146 lines
17 KiB
XML
146 lines
17 KiB
XML
<document id="230DF0CBF419938C26C13ED342B8EA97" ID-DOI="10.5281/zenodo.173361" ID-GBIF-Dataset="a6a235ee-2943-4164-89cc-0a79f173f2a0" ID-ISSN="11755326" ID-Zenodo-Dep="173361" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459773686406" checkinUser="plazi" docAuthor="Webb, J. M. & Mccafferty, W. P." docDate="2006" docId="040A87E8FFBA507EC629F981FCDB0DEE" docLanguage="en" docName="zt01277p068.pdf" docOrigin="Zootaxa 1277" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Epeorus aculeatus Braasch" docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="F833FF90FFBA507CC721FFBFFFC8093F" masterDocTitle="First description of the adult male of Epeorus aculeatus Braasch (Ephemeroptera: Heptageniidae)" masterLastPageNumber="68" masterPageNumber="65" pageNumber="65" updateTime="1698217512771" updateUser="plazi">
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<mods:title id="400432C5669FE3DC2A0368EA793AB49C">First description of the adult male of Epeorus aculeatus Braasch (Ephemeroptera: Heptageniidae)</mods:title>
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<mods:namePart id="0009069FDB2E91EC069A75FE4B2354A0">Webb, J. M.</mods:namePart>
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<mods:namePart id="AEF158AFC8FB60664E908C67D400C30F">Mccafferty, W. P.</mods:namePart>
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<mods:title id="4DBF1427FB682D26D6F725FADD055D32">Zootaxa</mods:title>
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<mods:part id="871AAE68568428EAF83CF248CA625608">
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<mods:date id="C21A34CF8886E825CA31F3307DA5E84B">2006</mods:date>
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<mods:detail id="D50BDC4820BB0DAF4C7EBBE186DE9D4E" type="volume">
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<mods:number id="7BAEE0DDCE6D08A3068012C888FFCE82">1277</mods:number>
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<treatment id="040A87E8FFBA507EC629F981FCDB0DEE" ID-DOI="http://doi.org/10.5281/zenodo.6256336" ID-GBIF-Taxon="119347525" ID-Zenodo-Dep="6256336" LSID="urn:lsid:plazi:treatment:040A87E8FFBA507EC629F981FCDB0DEE" httpUri="http://treatment.plazi.org/id/040A87E8FFBA507EC629F981FCDB0DEE" lastPageId="2" lastPageNumber="67" pageId="0" pageNumber="65">
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<subSubSection id="C4B96575FFBA507CC629F981FD9E0F67" box="[264,598,1598,1624]" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="8C1C36FEFFBA507CC629F981FD9E0F67" blockId="0.[264,598,1598,1624]" box="[264,598,1598,1624]" pageId="0" pageNumber="65">
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<heading id="D7548192FFBA507CC629F981FD9E0F67" bold="true" box="[264,598,1598,1624]" fontSize="11" level="1" pageId="0" pageNumber="65" reason="1">
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<taxonomicName id="4BA34D7DFFBA507CC629F981FD9E0F67" ID-CoL="6FNW4" authority="Braasch" authorityName="Braasch" box="[264,598,1598,1624]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFBA507CC629F981FD9E0F67" bold="true" box="[264,598,1598,1624]" pageId="0" pageNumber="65">
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<emphasis id="BED7EAECFFBA507CC629F981FE2F0F67" bold="true" box="[264,487,1598,1624]" italics="true" pageId="0" pageNumber="65">Epeorus aculeatus</emphasis>
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Braasch
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4B96575FFBA507CC629F936FDAB0FD4" pageId="0" pageNumber="65" type="reference_group">
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<paragraph id="8C1C36FEFFBA507CC629F936FC650F9F" blockId="0.[264,941,1673,1696]" box="[264,941,1673,1696]" pageId="0" pageNumber="65">
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<treatmentCitationGroup id="ACB311D0FFBA507CC629F936FC650F9F" box="[264,941,1673,1696]" pageId="0" pageNumber="65">
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<taxonomicName id="4BA34D7DFFBA507CC629F936FD4D0F9F" ID-CoL="6FNW4" authority="Braasch, 1990:7" authorityName="Braasch" authorityPageNumber="7" authorityYear="1990" box="[264,645,1673,1696]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFBA507CC629F936FE050F9F" box="[264,461,1673,1696]" italics="true" pageId="0" pageNumber="65">Epeorus aculeatus</emphasis>
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<treatmentCitation id="0D0210EFFFBA507CC6F5F936FD4D0F9F" author="Braasch" box="[468,645,1673,1696]" page="7" pageId="0" pageNumber="65" year="1990">
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<bibRefCitation id="E8324B0FFFBA507CC6F5F936FDBC0F9F" author="Braasch" box="[468,628,1673,1696]" pageId="0" pageNumber="67" refString="Braasch, D. (1990) Neue Eintagsfliegen aus Thailand nebst einigen Bemerkungen zu deren generischem Status (Insecta, Ephemeroptera, Heptageniidae). Reichenbachia, 28, 7 - 14." type="journal article" year="1990">Braasch, 1990</bibRefCitation>
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:7
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0D0210EFFFBA507CC5B1F936FC650F9F" author="Nguyen" box="[656,941,1673,1696]" page="19" pageId="0" pageNumber="65" year="2004">
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<bibRefCitation id="E8324B0FFFBA507CC5B1F936FC400F9F" author="Nguyen" box="[656,904,1673,1696]" pageId="0" pageNumber="67" refString="Nguyen, V. V. & Bae, Y. J. (2004) Larvae of the heptageniid mayfly genus Epeorus (Ephemeroptera: Heptageniidae) from Vietnam. Journal of Asia-Pacific Entomology, 7, 19 - 28." type="book chapter" year="2004">Nguyen and Bae, 2004</bibRefCitation>
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: 19
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8C1C36FEFFBA507CC629F96EFDAB0FD4" blockId="0.[264,1323,1745,1891]" box="[264,611,1745,1771]" pageId="0" pageNumber="65">Adult description (in alcohol)</paragraph>
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</subSubSection>
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<subSubSection id="C4B96575FFBA507EC619F946FCDB0DEE" lastPageId="2" lastPageNumber="67" pageId="0" pageNumber="65" type="description">
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<paragraph id="8C1C36FEFFBA507EC619F946FB8B0B46" blockId="0.[264,1323,1745,1891]" lastBlockId="2.[264,1323,284,633]" lastPageId="2" lastPageNumber="67" pageId="0" pageNumber="65">
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Body length
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<quantity id="4B5B9B1BFFBA507CC6F7F946FD8D0E2C" box="[470,581,1785,1811]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="65" unit="mm" value="10.6">10.6 mm</quantity>
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; forewing
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<quantity id="4B5B9B1BFFBA507CC5E8F946FC8A0E2C" box="[713,834,1785,1811]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="0" pageNumber="65" unit="mm" value="10.4">10.4 mm</quantity>
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; caudal filaments 27.0 mm. Eyes grayblack, contiguous dorsally. Ocelli black basally, white apically. Frons pale brown. Thorax brown; mesonotal transverse suture absent; medial depression of mesothoracic furcasternum convergent anteriorly; mesosternum with dark transverse band (
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<figureCitation id="14982A7BFFB8507EC3ECFEA3FAD30809" box="[1229,1307,284,310]" captionStart="FIGURES 1 – 5. 1" captionStartId="1.[264,383,1831,1855]" captionTargetBox="[377,1176,313,1812]" captionTargetId="figure@1.[353,1245,311,1823]" captionTargetPageId="1" captionText="FIGURES 1 – 5. 1, dorsal view of male; 2, ventral view of male abdomen; 3, thoracic sternum of male; 4, dorsal view of penes; 5, ventral view of penes." httpUri="https://zenodo.org/record/173362/files/figure.png" pageId="2" pageNumber="67">Fig. 3</figureCitation>
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). Forefemur pale brown with dark brown spot medially, dark apically; tibia and tarsus brown; ratio of foreleg segments (F:Ti:Ta1:Ta2:Ta3:Ta4:Ta5) 5.5:6.6:2.4:2.3:2.0:1.3:0.7; foreclaws dissimilar. Midleg light brown, femur with dark spot medially. Hindlegs missing. Wings with veins brown. Abdominal terga 2–7 yellowish brown with brown markings (
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<figureCitation id="14982A7BFFB8507EC6A4FE58FE030B3E" box="[389,459,487,513]" captionStart="FIGURES 1 – 5. 1" captionStartId="1.[264,383,1831,1855]" captionTargetBox="[377,1176,313,1812]" captionTargetId="figure@1.[353,1245,311,1823]" captionTargetPageId="1" captionText="FIGURES 1 – 5. 1, dorsal view of male; 2, ventral view of male abdomen; 3, thoracic sternum of male; 4, dorsal view of penes; 5, ventral view of penes." httpUri="https://zenodo.org/record/173362/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
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); terga 8–10 brown with dark markings; sterna (
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<figureCitation id="14982A7BFFB8507EC327FE58FB850B3E" box="[1030,1101,487,513]" captionStart="FIGURES 1 – 5. 1" captionStartId="1.[264,383,1831,1855]" captionTargetBox="[377,1176,313,1812]" captionTargetId="figure@1.[353,1245,311,1823]" captionTargetPageId="1" captionText="FIGURES 1 – 5. 1, dorsal view of male; 2, ventral view of male abdomen; 3, thoracic sternum of male; 4, dorsal view of penes; 5, ventral view of penes." httpUri="https://zenodo.org/record/173362/files/figure.png" pageId="2" pageNumber="67">Fig. 2</figureCitation>
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) yellowish brown, slightly darker medially. Genitalia (
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<figureCitation id="14982A7BFFB8507EC5E7FDB0FCA50B16" box="[710,877,527,553]" captionStart="FIGURES 1 – 5. 1" captionStartId="1.[264,383,1831,1855]" captionTargetBox="[377,1176,313,1812]" captionTargetId="figure@1.[353,1245,311,1823]" captionTargetPageId="1" captionText="FIGURES 1 – 5. 1, dorsal view of male; 2, ventral view of male abdomen; 3, thoracic sternum of male; 4, dorsal view of penes; 5, ventral view of penes." httpUri="https://zenodo.org/record/173362/files/figure.png" pageId="2" pageNumber="67">Figs. 4 and 5</figureCitation>
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) brown; base of forceps with small median flange; subgenital plate slightly excavated medially; penes fused basally, widely divergent apically and with titillators present. Caudal filaments brown.
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</paragraph>
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<caption id="D8DC6676FFBB507DC629F898FC9E0E5A" httpUri="https://zenodo.org/record/173362/files/figure.png" pageId="1" pageNumber="66" targetBox="[377,1176,313,1812]" targetPageId="1">
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<paragraph id="8C1C36FEFFBB507DC629F898FC9E0E5A" blockId="1.[264,1322,1831,1893]" pageId="1" pageNumber="66">
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<emphasis id="BED7EAECFFBB507DC629F898FE7D0E00" bold="true" box="[264,437,1831,1855]" pageId="1" pageNumber="66">FIGURES 1–5.</emphasis>
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1, dorsal view of male; 2, ventral view of male abdomen; 3, thoracic sternum of male; 4, dorsal view of penes; 5, ventral view of penes.
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</paragraph>
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</caption>
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<paragraph id="8C1C36FEFFB8507EC629FD10FDF80BF6" blockId="2.[264,1323,687,1033]" box="[264,560,687,713]" pageId="2" pageNumber="67">Diagnosis and discussion</paragraph>
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<paragraph id="8C1C36FEFFB8507EC619FD68FC400D36" blockId="2.[264,1323,687,1033]" pageId="2" pageNumber="67">
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Because no other
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<taxonomicName id="4BA34D7DFFB8507EC532FD68FDBC0BCF" box="[531,628,727,752]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
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<emphasis id="BED7EAECFFB8507EC532FD68FDBC0BCF" box="[531,628,727,752]" italics="true" pageId="2" pageNumber="67">Epeorus</emphasis>
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</taxonomicName>
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from southeast Asia are known from the adult stage, we cannot know how
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<taxonomicName id="4BA34D7DFFB8507EC520FD40FD550A27" box="[513,669,767,792]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFB8507EC520FD40FD550A27" box="[513,669,767,792]" italics="true" pageId="2" pageNumber="67">E. aculeatus</emphasis>
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</taxonomicName>
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differs from the other species. Based on larval characteristics,
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<bibRefCitation id="E8324B0FFFB8507EC69EFC98FD950A7E" author="Kluge" box="[447,605,807,833]" pageId="2" pageNumber="67" refString="Kluge, N. Ju. (2004) The Phylogenetic System of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 442 pp." type="book" year="2004">Kluge (2004)</bibRefCitation>
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indicated that
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<taxonomicName id="4BA34D7DFFB8507EC42EFC98FC690A7F" box="[783,929,807,832]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFB8507EC42EFC98FC690A7F" box="[783,929,807,832]" italics="true" pageId="2" pageNumber="67">E. aculeatus</emphasis>
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</taxonomicName>
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is closely related to the southern European
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<taxonomicName id="4BA34D7DFFB8507EC6A1FCF0FDA30A56" authority="Eaton" authorityName="Eaton" box="[384,619,847,873]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="torrentium">
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<emphasis id="BED7EAECFFB8507EC6A1FCF0FDD40A57" box="[384,540,847,872]" italics="true" pageId="2" pageNumber="67">E. torrentium</emphasis>
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Eaton
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</taxonomicName>
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, the
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<typeStatus id="5318885CFFB8507EC583FCF0FD1C0A56" box="[674,724,847,873]" pageId="2" pageNumber="67">type</typeStatus>
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species of
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<taxonomicName id="4BA34D7DFFB8507EC479FCF0FC720A57" box="[856,954,847,872]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
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<emphasis id="BED7EAECFFB8507EC479FCF0FC720A57" box="[856,954,847,872]" italics="true" pageId="2" pageNumber="67">Epeorus</emphasis>
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</taxonomicName>
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. The penes of
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<taxonomicName id="4BA34D7DFFB8507EC34FFCF0FB370A57" box="[1134,1279,847,872]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFB8507EC34FFCF0FB370A57" box="[1134,1279,847,872]" italics="true" pageId="2" pageNumber="67">E. aculeatus</emphasis>
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</taxonomicName>
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are similar to those of
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<taxonomicName id="4BA34D7DFFB8507EC6C8FCC8FD4F0AAF" box="[489,647,887,912]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="torrentium">
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<emphasis id="BED7EAECFFB8507EC6C8FCC8FD4F0AAF" box="[489,647,887,912]" italics="true" pageId="2" pageNumber="67">E. torrentium</emphasis>
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</taxonomicName>
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and closely related species in that they are fused basally and divergent apically, but the penis lobes are much more slender. It also differs in having titillators and dissimilar foretarsal claws. We do not feel it is prudent at this time to speculate on the phylogenetic position of
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<taxonomicName id="4BA34D7DFFB8507EC5D1FC50FC490D37" box="[752,897,1007,1032]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="BED7EAECFFB8507EC5D1FC50FC490D37" box="[752,897,1007,1032]" italics="true" pageId="2" pageNumber="67">E. aculeatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8C1C36FEFFB8507EC629FB80FE2D0D66" blockId="2.[264,1323,1087,1233]" box="[264,485,1087,1113]" pageId="2" pageNumber="67">Material examined</paragraph>
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<paragraph id="8C1C36FEFFB8507EC619FBD8FCDB0DEE" blockId="2.[264,1323,1087,1233]" pageId="2" pageNumber="67">
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<collectingCountry id="F4B4766EFFB8507EC619FBD8FE1B0DBE" box="[312,467,1127,1153]" name="Thailand" pageId="2" pageNumber="67">THAILAND</collectingCountry>
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, Chiengmai Province, small stream and waterfalls, Doi Sutep, W of Chiengmai,
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<quantity id="4B5B9B1BFFB8507EC6B9FB30FE390D96" box="[408,497,1167,1193]" metricMagnitude="2" metricUnit="m" metricValue="4.4196" pageId="2" pageNumber="67" unit="ft" value="1450.0">1450 ft</quantity>
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,
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<date id="F81D103EFFB8507EC6DCFB30FD430D96" box="[509,651,1167,1193]" pageId="2" pageNumber="67" value="1964-11-19">19XI1964</date>
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, WL & JG Peters, 2 reared male imagos with associated exuviae (penes missing from one specimen).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |