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<document id="82F4B2980D6E329B32C69E3D1A6F68C7" ID-DOI="10.5852/ejt.2024.935.2547" ID-ISSN="2118-9773" ID-Zenodo-Dep="11284619" ID-ZooBank="C6FAFB3A-1B4B-4245-A424-0E4EDDA8AF05" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1716568146815" checkinUser="plazi" docAuthor="Jaume-Schinkel, Santiago &amp; Kvifte, Gunnar Mikalsen" docDate="2024" docId="BB4F87F6FFDEF530FF1DFDFAFAE4FB70" docLanguage="en" docName="EJT.2024.935.81-100.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eugenys Quate 1996" docType="key" docVersion="1" lastPageNumber="97" masterDocId="4776FF8EFFCEF520FFA0FFCBFF93FFB1" masterDocTitle="Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species" masterLastPageNumber="100" masterPageNumber="81" pageNumber="97" updateTime="1717032848221" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6C8E7CC822F131B37AD158F8DBAE5DB9">Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species</mods:title>
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<mods:name id="FE2017EB39C426F9994D46C7AAF49E18" type="personal">
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<mods:namePart id="63E2A26EA6D190CE3BD8F035C7BD9332">Jaume-Schinkel, Santiago</mods:namePart>
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<mods:affiliation id="2C5C72CF7478B299137ED7FD365045BF">Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="F13EEBA083028693460407305246D256" type="email">santijaumes@hotmail.com &amp; s.jaume@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="9A6141350EBD562A9F83FE9E806E56B6">Kvifte, Gunnar Mikalsen</mods:namePart>
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<mods:affiliation id="93C42C2C6617A6F7FD00574F3EE1F7FE">Department of Biosciences and Aquaculture, Nord University, P. O. Box 2501, 7729 Steinkjer, Norway.</mods:affiliation>
<mods:nameIdentifier id="F6D782C6B6EA86D3CDD07BA141CF2A2F" type="email">gunnar.mikalsen-kvifte@nord.no</mods:nameIdentifier>
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<mods:title id="7AD1E95FE6593CE4617B1716954AE354">European Journal of Taxonomy</mods:title>
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<mods:part id="A55BA5E47B443429E5B47341191D4232">
<mods:date id="EAA203D570DEEB92EBE6804FEE256A51">2024</mods:date>
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<mods:number id="484A3ABC1A64C698781EE15258314D7A">2024-05-23</mods:number>
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<mods:number id="1194876AAAF2A9733C1BC08457BD7951">935</mods:number>
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<treatment id="BB4F87F6FFDEF530FF1DFDFAFAE4FB70" LSID="urn:lsid:plazi:treatment:BB4F87F6FFDEF530FF1DFDFAFAE4FB70" httpUri="http://treatment.plazi.org/id/BB4F87F6FFDEF530FF1DFDFAFAE4FB70" lastPageNumber="97" pageId="16" pageNumber="97">
<subSubSection id="7BFC656BFFDEF530FF1DFDFAFD4DFDFD" box="[189,734,560,589]" pageId="16" pageNumber="97" type="nomenclature">
<paragraph id="335936E0FFDEF530FF1DFDFAFD4DFDFD" blockId="16.[189,734,560,589]" box="[189,734,560,589]" pageId="16" pageNumber="97">
<emphasis id="0192EAF2FFDEF530FF1DFDFAFD4DFDFD" bold="true" box="[189,734,560,589]" italics="true" pageId="16" pageNumber="97">
Key to the males of
<taxonomicName id="F4E64D63FFDEF530FE63FDFAFD4DFDFD" ID-CoL="4FLT" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[451,734,560,589]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="genus">
Eugenys
<bibRefCitation id="57774B11FFDEF530FD9EFDFBFD4DFDFD" author="Quate L. W." box="[574,734,560,589]" pageId="16" pageNumber="97" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
</taxonomicName>
</emphasis>
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<subSubSection id="7BFC656BFFDEF530FF1DFDAFFAE4FB70" pageId="16" pageNumber="97" type="key">
<key id="477761D1FFDEF530FF1DFDAFFAE4FB70" pageId="16" pageNumber="97">
<keyStep id="88122E45FFDEF530FF1DFDAFFAE4FD13" pageId="16" pageNumber="97">
<paragraph id="335936E0FFDEF530FF1DFDAFFAE4FDCE" blockId="16.[189,1399,612,1217]" box="[189,1399,612,639]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF1DFDAFFAE4FDCE" box="[189,1399,612,639]" pageId="16" pageNumber="97">
1. Hypopods with a lateral digitiform and curved projection (see
<figureCitation id="ABDD2A65FFDEF530FC67FDAFFBD0FDCF" box="[967,1091,612,638]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[300,1338,296,1820]" captionTargetId="figure-12@11.[220,1367,260,1850]" captionTargetPageId="11" captionText="Fig. 6. Eugenys singularis sp. nov., ♂, holotype (INBIO-CRI001470242). A. Head. B. Aedeagus, gonocoxites and gonostyli (ventral view). C. Hypopods (ventral view). D. Aedeagus, gonocoxites and gonostyli (ventral view). E. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="16" pageNumber="97">Fig. 6C, E</figureCitation>
.) ....
<taxonomicName id="F4E64D63FFDEF530FBD8FDAEFA86FDCE" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[1144,1301,612,639]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="0192EAF2FFDEF530FBD8FDAEFA86FDCE" bold="true" box="[1144,1301,612,639]" italics="true" pageId="16" pageNumber="97">E. singularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFDEF530FABCFDAEFAE4FDCE" box="[1308,1399,613,639]" pageId="16" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="335936E0FFDEF530FF61FD43FAE4FD13" blockId="16.[189,1399,612,1217]" box="[193,1399,647,674]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF61FD43FAE4FD13" box="[193,1399,647,674]" pageId="16" pageNumber="97">
– Hypopods without a lateral digitiform projection (as in
<figureCitation id="ABDD2A65FFDEF530FCD4FD4CFC63FD13" box="[884,1008,647,674]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">Fig. 3A–E</figureCitation>
) ................................................ 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="88122E45FFDEF530FF1DFD02FAE4FCB7" pageId="16" pageNumber="97">
<paragraph id="335936E0FFDEF530FF1DFD02FAE4FD52" blockId="16.[189,1399,612,1217]" box="[189,1399,713,740]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF1DFD02FAE4FD52" box="[189,1399,713,740]" pageId="16" pageNumber="97">
2. Hypopods with 14–16 apical tenacula (
<figureCitation id="ABDD2A65FFDEF530FD15FD02FC86FD52" box="[693,789,713,739]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[409,1206,265,1842]" captionTargetId="figure-12@5.[361,1227,265,1852]" captionTargetPageId="5" captionText="Fig. 2. A. Eugenys clavellata Quate, 1996, ♂, paratype (USNMENT00857092), genitalia (ventral view). B. Eugenys cymosa Quate, 1999, ♂, paratype (BMNH(E)2001-8), genitalia (ventral view). Photos A by David Pecor, Smithsonian Institute Museum, B by Santiago Jaume-Schinkel. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284623" httpUri="https://zenodo.org/record/11284623/files/figure.png" pageId="16" pageNumber="97">Figs 2B</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FC81FD02FCD0FD52" box="[801,835,713,739]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">3E</figureCitation>
) ....................................
<taxonomicName id="F4E64D63FFDEF530FBC3FD02FAE4FD52" authority="Quate, 1999" authorityName="Quate" authorityYear="1999" box="[1123,1399,713,739]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="cymosa">
<emphasis id="0192EAF2FFDEF530FBC3FD02FB4CFD52" bold="true" box="[1123,1247,713,739]" italics="true" pageId="16" pageNumber="97">E. cymosa</emphasis>
<bibRefCitation id="57774B11FFDEF530FB47FD02FAE4FD52" author="Quate L. W." box="[1255,1399,713,739]" pageId="16" pageNumber="97" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate, 1999</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="335936E0FFDEF530FF61FD26FAE4FCB7" blockId="16.[189,1399,612,1217]" box="[193,1399,748,775]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF61FD26FAE4FCB7" box="[193,1399,748,775]" pageId="16" pageNumber="97">
– Hypopods with a single apical tenaculum (
<figureCitation id="ABDD2A65FFDEF530FD7FFD27FCEDFCB7" box="[735,894,748,774]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">Fig. 3B, D–E</figureCitation>
) ............................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="88122E45FFDEF530FF1DFCE5FAE4FCDA" pageId="16" pageNumber="97">
<paragraph id="335936E0FFDEF530FF1DFCE5FAE4FCF9" blockId="16.[189,1399,612,1217]" box="[189,1399,813,840]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF1DFCE5FAE4FCF9" box="[189,1399,813,840]" pageId="16" pageNumber="97">
3. Antennal flagellomeres without a patch of darkened sensilla (
<figureCitation id="ABDD2A65FFDEF530FC19FCE5FB86FCF9" box="[953,1045,814,840]" captionStart="Fig" captionStartId="7.[189,232,1891,1917]" captionTargetBox="[215,1375,280,1834]" captionTargetId="figure-11@7.[198,1390,263,1850]" captionTargetPageId="7" captionText="Fig. 4. Eugenys micra sp. nov., ♂, holotype (ZFMK-DIP-00097917). A. Head. B. Wing. C. Mouth parts. D. Scape, pedicel and first two flagellomeres. E. Aedeagus (ventral view). F. Epandrium and hypopods (ventral view). G. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284635" httpUri="https://zenodo.org/record/11284635/files/figure.png" pageId="16" pageNumber="97">Fig. 4A</figureCitation>
) ........................................... 5
</keyLead>
</paragraph>
<paragraph id="335936E0FFDEF530FF61FC9AFAE4FCDA" blockId="16.[189,1399,612,1217]" box="[193,1399,849,875]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF61FC9AFAE4FCDA" box="[193,1399,849,875]" pageId="16" pageNumber="97">
– Antennal flagellomeres with a patch of darkened sensilla (
<figureCitation id="ABDD2A65FFDEF530FC34FC9AFC63FCDA" box="[916,1008,849,875]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[249,1354,380,1923]" captionTargetId="figure-15@14.[208,1380,265,1923]" captionTargetPageId="14" captionText="Fig. 8. Eugenys upsilon sp. nov., ♀, paratype (ZFMK-DIP-00097916). A. Head. B. Wing. C. Mouth parts. D. Genitalia (ventral view). Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284649" httpUri="https://zenodo.org/record/11284649/files/figure.png" pageId="16" pageNumber="97">Fig. 8A</figureCitation>
) ................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="88122E45FFDEF530FF1DFC59FAE4FBA7" pageId="16" pageNumber="97">
<paragraph id="335936E0FFDEF530FF1DFC59FAE4FC61" blockId="16.[189,1399,612,1217]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF1DFC59FAE4FC61" pageId="16" pageNumber="97">
4. Aedeagus curved, without distinguishable paramere apart from the parameral sclerite (
<figureCitation id="ABDD2A65FFDEF530FB7AFC59FAA8FC1D" box="[1242,1339,914,940]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">Figs 3D</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FAE7FC58FE8DFC61" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[258,1333,330,1766]" captionTargetId="figure-12@13.[226,1345,302,1787]" captionTargetPageId="13" captionText="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="16" pageNumber="97">7E– F, H</figureCitation>
) ......................................................................................................................
<taxonomicName id="F4E64D63FFDEF530FB37FC7DFA86FC61" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[1175,1301,949,976]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
<emphasis id="0192EAF2FFDEF530FB37FC7DFA86FC61" bold="true" box="[1175,1301,949,976]" italics="true" pageId="16" pageNumber="97">E. upsilon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFDEF530FABCFC7DFAE4FC61" box="[1308,1399,950,976]" pageId="16" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="335936E0FFDEF530FF61FC12FAE4FBA7" blockId="16.[189,1399,612,1217]" pageId="16" pageNumber="97">
<keyLead id="881795D5FFDEF530FF61FC12FAE4FBA7" pageId="16" pageNumber="97">
– Aedeagus straight, with distinguishable paramere in addition to the parameral sclerite(
<figureCitation id="ABDD2A65FFDEF530FAAEFC12FAE1FC42" box="[1294,1394,985,1011]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">Figs 3B</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FF4AFC37FE9EFBA6" box="[234,269,1020,1047]" captionStart="Fig" captionStartId="10.[191,234,1928,1954]" captionTargetBox="[434,1189,339,1846]" captionTargetId="figure-14@10.[223,1370,265,1888]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia, ventral view. A. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284643" httpUri="https://zenodo.org/record/11284643/files/figure.png" pageId="16" pageNumber="97">5B</figureCitation>
) .........................................................................................................
<taxonomicName id="F4E64D63FFDEF530FB8CFC37FAE4FBA7" authority="Quate, 1999" authorityName="Quate" authorityYear="1999" box="[1068,1399,1020,1046]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="0192EAF2FFDEF530FB8CFC37FB73FBA7" bold="true" box="[1068,1248,1020,1046]" italics="true" pageId="16" pageNumber="97">E. panamensis</emphasis>
<bibRefCitation id="57774B11FFDEF530FB47FC37FAE4FBA7" author="Quate L. W." box="[1255,1399,1020,1046]" pageId="16" pageNumber="97" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate, 1999</bibRefCitation>
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<keyStep id="88122E45FFDEF530FF1DFBF6FAE4FB70" pageId="16" pageNumber="97">
<paragraph id="335936E0FFDEF530FF1DFBF6FAE4FBCA" blockId="16.[189,1399,612,1217]" pageId="16" pageNumber="97">
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5. Aedeagus short, less than half the length of the ejaculatory apodeme; parameral sclerite not Y-shaped, appears absent (
<figureCitation id="ABDD2A65FFDEF530FE04FBABFD91FBCA" box="[420,514,1120,1147]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">Figs 3E</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FDAEFBAAFDC5FBCA" box="[526,598,1121,1147]" captionStart="Fig" captionStartId="7.[189,232,1891,1917]" captionTargetBox="[215,1375,280,1834]" captionTargetId="figure-11@7.[198,1390,263,1850]" captionTargetPageId="7" captionText="Fig. 4. Eugenys micra sp. nov., ♂, holotype (ZFMK-DIP-00097917). A. Head. B. Wing. C. Mouth parts. D. Scape, pedicel and first two flagellomeres. E. Aedeagus (ventral view). F. Epandrium and hypopods (ventral view). G. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284635" httpUri="https://zenodo.org/record/11284635/files/figure.png" pageId="16" pageNumber="97">4E, G</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FDC1FBABFD15FBCA" box="[609,646,1120,1147]" captionStart="Fig" captionStartId="10.[191,234,1928,1954]" captionTargetBox="[434,1189,339,1846]" captionTargetId="figure-14@10.[223,1370,265,1888]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia, ventral view. A. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284643" httpUri="https://zenodo.org/record/11284643/files/figure.png" pageId="16" pageNumber="97">5A</figureCitation>
) ........................................................................
<taxonomicName id="F4E64D63FFDEF530FB0BFBAAFA86FBCB" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[1195,1301,1120,1147]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
<emphasis id="0192EAF2FFDEF530FB0BFBAAFA86FBCB" bold="true" box="[1195,1301,1120,1147]" italics="true" pageId="16" pageNumber="97">E. micra</emphasis>
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<taxonomicNameLabel id="1AA15789FFDEF530FABCFBAAFAE4FBCA" box="[1308,1399,1121,1147]" pageId="16" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="335936E0FFDEF530FF61FB4FFAE4FB70" blockId="16.[189,1399,612,1217]" pageId="16" pageNumber="97">
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– Aedeagus long, more than half the length of the ejaculatory apodeme; parameral sclerite Y-shaped and sclerotized (
<figureCitation id="ABDD2A65FFDEF530FE0CFB6CFD9EFB70" box="[428,525,1191,1217]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[409,1206,265,1842]" captionTargetId="figure-12@5.[361,1227,265,1852]" captionTargetPageId="5" captionText="Fig. 2. A. Eugenys clavellata Quate, 1996, ♂, paratype (USNMENT00857092), genitalia (ventral view). B. Eugenys cymosa Quate, 1999, ♂, paratype (BMNH(E)2001-8), genitalia (ventral view). Photos A by David Pecor, Smithsonian Institute Museum, B by Santiago Jaume-Schinkel. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284623" httpUri="https://zenodo.org/record/11284623/files/figure.png" pageId="16" pageNumber="97">Figs 2A</figureCitation>
,
<figureCitation id="ABDD2A65FFDEF530FDBAFB6CFDAFFB70" box="[538,572,1191,1217]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="16" pageNumber="97">3C</figureCitation>
) ....................................................................
<taxonomicName id="F4E64D63FFDEF530FBE8FB6CFAE4FB70" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[1096,1399,1190,1217]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="clavellata">
<emphasis id="0192EAF2FFDEF530FBE8FB6CFB73FB70" bold="true" box="[1096,1248,1190,1217]" italics="true" pageId="16" pageNumber="97">E. clavellata</emphasis>
<bibRefCitation id="57774B11FFDEF530FB47FB6CFAE4FB70" author="Quate L. W." box="[1255,1399,1191,1217]" pageId="16" pageNumber="97" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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