C570F3EE8124D6DC4527DC976DAAE364">2007</mods:date>
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<paragraph id="DD1D36D5C15AFF81FF3C745DFDBE032B" blockId="36.[248,512,1578,1605]" box="[248,512,1578,1605]" pageId="36" pageNumber="597">
<heading id="865581B9C15AFF81FF3C745DFDBE032B" box="[248,512,1578,1605]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="597" reason="2">
<taxonomicName id="1AA24D56C15AFF81FF3C745DFDBE032B" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[248,512,1578,1605]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="benguelae">
<emphasis id="EFD6EAC7C15AFF81FF3C745DFDBE032B" bold="true" box="[248,512,1578,1605]" italics="true" pageId="36" pageNumber="597">Pterocetus aff. benguelae</emphasis>
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<paragraph id="DD1D36D5C15AFF81FFB5742BFD3803CC" blockId="36.[113,647,1627,1698]" pageId="36" pageNumber="597">
REFERRED SPECIMENS. — SAM PQ 1770, partial skull including the rostrum and most of the dorsal surface of the cranium with vertex, trawled
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west of Slang-
</paragraph>
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<subSubSection id="95B8655EC15AFF83FD617619FE0D03CD" lastPageId="38" lastPageNumber="599" pageId="36" pageNumber="597" type="description">
<paragraph id="DD1D36D5C15AFF81FD617619FBE501F3" blockId="36.[677,1209,1134,1182]" pageId="36" pageNumber="597">
kop, Atlantic Ocean, depth of
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; SAM PQ 2751, rostrum, trawled off the South African coast.
</paragraph>
<paragraph id="DD1D36D5C15AFF81FD6176B9FC510188" blockId="36.[677,1210,1230,1699]" box="[677,1007,1230,1256]" pageId="36" pageNumber="597">DESCRIPTION AND DISCUSSION</paragraph>
<paragraph id="DD1D36D5C15AFF81FD617698FC6F0067" blockId="36.[677,1210,1230,1699]" box="[677,977,1262,1289]" pageId="36" pageNumber="597">
(FIGS 28; 29C, D;
<tableCitation id="9020036EC15AFF81FCAB7699FC760067" box="[879,968,1262,1289]" captionStart="TABLE" captionStartId="54.[113,122,220,237]" captionText="TABLE 5. — Measurements (in mm) on the skulls of Ziphiidae incertae sedis from the Neogene of South Africa. Abbreviations: e, estimate; +, nearly complete; -, no data." pageId="36" pageNumber="597">TABLE 5</tableCitation>
)
</paragraph>
<paragraph id="DD1D36D5C15AFF81FD61777AFB0403CD" blockId="36.[677,1210,1230,1699]" pageId="36" pageNumber="597">
The general shape and the anterior position of the premaxillary foramen cause us to refer the two specimens to
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<emphasis id="EFD6EAC7C15AFF81FCB3773AFC5A0008" box="[887,996,1357,1382]" italics="true" pageId="36" pageNumber="597">Pterocetus</emphasis>
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<taxonomicNameLabel id="F4E557BCC15AFF81FC29773AFB800009" box="[1005,1086,1357,1383]" pageId="36" pageNumber="597" rank="genus">n. gen.</taxonomicNameLabel>
Moreover, SAM PQ 1770 shows the deep and wide antorbital notch characteristic for
<taxonomicName id="1AA24D56C15AFF81FC3177FBFBDF00CB" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[1013,1121,1420,1445]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="36" pageNumber="597" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C15AFF81FC3177FBFBDF00CB" box="[1013,1121,1420,1445]" italics="true" pageId="36" pageNumber="597">Pterocetus</emphasis>
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<taxonomicNameLabel id="F4E557BCC15AFF81FBAE77FBFB0400C8" box="[1130,1210,1420,1446]" pageId="36" pageNumber="597" rank="genus">n. gen.</taxonomicNameLabel>
Nevertheless, both specimens differ from those referred above to
<taxonomicName id="1AA24D56C15AFF81FCB177BBFBE6008B" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[885,1112,1483,1509]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="benguelae" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C15AFF81FCB177BBFBE6008B" box="[885,1112,1483,1509]" italics="true" pageId="36" pageNumber="597">Pterocetus benguelae</emphasis>
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<taxonomicNameLabel id="F4E557BCC15AFF81FBA777BBFD5D036B" pageId="36" pageNumber="597" rank="species">n. gen., n. sp.</taxonomicNameLabel>
in their lower rostrum with an anteriorly flatter ventral surface. This difference in robustness might correspond to an intraspecific sexual dimorphism, as observed in the extant
<taxonomicName id="1AA24D56C15AFF81FBFB743DFCA403ED" baseAuthorityName="Blainville" baseAuthorityYear="1817" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="36" pageNumber="597" phylum="Chordata" rank="species" species="densirostris">
<emphasis id="EFD6EAC7C15AFF81FBFB743DFCA403ED" italics="true" pageId="36" pageNumber="597">Mesoplodon densirostris</emphasis>
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, whose rostrum is considerably more robust in the males than in the females (
<figureCitation id="45992A50C15AFF81FB9F74FEFB1203CD" box="[1115,1196,1673,1699]" captionStart="FIG" captionStartId="38.[113,124,1300,1317]" captionTargetBox="[196,1127,822,1254]" captionTargetId="figure-84@38.[123,1204,810,1260]" captionTargetPageId="38" captionText="FIG. 30. — A, B, rostrum of male of Mesoplodon densirostris (SAM ZM 40663); A, dorsal view; B, lateral view; C, D, rostrum of female of M. densirostris (SAM ZM 40836); C, dorsal view; D, lateral view. Reduced to the same lenght." figureDoi="http://doi.org/10.5281/zenodo.4651145" httpUri="https://zenodo.org/record/4651145/files/figure.png" pageId="36" pageNumber="597">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="DD1D36D5C15BFF80FCC977EDFB8A00C2" blockId="37.[113,1209,1433,1543]" box="[781,1076,1433,1453]" pageId="37" pageNumber="598">mesethmoid bony nares</paragraph>
<paragraph id="DD1D36D5C15BFF80FFB57797FCA50369" blockId="37.[113,1209,1433,1543]" pageId="37" pageNumber="598">
FIG. 28. — Skull of
<taxonomicName id="1AA24D56C15BFF80FED47797FDD1009F" authority="(SAM PQ 1770)" baseAuthorityName="SAM PQ" baseAuthorityYear="1770" box="[272,623,1504,1521]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="37" pageNumber="598" phylum="Chordata" rank="species" species="benguelae">
<emphasis id="EFD6EAC7C15BFF80FED47797FED4009F" box="[272,362,1504,1521]" italics="true" pageId="37" pageNumber="598">Pterocetus</emphasis>
aff.
<emphasis id="EFD6EAC7C15BFF80FE4B7797FE5B009F" box="[399,485,1504,1521]" italics="true" pageId="37" pageNumber="598">benguelae</emphasis>
(SAM PQ 1770)
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:
<emphasis id="EFD6EAC7C15BFF80FDB27797FD3D009F" bold="true" box="[630,643,1504,1521]" pageId="37" pageNumber="598">A</emphasis>
, dorsal view;
<emphasis id="EFD6EAC7C15BFF80FD337797FCBA009F" bold="true" box="[759,772,1504,1521]" pageId="37" pageNumber="598">B</emphasis>
, anterior view;
<emphasis id="EFD6EAC7C15BFF80FC417797FC2D009F" bold="true" box="[901,915,1504,1521]" pageId="37" pageNumber="598">C</emphasis>
, lateral view;
<emphasis id="EFD6EAC7C15BFF80FBC17797FBAD009F" bold="true" box="[1029,1043,1504,1521]" pageId="37" pageNumber="598">D</emphasis>
, detail of the vertex in dorsal view;
<emphasis id="EFD6EAC7C15BFF80FF2A7781FF440369" bold="true" box="[238,250,1526,1543]" pageId="37" pageNumber="598">E</emphasis>
, corresponding line drawing. Scale bars: A-C,
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; D, E,
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.
</paragraph>
<paragraph id="DD1D36D5C15BFF83FFB5743DFE0D03CD" blockId="37.[113,646,1610,1700]" lastBlockId="38.[113,647,1420,1700]" lastPageId="38" lastPageNumber="599" pageId="37" pageNumber="598">
However, in this case the difference in robustness of the rostrum is reinforced by the strong mesorostral ossification of the vomer, mostly present in the adult male, while both the robust and slender fossil rostra described above exhibit a complete mesorostral ossification. Other small differences are observed between SAM PQ 1770 and the
<typeStatus id="02198877C158FF83FF0977DCFE9100AB" box="[205,303,1451,1477]" pageId="38" pageNumber="599" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1AA24D56C158FF83FE9777DBFD9500AB" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[339,555,1451,1477]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="38" pageNumber="599" phylum="Chordata" rank="species" species="benguelae" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C158FF83FE9777DBFD9500AB" box="[339,555,1451,1477]" italics="true" pageId="38" pageNumber="599">Pterocetus benguelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC158FF83FDF777DBFF130088" pageId="38" pageNumber="599" rank="species">n. gen., n. sp.</taxonomicNameLabel>
at the level of the vertex: the nasals are narrower, the bony nares are smaller, the premaxillary crest seems laterally less developed, and the constriction of the ascending process is almost absent. Considering these differences, referral of SAM PQ 1770 and SAM PQ 2751 to
<taxonomicName id="1AA24D56C158FF83FDD9741DFEBD03CD" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="38" pageNumber="599" phylum="Chordata" rank="species" species="benguelae">
<emphasis id="EFD6EAC7C158FF83FDD9741DFD3803ED" box="[541,646,1642,1667]" italics="true" pageId="38" pageNumber="599">Pterocetus</emphasis>
aff.
<emphasis id="EFD6EAC7C158FF83FF5874FEFEBD03CD" box="[156,259,1673,1699]" italics="true" pageId="38" pageNumber="599">benguelae</emphasis>
</taxonomicName>
seems justified.
</paragraph>
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