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<document id="3C46C90D210D45687B0F26583A43CC3D" ID-DOI="10.5281/zenodo.274915" ID-GBIF-Dataset="801a1c0b-7aee-4ba6-a0a1-1e6b45aa5527" ID-ISSN="1175-5326" ID-Zenodo-Dep="274915" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461085262390" checkinUser="plazi" docAuthor="Onn, Chan Kin, Grismer, Lee, Ahmad, Norhayati &amp; Belabut, Daicus" docDate="2009" docId="F07587C66C7BA42BFF6AFF3854700B3B" docLanguage="en" docName="zt02124p068.pdf" docOrigin="Zootaxa 2124" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gastrophrynoides immaculatus" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="0C4CFFBE6C79A42FFFFDFFD9504C0A1D" masterDocTitle="A new species of Gastrophrynoides (Anura: Microhylidae): an addition to a previously monotypic genus and a new genus for Peninsular Malaysia" masterLastPageNumber="68" masterPageNumber="63" pageNumber="65" updateTime="1698588018186" updateUser="plazi">
<mods:mods id="953F087140FFF09383A021219DED1100" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E820CDDE5935100CCAAED5297488AAAA">
<mods:title id="2C738F54678A310E1336CB2CDC8A438E">A new species of Gastrophrynoides (Anura: Microhylidae): an addition to a previously monotypic genus and a new genus for Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="EB46A314B1A5DCE3AB0D9D9F7B63F275" type="personal">
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<mods:roleTerm id="B52EF7D028AB3C23E2CDEE76AB6DFB5D">Author</mods:roleTerm>
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<mods:namePart id="45BC7483C1564B34E4C08C8233214CDC">Onn, Chan Kin</mods:namePart>
</mods:name>
<mods:name id="80E54BF44296C573520A2E825B4D314E" type="personal">
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<mods:namePart id="9D25ED3256A485884A27088235FFDCF5">Grismer, Lee</mods:namePart>
</mods:name>
<mods:name id="41FB364BF1EA00E1C75486902F97743F" type="personal">
<mods:role id="0402DA1BF8E8564926B103033AFAE50A">
<mods:roleTerm id="C8634EF03200900E60F950880CD9BC2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E476308D117C3C20E8A222DAEC2E559A">Ahmad, Norhayati</mods:namePart>
</mods:name>
<mods:name id="2F2EF18FAC867B735B14EFCC6D2DC45B" type="personal">
<mods:role id="5B9D37B13B8E65C793AE6AD29C9AF532">
<mods:roleTerm id="8BD673B7E869028D987A1802A16065D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0684C03D3188BBD1477351E4FB8872C1">Belabut, Daicus</mods:namePart>
</mods:name>
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<mods:title id="B0D4C1CA66CEBF590A2FEC25751728E4">Zootaxa</mods:title>
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<mods:part id="083221D9B85CF7439318E5A15D39D6EC">
<mods:date id="122A0FD1ABCC9CD261E28C64638124CB">2009</mods:date>
<mods:detail id="705989596BE18360146C346B723FAA15" type="volume">
<mods:number id="FA51B0FFD1646D99180B5ED5D6661457">2124</mods:number>
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<mods:classification id="A6EAEB47663358C23022CD5BBFE3F997">journal article</mods:classification>
<mods:identifier id="3F7DBEC784A996E54DF282C81968DEB6" type="DOI">10.5281/zenodo.274915</mods:identifier>
<mods:identifier id="C90E6687A6CA1ACDEDACF2713AE2D480" type="GBIF-Dataset">801a1c0b-7aee-4ba6-a0a1-1e6b45aa5527</mods:identifier>
<mods:identifier id="8BB0B144573F865FA65F9F8E4A7103FA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8843941438A9FE8A6AC599D2E033D69D" type="Zenodo-Dep">274915</mods:identifier>
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<treatment id="F07587C66C7BA42BFF6AFF3854700B3B" ID-DOI="http://doi.org/10.5281/zenodo.5657717" ID-GBIF-Taxon="119618630" ID-Zenodo-Dep="5657717" LSID="urn:lsid:plazi:treatment:F07587C66C7BA42BFF6AFF3854700B3B" httpUri="http://treatment.plazi.org/id/F07587C66C7BA42BFF6AFF3854700B3B" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65">
<subSubSection id="30C6655B6C7BA42DFF6AFF3850B60B03" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="786336D06C7BA42DFF6AFF3852230AE7" blockId="2.[151,623,224,286]" box="[151,623,224,251]" pageId="2" pageNumber="65">
<heading id="232B81BC6C7BA42DFF6AFF3852230AE7" bold="true" box="[151,623,224,251]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="4AA8EAC26C7BA42DFF6AFF3852230AE7" bold="true" box="[151,623,224,251]" pageId="2" pageNumber="65">
<taxonomicName id="BFDC4D536C7BA42DFF6AFF38525D0AE6" ID-CoL="3FDW9" box="[151,529,225,251]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="4AA8EAC26C7BA42DFF6AFF38525D0AE6" bold="true" box="[151,529,225,251]" italics="true" pageId="2" pageNumber="65">Gastrophrynoides immaculatus</emphasis>
</taxonomicName>
sp nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="786336D06C7BA42DFF6AFEDD50B60B03" blockId="2.[151,623,224,286]" box="[151,250,260,286]" pageId="2" pageNumber="65">
<figureCitation id="E0E72A556C7BA42DFF6AFEDD50B60B03" box="[151,250,260,286]" captionStart="FIGURE 2" captionStartId="2.[151,255,1410,1434]" captionTargetBox="[188,1398,575,1384]" captionTargetId="figure@2.[188,1398,575,1385]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Gastrophrynoides immaculatus (HC 00279)." httpUri="https://zenodo.org/record/274917/files/figure.png" pageId="2" pageNumber="65">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="30C6655B6C7BA42DFF6AFE88537F0B76" box="[151,819,337,363]" pageId="2" pageNumber="65" type="description">
<paragraph id="786336D06C7BA42DFF6AFE88537F0B76" blockId="2.[151,1436,337,518]" box="[151,819,337,363]" pageId="2" pageNumber="65">Suggested common name: Tungs Narrow-mouthed Frog</paragraph>
</subSubSection>
<subSubSection id="30C6655B6C7BA42DFF38FEAE5172081B" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="786336D06C7BA42DFF38FEAE5172081B" blockId="2.[151,1436,337,518]" pageId="2" pageNumber="65">
<emphasis id="4AA8EAC26C7BA42DFF38FEAE51750B8C" bold="true" box="[197,313,375,401]" pageId="2" pageNumber="65">
<typeStatus id="A76788726C7BA42DFF38FEAE51750B8C" box="[197,313,375,401]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
. HC00279 (
<figureCitation id="E0E72A556C7BA42DFE33FEA152740B8F" box="[462,568,376,402]" captionStart="FIGURE 2" captionStartId="2.[151,255,1410,1434]" captionTargetBox="[188,1398,575,1384]" captionTargetId="figure@2.[188,1398,575,1385]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Gastrophrynoides immaculatus (HC 00279)." httpUri="https://zenodo.org/record/274917/files/figure.png" pageId="2" pageNumber="65">Figure 2</figureCitation>
), adult female, collected
<date id="0C6210106C7BA42DFC91FEA153BC0B8F" box="[876,1008,376,402]" pageId="2" pageNumber="65" value="2008-04">April 2008</date>
from Gunung Besar Hantu, Negeri Sembilan, Peninsular
<collectingCountry id="00CB76406C7BA42DFE6AFE47524E0BA5" box="[407,514,414,440]" name="Malaysia" pageId="2" pageNumber="65">Malaysia</collectingCountry>
(
<figureCitation id="E0E72A556C7BA42DFDEFFE4752380BA5" box="[530,628,414,440]" captionStart="FIGURE 1" captionStartId="1.[151,255,1101,1125]" captionTargetBox="[491,1096,622,1076]" captionTargetId="figure@1.[491,1096,622,1076]" captionTargetPageId="1" captionText="FIGURE 1. Type locality of Gastrophrynoides immaculatus and distribution of G. borneensis." httpUri="https://zenodo.org/record/274916/files/figure.png" pageId="2" pageNumber="65">Figure 1</figureCitation>
) by Orang Asli collectors at an elevation of 8001400 meters a.s.l.
<emphasis id="4AA8EAC26C7BA42DFF38FE1C510F0BC2" bold="true" box="[197,323,453,479]" pageId="2" pageNumber="65">
<typeStatus id="A76788726C7BA42DFF38FE1C510F0BC2" box="[197,323,453,479]" pageId="2" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
. HC00199, male;
<collectionCode id="1ECDAE156C7BA42DFDE2FE1C52CC0BC2" box="[543,640,453,479]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="2" pageNumber="65">LSUHC</collectionCode>
9166, female, collected from the same location as the
<typeStatus id="A76788726C7BA42DFAF3FE1C553A0BC2" box="[1294,1398,453,479]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
on the same date.
</paragraph>
</subSubSection>
<caption id="2CA366586C7BA42DFF6AFA5B53300F87" box="[151,892,1410,1434]" httpUri="https://zenodo.org/record/274917/files/figure.png" pageId="2" pageNumber="65" targetBox="[188,1398,575,1384]" targetPageId="2">
<paragraph id="786336D06C7BA42DFF6AFA5B53300F87" blockId="2.[151,892,1410,1434]" box="[151,892,1410,1434]" pageId="2" pageNumber="65">
<emphasis id="4AA8EAC26C7BA42DFF6AFA5B51550F87" bold="true" box="[151,281,1410,1434]" pageId="2" pageNumber="65">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="BFDC4D536C7BA42DFE5BFA5A52B90F87" box="[422,757,1411,1434]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="4AA8EAC26C7BA42DFE5BFA5A52B90F87" box="[422,757,1411,1434]" italics="true" pageId="2" pageNumber="65">Gastrophrynoides immaculatus</emphasis>
</taxonomicName>
(HC00279).
</paragraph>
</caption>
<subSubSection id="30C6655B6C7BA42DFF38FA1153AD0C32" pageId="2" pageNumber="65" type="diagnosis">
<paragraph id="786336D06C7BA42DFF38FA1153AD0C32" blockId="2.[151,1437,1480,2009]" pageId="2" pageNumber="65">
<emphasis id="4AA8EAC26C7BA42DFF38FA11510A0FFF" bold="true" box="[197,326,1480,1506]" pageId="2" pageNumber="65">Diagnosis.</emphasis>
<taxonomicName id="BFDC4D536C7BA42DFEB3FA11528D0FFC" box="[334,705,1480,1505]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="4AA8EAC26C7BA42DFEB3FA11528D0FFC" box="[334,705,1480,1505]" italics="true" pageId="2" pageNumber="65">Gastrophrynoides immaculatus</emphasis>
</taxonomicName>
is distinguished from all other microhylids by having a long, pointed snout that is 2.63.0 times the diameter of the eye; smooth skin; immaculate, grayish brown dorsum with no white spots; and a single, large, oval, outer metacarpal tubercle.
</paragraph>
</subSubSection>
<subSubSection id="30C6655B6C7BA42CFF38F9E553A4089C" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="description">
<paragraph id="786336D06C7BA42DFF38F9E554C70D23" blockId="2.[151,1437,1480,2009]" pageId="2" pageNumber="65">
<emphasis id="4AA8EAC26C7BA42DFF38F9E551B10C4B" bold="true" box="[197,509,1596,1622]" pageId="2" pageNumber="65">
Description of
<typeStatus id="A76788726C7BA42DFE7BF9E551B40C4B" box="[390,504,1596,1622]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female, SVL 29.3 mm; habitus robust; head slightly wider than long (HW/HL 1.06), triangular in dorsal profile; snout long, 3.0 times diameter of eye, half the length of the head, rounded in lateral profile, greatly projecting beyond lower jaw, ventral surface of projection concave; canthus rostralis indistinct; lores, flat, sloping; nostrils laterally placed, much closer to end of snout than to eye; internarial width less than interorbital width; interorbital width more than three times width of upper eyelid; eyes small, 25% of head length; pupil rounded, widely dilated; tympanum visible but obscured by skin, subcircular, as wide as high, slightly less than eye diameter; supratympanic fold weak.
</paragraph>
<paragraph id="786336D06C7BA42CFF3BF893519E0AAF" blockId="2.[151,1437,1480,2009]" lastBlockId="3.[151,1437,152,758]" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
Skin smooth on all dorsal and ventral surfaces. Forelimbs moderately short, somewhat robust; F3&gt;F2=F4&gt;F1; finger tips expanded into small, round disks bearing circum-marginal grooves; finger disks equal or slightly wider than penultimate phalanges; subarticular and metacarpal tubercles weak; inner metacarpal tubercle small, oval; outer metacarpal tubercle large, oval; fingers free of web.
<collectingCountry id="00CB76406C7BA42DFAECF86655030DC4" box="[1297,1359,1983,2009]" name="India" pageId="2" pageNumber="65">Hind</collectingCountry>
limbs moderate in length, robust;
</paragraph>
<paragraph id="786336D06C7AA42CFF38FF6753900B3B" blockId="3.[151,1437,152,758]" pageId="3" pageNumber="66">T4&gt;T2=T3=T5&gt;T1; toe tips expanded into small, round disks bearing circum-marginal grooves; toe disks slightly wider than penultimate phalanges; toes free of webbing; inner metatarsal tubercle ellipsoidal;outer metatarsal tubercle absent (See appendix for additional measurements).</paragraph>
<paragraph id="786336D06C7AA42CFF38FEEB51310B6E" blockId="3.[151,1437,152,758]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FEEB513D0B51" bold="true" box="[197,369,306,332]" pageId="3" pageNumber="66">Color in life.</emphasis>
Uniform grayish brown above; cream with fine brown stippling (only visible under microscope) below.
</paragraph>
<paragraph id="786336D06C7AA42CFF38FEA6530E0B87" blockId="3.[151,1437,152,758]" box="[197,834,383,410]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FEA651D80B84" bold="true" box="[197,404,383,409]" pageId="3" pageNumber="66">Color in alcohol.</emphasis>
Dark gray above, dirty white below.
</paragraph>
<paragraph id="786336D06C7AA42CFF38FE7F55760BFA" blockId="3.[151,1437,152,758]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FE7F51070BDD" bold="true" box="[197,331,422,448]" pageId="3" pageNumber="66">
<typeStatus id="A76788726C7AA42CFF38FE7F510A0BDD" box="[197,326,422,448]" pageId="3" pageNumber="66" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Resemble
<typeStatus id="A76788726C7AA42CFE2CFE7F52750BDD" box="[465,569,422,448]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
in coloration except the brown stippling beneath the limbs on HC00199 and LSUHC 9166 is denser than that of the
<typeStatus id="A76788726C7AA42CFD65FE1452B20BFA" box="[664,766,461,487]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
(See appendix for measurements of
<typeStatus id="A76788726C7AA42CFB51FE1454920BFA" box="[1196,1246,461,487]" pageId="3" pageNumber="66">type</typeStatus>
series).
</paragraph>
<paragraph id="786336D06C7AA42CFF38FE2A53A4089C" blockId="3.[151,1437,152,758]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FE2A51CD0810" bold="true" box="[197,385,499,525]" pageId="3" pageNumber="66">Comparisons.</emphasis>
<taxonomicName id="BFDC4D536C7AA42CFE77FE2D53520810" box="[394,798,500,525]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="immaculatus">
<emphasis id="4AA8EAC26C7AA42CFE77FE2D53520810" box="[394,798,500,525]" italics="true" pageId="3" pageNumber="66">Gastrophrynoides immaculatus</emphasis>
</taxonomicName>
is distinguished from
<emphasis id="4AA8EAC26C7AA42CFBBEFE2D54A30810" box="[1091,1263,500,525]" italics="true" pageId="3" pageNumber="66">G. b o r n e e n s i s</emphasis>
in having an immaculate, grayish brown dorsum as opposed to a spotted dorsum (Figure 2,3); a longer snout (2.63.0 vs. 2.02.5 times diameter of eye); and a single, large, oval, outer metacarpal tubercle beneath the hand (vs. smaller, paired, rectangular outer metacarpal tubercles in
<taxonomicName id="BFDC4D536C7AA42CFCC5FDB1539B089C" box="[824,983,616,641]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="4AA8EAC26C7AA42CFCC5FDB1539B089C" box="[824,983,616,641]" italics="true" pageId="3" pageNumber="66">G. borneensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="30C6655B6C7AA42CFF38FD57525108EB" pageId="3" pageNumber="66" type="distribution">
<paragraph id="786336D06C7AA42CFF38FD57553408B5" blockId="3.[151,1437,152,758]" box="[197,1400,654,680]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FD57512908B5" bold="true" box="[197,357,654,680]" pageId="3" pageNumber="66">Distribution.</emphasis>
Only known from the
<typeStatus id="A76788726C7AA42CFD8CFD5752EF08B5" box="[625,675,654,680]" pageId="3" pageNumber="66">type</typeStatus>
locality in Gunung Besar Hantu, Negeri Sembilan (
<figureCitation id="E0E72A556C7AA42CFAFEFD57552A08B5" box="[1283,1382,654,680]" captionStart="FIGURE 1" captionStartId="1.[151,255,1101,1125]" captionTargetBox="[491,1096,622,1076]" captionTargetId="figure@1.[491,1096,622,1076]" captionTargetPageId="1" captionText="FIGURE 1. Type locality of Gastrophrynoides immaculatus and distribution of G. borneensis." httpUri="https://zenodo.org/record/274916/files/figure.png" pageId="3" pageNumber="66">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="786336D06C7AA42CFF38FD6C525108EB" blockId="3.[151,1437,152,758]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF38FD6C51DF08D2" bold="true" box="[197,403,693,719]" pageId="3" pageNumber="66">Natural History.</emphasis>
The
<typeStatus id="A76788726C7AA42CFE2FFD6C524908D2" box="[466,517,693,719]" pageId="3" pageNumber="66">type</typeStatus>
series was found between
<quantity id="BF249B356C7AA42CFCB8FD6C539608D2" box="[837,986,693,719]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="3" pageNumber="66" unit="m" value="1100.0" valueMax="1400.0" valueMin="800.0">8001400 m</quantity>
elevation in water-filled bamboo cuts not more than
<quantity id="BF249B356C7AA42CFEBCFD05512208EB" box="[321,366,732,758]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="66" unit="m" value="1.0">1 m</quantity>
above ground.
</paragraph>
</subSubSection>
<caption id="2CA366586C7AA42CFF6AFB2354B20F0C" box="[151,1278,1274,1298]" httpUri="https://zenodo.org/record/274918/files/figure.png" pageId="3" pageNumber="66" targetBox="[468,1117,815,1248]" targetPageId="3">
<paragraph id="786336D06C7AA42CFF6AFB2354B20F0C" blockId="3.[151,1278,1274,1298]" box="[151,1278,1274,1298]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF6AFB2351550F0F" bold="true" box="[151,281,1274,1298]" pageId="3" pageNumber="66">FIGURE 3.</emphasis>
<taxonomicName id="BFDC4D536C7AA42CFEDDFB2352160F0C" box="[288,602,1274,1297]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="4AA8EAC26C7AA42CFEDDFB2352160F0C" box="[288,602,1274,1297]" italics="true" pageId="3" pageNumber="66">Gastrophrynoides borneensis</emphasis>
</taxonomicName>
from Tawau Hills, Sabah. Photograph by Mr. Kueh Boon Hee.
</paragraph>
</caption>
<caption id="2CA366586C7AA42CFF6AFAB354A60F9C" ID-Table-UUID="2CA366586C7AA42CFF6AFAB354A60F9C" box="[151,1258,1386,1410]" httpUri="http://table.plazi.org/id/2CA366586C7AA42CFF6AFAB354A60F9C" pageId="3" pageNumber="66" targetBox="[151,1258,1437,1972]" targetIsTable="true" targetPageId="3">
<paragraph id="786336D06C7AA42CFF6AFAB354A60F9C" blockId="3.[151,1258,1386,1410]" box="[151,1258,1386,1410]" pageId="3" pageNumber="66">
<emphasis id="4AA8EAC26C7AA42CFF6AFAB3514E0F9F" bold="true" box="[151,258,1386,1410]" pageId="3" pageNumber="66">TABLE 1</emphasis>
. Selected measurements (in mm) of type series. See materials and methods for abbreviations.
</paragraph>
</caption>
<paragraph id="786336D06C7AA42CFE44FA44547C0DA9" pageId="3" pageNumber="66">
<table id="0ADCC4706C7A5BD0FF6AFA4454A60DA9" box="[151,1258,1437,1972]" gridcols="4" gridrows="13" pageId="3" pageNumber="66">
<tr id="C6EC34926C7A5BD0FF6AFA4454A60FA9" box="[151,1258,1437,1460]" gridrow="0" pageId="3" pageNumber="66" rowspan-0="1">
<th id="853D5DEE6C7A5BD0FE44FA44526C0FA9" box="[441,544,1437,1460]" gridcol="1" gridrow="0" pageId="3" pageNumber="66">HC00279</th>
<th id="853D5DEE6C7A5BD0FD19FA4453070FA9" box="[740,843,1437,1460]" gridcol="2" gridrow="0" pageId="3" pageNumber="66">HC00199</th>
<th id="853D5DEE6C7A5BD0FBF2FA4454A60FA9" box="[1039,1258,1437,1460]" gridcol="3" gridrow="0" pageId="3" pageNumber="66">LSUHC 9166</th>
</tr>
<tr id="C6EC34926C7A5BD0FF6AFA1E54A60FC3" box="[151,1258,1479,1502]" gridrow="1" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AFA1E50AA0FC3" box="[151,230,1479,1502]" gridcol="0" gridrow="1" pageId="3" pageNumber="66">SVL</th>
<td id="853D5DEE6C7A5BD0FE44FA1E526C0FC3" box="[441,544,1479,1502]" gridcol="1" gridrow="1" pageId="3" pageNumber="66">29.3</td>
<td id="853D5DEE6C7A5BD0FD19FA1E53070FC3" box="[740,843,1479,1502]" gridcol="2" gridrow="1" pageId="3" pageNumber="66">24.4</td>
<td id="853D5DEE6C7A5BD0FBF2FA1E54A60FC3" box="[1039,1258,1479,1502]" gridcol="3" gridrow="1" pageId="3" pageNumber="66">26.2</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AFA2B54A60C14" box="[151,1258,1522,1545]" gridrow="2" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AFA2B50AA0C14" box="[151,230,1522,1545]" gridcol="0" gridrow="2" pageId="3" pageNumber="66">TiL</th>
<td id="853D5DEE6C7A5BD0FE44FA2B526C0C14" box="[441,544,1522,1545]" gridcol="1" gridrow="2" pageId="3" pageNumber="66">12.3</td>
<td id="853D5DEE6C7A5BD0FD19FA2B53070C14" box="[740,843,1522,1545]" gridcol="2" gridrow="2" pageId="3" pageNumber="66">9.4</td>
<td id="853D5DEE6C7A5BD0FBF2FA2B54A60C14" box="[1039,1258,1522,1545]" gridcol="3" gridrow="2" pageId="3" pageNumber="66">12.0</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF9C454A60C29" box="[151,1258,1565,1588]" gridrow="3" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF9C450AA0C29" box="[151,230,1565,1588]" gridcol="0" gridrow="3" pageId="3" pageNumber="66">ED</th>
<td id="853D5DEE6C7A5BD0FE44F9C4526C0C29" box="[441,544,1565,1588]" gridcol="1" gridrow="3" pageId="3" pageNumber="66">1.5</td>
<td id="853D5DEE6C7A5BD0FD19F9C453070C29" box="[740,843,1565,1588]" gridcol="2" gridrow="3" pageId="3" pageNumber="66">1.4</td>
<td id="853D5DEE6C7A5BD0FBF2F9C454A60C29" box="[1039,1258,1565,1588]" gridcol="3" gridrow="3" pageId="3" pageNumber="66">1.6</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF99154A60C42" box="[151,1258,1608,1631]" gridrow="4" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF99150AA0C42" box="[151,230,1608,1631]" gridcol="0" gridrow="4" pageId="3" pageNumber="66">HL</th>
<td id="853D5DEE6C7A5BD0FE44F991526C0C42" box="[441,544,1608,1631]" gridcol="1" gridrow="4" pageId="3" pageNumber="66">7.9</td>
<td id="853D5DEE6C7A5BD0FD19F99153070C42" box="[740,843,1608,1631]" gridcol="2" gridrow="4" pageId="3" pageNumber="66">7.3</td>
<td id="853D5DEE6C7A5BD0FBF2F99154A60C42" box="[1039,1258,1608,1631]" gridcol="3" gridrow="4" pageId="3" pageNumber="66">7.2</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF9AB54A60C94" box="[151,1258,1650,1673]" gridrow="5" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF9AB50AA0C94" box="[151,230,1650,1673]" gridcol="0" gridrow="5" pageId="3" pageNumber="66">HW</th>
<td id="853D5DEE6C7A5BD0FE44F9AB526C0C94" box="[441,544,1650,1673]" gridcol="1" gridrow="5" pageId="3" pageNumber="66">8.4</td>
<td id="853D5DEE6C7A5BD0FD19F9AB53070C94" box="[740,843,1650,1673]" gridcol="2" gridrow="5" pageId="3" pageNumber="66">6.1</td>
<td id="853D5DEE6C7A5BD0FBF2F9AB54A60C94" box="[1039,1258,1650,1673]" gridcol="3" gridrow="5" pageId="3" pageNumber="66">7.0</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF94454A60CA9" box="[151,1258,1693,1716]" gridrow="6" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF94450AA0CA9" box="[151,230,1693,1716]" gridcol="0" gridrow="6" pageId="3" pageNumber="66">SL</th>
<td id="853D5DEE6C7A5BD0FE44F944526C0CA9" box="[441,544,1693,1716]" gridcol="1" gridrow="6" pageId="3" pageNumber="66">4.6</td>
<td id="853D5DEE6C7A5BD0FD19F94453070CA9" box="[740,843,1693,1716]" gridcol="2" gridrow="6" pageId="3" pageNumber="66">4.1</td>
<td id="853D5DEE6C7A5BD0FBF2F94454A60CA9" box="[1039,1258,1693,1716]" gridcol="3" gridrow="6" pageId="3" pageNumber="66">4.2</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF91154A60CC2" box="[151,1258,1736,1759]" gridrow="7" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF91150AA0CC2" box="[151,230,1736,1759]" gridcol="0" gridrow="7" pageId="3" pageNumber="66">DETN</th>
<td id="853D5DEE6C7A5BD0FE44F911526C0CC2" box="[441,544,1736,1759]" gridcol="1" gridrow="7" pageId="3" pageNumber="66">2.8</td>
<td id="853D5DEE6C7A5BD0FD19F91153070CC2" box="[740,843,1736,1759]" gridcol="2" gridrow="7" pageId="3" pageNumber="66">2.2</td>
<td id="853D5DEE6C7A5BD0FBF2F91154A60CC2" box="[1039,1258,1736,1759]" gridcol="3" gridrow="7" pageId="3" pageNumber="66">2.1</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF92B54A60D14" box="[151,1258,1778,1801]" gridrow="8" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF92B50AA0D14" box="[151,230,1778,1801]" gridcol="0" gridrow="8" pageId="3" pageNumber="66">DNTS</th>
<td id="853D5DEE6C7A5BD0FE44F92B526C0D14" box="[441,544,1778,1801]" gridcol="1" gridrow="8" pageId="3" pageNumber="66">1.7</td>
<td id="853D5DEE6C7A5BD0FD19F92B53070D14" box="[740,843,1778,1801]" gridcol="2" gridrow="8" pageId="3" pageNumber="66">1.4</td>
<td id="853D5DEE6C7A5BD0FBF2F92B54A60D14" box="[1039,1258,1778,1801]" gridcol="3" gridrow="8" pageId="3" pageNumber="66">1.9</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF8C454A60D29" box="[151,1258,1821,1844]" gridrow="9" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF8C450AA0D29" box="[151,230,1821,1844]" gridcol="0" gridrow="9" pageId="3" pageNumber="66">ELW</th>
<td id="853D5DEE6C7A5BD0FE44F8C4526C0D29" box="[441,544,1821,1844]" gridcol="1" gridrow="9" pageId="3" pageNumber="66">1.1</td>
<td id="853D5DEE6C7A5BD0FD19F8C453070D29" box="[740,843,1821,1844]" gridcol="2" gridrow="9" pageId="3" pageNumber="66">1.0</td>
<td id="853D5DEE6C7A5BD0FBF2F8C454A60D29" box="[1039,1258,1821,1844]" gridcol="3" gridrow="9" pageId="3" pageNumber="66">1.1</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF89154A60D42" box="[151,1258,1864,1887]" gridrow="10" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF89150AA0D42" box="[151,230,1864,1887]" gridcol="0" gridrow="10" pageId="3" pageNumber="66">IOD</th>
<td id="853D5DEE6C7A5BD0FE44F891526C0D42" box="[441,544,1864,1887]" gridcol="1" gridrow="10" pageId="3" pageNumber="66">3.8</td>
<td id="853D5DEE6C7A5BD0FD19F89153070D42" box="[740,843,1864,1887]" gridcol="2" gridrow="10" pageId="3" pageNumber="66">3.1</td>
<td id="853D5DEE6C7A5BD0FBF2F89154A60D42" box="[1039,1258,1864,1887]" gridcol="3" gridrow="10" pageId="3" pageNumber="66">3.0</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF8AB54A60D94" box="[151,1258,1906,1929]" gridrow="11" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF8AB50AA0D94" box="[151,230,1906,1929]" gridcol="0" gridrow="11" pageId="3" pageNumber="66">TD</th>
<td id="853D5DEE6C7A5BD0FE44F8AB526C0D94" box="[441,544,1906,1929]" gridcol="1" gridrow="11" pageId="3" pageNumber="66">1.2</td>
<td id="853D5DEE6C7A5BD0FD19F8AB53070D94" box="[740,843,1906,1929]" gridcol="2" gridrow="11" pageId="3" pageNumber="66">1.0</td>
<td id="853D5DEE6C7A5BD0FBF2F8AB54A60D94" box="[1039,1258,1906,1929]" gridcol="3" gridrow="11" pageId="3" pageNumber="66">1.4</td>
</tr>
<tr id="C6EC34926C7A5BD0FF6AF84454A60DA9" box="[151,1258,1949,1972]" gridrow="12" pageId="3" pageNumber="66">
<th id="853D5DEE6C7A5BD0FF6AF84450AA0DA9" box="[151,230,1949,1972]" gridcol="0" gridrow="12" pageId="3" pageNumber="66">IND</th>
<td id="853D5DEE6C7A5BD0FE44F844526C0DA9" box="[441,544,1949,1972]" gridcol="1" gridrow="12" pageId="3" pageNumber="66">2.7</td>
<td id="853D5DEE6C7A5BD0FD19F84453070DA9" box="[740,843,1949,1972]" gridcol="2" gridrow="12" pageId="3" pageNumber="66">2.4</td>
<td id="853D5DEE6C7A5BD0FBF2F84454A60DA9" box="[1039,1258,1949,1972]" gridcol="3" gridrow="12" pageId="3" pageNumber="66">2.6</td>
</tr>
</table>
</paragraph>
<subSubSection id="30C6655B6C7DA42BFF38FF4E54700B3B" pageId="4" pageNumber="67" type="etymology">
<paragraph id="786336D06C7DA42BFF38FF4E54700B3B" blockId="4.[151,1437,151,294]" pageId="4" pageNumber="67">
<emphasis id="4AA8EAC26C7DA42BFF38FF4E511B0AAC" bold="true" box="[197,343,151,177]" pageId="4" pageNumber="67">Etymology.</emphasis>
The specific epithet is a latin masculine adjective in reference to the species uniformly colored dorsum, which differentiates it from its only congener,
<emphasis id="4AA8EAC26C7DA42BFC77FF66547F0AC5" box="[906,1075,191,216]" italics="true" pageId="4" pageNumber="67">
<taxonomicName id="BFDC4D536C7DA42BFC77FF6654630AC5" box="[906,1071,191,216]" class="Amphibia" family="Microhylidae" genus="Gastrophrynoides" kingdom="Animalia" order="Anura" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="borneensis">G. borneensis</taxonomicName>
.
</emphasis>
The proposed common name, Tungs Narrow-mouthed Frog honors Mr. Leong Sung Tung, who discovered this frog and has contributed immensely to the study and knowledge of herpetofauna in Peninsular
<collectingCountry id="00CB76406C7DA42BFC34FED554740B3B" box="[969,1080,268,294]" name="Malaysia" pageId="4" pageNumber="67">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>