996 lines
74 KiB
XML
996 lines
74 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59696" ID-GBIF-Dataset="b5267fd4-1620-46e2-a4df-ceb934834d8f" ID-GBIF-Taxon="183412856" ID-Pensoft-Pub="1860-0743-2-355" ID-Pensoft-UUID="59DE43E719E65A2DB08A18AA24ADEC0E" ID-ZooBank="FB92F5DF7FC74F01A1DD8E85B6F5FE67" LSID="urn:lsid:zoobank.org:act:D78F9340-1032-4D34-976A-B91C7001CC1C" ModsDocID="1860-0743-97-2-355" checkinTime="1625936817539" checkinUser="pensoft" docAuthor="Guenther, Rainer & Richards, Stephen" docDate="2021" docId="086356E267915A50A39C3E3C0070FABA" docLanguage="en" docName="ZoosystEvol 97(2): 355-382" docOrigin="Zoosystematics and Evolution 97 (2)" docPubDate="2021-07-09" docSource="http://dx.doi.org/10.3897/zse.97.59696" docTitle="Xenorhina lacrimosa Günther & Richards 2021, sp. nov." docType="treatment" docUuid="D78F9340-1032-4D34-976A-B91C7001CC1C" docUuidSource="ZooBank" docVersion="4" id="59DE43E719E65A2DB08A18AA24ADEC0E" lastPageNumber="355" masterDocId="59DE43E719E65A2DB08A18AA24ADEC0E" masterDocTitle="Description of six new species of Xenorhina Peters, 1863 from southern Papua New Guinea (Amphibia, Anura, Microhylidae)" masterLastPageNumber="382" masterPageNumber="355" pageId="0" pageNumber="355" updateTime="1634707979541" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of six new species of Xenorhina Peters, 1863 from southern Papua New Guinea (Amphibia, Anura, Microhylidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Guenther, Rainer</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9250-7658</mods:nameIdentifier>
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<mods:affiliation>Museum fuer Naturkunde, Herpetology Department, Invalidenstr. 43, 10115 Berlin, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">rainer.guenther@mfn-berlin.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Richards, Stephen</mods:namePart>
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<mods:affiliation>Herpetology Department, South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>355</mods:start>
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<mods:end>382</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59696</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59696</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-355</mods:identifier>
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<mods:identifier type="ZooBank">FB92F5DF7FC74F01A1DD8E85B6F5FE67</mods:identifier>
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<mods:identifier type="Pensoft-UUID">59DE43E719E65A2DB08A18AA24ADEC0E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="183412856" LSID="urn:lsid:zoobank.org:act:D78F9340-1032-4D34-976A-B91C7001CC1C" httpUri="http://treatment.plazi.org/id/086356E267915A50A39C3E3C0070FABA" lastPageNumber="355" pageId="0" pageNumber="355">
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<subSubSection pageId="0" pageNumber="355" type="nomenclature">
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<paragraph pageId="0" pageNumber="355">
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<taxonomicName LSID="http://zoobank.org/D78F9340-1032-4D34-976A-B91C7001CC1C" authority="Günther & Richards, 2021" authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa" status="sp. nov.">Xenorhina lacrimosa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="355">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="355" type="holotype">
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<paragraph pageId="0" pageNumber="355">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="355">
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SAMA R71648 (SJR 14203), adult male, from Rentoul River, Western Province, Papua New Guinea (
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<geoCoordinate degrees="6.4355" direction="south" orientation="latitude" precision="5" value="-6.4355">6.4355°S</geoCoordinate>
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,
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<geoCoordinate degrees="142.5615" direction="east" orientation="longitude" precision="5" value="142.5615">142.5615°E</geoCoordinate>
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; 380 m a.s.l.), collected on 10-08-2014 by S.J. Richards.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="355" type="paratypes">
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<paragraph pageId="0" pageNumber="355">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="355">
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SAMA R71647 (SJR10389), female with ripe eggs, ZMB 91129 (SJR10417) male, Camp 2, upper Strickland River basin, Western Province, Papua New Guinea (
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<geoCoordinate degrees="5.9018" direction="south" orientation="latitude" precision="5" value="-5.9018">5.9018°S</geoCoordinate>
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,
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<geoCoordinate degrees="142.4360" direction="east" orientation="longitude" precision="5" value="142.436">142.4360°E</geoCoordinate>
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; 950 m a.s.l.), collected by S.J. Richards on 18-02-2008 and 20-02-2008, respectively; ZMB 91130 (SJR10466) male, Camp 1, upper Strickland River basin, Western Province, Papua New Guinea (
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<geoCoordinate degrees="5.8078" direction="south" orientation="latitude" precision="5" value="-5.8078">5.8078°S</geoCoordinate>
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,
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<geoCoordinate degrees="142.3083" direction="east" orientation="longitude" precision="5" value="142.3083">142.3083°E</geoCoordinate>
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; 215 m a.s.l.), collected by S.J. Richards on 26-03-2008; SAMA R65069 (SJR10902) and R65070 (SJR10949), males, R65071 (SJR10963), (subadult?) female with scarcely developed eggs and R65072 (SJR10985), juvenile, Gugusu Camp, Muller Range, Western Province (
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<geoCoordinate degrees="5.7290" direction="south" orientation="latitude" precision="5" value="-5.729">5.7290°S</geoCoordinate>
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,
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<geoCoordinate degrees="142.2630" direction="east" orientation="longitude" precision="5" value="142.263">142.2630°E</geoCoordinate>
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; 515 m a.s.l.), all collected by S.J. Richards and C. Dahl between 7-9-09-2009.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="355" type="referred specimens">
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<paragraph pageId="0" pageNumber="355">Referred specimens.</paragraph>
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<paragraph pageId="0" pageNumber="355">
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SAMA R71649, R71650 (SJR2577, 2582), PNGNM (SJR2571), adult males, Herowana, Eastern Highlands Province, Papua New Guinea (
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<geoCoordinate degrees="6.6220" direction="south" orientation="latitude" precision="5" value="-6.622">6.6220°S</geoCoordinate>
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,
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<geoCoordinate degrees="145.1962" direction="east" orientation="longitude" precision="5" value="145.1962">145.1962°E</geoCoordinate>
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; 1,400 m a.s.l.), collected by S.J. Richards between 20 and 24-11-2001.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="355" type="diagnosis">
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<paragraph pageId="0" pageNumber="355">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="355">
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This species of
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<taxonomicName authorityName="Peters" authorityYear="1863" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina</emphasis>
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</taxonomicName>
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is characterised by the unique combination of: medium size (SUL of five males 34.5-41.0 mm); vomeropalatines each with one long and acuminate spike; legs moderately long (TL/SUL 0.42-0.46); all fingers without and all toes with expanded discs; eye-naris distance greater than internarial distance (END/IND 1.18-1.48); tympanum diameter smaller than or equal to that of eye (TyD/ED 0.75-1.00); dorsal surfaces in life different tones of brown or blue or a mixture of these colours; ventral surfaces different tones of orange with irregular whitish spots or mouse grey (RAL 7005) with whitish spots and reticulations; advertisement calls uttered in series containing 7-12 single, mournful
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<normalizedToken originalValue="“hoots”">"hoots"</normalizedToken>
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separated by long intervals of about five seconds.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="355" type="description">
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<paragraph pageId="0" pageNumber="355">Description of the holotype.</paragraph>
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<paragraph pageId="0" pageNumber="355">
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Measurements are summarised in Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Body measurements and body ratios of the type series of Xenorhina lacrimosa sp. nov. SAMA R 71648 is the male holotype, others are paratypes. All measurements in mm; for explanation of abbreviations see " Material and methods "; M = male, F = female, sa = subadult." httpUri="http://table.plazi.org/id/950F366140FAB12A3A26AD070F887FCE" pageId="0" pageNumber="355" tableUuid="950F366140FAB12A3A26AD070F887FCE">1</tableCitation>
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, a dorsolateral view in life is shown in Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1a</figureCitation>
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and ventral surfaces in life in Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1b</figureCitation>
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. Head broader than long (HL/HW 0.74); snout acuminate from above and below, distinctly protruding in profile; tongue very broad, only its lateral and posterior edges free; prepharyngeal ridge with five roundish denticles; left vomerine spike very well developed, right spike present, but malformed; loreal region oblique, no canthus rostralis; nostrils near tip of snout, directed more lateral than dorsal, visible from above, but not from below; eye-naris distance greater than internarial distance (END/IND 1.18); tympanum visible in life and preservative, its diameter slightly less than that of eye (TyD/ED 0.87); supratympanic fold weakly expressed, extending from behind eye to insertion of fore leg; shank short (TL/SUL 0.42); fingers moderately short, not webbed; tips of all fingers with circum-marginal grooves, all not wider than penultimate phalanges; relative lengths of fingers 3> 4 = 1 = 2 (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1c</figureCitation>
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); all toe tips with circum-marginal grooves and wider than penultimate phalanges; toes not webbed, relative lengths 4> 3> 5> 2> 1 (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1d</figureCitation>
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); plantar and palmar tubercles (with exception of prominent, oval inner metatarsal tubercle; Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1d</figureCitation>
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), as well as subarticular tubercles scarcely visible. Body laterally with numerous distinct tubercles in life, less prominent in preservative; dorsal surfaces of limbs and middle of dorsum with fewer tubercles, all ventral surfaces smooth; tip of snout with several tiny elevations (especially on underside).
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</paragraph>
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<caption doi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="355">
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<emphasis bold="true" pageId="0" pageNumber="355">Figure 1.</emphasis>
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Holotype (SAMA R71648) of
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<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
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<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
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</taxonomicName>
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sp. nov. in life: (
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<emphasis bold="true" pageId="0" pageNumber="355">a</emphasis>
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) Dorsolateral view; (
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<emphasis bold="true" pageId="0" pageNumber="355">b</emphasis>
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) Ventral view; (
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<emphasis bold="true" pageId="0" pageNumber="355">c</emphasis>
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) Volar view of right hand; (
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<emphasis bold="true" pageId="0" pageNumber="355">d</emphasis>
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) Thenar view of right foot.
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</paragraph>
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</caption>
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<caption ID-Table-UUID="950F366140FAB12A3A26AD070F887FCE" httpUri="http://table.plazi.org/id/950F366140FAB12A3A26AD070F887FCE" pageId="0" pageNumber="355" start="Table 1" startId="T1">
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<paragraph pageId="0" pageNumber="355">
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<emphasis bold="true" pageId="0" pageNumber="355">Table 1.</emphasis>
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Body measurements and body ratios of the type series of
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<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
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<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
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</taxonomicName>
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sp. nov. SAMA R71648 is the male holotype, others are paratypes. All measurements in mm; for explanation of abbreviations see "Material and methods"; M = male, F = female, sa = subadult.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="355">
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<table pageId="0" pageNumber="355">
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<tr pageId="0" pageNumber="355">
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">Reg.-No.</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">SAMA R71647</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">ZMB 91129</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">ZMB 91130</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">SAMA R71648</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">SAMA R65069</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">SAMA R65070</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">SAMA R65071</th>
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<th colspan="1" pageId="0" pageNumber="355" rowspan="1">
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Mean
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<normalizedToken originalValue="±">+/-</normalizedToken>
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SD
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</th>
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</tr>
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<tr pageId="0" pageNumber="355">
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">Sex</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">M</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">M</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">M</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">M</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">M</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">sa F</td>
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<td colspan="1" isEmpty="true" pageId="0" pageNumber="355" rowspan="1">-</td>
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</tr>
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<tr pageId="0" pageNumber="355">
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">SUL</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">34.3</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">36.9</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">41.0</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">36.2</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">37.3</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">34.5</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">35.1</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
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36.47
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<normalizedToken originalValue="±">+/-</normalizedToken>
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2.30
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</td>
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</tr>
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<tr pageId="0" pageNumber="355">
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TL</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.0</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">17.0</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">18.6</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.3</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.8</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.3</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.4</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
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16.06
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1.30
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</td>
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</tr>
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<tr pageId="0" pageNumber="355">
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TaL</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.0</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">11.8</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">12.6</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.2</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">11.0</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.3</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">9.9</td>
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
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10.83
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1.03
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</td>
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</tr>
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<tr pageId="0" pageNumber="355">
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<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T4L</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">16.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">17.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">19.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">16.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">15.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
16.61
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
1.46
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T4D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
1.41
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.12
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T1D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.93
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.076
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F3L</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">6.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">8.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">9.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">7.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">7.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">7.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
7.46
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.82
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F3D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.93
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.076
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F1D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.77
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.14
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">HL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">9.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">12.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">9.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">11.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">10.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
10.46
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.87
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">11.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">14.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">16.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">13.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">12.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">11.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">13.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
13.26
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
1.58
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">END</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
3.09
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.43
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">IND</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
2.24
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.28
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">SL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">5.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">5.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
4.57
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.42
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">EST</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">5.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">4.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
4.24
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.56
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">ED</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
2.41
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.25
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TyD</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
2.11
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.25
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TL/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.44
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.015
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TaL/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.30
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.015
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T4L/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.46
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.025
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T4D/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.038</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.043</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.037</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.036</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.040</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.041</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.037</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.039
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.003
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T1D/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.023</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.027</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.024</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.025</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.027</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.026</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.026</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.025
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.002
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F3L/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.184</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.220</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.222</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.193</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.209</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.194</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.205</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.204
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.014
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F3D/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.023</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.027</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.027</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.022</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.021</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.026</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.026</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.025
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.003
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">F1D/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.020</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.024</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.024</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.017</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.019</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.023</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.020</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.021
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.003
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T4D/F3D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
1.59
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.165
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">T1D/F1D</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
1.23
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.183
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">HL/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.29
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.009
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">HW/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.36
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.026
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">HL/HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.79</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.79
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.069
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">END/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.073</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.081</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.088</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.072</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.094</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.099</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.085</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.085
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.010
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">IND/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.050</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.062</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.063</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.061</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.064</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.067</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.063</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.061
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.005
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">END/IND</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
1.38
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.109
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">ED/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.070</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.073</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.068</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.064</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.064</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.064</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.060</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.066
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.004
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TyD/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.073</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.068</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.051</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.055</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.056</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.052</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.054</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.058
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.009
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">TyD/ED</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">1.00</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.88
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.079
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">SL/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.117</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.117</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.124</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.116</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.134</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.136</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.134</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.125
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.009
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="355">
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">EST/SUL</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.105</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.111</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.122</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.097</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.123</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.136</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">0.120</td>
|
||
<td colspan="1" pageId="0" pageNumber="355" rowspan="1">
|
||
0.116
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.013
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
In life, dorsal surfaces of head and anterior portion of body and fore limbs, uniform bluish-brown; remaining dorsal surfaces and flanks a mixture of saffron-yellow (RAL 1017) and blue-grey; tubercles with brown bases and whitish apices concentrated on flanks; body dorsally with light yellow mid-dorsal line that continues on to hind legs; lumbar region with light yellow semi-circular spot (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1a</figureCitation>
|
||
); vent and adjacent region enclosed within dark brown triangular patch; iris blackish with barely visible golden reticulation; plantar and ventral surfaces of toes predominantly brown, palms and ventral surfaces of fingers predominantly grey and cream; throat, chest, abdomen and ventral surfaces of extremities deep orange (RAL 2011), with some whitish spots (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype (SAMA R 71648) of Xenorhina lacrimosa sp. nov. in life: (a) Dorsolateral view; (b) Ventral view; (c) Volar view of right hand; (d) Thenar view of right foot." figureDoi="10.3897/zse.97.59696.figure1" httpUri="https://binary.pensoft.net/fig/564751" pageId="0" pageNumber="355">1b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">In preservative, dorsal surfaces of head, anterior back and fore limbs signal brown (RAL 8002); other dorsal surfaces ivory with diffuse brownish smears, tubercles with terra brown (RAL 8028) bases and whitish apices; ventral surfaces light ivory (RAL 1015); ivory lumbar spot on left side more clearly pronounced than on right.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="355" type="morphological variation">
|
||
<paragraph pageId="0" pageNumber="355">Morphological variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
Measurements and proportions of most paratypes show limited variation (Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Body measurements and body ratios of the type series of Xenorhina lacrimosa sp. nov. SAMA R 71648 is the male holotype, others are paratypes. All measurements in mm; for explanation of abbreviations see " Material and methods "; M = male, F = female, sa = subadult." httpUri="http://table.plazi.org/id/950F366140FAB12A3A26AD070F887FCE" pageId="0" pageNumber="355" tableUuid="950F366140FAB12A3A26AD070F887FCE">1</tableCitation>
|
||
). An exception is a juvenile (SAMA R65072) measuring 16.6 mm SUL that exhibits some major deviations in proportions from the remainder of the type series. As these differences are almost certainly due to allometry, measurements of this specimen are disregarded in Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Body measurements and body ratios of the type series of Xenorhina lacrimosa sp. nov. SAMA R 71648 is the male holotype, others are paratypes. All measurements in mm; for explanation of abbreviations see " Material and methods "; M = male, F = female, sa = subadult." httpUri="http://table.plazi.org/id/950F366140FAB12A3A26AD070F887FCE" pageId="0" pageNumber="355" tableUuid="950F366140FAB12A3A26AD070F887FCE">1</tableCitation>
|
||
. Males and females have the same body size, although some ratios of the adult female (SAMA R71647 differ to a negligible degree (Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Body measurements and body ratios of the type series of Xenorhina lacrimosa sp. nov. SAMA R 71648 is the male holotype, others are paratypes. All measurements in mm; for explanation of abbreviations see " Material and methods "; M = male, F = female, sa = subadult." httpUri="http://table.plazi.org/id/950F366140FAB12A3A26AD070F887FCE" pageId="0" pageNumber="355" tableUuid="950F366140FAB12A3A26AD070F887FCE">1</tableCitation>
|
||
). The smallest specimen in the series is the just-mentioned adult female with an SUL of
|
||
<normalizedToken originalValue="“only”">"only"</normalizedToken>
|
||
34.3 mm that contains ripe ovarian eggs; the largest specimen in the series is a male (ZMB 91130) with an SUL of 41.0 mm.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
Colour of paratypes in life varies considerably. Dorsal surfaces may be uniform blue-brown (SAMA R65070, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2a</figureCitation>
|
||
), uniform light red-orange similar to RAL 2008 (ZMB 91129, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2b</figureCitation>
|
||
), bluish on head and lower flanks, but reddish-brown on back and dorsal extremities (ZMB 91130, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2c</figureCitation>
|
||
) or dark brown with bluish hue on head, body and thighs, but beige on fore limbs, shanks and tarsi (SAMA R71647, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2d</figureCitation>
|
||
). Colour of ventral surfaces is also highly variable. Some specimens are uniform deep orange or traffic orange (RAL 2009) interspersed with scattered irregular whitish spots (SAMA R65070, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2e</figureCitation>
|
||
); others are more extensively spotted (ZMB 91129, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2f</figureCitation>
|
||
) or exhibit a mixture of whitish, orange and brown spots, but with throat and thighs more or less uniform traffic orange (SAMA R65071, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2g</figureCitation>
|
||
); others exhibit grey-brown ground colour with many irregular whitish spots, some of them interspersed with small irregular red patches (SAMA R71647, Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Xenorhina lacrimosa sp. nov. paratypes in life. Dorsolateral views: (a) SAMA R 65070; (b) ZMB 91129; (c) ZMB 91130; (d) SAMA R 71647. Ventral views: (e) SAMA R 65070 S; (f) ZMB 91129; (g) SAMA R 65071; (h) SAMA R 71647." figureDoi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355">2h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59696.figure2" httpUri="https://binary.pensoft.net/fig/564752" pageId="0" pageNumber="355" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<emphasis bold="true" pageId="0" pageNumber="355">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratypes in life. Dorsolateral views: (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">a</emphasis>
|
||
) SAMA R65070; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">b</emphasis>
|
||
) ZMB 91129; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">c</emphasis>
|
||
) ZMB 91130; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">d</emphasis>
|
||
) SAMA R71647. Ventral views: (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">e</emphasis>
|
||
) SAMA R65070S; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">f</emphasis>
|
||
) ZMB 91129; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">g</emphasis>
|
||
) SAMA R65071; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">h</emphasis>
|
||
) SAMA R71647.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="355">In preservative dorsal surfaces of three specimens predominantly violet, of two specimens copper brown, of one specimen beige and of the juvenile specimen beige-brown; ventral surfaces of three specimens almost completely light ivory, of the four other specimens a light ivory ground colour with a brown-beige pattern of various extent. All paratypes, except SAMA R71647, have a light ivory mid-dorsal line and all specimens including the juvenile have a greyish snout tip. None of the paratypes has a clearly pronounced lumbar spot in life or in preservative.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="355" type="distribution">
|
||
<paragraph pageId="0" pageNumber="355">Distribution and ecological notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
Most records of
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are from lowland and foothill forest in south-central Papua New Guinea (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of Papua New Guinea showing the known distributions of X. lacrimosa sp. nov. (blue circles), X. perexigua sp. nov. (yellow triangle) and X. pohleorum sp. nov. (red squares). Arrows indicate the type localities." figureDoi="10.3897/zse.97.59696.figure8" httpUri="https://binary.pensoft.net/fig/564758" pageId="0" pageNumber="355">8</figureCitation>
|
||
), where this species appears to have a broad distribution at altitudes ranging from near sea level around Kopi to at least 950 m a.s.l. We also refer several specimens from Herowana Village at 1,400 m a.s.l. (the most easterly location in Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of Papua New Guinea showing the known distributions of X. lacrimosa sp. nov. (blue circles), X. perexigua sp. nov. (yellow triangle) and X. pohleorum sp. nov. (red squares). Arrows indicate the type localities." figureDoi="10.3897/zse.97.59696.figure8" httpUri="https://binary.pensoft.net/fig/564758" pageId="0" pageNumber="355">8</figureCitation>
|
||
) to this species pending confirmation of genetic relationships. Males called at night, normally after rain, either from within the leaf litter on the forest floor or down to several centimetres depth in the humus layer beneath the litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="355" type="vocalisation">
|
||
<paragraph pageId="0" pageNumber="355">Vocalisation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
One call series from SAMA R71648 (holotype), one from ZMB 91129 (paratype), one from SAMA R65069 (paratype) and four from ZMB 91130 (paratype) recorded at air temperatures of 21.2-25.5 °C were analysed. Each call is a single, unpulsed mournful note that is always produced in a series, during which both volume and pitch increase gradually between first and last call (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3a</figureCitation>
|
||
). Despite some variation in recording temperature, all recorded calls are extremely similar, so they were pooled for analyses. Call series last 26.4-60.4 s (mean 40.0
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
11.8 s, n = 7), with 7-12 calls/series (mean 9.0
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
2.2, n = 7) produced at a rate of 0.20-0.27 calls/s (mean 0.23
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.02). Call length is 141-231 ms (mean 193.5
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
19.1 ms, n = 63) and first and last call in a series are often the shortest; inter-call interval length is 2.8-8.0 s. (mean 4.8
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
1.0 s, n = 56). Calls start abruptly at high amplitude that rises quickly to a maximum, then decreases gradually until termination of call (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3b</figureCitation>
|
||
). All calls have 5-7 harmonics (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3c</figureCitation>
|
||
and
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3d</figureCitation>
|
||
)). Dominant frequency may be carried by a second harmonic (i.e. first two calls of series from holotype, with peak at 1.2 kHz) or by first harmonic (all other calls, with peaks increasing from 0.7 kHz in third call to 0.8 kHz in last call in the series. Frequency declines at end of each call in second half of series (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<emphasis bold="true" pageId="0" pageNumber="355">Figure 3.</emphasis>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="355">a</emphasis>
|
||
) Oscillogram of a complete advertisement call series with seven calls from the holotype of
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">b</emphasis>
|
||
) oscillogram and (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">c</emphasis>
|
||
) spectrogram of the penultimate call of the call series shown on Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. (a) Oscillogram of a complete advertisement call series with seven calls from the holotype of Xenorhina lacrimosa sp. nov.; (b) oscillogram and (c) spectrogram of the penultimate call of the call series shown on Fig. 3 a; (d) amplitude spectrum of the call on (b.) and (c.)." figureDoi="10.3897/zse.97.59696.figure3" httpUri="https://binary.pensoft.net/fig/564753" pageId="0" pageNumber="355">3a</figureCitation>
|
||
; (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">d</emphasis>
|
||
) amplitude spectrum of the call on (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">b.</emphasis>
|
||
) and (
|
||
<emphasis bold="true" pageId="0" pageNumber="355">c.</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="355" type="etymology">
|
||
<paragraph pageId="0" pageNumber="355">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
The specific epithet
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
is a Latin adjective in female gender; translated literally it means
|
||
<normalizedToken originalValue="“tearful”">"tearful"</normalizedToken>
|
||
, but it is also translated as "lamentable voice" and refers to the mournful sounding advertisement call of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="355" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="355">Comparisons with other species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
We compare
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. with all congeners of a similar size (SUL 30-43 mm) that have a single spike on each vomeropalatine bone.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<taxonomicName baseAuthorityName="Blum & Menzies" baseAuthorityYear="1989" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina fuscigula" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="fuscigula">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina fuscigula</emphasis>
|
||
</taxonomicName>
|
||
(Blum & Menzies, 1989) has hind legs shorter (TL/SVL <0.40 vs.> 0.40 in
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.), eye-naris distance shorter (END/SVL 0.064-0.074 vs. 0.072-0.099), inner metatarsal tubercle absent (vs. present), ventral surfaces black (vs. orange-red or grey-brown) and call consisting of a single long note (vs. a series of 7-12 notes = calls).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<taxonomicName baseAuthorityName="Blum & Menzies" baseAuthorityYear="1989" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina huon" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="huon">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina huon</emphasis>
|
||
</taxonomicName>
|
||
(Blum & Menzies, 1989) is smaller (SUL to 32 mm vs. 34.3-41.0 mm), with hind legs shorter (TL/SUL <0.40 vs.> 0.40), internarial distance greater (0.064-0.081 vs. 0.050-0.067), eyes larger (ED/SVL 0.070-0.091 vs. 0.060-0.073) and ventral surfaces with dark flecking (vs. ventral surfaces with no or sparse brownish reticulation).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<taxonomicName baseAuthorityName="Menzies & Tyler" baseAuthorityYear="1977" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina subcrocea" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="subcrocea">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina subcrocea</emphasis>
|
||
</taxonomicName>
|
||
(Menzies & Tyler, 1977) is smaller (SUL 30.5-33.3 mm vs. 34.3-41.0 mm), with hind legs longer (TL/SVL> 0.46 vs. <0.46 in
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) and ventral surfaces with dark reticulation (vs. without dark reticulation); call length is shorter 64-69 ms (vs. 141-231 ms), with inter-call interval also much shorter (154-285 ms vs. 2.8-8.0 s).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="355">
|
||
<taxonomicName baseAuthorityName="Kraus & Allison" baseAuthorityYear="2002" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina zweifeli" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina zweifeli</emphasis>
|
||
</taxonomicName>
|
||
(Kraus & Allison, 2002) is about the same size and has similar body ratios. It differs by having a conspicuous dark brown supratympanic stripe (vs. absent in
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) and in several aspects of its advertisement calls.
|
||
<taxonomicName baseAuthorityName="Kraus & Allison" baseAuthorityYear="2002" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina zweifeli" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina zweifeli</emphasis>
|
||
</taxonomicName>
|
||
utters single calls at irregular intervals, with two or three calls sometimes produced in quick succession (
|
||
<bibRefCitation DOI="https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" author="Kraus, F" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="355" refId="B29" refString="Kraus, F, Allison, A, 2002. A new species of Xenobatrachus (Anura: Microhylidae) from northern Papua New Guinea. Herpetologica 58(1): 56-66. https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" title="A new species of Xenobatrachus (Anura: Microhylidae) from northern Papua New Guinea. Herpetologica 58 (1): 56 - 66." url="https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" year="2002">Kraus and Allison 2002</bibRefCitation>
|
||
), during both day and early evening. In contrast,
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. always produces calls in discrete series of at least seven relatively evenly spaced calls of increasing pitch and volume; calls are never produced in quick succession and males always call at night. Other differences include: mean length of calls produced by holotype of
|
||
<taxonomicName genus="X." lsidName="X. zweifeli" pageId="0" pageNumber="355" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">X. zweifeli</emphasis>
|
||
</taxonomicName>
|
||
is 310 ms (
|
||
<bibRefCitation DOI="https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" author="Kraus, F" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="355" refId="B29" refString="Kraus, F, Allison, A, 2002. A new species of Xenobatrachus (Anura: Microhylidae) from northern Papua New Guinea. Herpetologica 58(1): 56-66. https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" title="A new species of Xenobatrachus (Anura: Microhylidae) from northern Papua New Guinea. Herpetologica 58 (1): 56 - 66." url="https://doi.org/10.1655/0018-0831(2002)058[0056:ANSOXA]2.0.CO;2" year="2002">Kraus and Allison 2002</bibRefCitation>
|
||
) (vs. mean length of calls from
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. 194 ms); the fundamental frequency of
|
||
<taxonomicName baseAuthorityName="Kraus & Allison" baseAuthorityYear="2002" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina zweifeli" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina zweifeli</emphasis>
|
||
</taxonomicName>
|
||
calls is at 610 Hz and dominant frequency at 1910 Hz (third harmonic), (vs. fundamental and dominant frequency of
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., both at 800 Hz); amplitude of
|
||
<taxonomicName genus="X." lsidName="X. zweifeli" pageId="0" pageNumber="355" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">X. zweifeli</emphasis>
|
||
</taxonomicName>
|
||
calls rises more slowly than that of
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
calls and all harmonics are frequency modulated, with pitch decreasing during entire length of call (vs. frequency modulation only occurring at end of harmonics in
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. calls). Moreover,
|
||
<taxonomicName genus="X." lsidName="X. zweifeli" pageId="0" pageNumber="355" rank="species" species="zweifeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">X. zweifeli</emphasis>
|
||
</taxonomicName>
|
||
occurs only on two mountain ranges in northern Papua New Guinea, while
|
||
<taxonomicName authorityName="Günther & Richards" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Xenorhina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenorhina lacrimosa" order="Anura" pageId="0" pageNumber="355" phylum="Chordata" rank="species" species="lacrimosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="355">Xenorhina lacrimosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. lives predominantly in the lowlands and foothills of southern Papua New Guinea. Therefore, the known distributions of the two species are separated by a major biogeographic barrier, New
|
||
<normalizedToken originalValue="Guinea’s">Guinea's</normalizedToken>
|
||
central cordillera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |