207 lines
28 KiB
XML
207 lines
28 KiB
XML
<document id="DB2975F9EE65E307B1D32056C48920F5" ID-CLB-Dataset="27761" ID-DOI="10.11646/zootaxa.4532.1.3" ID-GBIF-Dataset="114a9cd5-ae28-4133-a31e-e847c3e4f316" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615023" ID-ZooBank="A6128B92-0B20-4D4D-AE8B-483D39BB2C04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787707609" checkinUser="plazi" docAuthor="Thandar, Ahmed S." docDate="2018" docId="2A0887A599095E35BCF8FC8EFF3CFE22" docLanguage="en" docName="zootaxa.4532.1.3.pdf" docOrigin="Zootaxa 4532 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psolus imperfectus H. L. Clark 1923" docType="treatment" docVersion="18" lastPageNumber="79" masterDocId="D631FFDD991F5E22BC6FFFCAFFE4FF8E" masterDocTitle="On some miscellaneous sea cucumbers (Echinodermata: Holothuroidea) in the collections of the South African Museum with three new species" masterLastPageNumber="85" masterPageNumber="57" pageNumber="79" updateTime="1701998753008" updateUser="plazi">
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<mods:title id="1D15F8F826FD967C36E0853033298FC3">On some miscellaneous sea cucumbers (Echinodermata: Holothuroidea) in the collections of the South African Museum with three new species</mods:title>
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<mods:namePart id="CBACE8260B4799D3A48563F598C9497F">Thandar, Ahmed S.</mods:namePart>
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<mods:title id="F3635088B7A5D3F866921D08F6102066">Zootaxa</mods:title>
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<mods:part id="8D5993E8AB1CC84BF7569A2B32056BF8">
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<mods:date id="9246833F82B66798FE8BF217A7CE6861">2018</mods:date>
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<mods:number id="3326DD48A354C5B3D6A0E058994F5288">2018-12-17</mods:number>
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<treatment id="2A0887A599095E35BCF8FC8EFF3CFE22" ID-DOI="http://doi.org/10.5281/zenodo.5107320" ID-GBIF-Taxon="154521184" ID-Zenodo-Dep="5107320" LSID="urn:lsid:plazi:treatment:2A0887A599095E35BCF8FC8EFF3CFE22" httpUri="http://treatment.plazi.org/id/2A0887A599095E35BCF8FC8EFF3CFE22" lastPageId="23" lastPageNumber="79" pageId="22" pageNumber="79">
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<subSubSection id="EABB653899095E34BCF8FC8EFDF2FCD0" box="[151,534,836,862]" pageId="22" pageNumber="79" type="nomenclature">
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<paragraph id="A21E36B399095E34BCF8FC8EFDF2FCD0" blockId="22.[151,534,836,895]" box="[151,534,836,862]" pageId="22" pageNumber="79">
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<heading id="F95681DF99095E34BCF8FC8EFDF2FCD0" bold="true" box="[151,534,836,862]" fontSize="11" level="1" pageId="22" pageNumber="79" reason="1">
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<taxonomicName id="65A14D3099095E34BCF8FC8EFDF2FCD0" ID-CoL="4PL99" authority="Clark, 1923" authorityName="H. L. Clark" authorityYear="1923" box="[151,534,836,862]" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="79" phylum="Echinodermata" rank="species" species="imperfectus">
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<emphasis id="90D5EAA199095E34BCF8FC8EFDF2FCD0" bold="true" box="[151,534,836,862]" pageId="22" pageNumber="79">
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<emphasis id="90D5EAA199095E34BCF8FC8EFE98FCD0" bold="true" box="[151,380,836,862]" italics="true" pageId="22" pageNumber="79">Psolus imperfectus</emphasis>
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<bibRefCitation id="C6304B4299095E34BDECFC8EFDF2FCD0" author="Clark, H. L." box="[387,534,836,862]" pageId="22" pageNumber="79" pagination="221 - 435" refId="ref14325" refString="Clark, H. L. (1923) The echinoderm fauna of South Africa. Annals of the South African Museum, 13, 221 - 435. Available from: https: // www. biodiversitylibrary. org / page / 1541222 # page / 1 / mode / 1 up (Accessed 14 Dec. 2018)" type="journal article" year="1923">Clark, 1923</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EABB653899095E34BCF8FCACFEF2FCF1" box="[151,278,870,895]" pageId="22" pageNumber="79" type="description">
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<paragraph id="A21E36B399095E34BCF8FCACFEF2FCF1" blockId="22.[151,534,836,895]" box="[151,278,870,895]" pageId="22" pageNumber="79">
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(
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<figureCitation id="3A9A2A3699095E34BCF1FCACFEE8FCF1" box="[158,268,870,895]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="EABB653899095E34BCF8FC64FD3DFC6D" pageId="22" pageNumber="79" type="reference_group">
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<paragraph id="A21E36B399095E34BCF8FC64FDDAFC4A" blockId="22.[151,729,942,995]" box="[151,574,942,964]" pageId="22" pageNumber="79">
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<treatmentCitationGroup id="82B1119D99095E34BCF8FC64FDDAFC4A" box="[151,574,942,964]" pageId="22" pageNumber="79">
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<taxonomicName id="65A14D3099095E34BCF8FC64FDDDFC4A" ID-CoL="4PL99" authority="H. L. Clark, 1923: 418" authorityName="H. L. Clark" authorityPageNumber="418" authorityYear="1923" box="[151,569,942,964]" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="79" phylum="Echinodermata" rank="species" species="imperfectus">
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<emphasis id="90D5EAA199095E34BCF8FC64FEB3FC4A" box="[151,343,942,964]" italics="true" pageId="22" pageNumber="79">Psolus imperfectus</emphasis>
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<treatmentCitation id="230010A299095E34BD32FC64FDDDFC4A" author="Clark, H. L." box="[349,569,942,964]" page="418" pageId="22" pageNumber="79" year="1923">
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<bibRefCitation id="C6304B4299095E34BD32FC64FDE2FC4A" author="Clark, H. L." box="[349,518,942,964]" pageId="22" pageNumber="79" pagination="221 - 435" refId="ref14325" refString="Clark, H. L. (1923) The echinoderm fauna of South Africa. Annals of the South African Museum, 13, 221 - 435. Available from: https: // www. biodiversitylibrary. org / page / 1541222 # page / 1 / mode / 1 up (Accessed 14 Dec. 2018)" type="journal article" year="1923">H.L.Clark, 1923</bibRefCitation>
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: 418
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A21E36B399095E34BCF8FC07FD3DFC6D" blockId="22.[151,729,942,995]" box="[151,729,973,995]" pageId="22" pageNumber="79">
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<treatmentCitationGroup id="82B1119D99095E34BCF8FC07FD3DFC6D" box="[151,729,973,995]" pageId="22" pageNumber="79">
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<taxonomicName id="65A14D3099095E34BCF8FC07FDA2FC6D" ID-CoL="4PL99" authority="Deichmann, 1948: 363" authorityName="Deichmann" authorityPageNumber="363" authorityYear="1948" box="[151,582,973,995]" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="79" phylum="Echinodermata" rank="species" species="imperfectus">
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<emphasis id="90D5EAA199095E34BCF8FC07FEB3FC6D" box="[151,343,973,995]" italics="true" pageId="22" pageNumber="79">Psolus imperfectus</emphasis>
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<treatmentCitation id="230010A299095E34BD32FC07FDA2FC6D" author="Deichmann, E." box="[349,582,973,995]" page="363" pageId="22" pageNumber="79" year="1948">
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<bibRefCitation id="C6304B4299095E34BD32FC07FDF0FC6D" author="Deichmann, E." box="[349,532,973,995]" pageId="22" pageNumber="79" pagination="325 - 376" refId="ref14539" refString="Deichmann, E. (1948) The holothurian fauna of South Africa. Annals of the Natal Museum, 11 (2), 325 - 376." type="journal article" year="1948">Deichmann, 1948</bibRefCitation>
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: 363
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</treatmentCitation>
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</taxonomicName>
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, fig. 21, 1–10.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="EABB653899095E34BCF8FBC5FDF2FBA6" box="[151,534,1039,1064]" pageId="22" pageNumber="79" type="diagnosis">
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<paragraph id="A21E36B399095E34BCF8FBC5FDF2FBA6" blockId="22.[151,1437,1039,2037]" box="[151,534,1039,1064]" pageId="22" pageNumber="79">
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<emphasis id="90D5EAA199095E34BCF8FBC5FEEEFBA6" bold="true" box="[151,266,1039,1064]" pageId="22" pageNumber="79">Diagnosis</emphasis>
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(see
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<bibRefCitation id="C6304B4299095E34BD2AFBC5FDEDFBA6" author="Deichmann, E." box="[325,521,1039,1064]" pageId="22" pageNumber="79" pagination="325 - 376" refId="ref14539" refString="Deichmann, E. (1948) The holothurian fauna of South Africa. Annals of the Natal Museum, 11 (2), 325 - 376." type="journal article" year="1948">Deichmann 1948</bibRefCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EABB653899095E34BCA8FBFEFE97FBE1" pageId="22" pageNumber="79" type="materials_examined">
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<paragraph id="A21E36B399095E34BCA8FBFEFE97FBE1" blockId="22.[151,1437,1039,2037]" pageId="22" pageNumber="79">
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||
<emphasis id="90D5EAA199095E34BCA8FBFEFE56FBC3" bold="true" box="[199,434,1076,1101]" pageId="22" pageNumber="79">Material examined.</emphasis>
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<materialsCitation id="12C93CEE99095E34BDD5FBFEFD50FBC2" ID-GBIF-Occurrence="2117766347" box="[442,692,1076,1100]" collectingDate="28-09-09" collectionCode="SM" pageId="22" pageNumber="79" specimenCode="SAMC-A090928" specimenCount="1">
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||
<specimenCode id="F2079EC899095E34BDD5FBFEFD9AFBC2" box="[442,638,1076,1100]" collectionCode="SAMC-A" pageId="22" pageNumber="79">
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SAMC-A
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<date id="D61F107399095E34BE46FBFEFD9AFBC2" box="[553,638,1076,1100]" pageId="22" pageNumber="79" value="28-09-09">090928</date>
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</specimenCode>
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||
,
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<collectionCode id="C4B0AE7699095E34BEE4FBFEFD50FBC2" box="[651,692,1076,1100]" country="Malaysia" name="Sarawak Museum" pageId="22" pageNumber="79">SM</collectionCode>
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</materialsCitation>
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232,
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<materialsCitation id="12C93CEE99095E34BE82FBFEFF5CFBFE" ID-GBIF-Occurrence="2117766360" collectingDate="1979-06-25" collectionCode="SM" elevation="560" latitude="-32.181667" location="Port St. Johns" longLatPrecision="121" longitude="29.173334" pageId="22" pageNumber="79" specimenCount="1">
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south of
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<location id="A77E606899095E34BF3DFBFFFC17FBC3" LSID="urn:lsid:plazi:treatment:2A0887A599095E35BCF8FC8EFF3CFE22:A77E606899095E34BF3DFBFFFC17FBC3" box="[850,1011,1076,1101]" latitude="-32.181667" longLatPrecision="121" longitude="29.173334" name="Port St. Johns" pageId="22" pageNumber="79">Port St. Johns</location>
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,
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<geoCoordinate id="C795507499095E34B86FFBFEFB9FFBC2" box="[1024,1147,1076,1100]" direction="south" orientation="latitude" pageId="22" pageNumber="79" precision="92" value="-32.181667">32°10.9’ S</geoCoordinate>
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,
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<geoCoordinate id="C795507499095E34B8E6FBFEFAE3FBC2" box="[1161,1287,1076,1100]" direction="east" orientation="longitude" pageId="22" pageNumber="79" precision="92" value="29.173334">29°10.4’ E</geoCoordinate>
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, 620–
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<quantity id="65599B5699095E34B923FBFEFA7CFBC2" box="[1356,1432,1076,1101]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="22" pageNumber="79" unit="m" value="560.0">
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<elevation id="298CD18099095E34B923FBFEFA7CFBC2" box="[1356,1432,1076,1101]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="22" pageNumber="79" unit="m" value="560.0">560 m</elevation>
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</quantity>
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,
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<date id="D61F107399095E34BCF8FB9DFF5CFBFE" box="[151,184,1111,1136]" pageId="22" pageNumber="79" value="1979-06-25">25.</date>
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</materialsCitation>
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VI.1979, 1 spec.
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</paragraph>
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</subSubSection>
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<subSubSection id="EABB653899095E34BCA8FBB6FCEFF9E6" pageId="22" pageNumber="79" type="description">
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<paragraph id="A21E36B399095E34BCA8FBB6FCEFF9E6" blockId="22.[151,1437,1039,2037]" pageId="22" pageNumber="79">
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<emphasis id="90D5EAA199095E34BCA8FBB6FEBDFB1B" bold="true" box="[199,345,1148,1173]" pageId="22" pageNumber="79">Description.</emphasis>
|
||
Specimen minute, perhaps immature (
|
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<figureCitation id="3A9A2A3699095E34BF7EFBB6FC7CFB1A" box="[785,920,1148,1173]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17A</figureCitation>
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||
). Length
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||
<quantity id="65599B5699095E34B869FBB6FBADFB1A" box="[1030,1097,1148,1173]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="79" unit="mm" value="5.0">5 mm</quantity>
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||
(including tentacles), breadth about
|
||
<quantity id="65599B5699095E34BC8BFB55FEA5FB39" box="[228,321,1183,1208]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="79" unit="mm" value="2.5">2.5 mm</quantity>
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in mid-body. Colour off-white. Mouth terminal, slightly dorsally directed; anus not visible. Specimen flat, dorsal surface only slightly arched, ventral surface flat, left margin folded over a rather indistinct sole, hence ventral tube feet displaced. Only eight tube feet on the right side, about 10 on the left side, few midventrally. Because of deformed nature of the body the lateral tube feet do not appear in single series as some are displaced medially; those on left lateral side protrude, including the two most posterior ones. Dorsal surface without tube feet. Tentacles 10, clearly dendritic, two ventral slightly reduced. Oral valves absent. Dorsal and ventral body wall covered by smooth imbricating plates (
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<figureCitation id="3A9A2A3699095E34BF74FABDFC3CFA1E" box="[795,984,1399,1424]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17B & D</figureCitation>
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), dorsally up to 80 µm in size, ventrally up to 58 µm. Baskets (
|
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<figureCitation id="3A9A2A3699095E34BDFBFA56FDFCFA3A" box="[404,536,1436,1461]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17G</figureCitation>
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) common in body wall, minute, with four holes piercing a rather thick base and about 15–16 spines on the rim, pointed in all directions; dorsal baskets up to 40 µm, ventral baskets up to 35 µm. Ossicles of tube feet include large, thick perforated plates and rods (
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<figureCitation id="3A9A2A3699095E34BFC5FA2EFBD6FA72" box="[938,1074,1508,1533]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17H</figureCitation>
|
||
), up to 175 µm, plus numerous baskets; the latter apparently larger than those of body wall, and often with more than four basal holes. End-plates reduced (
|
||
<figureCitation id="3A9A2A3699095E34BD6DF9E6FE6DF9CA" box="[258,393,1580,1605]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17E</figureCitation>
|
||
), up to 150 µm, not always present. Tentacle ossicles include large thick rods and small smooth plates similar to those of tube feet (
|
||
<figureCitation id="3A9A2A3699095E34BE15F985FD1AF9E6" box="[634,766,1615,1640]" captionStart="FIGURE 17" captionStartId="23.[151,250,1130,1152]" captionTargetBox="[151,1436,481,1109]" captionTargetId="figure@23.[151,1436,481,1109]" captionTargetPageId="23" captionText="FIGURE 17. Psolus imperfectus Clark, 1923. SAMC-A090928. A. Specimen (entire); B. Plates from dorsal body wall; C. Rods and plates from tentacles; D. Plates from ventral body wall; E. Developing end-plate from tube foot; F. Developing plate from ventral body wall; G. Baskets from ventral body wall; H. Rods from tube feet of ventral body wall. (B & D same scale; E & F same scale)." httpUri="https://zenodo.org/record/2615058/files/figure.png" pageId="22" pageNumber="79">Figure 17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EABB653899095E34BCA8F9BEFC5AF902" box="[199,958,1652,1677]" pageId="22" pageNumber="79" type="distribution">
|
||
<paragraph id="A21E36B399095E34BCA8F9BEFC5AF902" blockId="22.[151,1437,1039,2037]" box="[199,958,1652,1677]" pageId="22" pageNumber="79">
|
||
<emphasis id="90D5EAA199095E34BCA8F9BEFE84F903" bold="true" box="[199,352,1652,1677]" pageId="22" pageNumber="79">Distribution.</emphasis>
|
||
South of Port St. Johns to
|
||
<collectingRegion id="6065F85199095E34BEE3F9BEFD21F902" box="[652,709,1652,1676]" country="South Africa" name="Western Cape" pageId="22" pageNumber="79">Cape</collectingRegion>
|
||
Agulhas,
|
||
<quantity id="65599B5699095E34BF57F9BEFC5EF902" box="[824,954,1652,1677]" metricMagnitude="2" metricUnit="m" metricValue="4.9799999999999995" metricValueMax="6.2" metricValueMin="3.76" pageId="22" pageNumber="79" unit="m" value="498.0" valueMax="620.0" valueMin="376.0">
|
||
376–
|
||
<specimenCount id="B4A7FD3A99095E34BF1FF9BEFC5EF902" box="[880,954,1652,1677]" pageId="22" pageNumber="79" type="generic">620 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EABB653899095E35BCA8F95DFF3CFE22" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79" type="discussion">
|
||
<paragraph id="A21E36B399095E35BCA8F95DFF3CFE22" blockId="22.[151,1437,1039,2037]" lastBlockId="23.[151,1437,151,428]" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">
|
||
<emphasis id="90D5EAA199095E34BCA8F95DFEDFF93E" bold="true" box="[199,315,1687,1712]" pageId="22" pageNumber="79">Remarks.</emphasis>
|
||
The size and form of the animal, the number and distribution of tube feet and the type of body wall ossicles, confirm that this specimen as a juvenile of
|
||
<taxonomicName id="65A14D3099095E34BE98F976FC2AF95A" authorityName="Deichmann" authorityYear="1948" box="[759,974,1724,1749]" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="79" phylum="Echinodermata" rank="species" species="imperfectus">
|
||
<emphasis id="90D5EAA199095E34BE98F976FC2AF95A" box="[759,974,1724,1749]" italics="true" pageId="22" pageNumber="79">Psolus imperfectus</emphasis>
|
||
</taxonomicName>
|
||
. It is regrettable that
|
||
<bibRefCitation id="C6304B4299095E34B8A3F977FA78F95A" author="Clark, H. L." box="[1228,1436,1724,1749]" pageId="22" pageNumber="79" pagination="221 - 435" refId="ref14325" refString="Clark, H. L. (1923) The echinoderm fauna of South Africa. Annals of the South African Museum, 13, 221 - 435. Available from: https: // www. biodiversitylibrary. org / page / 1541222 # page / 1 / mode / 1 up (Accessed 14 Dec. 2018)" type="journal article" year="1923">H.L. Clark (1923)</bibRefCitation>
|
||
neither figured the ossicles nor the animal itself, and perhaps failed to detect any baskets found in association with the plates. His material comprised two juveniles collected off south-east of
|
||
<collectingRegion id="6065F85199095E34B876F8CEFBB7F892" box="[1049,1107,1796,1820]" country="South Africa" name="Western Cape" pageId="22" pageNumber="79">Cape</collectingRegion>
|
||
Agulhas, at
|
||
<quantity id="65599B5699095E34B881F8CEFAB9F892" box="[1262,1373,1796,1821]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="22" pageNumber="79" unit="m" value="365.76">365.76 m</quantity>
|
||
(200 fathoms). The
|
||
<typeStatus id="7D1A881199095E34BD51F8EDFE44F8CE" box="[318,416,1831,1856]" pageId="22" pageNumber="79" type="holotype">holotype</typeStatus>
|
||
which was presumably in the South African Museum could not be located. The
|
||
<typeStatus id="7D1A881199095E34B953F8E3FA78F8CF" box="[1340,1436,1833,1857]" pageId="22" pageNumber="79" type="paratype">paratype</typeStatus>
|
||
from the Museum of Comparative Zoology (Harvard) was studied by
|
||
<bibRefCitation id="C6304B4299095E34BFF0F886FB90F8EA" author="Deichmann, E." box="[927,1140,1868,1893]" pageId="22" pageNumber="79" pagination="325 - 376" refId="ref14539" refString="Deichmann, E. (1948) The holothurian fauna of South Africa. Annals of the Natal Museum, 11 (2), 325 - 376." type="journal article" year="1948">Deichmann (1948)</bibRefCitation>
|
||
. Deichmann was therefore the first to find minute baskets, in addition to the plates. Except for the fewer number of holes in the plates of the current material, its baskets are identical to those described by
|
||
<bibRefCitation id="C6304B4299095E34BF33F85EFBD7F822" author="Deichmann, E." box="[860,1075,1940,1965]" pageId="22" pageNumber="79" pagination="325 - 376" refId="ref14539" refString="Deichmann, E. (1948) The holothurian fauna of South Africa. Annals of the Natal Museum, 11 (2), 325 - 376." type="journal article" year="1948">Deichmann (1948)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="C6304B4299095E34B852F85EFA78F823" author="Samyn, Y. & VandenSpiegel, D." box="[1085,1436,1940,1965]" pageId="22" pageNumber="79" pagination="451 - 497" refId="ref15649" refString="Samyn, Y. and VandenSpiegel, D., 2016. Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species. Zootaxa, 4196 (4), pp. 451 - 497." type="journal article" year="2016">Samyn & Vandenspigel (2016)</bibRefCitation>
|
||
show some spired scales which were not observed in the South African material. A point of interest is that Deichmann illustrated both smooth and knobbed plates. Despite this the specimen is identified as a juvenile
|
||
<taxonomicName id="65A14D3099095E35B93EF816FEFEFF3E" authorityName="Deichmann" authorityYear="1948" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="23" lastPageNumber="80" order="Dendrochirotida" pageId="22" pageNumber="79" phylum="Echinodermata" rank="species" species="imperfectus">
|
||
<emphasis id="90D5EAA199095E35B93EF816FEFEFF3E" italics="true" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">Psolus imperfectus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="C6304B4299085E35BD46FF5DFE19FF3E" author="Deichmann, E." box="[297,509,151,176]" pageId="23" pageNumber="80" pagination="325 - 376" refId="ref14539" refString="Deichmann, E. (1948) The holothurian fauna of South Africa. Annals of the Natal Museum, 11 (2), 325 - 376." type="journal article" year="1948">Deichmann (1948)</bibRefCitation>
|
||
mentioned the presence of about 20–25 holes in the plates which have a slightly knobbed surface.
|
||
<bibRefCitation id="C6304B4299085E35BD33FF76FE0FFF5A" author="Clark, H. L." box="[348,491,188,213]" pageId="23" pageNumber="80" pagination="221 - 435" refId="ref14325" refString="Clark, H. L. (1923) The echinoderm fauna of South Africa. Annals of the South African Museum, 13, 221 - 435. Available from: https: // www. biodiversitylibrary. org / page / 1541222 # page / 1 / mode / 1 up (Accessed 14 Dec. 2018)" type="journal article" year="1923">Clark (1923)</bibRefCitation>
|
||
also recorded about 20–24 holes, but did not comment on the knobbed plate surface. There is a possibility that the two specimens studied by Clark are perhaps not conspecific with that described by Deichmann and herein. However, the current material with fewer holes in the otherwise smooth plates, appears intermediate in form, perhaps because of its small size. In the absence of the
|
||
<typeStatus id="7D1A881199085E35B844FEEDFB74FECE" box="[1067,1168,295,320]" pageId="23" pageNumber="80" type="holotype">holotype</typeStatus>
|
||
, the specimen must be regarded as a juvenile of
|
||
<taxonomicName id="65A14D3099085E35BDC1FE87FDA8FEEA" authorityName="Deichmann" authorityYear="1948" box="[430,588,332,356]" class="Holothuroidea" family="Psolidae" genus="Psolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="80" phylum="Echinodermata" rank="species" species="imperfectus">
|
||
<emphasis id="90D5EAA199085E35BDC1FE87FDA8FEEA" box="[430,588,332,356]" italics="true" pageId="23" pageNumber="80">P. imperfectus</emphasis>
|
||
</taxonomicName>
|
||
. The type was described from south of
|
||
<collectingRegion id="6065F85199085E35B86CFE86FBD9FEEA" box="[1027,1085,332,356]" country="South Africa" name="Western Cape" pageId="23" pageNumber="80">Cape</collectingRegion>
|
||
Agulhas at about
|
||
<quantity id="65599B5699085E35B969FE86FAB4FEEA" box="[1286,1360,332,357]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="23" pageNumber="80" unit="m" value="366.0">366 m</quantity>
|
||
; hence the current specimen indicates a noteworthy extension of its range eastwards into the warmer waters of the
|
||
<collectingRegion id="6065F85199085E35B927FEBAFF30FE22" country="South Africa" name="Eastern Cape" pageId="23" pageNumber="80">Eastern Cape</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |